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<document id="787CF53BF964143224AD489CDE9CA0B9" ID-CLB-Dataset="202083" ID-DOI="10.11646/zootaxa.5211.1.1" ID-GBIF-Dataset="bb97b9dd-2acc-4244-8f30-72bef07b9ed9" ID-ISSN="1175-5326" ID-Zenodo-Dep="7345007" ID-ZooBank="5754769C-B747-4714-BDD9-7D5509D48BEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669101340847" checkinUser="plazi" docAuthor="López-García, Margarita M. &amp; Deloya, Cuauhtémoc" docDate="2022" docId="A209493BFD79FFA7AFA619C1FAA1BA1B" docLanguage="en" docName="zootaxa.5211.1.1.pdf" docOrigin="Zootaxa 5211 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5211.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ligyrus (Ligyrus) bidentulus Fairmaire 1892" docType="treatment" docVersion="9" lastPageNumber="39" masterDocId="5E303143FD5FFF81AF311956FFBEBC61" masterDocTitle="Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision" masterLastPageNumber="119" masterPageNumber="1" pageNumber="39" updateTime="1699396337306" updateUser="plazi">
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<mods:title id="918E2DAD4EBEA967369577B6524ABBEB">Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision</mods:title>
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<mods:namePart id="F6C31BEC7F1295AAA5974A15539751E2">López-García, Margarita M.</mods:namePart>
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<mods:namePart id="BC330600B0692EFCB7209A69E9E18934">Deloya, Cuauhtémoc</mods:namePart>
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<treatment id="A209493BFD79FFA7AFA619C1FAA1BA1B" ID-DOI="http://doi.org/10.5281/zenodo.7345037" ID-GBIF-Taxon="205794474" ID-Zenodo-Dep="7345037" LSID="urn:lsid:plazi:treatment:A209493BFD79FFA7AFA619C1FAA1BA1B" httpUri="http://treatment.plazi.org/id/A209493BFD79FFA7AFA619C1FAA1BA1B" lastPageNumber="39" pageId="38" pageNumber="39">
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<heading id="71574F41FD79FFA7AFA619C1FD7CBCD0" bold="true" box="[151,706,151,178]" fontSize="11" level="2" pageId="38" pageNumber="39" reason="4">
<taxonomicName id="EDA083AEFD79FFA7AFA619C1FD7CBCD0" authority="Fairmaire, 1892" authorityName="Fairmaire" authorityYear="1892" box="[151,706,151,178]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="bidentulus" subGenus="Ligyrus">
<emphasis id="18D4243FFD79FFA7AFA619C1FD7CBCD0" bold="true" box="[151,706,151,178]" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7AFA619C1FF4ABCD0" bold="true" box="[151,244,151,177]" italics="true" pageId="38" pageNumber="39">Ligyrus</emphasis>
(
<emphasis id="18D4243FFD79FFA7AE3419C1FEDCBCD0" bold="true" box="[261,354,151,177]" italics="true" pageId="38" pageNumber="39">Ligyrus</emphasis>
)
<emphasis id="18D4243FFD79FFA7AE4219C1FE4CBCD3" bold="true" box="[371,498,151,178]" italics="true" pageId="38" pageNumber="39">bidentulus</emphasis>
<bibRefCitation id="4E3185DCFD79FFA7AECB19C1FD7CBCD0" author="Fairmaire, L." box="[506,706,151,177]" pageId="38" pageNumber="39" pagination="242 - 253" refId="ref90130" refString="Fairmaire, L. (1892) Descriptions de quelques coleopteres argentins. Annales de la Societe Entomologique de Belgique, 36, 242 - 253." type="journal article" year="1892">Fairmaire, 1892</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="62BAABA6FD79FFA7AFA619EDFDA2BCB7" box="[151,540,187,214]" pageId="38" pageNumber="39" type="description">
<paragraph id="2A1FF82DFD79FFA7AFA619EDFDA2BCB7" blockId="38.[151,706,151,214]" box="[151,540,187,214]" pageId="38" pageNumber="39">
(
<figureCitation id="B29BE4A8FD79FFA7AFAE19EDFEBCBCB4" box="[159,258,187,213]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="38" pageNumber="39">Figs. 1G</figureCitation>
,
<figureCitation id="B29BE4A8FD79FFA7AE3E19EDFE85BCB4" box="[271,315,187,213]" captionStart="FIGURE 13" captionStartId="89.[151,250,1041,1065]" captionTargetBox="[154,1432,183,1016]" captionTargetId="figure-17@89.[151,1436,181,1017]" captionTargetPageId="89" captionText="FIGURE 13. Female genital plates. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) bidentulus, G) L. (Ligyrus) neglectus, H) Tomarus laticaudus, I) T. maternus, J) T. rostratus, K) T. subtropicus. Abbreviations: SCx: Subcoxite, Cx: Coxite." figureDoi="http://doi.org/10.5281/zenodo.7345057" httpUri="https://zenodo.org/record/7345057/files/figure.png" pageId="38" pageNumber="39">13F</figureCitation>
,
<figureCitation id="B29BE4A8FD79FFA7AE7719EDFED1BCB7" box="[326,367,187,214]" captionStart="FIGURE 15" captionStartId="90.[151,250,1535,1559]" captionTargetBox="[154,1430,183,1509]" captionTargetId="figure-24@90.[151,1436,181,1512]" captionTargetPageId="90" captionText="FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345063" httpUri="https://zenodo.org/record/7345063/files/figure.png" pageId="38" pageNumber="39">15J</figureCitation>
,
<figureCitation id="B29BE4A8FD79FFA7AE4A19EDFE12BCB4" box="[379,428,187,213]" captionStart="FIGURE 18" captionStartId="93.[151,250,1597,1621]" captionTargetBox="[152,1436,184,1567]" captionTargetId="figure-17@93.[151,1436,181,1573]" captionTargetPageId="93" captionText="FIGURE 18. Parameres in lateral view.A) L. (Ligyrus.) allonasutus, B) L. (Ligyrus) bidentulus, C) L. (Ligyrus) burmeisteri, D) Ligyrus (Ligyrus) fossor, E) L. (Ligyrus) gibbosus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) neglectus, H) L. (Ligyrus) paranaensis, I) L. (Ligyrus) peninsularis, J) L. (Ligyrus) rubripes, K) L. (Ligyrus) ruginasus, L) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345077" httpUri="https://zenodo.org/record/7345077/files/figure.png" pageId="38" pageNumber="39">18B</figureCitation>
