treatments-xml/data/86/E0/42/86E042B52E1B53F3A40D7320B7148660.xml
2024-06-21 12:42:39 +02:00

251 lines
21 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://dx.doi.org/10.3897/dez.68.60193" ID-GBIF-Dataset="416195dc-df47-4f90-abbd-45eca9a23796" ID-GBIF-Taxon="182223388" ID-Pensoft-Pub="1860-1324-1-45" ID-Pensoft-UUID="EE57E5591FE3500892459D0CF4FBB496" ID-Zenodo-Dep="4435456" ID-ZooBank="227B739469C5447FB984250DF1ADFBE1" LSID="urn:lsid:zoobank.org:act:EEE36B7E-C218-44AB-A00B-2DEA756A6460" ModsDocID="1860-1324-68-1-45" checkinTime="1621372298573" checkinUser="pensoft" docAuthor="Hemp, Claudia &amp; Massa, Bruno" docDate="2021" docId="86E042B52E1B53F3A40D7320B7148660" docLanguage="en" docName="DeutEntomolZeit 68(1): 45-79" docOrigin="Deutsche Entomologische Zeitschrift 68 (1)" docPubDate="2021-01-05" docSource="http://dx.doi.org/10.3897/dez.68.60193" docTitle="Poreuomena tshuapa Hemp &amp; Massa 2021, sp. nov." docType="treatment" docUuid="EEE36B7E-C218-44AB-A00B-2DEA756A6460" docUuidSource="ZooBank" docVersion="3" id="EE57E5591FE3500892459D0CF4FBB496" lastPageNumber="45" masterDocId="EE57E5591FE3500892459D0CF4FBB496" masterDocTitle="Biogeographical and evolutionary aspects of a Guineo-Congolian bushcricket tribe: Revision of the genera Cestromoecha Karsch, 1893 and Poreuomena Brunner von Wattenwyl, 1878, with the description of new species (Orthoptera, Tettigoniidae, Phaneropterinae)" masterLastPageNumber="79" masterPageNumber="45" pageId="0" pageNumber="45" updateTime="1632023681013" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Biogeographical and evolutionary aspects of a Guineo-Congolian bushcricket tribe: Revision of the genera Cestromoecha Karsch, 1893 and Poreuomena Brunner von Wattenwyl, 1878, with the description of new species (Orthoptera, Tettigoniidae, Phaneropterinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hemp, Claudia</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9170-7113</mods:nameIdentifier>
<mods:affiliation>Dept Plant Systematics, Univ. of Bayreuth, Germany</mods:affiliation>
<mods:nameIdentifier type="email">claudia.hemp@uni-bayreuth.de</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Massa, Bruno</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2127-0715</mods:nameIdentifier>
<mods:affiliation>Dept of Agriculture, Food and Forest Sciences, University of Palermo, Italy</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-01-05</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>68</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>45</mods:start>
<mods:end>79</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/dez.68.60193</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.68.60193</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-1-45</mods:identifier>
<mods:identifier type="ZooBank">227B739469C5447FB984250DF1ADFBE1</mods:identifier>
<mods:identifier type="Pensoft-UUID">EE57E5591FE3500892459D0CF4FBB496</mods:identifier>
<mods:identifier type="Zenodo-Dep">4435456</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182223388" LSID="urn:lsid:zoobank.org:act:EEE36B7E-C218-44AB-A00B-2DEA756A6460" httpUri="http://treatment.plazi.org/id/86E042B52E1B53F3A40D7320B7148660" lastPageNumber="45" pageId="0" pageNumber="45">
<subSubSection pageId="0" pageNumber="45" type="nomenclature">
<paragraph pageId="0" pageNumber="45">
<taxonomicName LSID="http://zoobank.org/EEE36B7E-C218-44AB-A00B-2DEA756A6460" authority="Hemp &amp; Massa, 2021" authorityName="Hemp &amp; Massa" authorityYear="2021" class="Insecta" family="Tettigoniidae" genus="Poreuomena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Poreuomena tshuapa" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="tshuapa" status="sp. nov.">Poreuomena tshuapa</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="45">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 97100" captionStartId="F19" captionText="Figures 97 - 100. Morphological details of male Poreuomena tshuapa sp. nov. Habitus (97), stridulatory file on the underside of the left tegminal flap (98), different views on apex with cerci (99, 100)." figureDoi="10.3897/dez.68.60193.figures97-100" httpUri="https://binary.pensoft.net/fig/499198" pageId="0" pageNumber="45">Figs 97-100</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="materials_examined">
<paragraph pageId="0" pageNumber="45">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="45">
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="M. Dupuis" country="Democratic Republic of the Congo" location="Tshuapa" municipality="Bokungu" specimenCount="♂" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="45">
<collectingCountry name="Democratic Republic of the Congo">Democratic Republic of the Congo</collectingCountry>
</emphasis>
,
<location LSID="urn:lsid:plazi:treatment:86E042B52E1B53F3A40D7320B7148660:0FC9372207BAE4D682F50691346E26CD" country="Democratic Republic of the Congo" municipality="Bokungu" name="Tshuapa">Tshuapa</location>
,
<collectingMunicipality>Bokungu</collectingMunicipality>
1949,
<collectorName>M. Dupuis</collectorName>
(
<typeStatus>Holotype</typeStatus>
<specimenCount></specimenCount>
)
</materialsCitation>
;
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="M. Dupuis" country="Democratic Republic of the Congo" location="Tshuapa" municipality="Bokungu" specimenCount="♂" typeStatus="Paratype">
same data as holotype (
<typeStatus>Paratype</typeStatus>
<specimenCount></specimenCount>
)
</materialsCitation>
;
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Ghesquiere" country="Democratic Republic of the Congo" location="Haut-Lopori" specimenCount="♂" typeStatus="Paratype">
<emphasis bold="true" pageId="0" pageNumber="45">
<collectingCountry name="Democratic Republic of the Congo">Democratic Republic of the Congo</collectingCountry>
</emphasis>
,
<location LSID="urn:lsid:plazi:treatment:86E042B52E1B53F3A40D7320B7148660:0031DA9DFA520CDE7D0306C76F3EE638" country="Democratic Republic of the Congo" name="Haut-Lopori">Haut-Lopori</location>
V-VI 1927,
<collectorName>
J.
<normalizedToken originalValue="Ghesquiére">Ghesquiere</normalizedToken>
</collectorName>
(
<typeStatus>Paratype</typeStatus>
<specimenCount></specimenCount>
)
</materialsCitation>
(MRAC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="diagnosis">
<paragraph pageId="0" pageNumber="45">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="45">
Similar in the outer male genitalic apparatus to
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. magnicerca" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="magnicerca">
<emphasis italics="true" pageId="0" pageNumber="45">P. magnicerca</emphasis>
</taxonomicName>
from further west in the Congo Basin. Differentiated from
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. forcipata" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="forcipata">
<emphasis italics="true" pageId="0" pageNumber="45">P. forcipata</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. laeglae" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="laeglae">
<emphasis italics="true" pageId="0" pageNumber="45">P. laeglae</emphasis>
</taxonomicName>
by a different 10th abdominal tergite which is clearly divided into two lobes in
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. forcipata" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="forcipata">
<emphasis italics="true" pageId="0" pageNumber="45">P. forcipata</emphasis>
</taxonomicName>
, while
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. tshuapa" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="tshuapa">
<emphasis italics="true" pageId="0" pageNumber="45">P. tshuapa</emphasis>
</taxonomicName>
sp. nov. only has small humps and the posterior margin of the 10th abdominal tergite of
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. laeglae" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="laeglae">
<emphasis italics="true" pageId="0" pageNumber="45">P. laeglae</emphasis>
</taxonomicName>
is more-or-less straight.
