150 lines
14 KiB
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150 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.443.7880" ID-GBIF-Dataset="33ef48a3-7145-4103-9159-e11be2ba9adc" ID-PMC="PMC4205504" ID-Pensoft-Pub="1313-2970-443-61" ID-PubMed="25349497" ID-ZBK="E33841396A6E426C840D85BC32A12E78" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-443-61" ModsDocOrigin="ZooKeys 443" ModsDocTitle="The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation" checkinTime="1451245273153" checkinUser="pensoft" docAuthor="Sokolov, Igor M. & Kavanaugh, David H." docDate="2014" docId="801363B8A681AC9505AB437D69FCF25A" docLanguage="en" docName="ZooKeys 443: 61-118" docOrigin="ZooKeys 443" docSource="http://dx.doi.org/10.3897/zookeys.443.7880" docTitle="Geocharidius gimlii Erwin" docType="treatment" docVersion="3" lastPageNumber="77" masterDocId="3546FFA8CF78F064801706227F76A615" masterDocTitle="The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation" masterLastPageNumber="118" masterPageNumber="61" pageNumber="76" updateTime="1668159453511" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sokolov, Igor M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kavanaugh, David H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>443</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>61</mods:start>
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<mods:end>118</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.443.7880</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.443.7880</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-443-61</mods:identifier>
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<mods:identifier type="ZBK">E33841396A6E426C840D85BC32A12E78</mods:identifier>
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<mods:identifier type="ZooBank">E33841396A6E426C840D85BC32A12E78</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152055692" LSID="urn:lsid:plazi:treatment:801363B8A681AC9505AB437D69FCF25A" httpUri="http://treatment.plazi.org/id/801363B8A681AC9505AB437D69FCF25A" lastPageId="16" lastPageNumber="77" pageId="15" pageNumber="76">
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<subSubSection pageId="15" pageNumber="76" type="multiple">
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<paragraph pageId="15" pageNumber="76">Taxon classification Animalia Coleoptera Carabidae</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="76" type="nomenclature">
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<paragraph pageId="15" pageNumber="76">
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<taxonomicName authority="Erwin" class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius gimlii" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="species" species="gimlii">Geocharidius gimlii Erwin</taxonomicName>
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Figs 12A, 13
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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, 14A, 15B, 16A, 22
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="77" type="reference_group">
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<paragraph pageId="16" pageNumber="77">
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius gimlii" order="Coleoptera" pageId="16" pageNumber="77" phylum="Arthropoda" rank="species" species="gimlii">
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<pageBreakToken pageId="16" pageNumber="77" start="start">Geocharidius</pageBreakToken>
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gimlii
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</taxonomicName>
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Erwin, 1982: 488.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="77" type="holotype">
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<paragraph pageId="16" pageNumber="77">Holotype.</paragraph>
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<paragraph pageId="16" pageNumber="77">A male, deposited in NMNH, point-mounted, dissected, labeled (Fig. 14A): \ ♂ -wing \ ADP 26556 \ GUATEMALA: Hue. 7.7km S SanJuan Ixcoy, 2780 MS 15 35'N, 091 27'W 11 August 1974 \ G. E. Ball, H. Frania, D.R. Whitehead Colls. leaf litter \. Type locality. Guatemala, Huehuetenango Department, 7.7 km S of San Juan Ixcoy.</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="77" type="recognition">
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<paragraph pageId="16" pageNumber="77">Recognition.</paragraph>
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<paragraph pageId="16" pageNumber="77">Males of this species are distinguished from those of other members of the integripennis species group by the following combination of characters: pronotum small, transverse, elytra comparatively wide and structure of median lobe of male as in Fig. 13A.</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="77" type="description">
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<paragraph pageId="16" pageNumber="77">Description.</paragraph>
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<paragraph pageId="16" pageNumber="77">Size. Medium for genus (SBL 1.42 mm).</paragraph>
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<paragraph pageId="16" pageNumber="77">Habitus. Body form (Fig. 12A) moderately convex, broadly ovoid; general proportions (WE/SBL 0.41) rather wide; proportions of head (WH/WPm 0.72) average for group; pronotum narrow (WPm/WE 0.74) relative to elytra.</paragraph>
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<paragraph pageId="16" pageNumber="77">Color. Body rufotestaceous, appendages testaceous.</paragraph>
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<paragraph pageId="16" pageNumber="77">Microsculpture. Mesh pattern of irregularly isodiametric sculpticells present over all dorsal surfaces of head and elytra. Pronotum smooth (without evident microsculpture). Proepisternum with microsculpture.</paragraph>
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<paragraph pageId="16" pageNumber="77">Prothorax. Pronotum transverse (WPm/LP 1.32), with lateral margins markedly constricted posteriorly (WPm/WPp 1.38). Posterior angles obtuse (112°). Widths between anterior and posterior angles equal (WPa/WPp 1.01).</paragraph>
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<paragraph pageId="16" pageNumber="77">Elytra. Moderately convex, slightly depressed along suture, markedly wide (WE/LE 0.70), without traces of striae. Humeri broadly rounded, in outline forming slightly obtuse angle with longitudinal axis of body. Lateral margins convex, evenly divergent at basal half, evenly rounded to apex in apical half.</paragraph>
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<paragraph pageId="16" pageNumber="77">Male genitalia. Median lobe of aedeagus (Fig. 13A, 15B) with shaft long, widened apically, and apex small and acutely rounded. Ventral margin straight. Dorsal sclerites of internal sac in form of very long fig, protruding from basal orifice, and tapered apically in rather long flagellum, abruptly widened basally as a semicircular dilation, bent ventrally and surrounded by complex of semisclerotized sheaths of peculiar shape. Right paramere with long, narrow apical constriction (Fig. 13C). Left paramere with long, narrow and curved apical constriction (Fig. 13B). Ring sclerite of triangular shape, with sinuations on both sides of long basal handle (Fig. 16A).</paragraph>
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<paragraph pageId="16" pageNumber="77">Female internal genitalia. Females unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="77" type="geographical distribution">
