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<mods:title id="6E929272F2A39BFF8BDB06181A04B79E">Singapores herpetofauna: updated and annotated checklist, history, conservation, and distribution</mods:title>
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<treatment id="88502B73FE90B97CFF6B46447D5B0AD3" ID-DOI="http://doi.org/10.5281/zenodo.7968263" ID-GBIF-Taxon="209661652" ID-Zenodo-Dep="7968263" LSID="urn:lsid:plazi:treatment:88502B73FE90B97CFF6B46447D5B0AD3" httpUri="http://treatment.plazi.org/id/88502B73FE90B97CFF6B46447D5B0AD3" lastPageNumber="267" pageId="266" pageNumber="267">
<subSubSection id="48E3C9EEFE90B97CFF6B464478580E08" pageId="266" pageNumber="267" type="nomenclature">
<paragraph id="00469A65FE90B97CFF6B46447B840E64" blockId="266.[151,1437,285,1284]" box="[151,650,285,312]" pageId="266" pageNumber="267">
<heading id="5B0E2D09FE90B97CFF6B46447B840E64" bold="true" box="[151,650,285,312]" fontSize="11" level="2" pageId="266" pageNumber="267" reason="4">
<emphasis id="328D4677FE90B97CFF6B46447B840E64" bold="true" box="[151,650,285,312]" pageId="266" pageNumber="267">
<taxonomicName id="C7F9E1E6FE90B97CFF6B464478E70E6B" ID-CoL="WRWN" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[151,489,285,312]" class="Reptilia" family="Colubridae" genus="Coelognathus" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="266" pageNumber="267" phylum="Chordata" rank="species" species="subradiatus">
<emphasis id="328D4677FE90B97CFF6B464478390E6B" bold="true" box="[151,311,285,312]" italics="true" pageId="266" pageNumber="267">Coelognathus</emphasis>
cf.
<emphasis id="328D4677FE90B97CFE9E464478E70E6B" bold="true" box="[354,489,285,312]" italics="true" pageId="266" pageNumber="267">subradiatus</emphasis>
</taxonomicName>
—Erroneous.
</emphasis>
</heading>
</paragraph>
<paragraph id="00469A65FE90B97CFF6B461878580E08" blockId="266.[151,1437,285,1284]" box="[151,342,321,348]" pageId="266" pageNumber="267">Sunda Rat Snake</paragraph>
</subSubSection>
<subSubSection id="48E3C9EEFE90B97CFF6B463C78730E2C" box="[151,381,357,383]" pageId="266" pageNumber="267" type="description">
<paragraph id="00469A65FE90B97CFF6B463C78730E2C" blockId="266.[151,1437,285,1284]" box="[151,381,357,383]" pageId="266" pageNumber="267">
(
<figureCitation id="98C286E0FE90B97CFF63463C787B0E2C" box="[159,373,357,383]" captionStart="FIGURE 27" captionStartId="371.[152,255,1079,1103]" captionTargetBox="[151,1436,181,1051]" captionTargetId="figure-23@371.[151,1436,181,1051]" captionTargetPageId="371" captionText="FIGURE 27. A) Indotyphlops albiceps, B) Indotyphlops porrectus, and CD) Coelognathus cf. subradiatus. Photographs by: Evan S.H. Quah [A], Dan Rosenberg [B], and Kelvin K.P. Lim [CD]." figureDoi="http://doi.org/10.5281/zenodo.7960375" httpUri="https://zenodo.org/record/7960375/files/figure.png" pageId="266" pageNumber="267">Figure 27C &amp; 27D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="48E3C9EEFE90B97CFF6B46D078760EF0" box="[151,376,393,419]" pageId="266" pageNumber="267" type="materials_examined">
<paragraph id="00469A65FE90B97CFF6B46D078760EF0" blockId="266.[151,1437,285,1284]" box="[151,376,393,419]" pageId="266" pageNumber="267">
<emphasis id="328D4677FE90B97CFF6B46D078760EF0" bold="true" box="[151,376,393,419]" pageId="266" pageNumber="267">
<collectingCountry id="78EEDAF5FE90B97CFF6B46D0781E0EF0" box="[151,272,393,419]" name="Singapore" pageId="266" pageNumber="267">Singapore</collectingCountry>
records.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="48E3C9EEFE90B97CFF0B46F47BBC0EB8" pageId="266" pageNumber="267" type="reference_group">
<paragraph id="00469A65FE90B97CFF0B46F47DBB0E94" blockId="266.[151,1437,285,1284]" box="[247,1205,429,455]" pageId="266" pageNumber="267">