,
<figureCitation id="B29BE4A8FD79FFA7AE8919EDFE55BCB7" box="[440,491,187,214]" captionStart="FIGURE 25" captionStartId="100.[151,250,1841,1865]" captionTargetBox="[163,1432,189,1806]" captionTargetId="figure-24@100.[151,1436,181,1817]" captionTargetPageId="100" captionText="FIGURE 25. Habitus of the species of Ligyrus. Subgenus Anagrylius: A) Ligyrus cuniculus, B) L. moroni. Subgenus Ligyrodes: C) L. peruvianus, D) L. relictus, E) L. sallaei. Subgenus Ligyrus: F) L. allonasutus, G) L. bidentulus, H) L. burmeisteri. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345097" httpUri="https://zenodo.org/record/7345097/files/figure.png" pageId="38" pageNumber="39">25G</figureCitation>
;
<figureCitation id="B29BE4A8FD79FFA7AEC619EDFDAABCB7" box="[503,532,187,214]" captionStart="FIGURE 35" captionStartId="108.[151,250,1066,1090]" captionTargetBox="[215,1380,181,1033]" captionTargetId="figure-24@108.[207,1380,181,1043]" captionTargetPageId="108" captionText="FIGURE 35. Distributional records of Ligyrus (Ligyrus) villosus, L. bidentulus, and L. rubripes." figureDoi="http://doi.org/10.5281/zenodo.7345131" httpUri="https://zenodo.org/record/7345131/files/figure.png" pageId="38" pageNumber="39">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD79FFA7AFA619ADFD8ABD73" box="[151,564,250,274]" pageId="38" pageNumber="39" type="reference_group">
<paragraph id="2A1FF82DFD79FFA7AFA619ADFD8ABD73" blockId="38.[151,1437,250,336]" box="[151,564,250,274]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="0AB0DF03FD79FFA7AFA619ADFD8ABD73" box="[151,564,250,274]" pageId="38" pageNumber="39">
<treatmentCitation id="AB01DE3CFD79FFA7AFA619ADFD8EBD73" author="Fairmaire, L." box="[151,560,250,274]" page="244" pageId="38" pageNumber="39" year="1892">
<taxonomicName id="EDA083AEFD79FFA7AFA619ADFD8EBD73" authority="Fairmaire, 1892: 244" authorityName="Fairmaire" authorityPageNumber="244" authorityYear="1892" box="[151,560,250,274]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="bidentulus">
<emphasis id="18D4243FFD79FFA7AFA619ADFEEABD73" box="[151,340,250,274]" italics="true" pageId="38" pageNumber="39">Ligyrus bidentulus</emphasis>
<bibRefCitation id="4E3185DCFD79FFA7AE6819ACFD8EBD73" author="Fairmaire, L." box="[345,560,250,274]" pageId="38" pageNumber="39" pagination="242 - 253" refId="ref90130" refString="Fairmaire, L. (1892) Descriptions de quelques coleopteres argentins. Annales de la Societe Entomologique de Belgique, 36, 242 - 253." type="journal article" year="1892">Fairmaire, 1892: 244</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD79FFA7AD0819ACFD97BD31" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="2A1FF82DFD79FFA7AD0819ACFD97BD31" blockId="38.[151,1437,250,336]" pageId="38" pageNumber="39">
Male
<typeStatus id="F51B468FFD79FFA7AD4319ADFD7CBD72" box="[626,706,251,275]" pageId="38" pageNumber="39" type="neotype">neotype</typeStatus>
designated by
<treatmentCitation id="AB01DE3CFD79FFA7AC6819ACFB76BD73" author="Lopez-Garcia, M. M. &amp; Deloya, C." box="[857,1224,250,274]" page="128" pageId="38" pageNumber="39" year="2019">
<bibRefCitation id="4E3185DCFD79FFA7AC6819ACFB76BD73" author="Lopez-Garcia, M. M. &amp; Deloya, C." box="[857,1224,250,274]" pageId="38" pageNumber="39" pagination="127 - 141" refId="ref91328" refString="Lopez-Garcia, M. M. &amp; Deloya, C. (2019) Five new species of the dynastine genus Tomarus (Coleoptera: Scarabaeidae), with an illustrated key to species. The Coleopterists Bulletin, 73, 127 - 141. https: // doi. org / 10.1649 / 0010 - 065 X- 73.1.127" type="journal article" year="2019">López-García &amp; Deloya (2019: 128)</bibRefCitation>
</treatmentCitation>
(USNM) “
<collectingRegion id="E86436CFFD79FFA7AA0719ACFA2EBD73" box="[1334,1424,250,274]" country="Argentina" name="La Rioja" pageId="38" pageNumber="39">La Rioja</collectingRegion>
/ Tinogasta / v.965 Kohler //
<collectingCountry id="52B7B8BDFD79FFA7AEE9184FFDD4BD50" box="[472,618,281,305]" name="Argentina" pageId="38" pageNumber="39">ARGENTINA</collectingCountry>
/ 1968 Colln. / J. Daguerre //
<taxonomicName id="EDA083AEFD79FFA7ACA2184FFBECBD50" authorityName="Fairmaire" authorityYear="1892" box="[915,1106,281,305]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="bidentulus">Ligyrus bidentulus</taxonomicName>
/
<bibRefCitation id="4E3185DCFD79FFA7AB55184FFABBBD50" author="Fairmaire, L." box="[1124,1285,281,305]" pageId="38" pageNumber="39" pagination="242 - 253" refId="ref90130" refString="Fairmaire, L. (1892) Descriptions de quelques coleopteres argentins. Annales de la Societe Entomologique de Belgique, 36, 242 - 253." type="journal article" year="1892">Fairmaire, 1892</bibRefCitation>
/
<typeStatus id="F51B468FFD79FFA7AA26184FFA2EBD50" box="[1303,1424,281,305]" pageId="38" pageNumber="39" type="neotype">NEOTYPE</typeStatus>
”. Type locality:
<collectingRegion id="E86436CFFD79FFA7AE68186EFE0ABD31" box="[345,436,312,336]" country="Argentina" name="La Rioja" pageId="38" pageNumber="39">La Rioja</collectingRegion>
, Argentina.
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD79FFA7AFA6182EFE79BFB3" pageId="38" pageNumber="39" type="description">
<paragraph id="2A1FF82DFD79FFA7AFA6182EFE79BFB3" blockId="38.[151,1437,376,1658]" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7AFA6182EFE97BDF3" bold="true" box="[151,297,376,402]" pageId="38" pageNumber="39">Description.</emphasis>
Habitus as in
<figureCitation id="B29BE4A8FD79FFA7AEF3182EFD96BDF2" box="[450,552,376,403]" captionStart="FIGURE 25" captionStartId="100.[151,250,1841,1865]" captionTargetBox="[163,1432,189,1806]" captionTargetId="figure-24@100.[151,1436,181,1817]" captionTargetPageId="100" captionText="FIGURE 25. Habitus of the species of Ligyrus. Subgenus Anagrylius: A) Ligyrus cuniculus, B) L. moroni. Subgenus Ligyrodes: C) L. peruvianus, D) L. relictus, E) L. sallaei. Subgenus Ligyrus: F) L. allonasutus, G) L. bidentulus, H) L. burmeisteri. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345097" httpUri="https://zenodo.org/record/7345097/files/figure.png" pageId="38" pageNumber="39">Fig. 25G</figureCitation>
. Length
<quantity id="ED5855C8FD79FFA7ADB7182FFC94BDF2" box="[646,810,377,403]" metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" metricValueMax="1.93" metricValueMin="1.21" pageId="38" pageNumber="39" unit="mm" value="15.7" valueMax="19.3" valueMin="12.1">12.1–19.3 mm</quantity>
; humeral width
<quantity id="ED5855C8FD79FFA7ACEB182EFBCFBDF2" box="[986,1137,376,403]" metricMagnitude="-3" metricUnit="m" metricValue="8.25" metricValueMax="10.7" metricValueMin="5.8" pageId="38" pageNumber="39" unit="mm" value="8.25" valueMax="10.7" valueMin="5.8">5.8–10.7 mm</quantity>
. Color dark reddish brown.
<emphasis id="18D4243FFD79FFA7AFA618CBFF5EBDD6" bold="true" box="[151,224,413,439]" pageId="38" pageNumber="39">Head:</emphasis>
Frons deeply rugopunctate, with large and sparse punctures, nearly smooth on vertex and with only sparse, small punctures. Frontoclypeal region with a carina, narrowly interrupted at middle (
<figureCitation id="B29BE4A8FD79FFA7AB711896FB26BDBA" box="[1088,1176,448,475]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="38" pageNumber="39">Fig. 1G</figureCitation>
). Ocular canthus acute; with 8 ventral setae. Clypeal surface rugopunctate to transversely rugose, striae denser than those on frons. Clypeus triangular, base 3.5 times wider than apex (
<figureCitation id="B29BE4A8FD79FFA7AD421B5EFD72BE42" box="[627,716,520,547]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="38" pageNumber="39">Fig. 1G</figureCitation>
). Clypeal teeth triangular, slightly separated (
<figureCitation id="B29BE4A8FD79FFA7ABF41B5EFAA3BE42" box="[1221,1309,520,547]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="38" pageNumber="39">Fig. 1G</figureCitation>
). Mandible with 2 apical and 1 lateral, rounded tooth, tooth 1 longer and narrower than tooth 2. Maxilla subrectangular; galea with 3 dorsal and 2 ventral teeth. Apex of labrum straight to rounded. Interocular distance equal to 3.5 times an eye width. Antennal club very long.
<emphasis id="18D4243FFD79FFA7AD3F1B23FD2EBEEE" bold="true" box="[526,656,629,655]" pageId="38" pageNumber="39">Pronotum:</emphasis>
Surface with large, sparse punctures. Apex with small tubercle and shallow fovea.
<emphasis id="18D4243FFD79FFA7AE751BCFFE24BED2" bold="true" box="[324,410,665,691]" pageId="38" pageNumber="39">Elytra:</emphasis>
First interval punctate, punctures similar to the other intervals. Inner surface of apex with small, transverse tubercles forming about 115 parallel lines.
<emphasis id="18D4243FFD79FFA7AC701BEAFC01BEB7" bold="true" box="[833,959,700,726]" pageId="38" pageNumber="39">Abdomen:</emphasis>
Apex of tergite IV with a diagonal line of 10 transverse, parallel, short carinae.
<emphasis id="18D4243FFD79FFA7AD081BB6FD0BBE9B" bold="true" box="[569,693,736,762]" pageId="38" pageNumber="39">Pygidium:</emphasis>
Surface with small to large punctures, denser on base and anterior corners. Strongly (male) to slightly (female) convex in lateral view.
<emphasis id="18D4243FFD79FFA7ACAE1A53FC5FBF7E" bold="true" box="[927,993,773,799]" pageId="38" pageNumber="39">Legs:</emphasis>
Protibia tridentate; basal tooth slightly distant from others. Protarsus of male simple, inner claw simple. Metatibia triangular. Apex of metatibia entire, with 25–29 spinules.
<emphasis id="18D4243FFD79FFA7AEB51A1BFDEFBF07" bold="true" box="[388,593,844,871]" pageId="38" pageNumber="39">Female genitalia:</emphasis>
Subcoxite subrectangular (2 times wider than long). Coxite subquadrate (as long as wide); surface flat. Subcoxite wider and subequal in length to coxite (
<figureCitation id="B29BE4A8FD79FFA7ACCF1A26FBDCBFEB" box="[1022,1122,880,906]" captionStart="FIGURE 13" captionStartId="89.[151,250,1041,1065]" captionTargetBox="[154,1432,183,1016]" captionTargetId="figure-17@89.[151,1436,181,1017]" captionTargetPageId="89" captionText="FIGURE 13. Female genital plates. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) bidentulus, G) L. (Ligyrus) neglectus, H) Tomarus laticaudus, I) T. maternus, J) T. rostratus, K) T. subtropicus. Abbreviations: SCx: Subcoxite, Cx: Coxite." figureDoi="http://doi.org/10.5281/zenodo.7345057" httpUri="https://zenodo.org/record/7345057/files/figure.png" pageId="38" pageNumber="39">Fig. 13F</figureCitation>
).
<emphasis id="18D4243FFD79FFA7AB471A26FA94BFEB" bold="true" box="[1142,1322,880,906]" pageId="38" pageNumber="39">Male genitalia:</emphasis>
Spiculum gastrale thin, apex truncate. Parameres with apical 4th narrowed, without minute spines; lateral ventral teeth short, triangular (
<figureCitation id="B29BE4A8FD79FFA7AE221AEEFEC5BFB3" box="[275,379,951,978]" captionStart="FIGURE 15" captionStartId="90.[151,250,1535,1559]" captionTargetBox="[154,1430,183,1509]" captionTargetId="figure-24@90.[151,1436,181,1512]" captionTargetPageId="90" captionText="FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345063" httpUri="https://zenodo.org/record/7345063/files/figure.png" pageId="38" pageNumber="39">Figs. 15J</figureCitation>
,
<figureCitation id="B29BE4A8FD79FFA7AEB91AEEFE07BFB3" box="[392,441,952,978]" captionStart="FIGURE 18" captionStartId="93.[151,250,1597,1621]" captionTargetBox="[152,1436,184,1567]" captionTargetId="figure-17@93.[151,1436,181,1573]" captionTargetPageId="93" captionText="FIGURE 18. Parameres in lateral view.A) L. (Ligyrus.) allonasutus, B) L. (Ligyrus) bidentulus, C) L. (Ligyrus) burmeisteri, D) Ligyrus (Ligyrus) fossor, E) L. (Ligyrus) gibbosus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) neglectus, H) L. (Ligyrus) paranaensis, I) L. (Ligyrus) peninsularis, J) L. (Ligyrus) rubripes, K) L. (Ligyrus) ruginasus, L) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345077" httpUri="https://zenodo.org/record/7345077/files/figure.png" pageId="38" pageNumber="39">18B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD79FFA7AFF61A8DFEEDB8E5" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="2A1FF82DFD79FFA7AFF61A8DFEEDB8E5" blockId="38.[151,1437,376,1658]" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7AFF61A8DFEFCBF94" bold="true" box="[199,322,987,1013]" pageId="38" pageNumber="39">Diagnosis.</emphasis>
<taxonomicName id="EDA083AEFD79FFA7AE7B1A8AFDA0BF94" authorityName="Fairmaire" authorityYear="1892" box="[330,542,987,1013]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="bidentulus">
<emphasis id="18D4243FFD79FFA7AE7B1A8AFDA0BF94" box="[330,542,987,1013]" italics="true" pageId="38" pageNumber="39">Ligyrus bidentulus</emphasis>
</taxonomicName>
can be recognized by the very long antennal club (2 times longer than antennomeres 2–7); clypeus long, strongly contracted towards apex (
<figureCitation id="B29BE4A8FD79FFA7AC471AA9FC6CB878" box="[886,978,1023,1049]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="38" pageNumber="39">Fig. 1G</figureCitation>
); clypeal teeth triangular, long, and not separated; frontal carinae narrowly interrupted at middle (
<figureCitation id="B29BE4A8FD79FFA7AC2D1D75FCCBB85C" box="[796,885,1059,1085]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="38" pageNumber="39">Fig. 1G</figureCitation>
); pronotum with small tubercle and narrow fovea (
<figureCitation id="B29BE4A8FD79FFA7AFAE1D10FEB6B800" box="[159,264,1094,1121]" captionStart="FIGURE 25" captionStartId="100.[151,250,1841,1865]" captionTargetBox="[163,1432,189,1806]" captionTargetId="figure-24@100.[151,1436,181,1817]" captionTargetPageId="100" captionText="FIGURE 25. Habitus of the species of Ligyrus. Subgenus Anagrylius: A) Ligyrus cuniculus, B) L. moroni. Subgenus Ligyrodes: C) L. peruvianus, D) L. relictus, E) L. sallaei. Subgenus Ligyrus: F) L. allonasutus, G) L. bidentulus, H) L. burmeisteri. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345097" httpUri="https://zenodo.org/record/7345097/files/figure.png" pageId="38" pageNumber="39">Fig. 25G</figureCitation>
); parameres with apical 4th narrowed, without minute spines; lateral teeth of parameres short, triangular (
<figureCitation id="B29BE4A8FD79FFA7AFAE1D3CFEB9B8E4" box="[159,263,1130,1157]" captionStart="FIGURE 15" captionStartId="90.[151,250,1535,1559]" captionTargetBox="[154,1430,183,1509]" captionTargetId="figure-24@90.[151,1436,181,1512]" captionTargetPageId="90" captionText="FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345063" httpUri="https://zenodo.org/record/7345063/files/figure.png" pageId="38" pageNumber="39">Figs. 15J</figureCitation>
,
<figureCitation id="B29BE4A8FD79FFA7AE251D3CFEFBB8E5" box="[276,325,1130,1156]" captionStart="FIGURE 18" captionStartId="93.[151,250,1597,1621]" captionTargetBox="[152,1436,184,1567]" captionTargetId="figure-17@93.[151,1436,181,1573]" captionTargetPageId="93" captionText="FIGURE 18. Parameres in lateral view.A) L. (Ligyrus.) allonasutus, B) L. (Ligyrus) bidentulus, C) L. (Ligyrus) burmeisteri, D) Ligyrus (Ligyrus) fossor, E) L. (Ligyrus) gibbosus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) neglectus, H) L. (Ligyrus) paranaensis, I) L. (Ligyrus) peninsularis, J) L. (Ligyrus) rubripes, K) L. (Ligyrus) ruginasus, L) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345077" httpUri="https://zenodo.org/record/7345077/files/figure.png" pageId="38" pageNumber="39">18B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD79FFA7AFF61DD8FAA1BA1B" pageId="38" pageNumber="39" type="distribution">
<paragraph id="2A1FF82DFD79FFA7AFF61DD8FB18B8C9" blockId="38.[151,1437,376,1658]" box="[199,1190,1166,1193]" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7AFF61DD8FEDFB8C9" bold="true" box="[199,353,1166,1192]" pageId="38" pageNumber="39">Distribution.</emphasis>
<collectingCountry id="52B7B8BDFD79FFA7AE561DD8FE68B8C9" box="[359,470,1166,1192]" name="Argentina" pageId="38" pageNumber="39">Argentina</collectingCountry>
(
<bibRefCitation id="4E3185DCFD79FFA7AED41DD8FD3FB8C8" author="Endrodi, S." box="[485,641,1166,1193]" pageId="38" pageNumber="39" refId="ref89854" refString="Endrodi, S. (1985) The Dynastinae of the World. W. Junk, Dordrecht, 800 pp." type="book" year="1985">Endrödi 1985</bibRefCitation>
) and
<collectingCountry id="52B7B8BDFD79FFA7ADF11DD9FC9CB8C8" box="[704,802,1167,1193]" name="Uruguay" pageId="38" pageNumber="39">Uruguay</collectingCountry>
(
<bibRefCitation id="4E3185DCFD79FFA7AC001DD8FB24B8C9" author="Neita-Moreno, J. C. &amp; Ratcliffe, B. C." box="[817,1178,1166,1192]" pageId="38" pageNumber="39" pagination="1 - 36" refId="ref91798" refString="Neita-Moreno, J. C. &amp; Ratcliffe, B. C. (2017) The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay. Insecta Mundi, 547, 1 - 36." type="journal article" year="2017">Neita-Moreno &amp; Ratcliffe 2017</bibRefCitation>
).
</paragraph>
<paragraph id="2A1FF82DFD79FFA7AFF61DE7FC07BA37" blockId="38.[151,1437,376,1658]" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7AFF61DE7FE40B8AA" bold="true" box="[199,510,1201,1228]" pageId="38" pageNumber="39">
Locality records (
<figureCitation id="B29BE4A8FD79FFA7AEAA1DE7FE4CB8AA" box="[411,498,1201,1227]" captionStart="FIGURE 35" captionStartId="108.[151,250,1066,1090]" captionTargetBox="[215,1380,181,1033]" captionTargetId="figure-24@108.[207,1380,181,1043]" captionTargetPageId="108" captionText="FIGURE 35. Distributional records of Ligyrus (Ligyrus) villosus, L. bidentulus, and L. rubripes." figureDoi="http://doi.org/10.5281/zenodo.7345131" httpUri="https://zenodo.org/record/7345131/files/figure.png" pageId="38" pageNumber="39">Fig. 35</figureCitation>
).
</emphasis>
58 examined specimens from MERKB, MNHM, MNHN, USNM, ZMUB. Some records from
<bibRefCitation id="4E3185DCFD79FFA7AE021D83FD0AB891" author="Neita-Moreno, J. C. &amp; Ratcliffe, B. C." box="[307,692,1237,1264]" pageId="38" pageNumber="39" pagination="1 - 36" refId="ref91798" refString="Neita-Moreno, J. C. &amp; Ratcliffe, B. C. (2017) The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay. Insecta Mundi, 547, 1 - 36." type="journal article" year="2017">Neita-Moreno &amp; Ratcliffe (2017)</bibRefCitation>
.
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<emphasis id="18D4243FFD79FFA7ADF31D83FB2AB88E" bold="true" box="[706,1172,1237,1264]" pageId="38" pageNumber="39">
<collectingCountry id="52B7B8BDFD79FFA7ADF31D83FCD2B88E" box="[706,876,1237,1263]" name="Argentina" pageId="38" pageNumber="39">ARGENTINA</collectingCountry>
(101). Buenos Aires (9):
</emphasis>
</materialsCitation>
<materialsCitation id="9AC8F270FD79FFA7ABAF1D83FA84B88E" ID-GBIF-Occurrence="3986696397" box="[1182,1338,1237,1263]" pageId="38" pageNumber="39">
<collectingCounty id="C37E80A1FD79FFA7ABAF1D83FA84B88E" box="[1182,1338,1237,1263]" pageId="38" pageNumber="39">Bahía Blanca</collectingCounty>
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AA791D80FF69B972" ID-GBIF-Occurrence="3986696352" pageId="38" pageNumber="39">Buenos Aires</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AFD31DAFFE68B975" ID-GBIF-Occurrence="3986696897" box="[226,470,1273,1300]" pageId="38" pageNumber="39">
<collectingMunicipality id="CA7B6257FD79FFA7AFD31DAFFE68B975" box="[226,470,1273,1300]" pageId="38" pageNumber="39">
Carmen
<collectorName id="87559DFBFD79FFA7AE711DAFFE68B975" box="[320,470,1273,1300]" pageId="38" pageNumber="39">de Patagones</collectorName>
</collectingMunicipality>
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AED11DAFFDECB972" ID-GBIF-Occurrence="3986696469" box="[480,594,1273,1299]" pageId="38" pageNumber="39">Necochea</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AD6C1DAFFD74B972" ID-GBIF-Occurrence="3986696741" box="[605,714,1273,1299]" pageId="38" pageNumber="39">Olavarria</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7ADE51DAFFC87B972" ID-GBIF-Occurrence="3986696830" box="[724,825,1273,1299]" pageId="38" pageNumber="39">San Blas</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AC751DAFFB56B972" ID-GBIF-Occurrence="3986696710" box="[836,1256,1273,1299]" pageId="38" pageNumber="39">
Villa Gesell.
<emphasis id="18D4243FFD79FFA7ACE41DAFFB34B972" bold="true" box="[981,1162,1273,1299]" pageId="38" pageNumber="39">Catamarca (3):</emphasis>
Ancafri
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7ABC31DAFFEF1B956" ID-GBIF-Occurrence="3986696715" pageId="38" pageNumber="39">
Sinquil.
<emphasis id="18D4243FFD79FFA7AA611DAFFF7FB956" bold="true" pageId="38" pageNumber="39">Chaco (1):</emphasis>
Resistencia.
</materialsCitation>
<materialsCitation id="9AC8F270FD79FFA7AE641C4BFDF3B956" ID-GBIF-Occurrence="3986696929" box="[341,589,1309,1335]" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7AE641C4BFE5CB956" bold="true" box="[341,482,1309,1335]" pageId="38" pageNumber="39">Chubut (6):</emphasis>
Dolavon
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AD691C4BFC8EB956" ID-GBIF-Occurrence="3986696764" box="[600,816,1309,1335]" pageId="38" pageNumber="39">
<location id="2F7FAEF6FD79FFA7AD691C4BFC8EB956" LSID="urn:lsid:plazi:treatment:A209493BFD79FFA7AFA619C1FAA1BA1B:2F7FAEF6FD79FFA7AD691C4BFC8EB956" box="[600,816,1309,1335]" country="Argentina" county="Bahia Blanca" municipality="Carmen de Patagones" name="Estacion de Esquel" pageId="38" pageNumber="39" stateProvince="Canelones">
Estación
<collectorName id="87559DFBFD79FFA7AD8E1C4BFC8EB956" box="[703,816,1309,1335]" pageId="38" pageNumber="39">de Esquel</collectorName>
</location>
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AC0A1C4BFBABB956" ID-GBIF-Occurrence="3986696331" box="[827,1045,1309,1335]" pageId="38" pageNumber="39">
<location id="2F7FAEF6FD79FFA7AC0A1C4BFBABB956" LSID="urn:lsid:plazi:treatment:A209493BFD79FFA7AFA619C1FAA1BA1B:2F7FAEF6FD79FFA7AC0A1C4BFBABB956" box="[827,1045,1309,1335]" country="Argentina" county="Bahia Blanca" municipality="Carmen de Patagones" name="Golfo de San Jorge" pageId="38" pageNumber="39" stateProvince="Canelones">Golfo de San Jorge</location>
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AB111C4BFB58B956" ID-GBIF-Occurrence="3986696749" box="[1056,1254,1309,1335]" pageId="38" pageNumber="39">Península Valdés.</materialsCitation>
<materialsCitation id="9AC8F270FD79FFA7ABDD1C4BFEFFB93A" ID-GBIF-Occurrence="3986696701" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7ABDD1C4BFA22B956" bold="true" box="[1260,1436,1308,1335]" pageId="38" pageNumber="39">Entre Ríos (3):</emphasis>
Gualeguaychu.
</materialsCitation>
<materialsCitation id="9AC8F270FD79FFA7AE7B1C16FD2DB93A" ID-GBIF-Occurrence="3986696429" box="[330,659,1344,1371]" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7AE7B1C16FE7DB93B" bold="true" box="[330,451,1344,1370]" pageId="38" pageNumber="39">Jujuy (1):</emphasis>
<location id="2F7FAEF6FD79FFA7AEFD1C17FD2DB93A" LSID="urn:lsid:plazi:treatment:A209493BFD79FFA7AFA619C1FAA1BA1B:2F7FAEF6FD79FFA7AEFD1C17FD2DB93A" box="[460,659,1345,1371]" country="Argentina" county="Bahia Blanca" municipality="Carmen de Patagones" name="Posta de Lozano." pageId="38" pageNumber="39" stateProvince="Canelones">Posta de Lozano.</location>
</materialsCitation>
<materialsCitation id="9AC8F270FD79FFA7ADAC1C17FBA4B93A" ID-GBIF-Occurrence="3986697052" box="[669,1050,1344,1371]" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7ADAC1C17FCEEB93B" bold="true" box="[669,848,1344,1371]" pageId="38" pageNumber="39">La Pampa (1):</emphasis>
Chacharramendi.
</materialsCitation>
<materialsCitation id="9AC8F270FD79FFA7AB121C17FAA7B93A" ID-GBIF-Occurrence="3986696821" box="[1059,1305,1344,1371]" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7AB121C17FB7DB93B" bold="true" box="[1059,1219,1344,1371]" pageId="38" pageNumber="39">La Rioja (9):</emphasis>
Jaguel
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AA161C17FA26B93A" ID-GBIF-Occurrence="3986696502" box="[1319,1432,1345,1371]" pageId="38" pageNumber="39">
<collectorName id="87559DFBFD79FFA7AA161C17FA26B93A" box="[1319,1432,1345,1371]" pageId="38" pageNumber="39">La Unión</collectorName>
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AFA61C32FEBAB91F" ID-GBIF-Occurrence="3986696542" box="[151,260,1380,1406]" pageId="38" pageNumber="39">Mascasin</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AE3E1C33FED9B91E" ID-GBIF-Occurrence="3986697051" box="[271,359,1381,1407]" pageId="38" pageNumber="39">Patquía</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AE401C32FE5CB91F" ID-GBIF-Occurrence="3986696952" box="[369,482,1380,1406]" pageId="38" pageNumber="39">Tinogasta</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AEDC1C32FC5CB91F" ID-GBIF-Occurrence="3986696746" box="[493,994,1380,1407]" pageId="38" pageNumber="39">
Villa Unión.
<emphasis id="18D4243FFD79FFA7AD4C1C33FC95B91F" bold="true" box="[637,811,1380,1407]" pageId="38" pageNumber="39">Mendoza (19):</emphasis>
Bardas Blancas
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7ACDC1C33FB39B91F" ID-GBIF-Occurrence="3986696862" box="[1005,1159,1380,1407]" pageId="38" pageNumber="39">Cerro Guadal</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7ABA31C32FB40B91F" ID-GBIF-Occurrence="3986696827" box="[1170,1278,1380,1406]" pageId="38" pageNumber="39">Mendoza</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AA381C32FF60B9C3" ID-GBIF-Occurrence="3986696922" pageId="38" pageNumber="39">
<collectorName id="87559DFBFD79FFA7AA381C32FACFB91F" box="[1289,1393,1380,1406]" pageId="38" pageNumber="39">El Nihuil</collectorName>
(
<quantity id="ED5855C8FD79FFA7AA4E1C32FF05B9C3" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="38" pageNumber="39" unit="km" value="60.0">60 km</quantity>
S)
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AFDB1CDFFEE1B9C2" ID-GBIF-Occurrence="3986696589" box="[234,351,1417,1443]" pageId="38" pageNumber="39">Las Heras</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AE5B1CDFFE71B9C3" ID-GBIF-Occurrence="3986696484" box="[362,463,1416,1443]" pageId="38" pageNumber="39">
<collectorName id="87559DFBFD79FFA7AE5B1CDFFE71B9C3" box="[362,463,1416,1443]" pageId="38" pageNumber="39">La Valle.</collectorName>
</materialsCitation>
<materialsCitation id="9AC8F270FD79FFA7AEE71CDEFD64B9C3" ID-GBIF-Occurrence="3986696394" box="[470,730,1416,1442]" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7AEE71CDEFDCFB9C3" bold="true" box="[470,625,1416,1442]" pageId="38" pageNumber="39">Misiones (1):</emphasis>
Posadas.