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. forcipata" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="forcipata">
<emphasis italics="true" pageId="0" pageNumber="45">P. forcipata</emphasis>
</taxonomicName>
has male cerci differentiated into two branches, an outer rather blunt, finger-like part and an inner blade-like expanded branch with an acute tip, similar to the inner branch of
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. tshuapa" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="tshuapa">
<emphasis italics="true" pageId="0" pageNumber="45">P. tshuapa</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. magnicerca" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="magnicerca">
<emphasis italics="true" pageId="0" pageNumber="45">P. magnicerca</emphasis>
</taxonomicName>
. However,
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. tshuapa" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="tshuapa">
<emphasis italics="true" pageId="0" pageNumber="45">P. tshuapa</emphasis>
</taxonomicName>
sp. nov. has an additional branch just below the finger-like part of the cerci, absent in
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. forcipata" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="forcipata">
<emphasis italics="true" pageId="0" pageNumber="45">P. forcipata</emphasis>
</taxonomicName>
, but also present in
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. magnicerca" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="magnicerca">
<emphasis italics="true" pageId="0" pageNumber="45">P. magnicerca</emphasis>
</taxonomicName>
(also see diagnosis at
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. laeglae" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="laeglae">
<emphasis italics="true" pageId="0" pageNumber="45">P. laeglae</emphasis>
</taxonomicName>
). In
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. magnicerca" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="magnicerca">
<emphasis italics="true" pageId="0" pageNumber="45">P. magnicerca</emphasis>
</taxonomicName>
, the blade-like subapical section or the
<normalizedToken originalValue="“third”">&quot;third&quot;</normalizedToken>
branch is more roundish and larger than in
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. tshuapa" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="tshuapa">
<emphasis italics="true" pageId="0" pageNumber="45">P. tshuapa</emphasis>
</taxonomicName>
sp. nov. and the subgenital plate differs between these two species. In
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. tshuapa" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="tshuapa">
<emphasis italics="true" pageId="0" pageNumber="45">P. tshuapa</emphasis>
</taxonomicName>
sp. nov., the subgenital plate is bilobate with the processes clearly separated, while in
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. magnicerca" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="magnicerca">
<emphasis italics="true" pageId="0" pageNumber="45">P. magnicerca</emphasis>
</taxonomicName>
, the lobes of the subgenital plate are shorter and situated more closely to each other. Further, in
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. forcipata" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="forcipata">
<emphasis italics="true" pageId="0" pageNumber="45">P. forcipata</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. laeglae" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="laeglae">
<emphasis italics="true" pageId="0" pageNumber="45">P. laeglae</emphasis>
</taxonomicName>
, asymmetrical spines are present on the left side of the supra-anal plates.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">Description.</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Male.</emphasis>
Probably predominantly green when alive, preserved specimens of tawny colour (Fig.
<figureCitation captionStart="Figures 97100" captionStartId="F19" captionText="Figures 97 - 100. Morphological details of male Poreuomena tshuapa sp. nov. Habitus (97), stridulatory file on the underside of the left tegminal flap (98), different views on apex with cerci (99, 100)." figureDoi="10.3897/dez.68.60193.figures97-100" httpUri="https://binary.pensoft.net/fig/499198" pageId="0" pageNumber="45">97</figureCitation>
). Where tegmina meet when folded, interior part of cells of dark colour while surrounding and elevated veins green/tawny. Typical
<taxonomicName authorityName="Brunner von Wattenwyl" authorityYear="1878" class="Insecta" family="Tettigoniidae" genus="Poreuomena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Poreuomena" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Poreuomena</emphasis>
</taxonomicName>
species with Rs branching off from the radius near the base. The tegmina have two typical well-developed flaps at their base, the flap on the left side smaller and more pointed than the flap on the right tegmen. Beneath the flaps, tegmina with narrow longish brown markings are typical for several
<taxonomicName authorityName="Brunner von Wattenwyl" authorityYear="1878" class="Insecta" family="Tettigoniidae" genus="Poreuomena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Poreuomena" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Poreuomena</emphasis>
</taxonomicName>
species. The stridulatory file on the left tegminal flap is about 1.5 mm long with small broadly-spaced inner teeth becoming gradually larger and are more densely set (Fig.
<figureCitation captionStart="Figures 97100" captionStartId="F19" captionText="Figures 97 - 100. Morphological details of male Poreuomena tshuapa sp. nov. Habitus (97), stridulatory file on the underside of the left tegminal flap (98), different views on apex with cerci (99, 100)." figureDoi="10.3897/dez.68.60193.figures97-100" httpUri="https://binary.pensoft.net/fig/499198" pageId="0" pageNumber="45">98</figureCitation>
). The last third consists of very densely packed teeth becoming smaller again; the apical part is strongly curved. All-in-all, the stridulatory file consists of about 70-80 teeth. Stridulatory file on the underside of the right tegmen not as well-developed, but similar in shape and size and arrangement of the teeth. The 10th abdominal tergite has almost a straight posterior margin with only two bump-like structures laterally (Fig.
<figureCitation captionStart="Figures 97100" captionStartId="F19" captionText="Figures 97 - 100. Morphological details of male Poreuomena tshuapa sp. nov. Habitus (97), stridulatory file on the underside of the left tegminal flap (98), different views on apex with cerci (99, 100)." figureDoi="10.3897/dez.68.60193.figures97-100" httpUri="https://binary.pensoft.net/fig/499198" pageId="0" pageNumber="45">99</figureCitation>
). The cerci are stout, divided at their apical part into three branches: a subapical blade located just beneath the second elongated, finger-like branch and an inner blade-like or leaf-like expanded branch with sclerotised margins (Figs
<figureCitation captionStart="Figures 97100" captionStartId="F19" captionText="Figures 97 - 100. Morphological details of male Poreuomena tshuapa sp. nov. Habitus (97), stridulatory file on the underside of the left tegminal flap (98), different views on apex with cerci (99, 100)." figureDoi="10.3897/dez.68.60193.figures97-100" httpUri="https://binary.pensoft.net/fig/499198" pageId="0" pageNumber="45">99</figureCitation>
,
<figureCitation captionStart="Figures 97100" captionStartId="F19" captionText="Figures 97 - 100. Morphological details of male Poreuomena tshuapa sp. nov. Habitus (97), stridulatory file on the underside of the left tegminal flap (98), different views on apex with cerci (99, 100)." figureDoi="10.3897/dez.68.60193.figures97-100" httpUri="https://binary.pensoft.net/fig/499198" pageId="0" pageNumber="45">100</figureCitation>
). The subgenital plate is elongated and deeply divided into two lobes; without styli (Fig.
<figureCitation captionStart="Figures 97100" captionStartId="F19" captionText="Figures 97 - 100. Morphological details of male Poreuomena tshuapa sp. nov. Habitus (97), stridulatory file on the underside of the left tegminal flap (98), different views on apex with cerci (99, 100)." figureDoi="10.3897/dez.68.60193.figures97-100" httpUri="https://binary.pensoft.net/fig/499198" pageId="0" pageNumber="45">100</figureCitation>
). Between the subgenital plate and the cerci, two longish internal structures are present (titillators).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">Measurements</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">(mm)</emphasis>
. Males (n = 3). Body length: 17.7-18.2; pronotum length: 3.8-4.1; length hind femora: 20.3-21.2; length of tegmina 29.6-30.4; width of tegmina: 4.6-5.0.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="distribution">
<paragraph pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="45">Democratic Republic of the Congo, Tshuapa Province.</paragraph>
<caption ID-Zenodo-Dep="4435480" doi="10.3897/dez.68.60193.figures97-100" httpUri="https://binary.pensoft.net/fig/499198" pageId="0" pageNumber="45" start="Figures 97100" startId="F19">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figures 97-100.</emphasis>
Morphological details of male
<taxonomicName authorityName="Hemp &amp; Massa" authorityYear="2021" class="Insecta" family="Tettigoniidae" genus="Poreuomena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Poreuomena tshuapa" order="Orthoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="tshuapa">
<emphasis italics="true" pageId="0" pageNumber="45">Poreuomena tshuapa</emphasis>
</taxonomicName>
sp. nov. Habitus (
<emphasis bold="true" pageId="0" pageNumber="45">97</emphasis>
), stridulatory file on the underside of the left tegminal flap (
<emphasis bold="true" pageId="0" pageNumber="45">98</emphasis>
), different views on apex with cerci (
<emphasis bold="true" pageId="0" pageNumber="45">99, 100</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>