<treatmentCitationGroup id="20E9BD4BFE90B97CFF0B46F47DBB0E94" box="[247,1205,429,455]" pageId="266" pageNumber="267">
<taxonomicName id="C7F9E1E6FE90B97CFF0B46F478C60E94" authorityName="Boulenger" authorityYear="1894" box="[247,456,429,455]" class="Reptilia" family="Colubridae" genus="Coluber" kingdom="Animalia" order="Squamata" pageId="266" pageNumber="267" phylum="Chordata" rank="species" species="hodgsonii">
<emphasis id="328D4677FE90B97CFF0B46F478C60E94" box="[247,456,429,455]" italics="true" pageId="266" pageNumber="267">Coluber hodgsonii</emphasis>
</taxonomicName>
<treatmentCitation id="8158BC74FE90B97CFE1A46F47BB90E94" author="Hanitsch, R." box="[486,695,429,455]" page="10" pageId="266" pageNumber="267" year="1898">
<bibRefCitation id="6468E794FE90B97CFE1A46F47BB90E94" author="Hanitsch, R." box="[486,695,429,455]" pageId="266" pageNumber="267" pagination="11 - 22" refId="ref312325" refString="Hanitsch, R. (1898) Annual report of the curator and librarian on the Raffles Library and Museum, for the Year Ending 31 st December, 1897. In: Straits Settlements, Annual Reports for the Year 1898. Published by Authority, Singapore, pp. 11 - 22." type="book chapter" year="1898">Hanitsch, 1898: 10</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="8158BC74FE90B97CFD2546F47AA70E94" author="Flower, S. S." box="[729,937,429,455]" page="668" pageId="266" pageNumber="267" year="1899">
<bibRefCitation id="6468E794FE90B97CFD2546F47AA70E94" author="Flower, S. S." box="[729,937,429,455]" pageId="266" pageNumber="267" pagination="600 - 696" refId="ref305863" refString="Flower, S. S. (1899 b) Notes on a second collection of reptiles made in the Malay Peninsula and Siam, from November 1896 - September 1898, with a list of the species recorded from those countries. Proceedings of the Zoological Society of London, 1899 (4), 600 - 696, pls. 36 - 37." type="journal article" year="1899">Flower, 1899: 668</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="8158BC74FE90B97CFC3646F47DBF0E94" author="Hanitsch, R." box="[970,1201,429,455]" page="16" pageId="266" pageNumber="267" year="1912">
<bibRefCitation id="6468E794FE90B97CFC3646F47DBF0E94" author="Hanitsch, R." box="[970,1201,429,455]" pageId="266" pageNumber="267" refId="ref312636" refString="Hanitsch, R. (1912 b) List of the Birds, Reptiles and Amphibians in the Raffles Museum, Singapore. Raffles Library and Museum, Singapore, 19 pp." type="book" year="1912">Hanitsch, 1912b: 16</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="00469A65FE90B97CFF0B46887BBC0EB8" blockId="266.[151,1437,285,1284]" box="[247,690,465,492]" pageId="266" pageNumber="267">
<taxonomicName id="C7F9E1E6FE90B97CFF0B468878C90EB8" authorityName="Boulenger" authorityYear="1890" box="[247,455,465,491]" class="Reptilia" family="Colubridae" genus="Coluber" kingdom="Animalia" order="Squamata" pageId="266" pageNumber="267" phylum="Chordata" rank="species" species="hodgsoni">
<emphasis id="328D4677FE90B97CFF0B468878C90EB8" box="[247,455,465,491]" italics="true" pageId="266" pageNumber="267">Coluber Hodgsoni</emphasis>
</taxonomicName>
—Ridley, 1899: 208.
</paragraph>
</subSubSection>
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<paragraph id="00469A65FE90B97CFF0B46AC7B900D5C" blockId="266.[151,1437,285,1284]" box="[247,670,501,527]" pageId="266" pageNumber="267">
<taxonomicName id="C7F9E1E6FE90B97CFF0B46AC78A30D5C" baseAuthorityName="Cope" baseAuthorityYear="1861" box="[247,429,501,527]" class="Reptilia" family="Colubridae" genus="Elaphe" kingdom="Animalia" order="Squamata" pageId="266" pageNumber="267" phylum="Chordata" rank="species" species="taeniura">
<emphasis id="328D4677FE90B97CFF0B46AC78A30D5C" box="[247,429,501,527]" italics="true" pageId="266" pageNumber="267">Elaphe taeniura</emphasis>
</taxonomicName>
—Sworder, 1923: 63.
</paragraph>
</subSubSection>
<subSubSection id="48E3C9EEFE90B97CFF6B45407DCB0B8C" pageId="266" pageNumber="267" type="discussion">
<paragraph id="00469A65FE90B97CFF6B45407B750C7C" blockId="266.[151,1437,285,1284]" pageId="266" pageNumber="267">
<emphasis id="328D4677FE90B97CFF6B454078050D60" bold="true" box="[151,267,537,563]" pageId="266" pageNumber="267">Remarks.</emphasis>
A specimen in the ZRC collections (no catalogue data) identified as “
<taxonomicName id="C7F9E1E6FE90B97CFBAE45407C2E0D60" authorityName="Boulenger" authorityYear="1890" box="[1106,1312,537,563]" class="Reptilia" family="Colubridae" genus="Coluber" kingdom="Animalia" order="Squamata" pageId="266" pageNumber="267" phylum="Chordata" rank="species" species="hodgsoni">
<emphasis id="328D4677FE90B97CFBAE45407C2E0D60" box="[1106,1312,537,563]" italics="true" pageId="266" pageNumber="267">Coluber hodgsoni</emphasis>
</taxonomicName>
” was first reported by
<bibRefCitation id="6468E794FE90B97CFE8345647B3B0D04" author="Hanitsch, R." box="[383,565,573,599]" pageId="266" pageNumber="267" pagination="11 - 22" refId="ref312325" refString="Hanitsch, R. (1898) Annual report of the curator and librarian on the Raffles Library and Museum, for the Year Ending 31 st December, 1897. In: Straits Settlements, Annual Reports for the Year 1898. Published by Authority, Singapore, pp. 11 - 22." type="book chapter" year="1898">Hanitsch (1898)</bibRefCitation>
in an annual report of the museums holdings.
<bibRefCitation id="6468E794FE90B97CFBB545647DE20D04" author="Flower, S. S." box="[1097,1260,573,599]" pageId="266" pageNumber="267" pagination="600 - 696" refId="ref305863" refString="Flower, S. S. (1899 b) Notes on a second collection of reptiles made in the Malay Peninsula and Siam, from November 1896 - September 1898, with a list of the species recorded from those countries. Proceedings of the Zoological Society of London, 1899 (4), 600 - 696, pls. 36 - 37." type="journal article" year="1899">Flower (1899)</bibRefCitation>
, Ridley (1899), and
<bibRefCitation id="6468E794FE90B97CFED4453878E30D28" author="Hanitsch, R." box="[296,493,609,635]" pageId="266" pageNumber="267" refId="ref312636" refString="Hanitsch, R. (1912 b) List of the Birds, Reptiles and Amphibians in the Raffles Museum, Singapore. Raffles Library and Museum, Singapore, 19 pp." type="book" year="1912">Hanitsch (1912b)</bibRefCitation>
all refer to the same specimen. Sworder (1923) later amended the identification to
<taxonomicName id="C7F9E1E6FE90B97CFF0B45DC78A20DCC" baseAuthorityName="Cope" baseAuthorityYear="1861" box="[247,428,645,671]" class="Reptilia" family="Colubridae" genus="Elaphe" kingdom="Animalia" order="Squamata" pageId="266" pageNumber="267" phylum="Chordata" rank="species" species="taeniura">
<emphasis id="328D4677FE90B97CFF0B45DC78A20DCC" box="[247,428,645,671]" italics="true" pageId="266" pageNumber="267">Elaphe taeniura</emphasis>
</taxonomicName>
after Malcolm Smith examined the specimen and stated, “that this specimen appears to be an intermediate form between the common form and the variety (ridleyi) which inhabits caves.” Sworder (1923) also noted that the identification as “
<taxonomicName id="C7F9E1E6FE90B97CFD0F45947AB10DB4" authorityName="Boulenger" authorityYear="1890" box="[755,959,717,743]" class="Reptilia" family="Colubridae" genus="Coluber" kingdom="Animalia" order="Squamata" pageId="266" pageNumber="267" phylum="Chordata" rank="species" species="hodgsoni">
<emphasis id="328D4677FE90B97CFD0F45947AB10DB4" box="[755,959,717,743]" italics="true" pageId="266" pageNumber="267">Coluber hodgsoni</emphasis>
</taxonomicName>
” must be incorrect given that the species “is only known from a limited range, in the Himalayan region”, making “the record of a specimen from
<collectingCountry id="78EEDAF5FE90B97CFF0B444C78670C7C" box="[247,361,789,815]" name="Singapore" pageId="266" pageNumber="267">Singapore</collectingCountry>
… somewhat startling”.
</paragraph>
<paragraph id="00469A65FE90B97CFEAB44607B720BE8" blockId="266.[151,1437,285,1284]" pageId="266" pageNumber="267">
After examining specimen ZRC.2.4537, we find that it closely identifies with
<taxonomicName id="C7F9E1E6FE90B97CFAFD446078750C24" baseAuthorityName="Schlegel" baseAuthorityYear="1837" class="Reptilia" family="Colubridae" genus="Coelognathus" kingdom="Animalia" order="Squamata" pageId="266" pageNumber="267" phylum="Chordata" rank="species" species="subradiatus">
<emphasis id="328D4677FE90B97CFAFD446078750C24" italics="true" pageId="266" pageNumber="267">Coelognathus subradiatus</emphasis>
</taxonomicName>
(
<figureCitation id="98C286E0FE90B97CFE7644047B1F0C24" box="[394,529,861,887]" captionStart="FIGURE 27" captionStartId="371.[152,255,1079,1103]" captionTargetBox="[151,1436,181,1051]" captionTargetId="figure-23@371.[151,1436,181,1051]" captionTargetPageId="371" captionText="FIGURE 27. A) Indotyphlops albiceps, B) Indotyphlops porrectus, and CD) Coelognathus cf. subradiatus. Photographs by: Evan S.H. Quah [A], Dan Rosenberg [B], and Kelvin K.P. Lim [CD]." figureDoi="http://doi.org/10.5281/zenodo.7960375" httpUri="https://zenodo.org/record/7960375/files/figure.png" pageId="266" pageNumber="267">Fig. 27C, D</figureCitation>
). One of the main characters differentiating
<taxonomicName id="C7F9E1E6FE90B97CFC0644047D980C24" authorityName="Fitzinger" authorityYear="1843" box="[1018,1174,861,887]" class="Reptilia" family="Colubridae" genus="Coelognathus" kingdom="Animalia" order="Squamata" pageId="266" pageNumber="267" phylum="Chordata" rank="genus">
<emphasis id="328D4677FE90B97CFC0644047D980C24" box="[1018,1174,861,887]" italics="true" pageId="266" pageNumber="267">Coelognathus</emphasis>
</taxonomicName>
from species of
<taxonomicName id="C7F9E1E6FE90B97CFAB144047C920C24" box="[1357,1436,861,887]" class="Reptilia" family="Colubridae" genus="Elaphe" kingdom="Animalia" order="Squamata" pageId="266" pageNumber="267" phylum="Chordata" rank="genus">
<emphasis id="328D4677FE90B97CFAB144047C920C24" box="[1357,1436,861,887]" italics="true" pageId="266" pageNumber="267">Elaphe</emphasis>
</taxonomicName>
formerly classified under
<taxonomicName id="C7F9E1E6FE90B97CFDEF44D87B990CC8" authorityName="Utiger, Helfenberger, Schätti, Schmidt, Ruf &amp; Ziswiler" authorityYear="2002" box="[531,663,897,923]" class="Reptilia" family="Colubridae" genus="Orthriophis" kingdom="Animalia" order="Squamata" pageId="266" pageNumber="267" phylum="Chordata" rank="genus">
<emphasis id="328D4677FE90B97CFDEF44D87B990CC8" box="[531,663,897,923]" italics="true" pageId="266" pageNumber="267">Orthriophis</emphasis>
</taxonomicName>
, is that the last supralabial touching the eye also touches the temporals (Smith 1943). ZRC.2.4537 possesses a presubocular, which
<taxonomicName id="C7F9E1E6FE90B97CFC6444FF7D320CEC" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[920,1084,933,960]" class="Reptilia" family="Colubridae" genus="Coelognathus" kingdom="Animalia" order="Squamata" pageId="266" pageNumber="267" phylum="Chordata" rank="species" species="subradiatus">
<emphasis id="328D4677FE90B97CFC6444FF7D320CEC" box="[920,1084,933,960]" italics="true" pageId="266" pageNumber="267">C. subradiatus</emphasis>
</taxonomicName>
is the only species in the genus having this trait (Utiger
<emphasis id="328D4677FE90B97CFDF444937B4F0CB0" box="[520,577,969,995]" italics="true" pageId="266" pageNumber="267">et al.</emphasis>
2005). Diagnosis of ZRC.2.4537 is as follows: snout-vent-length 125.0 cm; tail length 37.0 cm; 230 ventrals; 111 subcaudals; anal plate divided; dorsals 27:25:19; 1 loreal; 1 preocular; 2 postoculars; 1 presubocular present; 8 supralabials (IV and V touching the eye, and V touching the first two temporals); 9 infralabials (IIIV touching the anterior chin shield); 2+3+4 temporals; dorsal dark brown above with irregular light spots; venter yellowish anteriorly with brown mottling transitioning to uniform dark brown posteriorly.
<taxonomicName id="C7F9E1E6FE90B97CFDF443247A210BC4" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[520,815,1149,1175]" class="Reptilia" family="Colubridae" genus="Coelognathus" kingdom="Animalia" order="Squamata" pageId="266" pageNumber="267" phylum="Chordata" rank="species" species="subradiatus">
<emphasis id="328D4677FE90B97CFDF443247A210BC4" box="[520,815,1149,1175]" italics="true" pageId="266" pageNumber="267">Coelognathus subradiatus</emphasis>
</taxonomicName>
is native to the
<collectingRegion id="C23D5487FE90B97CFC1443277DD90BC4" box="[1000,1239,1149,1176]" country="Indonesia" name="Nusa Tenggara Timur" pageId="266" pageNumber="267">Lesser Sunda Islands</collectingRegion>
of
<collectingCountry id="78EEDAF5FE90B97CFB0343247C620BC4" box="[1279,1388,1149,1175]" name="Indonesia" pageId="266" pageNumber="267">Indonesia</collectingCountry>
and
<collectingCountry id="78EEDAF5FE90B97CFF0B43F8788F0BE8" box="[247,385,1185,1211]" name="Timor-Leste" pageId="266" pageNumber="267">Timor-Leste</collectingCountry>
(Wallach
<emphasis id="328D4677FE90B97CFE0843FB7B220BE8" box="[500,556,1185,1211]" italics="true" pageId="266" pageNumber="267">et al.</emphasis>
2014).
</paragraph>
<paragraph id="00469A65FE90B97CFF6B439C7DCB0B8C" blockId="266.[151,1437,285,1284]" box="[151,1221,1221,1248]" pageId="266" pageNumber="267">
<emphasis id="328D4677FE90B97CFF6B439C7ACF0B8C" bold="true" box="[151,961,1221,1247]" pageId="266" pageNumber="267">
LKCNHM &amp; NHMUK Museum specimens.
<collectingCountry id="78EEDAF5FE90B97CFD56439C7A2D0B8C" box="[682,803,1221,1247]" name="Singapore" pageId="266" pageNumber="267">Singapore</collectingCountry>
(no locality):
</emphasis>
ZRC.2.4537 (no date).
</paragraph>
</subSubSection>
<subSubSection id="48E3C9EEFE90B97CFF6B43B07D5B0AD3" pageId="266" pageNumber="267" type="materials_examined">
<paragraph id="00469A65FE90B97CFF6B43B07A390A50" blockId="266.[151,1437,285,1284]" box="[151,823,1257,1284]" pageId="266" pageNumber="267">
<emphasis id="328D4677FE90B97CFF6B43B07B840A50" bold="true" box="[151,650,1257,1283]" pageId="266" pageNumber="267">
Additional
<collectingCountry id="78EEDAF5FE90B97CFEE243B078990A50" box="[286,407,1257,1283]" name="Singapore" pageId="266" pageNumber="267">Singapore</collectingCountry>
museum specimens.
</emphasis>
No specimens.
</paragraph>
<paragraph id="00469A65FE90B97CFE23423F7D5B0AD3" blockId="266.[479,1109,1382,1409]" box="[479,1109,1382,1409]" pageId="266" pageNumber="267">
<emphasis id="328D4677FE90B97CFE23423F7D5B0AD3" bold="true" box="[479,1109,1382,1409]" pageId="266" pageNumber="267">
Genus
<taxonomicName id="C7F9E1E6FE90B97CFDCA423F7A0B0AD3" authority="Fitzinger" authorityName="Fitzinger" box="[566,773,1382,1409]" class="Reptilia" family="Colubridae" genus="Elaphe" kingdom="Animalia" order="Squamata" pageId="266" pageNumber="267" phylum="Chordata" rank="genus">
<emphasis id="328D4677FE90B97CFDCA423F7B820AD2" bold="true" box="[566,652,1382,1409]" italics="true" pageId="266" pageNumber="267">Elaphe</emphasis>
Fitzinger
</taxonomicName>
in Wagler, 1833 (1 species)
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>