</materialsCitation>
<materialsCitation id="9AC8F270FD79FFA7ADD11CD1FC67B9C3" ID-GBIF-Occurrence="3986696553" box="[736,985,1415,1442]" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7ADD11CD1FC35B9C3" bold="true" box="[736,907,1415,1442]" pageId="38" pageNumber="39">Neuquén (14):</emphasis>
Añelo
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7ACD51CDEFB75B9C3" ID-GBIF-Occurrence="3986696414" box="[996,1227,1416,1442]" pageId="38" pageNumber="39">
<location id="2F7FAEF6FD79FFA7ACD51CDEFB75B9C3" LSID="urn:lsid:plazi:treatment:A209493BFD79FFA7AFA619C1FAA1BA1B:2F7FAEF6FD79FFA7ACD51CDEFB75B9C3" box="[996,1227,1416,1442]" country="Argentina" county="Bahia Blanca" municipality="Carmen de Patagones" name="Bajada del Marucho" pageId="38" pageNumber="39" stateProvince="Canelones">Bajada del Marucho</location>
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7ABE71CDEFA26B9C3" ID-GBIF-Occurrence="3986697056" box="[1238,1432,1416,1442]" pageId="38" pageNumber="39">
<location id="2F7FAEF6FD79FFA7ABE71CDEFA26B9C3" LSID="urn:lsid:plazi:treatment:A209493BFD79FFA7AFA619C1FAA1BA1B:2F7FAEF6FD79FFA7ABE71CDEFA26B9C3" box="[1238,1432,1416,1442]" country="Argentina" county="Bahia Blanca" municipality="Carmen de Patagones" name="Bajada del Agrio" pageId="38" pageNumber="39" stateProvince="Canelones">Bajada del Agrio</location>
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AFA61CFAFE99B9A7" ID-GBIF-Occurrence="3986696353" box="[151,295,1452,1478]" pageId="38" pageNumber="39">Collón-Curá</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AE031CFAFE25B9A7" ID-GBIF-Occurrence="3986696550" box="[306,411,1452,1478]" pageId="38" pageNumber="39">Covunco</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AE971CFAFE44B9A7" ID-GBIF-Occurrence="3986696596" box="[422,506,1452,1478]" pageId="38" pageNumber="39">Huncal</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AD341CFAFDCCB9A7" ID-GBIF-Occurrence="3986696609" box="[517,626,1452,1478]" pageId="38" pageNumber="39">
<collectorName id="87559DFBFD79FFA7AD341CFAFDCCB9A7" box="[517,626,1452,1478]" pageId="38" pageNumber="39">La Negra</collectorName>
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AD4C1CFAFD47B9A7" ID-GBIF-Occurrence="3986696726" box="[637,761,1452,1478]" pageId="38" pageNumber="39">
<collectorName id="87559DFBFD79FFA7AD4C1CFAFD47B9A7" box="[637,761,1452,1478]" pageId="38" pageNumber="39">La Pintada</collectorName>
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AC351CFAFC2CB9A7" ID-GBIF-Occurrence="3986696947" box="[772,914,1452,1478]" pageId="38" pageNumber="39">Picún Leufú</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7ACAC1CFAFBDAB9A9" ID-GBIF-Occurrence="3986696495" box="[925,1124,1452,1480]" pageId="38" pageNumber="39">
<location id="2F7FAEF6FD79FFA7ACAC1CFAFBDAB9A9" LSID="urn:lsid:plazi:treatment:A209493BFD79FFA7AFA619C1FAA1BA1B:2F7FAEF6FD79FFA7ACAC1CFAFBDAB9A9" box="[925,1124,1452,1480]" country="Argentina" county="Bahia Blanca" municipality="Carmen de Patagones" name="Piedra del Aguila" pageId="38" pageNumber="39" stateProvince="Canelones">Piedra del Águila</location>
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AB5E1CFAFB68B9A7" ID-GBIF-Occurrence="3986696777" box="[1135,1238,1452,1478]" pageId="38" pageNumber="39">Pilmatué</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7ABD01CFAFF08B98B" ID-GBIF-Occurrence="3986696506" pageId="38" pageNumber="39">Planicie Banderita</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AFF21C86FEFAB98B" ID-GBIF-Occurrence="3986696643" box="[195,324,1488,1514]" pageId="38" pageNumber="39">
<location id="2F7FAEF6FD79FFA7AFF21C86FEFAB98B" LSID="urn:lsid:plazi:treatment:A209493BFD79FFA7AFA619C1FAA1BA1B:2F7FAEF6FD79FFA7AFF21C86FEFAB98B" box="[195,324,1488,1514]" country="Argentina" county="Bahia Blanca" municipality="Carmen de Patagones" name="Rio Salado" pageId="38" pageNumber="39" stateProvince="Canelones">Río Salado</location>
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AE601C86FE47B98B" ID-GBIF-Occurrence="3986696772" box="[337,505,1488,1514]" pageId="38" pageNumber="39">Zalapa-Catuto.</materialsCitation>
<materialsCitation id="9AC8F270FD79FFA7AD301C99FCBBB98B" ID-GBIF-Occurrence="3986696548" box="[513,773,1487,1514]" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7AD301C99FD02B988" bold="true" box="[513,700,1487,1514]" pageId="38" pageNumber="39">Río Negro (15):</emphasis>
Allen
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AC231C86FCDCB98B" ID-GBIF-Occurrence="3986696832" box="[786,866,1488,1514]" pageId="38" pageNumber="39">Catriel</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AC5F1C86FC65B98B" ID-GBIF-Occurrence="3986696556" box="[878,987,1488,1514]" pageId="38" pageNumber="39">Cipolletti</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7ACD91C86FB80B98B" ID-GBIF-Occurrence="3986696699" box="[1000,1086,1488,1514]" pageId="38" pageNumber="39">Conesa</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AB7A1C86FB01B98B" ID-GBIF-Occurrence="3986697015" box="[1099,1215,1488,1514]" pageId="38" pageNumber="39">
<collectorName id="87559DFBFD79FFA7AB7A1C86FB01B98B" box="[1099,1215,1488,1514]" pageId="38" pageNumber="39">El Bolsón</collectorName>
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7ABFD1C86FA26B98B" ID-GBIF-Occurrence="3986696766" box="[1228,1432,1488,1514]" pageId="38" pageNumber="39">Fray Luís Beltrán</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AFA61CA2FEF3BA6F" ID-GBIF-Occurrence="3986696976" box="[151,333,1524,1550]" pageId="38" pageNumber="39">General Gómez</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AE6B1CA2FE46BA6F" ID-GBIF-Occurrence="3986696405" box="[346,504,1524,1550]" pageId="38" pageNumber="39">General Roca</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AD341CA2FD33BA6F" ID-GBIF-Occurrence="3986696812" box="[517,653,1524,1550]" pageId="38" pageNumber="39">Paso Flores</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7ADAB1CA2FCDBBA71" ID-GBIF-Occurrence="3986696958" box="[666,869,1524,1552]" pageId="38" pageNumber="39">
<location id="2F7FAEF6FD79FFA7ADAB1CA2FCDBBA71" LSID="urn:lsid:plazi:treatment:A209493BFD79FFA7AFA619C1FAA1BA1B:2F7FAEF6FD79FFA7ADAB1CA2FCDBBA71" box="[666,869,1524,1552]" country="Argentina" county="Bahia Blanca" municipality="Carmen de Patagones" name="Piedra del Aguila" pageId="38" pageNumber="39" stateProvince="Canelones">Piedra del Águila</location>
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AC431CA2FC53BA6F" ID-GBIF-Occurrence="3986696386" box="[882,1005,1524,1550]" pageId="38" pageNumber="39">Río Negro</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7ACC81CA2FB34BA6F" ID-GBIF-Occurrence="3986696346" box="[1017,1162,1524,1550]" pageId="38" pageNumber="39">San Antonio</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7ABA71CA2FB44BA6F" ID-GBIF-Occurrence="3986696654" box="[1174,1274,1524,1550]" pageId="38" pageNumber="39">Valcheta</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AA361CA2FA23BA6F" ID-GBIF-Occurrence="3986696740" box="[1287,1437,1524,1550]" pageId="38" pageNumber="39">Villa Regina.</materialsCitation>
<materialsCitation id="9AC8F270FD79FFA7AFA61F41FE4BBA53" ID-GBIF-Occurrence="3986696428" box="[151,501,1559,1586]" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7AFA61F41FEAABA50" bold="true" box="[151,276,1559,1585]" pageId="38" pageNumber="39">Salta (14):</emphasis>
Cafayate (desierto)
</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AD301F4EFD07BA53" ID-GBIF-Occurrence="3986696487" box="[513,697,1560,1586]" pageId="38" pageNumber="39">Coronel Olleros</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7ADF41F4EFCFBBA53" ID-GBIF-Occurrence="3986696309" box="[709,837,1560,1586]" pageId="38" pageNumber="39">Las Lajitas</materialsCitation>
;
<materialsCitation id="9AC8F270FD79FFA7AC611F4EFB01BA53" ID-GBIF-Occurrence="3986696974" box="[848,1215,1559,1586]" pageId="38" pageNumber="39">
Salta.
<emphasis id="18D4243FFD79FFA7ACA61F41FB8DBA50" bold="true" box="[919,1075,1559,1586]" pageId="38" pageNumber="39">San Luís (1):</emphasis>
Paso Patria.
</materialsCitation>
<materialsCitation id="9AC8F270FD79FFA7ABF71F41FF47BA37" ID-GBIF-Occurrence="3986696953" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7ABF71F41FAD7BA50" bold="true" box="[1222,1385,1559,1586]" pageId="38" pageNumber="39">Tucumán (1):</emphasis>
<location id="2F7FAEF6FD79FFA7AA411F4EFF47BA37" LSID="urn:lsid:plazi:treatment:A209493BFD79FFA7AFA619C1FAA1BA1B:2F7FAEF6FD79FFA7AA411F4EFF47BA37" country="Argentina" county="Bahia Blanca" municipality="Carmen de Patagones" name="Tafi de Valle. No" pageId="38" pageNumber="39" stateProvince="Canelones">Tafi de Valle.</location>
</materialsCitation>
<materialsCitation id="9AC8F270FD79FFA7AE311F6AFE33BA34" ID-GBIF-Occurrence="3986696679" box="[256,397,1595,1622]" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7AE311F6AFE9CBA37" bold="true" box="[256,290,1596,1622]" pageId="38" pageNumber="39">No</emphasis>
data (3).
</materialsCitation>
<materialsCitation id="9AC8F270FD79FFA7AEA21F6DFC0BBA37" ID-GBIF-Occurrence="3986696344" box="[403,949,1595,1622]" pageId="38" pageNumber="39">
<collectingCountry id="52B7B8BDFD79FFA7AEA21F6DFDA0BA34" box="[403,542,1595,1621]" name="Uruguay" pageId="38" pageNumber="39">URUGUAY</collectingCountry>
(1).
<collectingRegion id="E86436CFFD79FFA7AD641F6DFD6EBA34" box="[597,720,1595,1621]" country="Uruguay" name="Canelones" pageId="38" pageNumber="39">Canelones</collectingRegion>
(1): Banda Oriental
</materialsCitation>
.
</paragraph>
<paragraph id="2A1FF82DFD79FFA7AFF61F36FAA1BA1B" blockId="38.[151,1437,376,1658]" box="[199,1311,1631,1658]" pageId="38" pageNumber="39">
<emphasis id="18D4243FFD79FFA7AFF61F36FE3BBA18" bold="true" box="[199,389,1631,1658]" pageId="38" pageNumber="39">Natural history.</emphasis>
Adults are attracted to lights at night. Most records are from arid and desert areas.
</paragraph>
</subSubSection>
</treatment>
</document>
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