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<paragraph pageId="16" pageNumber="77">Geographical distribution.</paragraph>
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<paragraph pageId="16" pageNumber="77">This species is known only from the type locality in the mountains of the Sierra de los Cuchumatanes, located in the Huehuetenango Department of Guatemala (Fig. 22, white triangle).</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="77" type="way of life">
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<paragraph pageId="16" pageNumber="77">Way of life.</paragraph>
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<paragraph pageId="16" pageNumber="77">
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The unique type specimen was sifted from leaf litter in Lower Montane Wet Forest (
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<bibRefCitation author="Erwin, TL" journalOrPublisher="Proceedings of the California Academy of Sciences" pageId="32" pageNumber="93" pagination="455 - 496" title="Small terrestrial ground-beetles of Central America (Carabidae: Bembidiina and Anillina)." url="http://biostor.org/reference/3321" volume="42" year="1982">Erwin 1982</bibRefCitation>
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) at an elevation of 2780 m.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="77" type="relationships">
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<paragraph pageId="16" pageNumber="77">Relationships.</paragraph>
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<paragraph pageId="16" pageNumber="77">
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The shape of the median lobe in the holotype of
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius gimlii" order="Coleoptera" pageId="16" pageNumber="77" phylum="Arthropoda" rank="species" species="gimlii">Geocharidius gimlii</taxonomicName>
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(Fig. 15B) is almost identical to that of the male holotype of
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius integripennis" order="Coleoptera" pageId="16" pageNumber="77" phylum="Arthropoda" rank="species" species="integripennis">Geocharidius integripennis</taxonomicName>
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(Fig. 15A); hence, at least for now, the latter can be considered as its closest relative. The general shape of the dorsal sclerites of the internal sac (namely the apically tapered fig, widened and ventrally bent at the basal end) is also similar to that in G andersoni (Fig. 9H) males described above.
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</paragraph>
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<caption pageId="16" pageNumber="77">
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<paragraph pageId="16" pageNumber="77">
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Figures 15. Schematic line drawings of median lobe and dorsal sclerites of internal sac of two previously described Guatemalan
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius" order="Coleoptera" pageId="16" pageNumber="77" phylum="Arthropoda" rank="genus">Geocharidius</taxonomicName>
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species. A
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius integripennis" order="Coleoptera" pageId="16" pageNumber="77" phylum="Arthropoda" rank="species" species="integripennis">Geocharidius integripennis</taxonomicName>
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(GUATEMALA,
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<normalizedToken originalValue="Totonicapán">Totonicapan</normalizedToken>
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,
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<normalizedToken originalValue="“Totonicapam”">"Totonicapam"</normalizedToken>
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), holotype B
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius gimlii" order="Coleoptera" pageId="16" pageNumber="77" phylum="Arthropoda" rank="species" species="gimlii">Geocharidius gimlii</taxonomicName>
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(GUATEMALA, Huehuetenango, San Juan Ixcoy), holotype, both left lateral aspect.
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</paragraph>
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</caption>
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<caption pageId="16" pageNumber="77">
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<paragraph pageId="16" pageNumber="77">
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Figures 16. Line drawings of ring sclerite of Guatemalan
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius" order="Coleoptera" pageId="16" pageNumber="77" phylum="Arthropoda" rank="genus">Geocharidius</taxonomicName>
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species, male genitalia, dorsal aspect. A
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius gimlii" order="Coleoptera" pageId="16" pageNumber="77" phylum="Arthropoda" rank="species" species="gimlii">Geocharidius gimlii</taxonomicName>
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(GUATEMALA, Huehuetenango, San Juan Ixcoy), holotype B
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius erwini" order="Coleoptera" pageId="16" pageNumber="77" phylum="Arthropoda" rank="species" species="erwini">Geocharidius erwini</taxonomicName>
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(GUATEMALA,
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<normalizedToken originalValue="Quiché">Quiche</normalizedToken>
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, Los Encuentros), paratype C
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius minimus" order="Coleoptera" pageId="16" pageNumber="77" phylum="Arthropoda" rank="species" species="minimus">Geocharidius minimus</taxonomicName>
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(GUATEMALA,
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<normalizedToken originalValue="Sacatepéquez">Sacatepequez</normalizedToken>
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, Antigua), paratype D
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius jalapensis" order="Coleoptera" pageId="16" pageNumber="77" phylum="Arthropoda" rank="species" species="jalapensis">Geocharidius jalapensis</taxonomicName>
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(GUATEMALA, Jalapa, Mataquescuintla), paratype E
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius balini" order="Coleoptera" pageId="16" pageNumber="77" phylum="Arthropoda" rank="species" species="balini">Geocharidius balini</taxonomicName>
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(GUATEMALA,
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<normalizedToken originalValue="Suchitepéquez">Suchitepequez</normalizedToken>
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, Volcano
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<normalizedToken originalValue="Atitlán">Atitlan</normalizedToken>
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), paratype F
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius longinoi" order="Coleoptera" pageId="16" pageNumber="77" phylum="Arthropoda" rank="species" species="longinoi">Geocharidius longinoi</taxonomicName>
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(GUATEMALA, El Progreso, Cerro
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<normalizedToken originalValue="Pinalón">Pinalon</normalizedToken>
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), paratype. Scale = 0.1mm.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |