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<document id="893A8BEBF3794D5A43E71D66AFB27261" ID-DOI="10.1051/acarologia/20162188" ID-ISSN="2107-7207" ID-Zenodo-Dep="5404290" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630342953770" checkinUser="marcus" docAuthor="Ács, A., Sutak, A. &amp; Kontschan, J." docDate="2016" docId="77605D0E35086E1EFE80FCDFFDFFF9A8" docLanguage="en" docName="Acarologia.2016.56.1.63-71.pdf" docOrigin="Acarologia 56 (1)" docSource="http://dx.doi.org/10.1051/acarologia/20162188" docStyle="DocumentStyle:8E0DCBB23C2CB3FD20935B7794B24F85.3:Acarologia.2010-2017.journal_article" docStyleId="8E0DCBB23C2CB3FD20935B7794B24F85" docStyleName="Acarologia.2010-2017.journal_article" docStyleVersion="3" docTitle="Neopodocinum longisetum Ács &amp; Sutak &amp; Kontschan 2016, n. sp." docType="treatment" docVersion="3" lastPageNumber="68" masterDocId="8B592576350D6E18FF98FF92FF8CFFA4" masterDocTitle="New records of macrochelid mites and description of a new phoretic species (Acari: Mesostigmata: Macrochelidae) from Greece" masterLastPageNumber="71" masterPageNumber="63" pageNumber="67" updateTime="1699236263772" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-ND-4.0">
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<mods:title id="0D665D9F8538E3AF8F3551BF779F8142">New records of macrochelid mites and description of a new phoretic species (Acari: Mesostigmata: Macrochelidae) from Greece</mods:title>
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<taxonomicName id="38C9979B35086E1DFE80FCDFFDB4FCC1" authority="Ács &amp; Sutak &amp; Kontschan, 2016" authorityName="Ács &amp; Sutak &amp; Kontschan" authorityYear="2016" box="[280,568,845,869]" class="Arachnida" family="Macrochelidae" genus="Neopodocinum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="longisetum" status="sp. nov.">
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(
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)
</paragraph>
<paragraph id="FF76EC1835086E1DFF07FC32FD76FB0A" blockId="5.[159,762,845,1948]" pageId="5" pageNumber="67">Diagnosis — Dorsal shield bears long, slightly bent setae marginally and short needle-like setae on median part. Surface of dorsum with 11 pairs of pores and with 8 pairs of microspicules. Posterior margin of sternal shield with a pair of spur-like structure. Anal shield small, egg-shaped with short post-anal seta and a pair of para-anal setae. Tectum typical for the genus, unipartite with small denticles along.</paragraph>
<paragraph id="FF76EC1835086E1DFF5DFB52FE66F993" blockId="5.[159,762,845,1948]" pageId="5" pageNumber="67">
<materialsCitation id="4FA1E64535086E1DFF5DFB52FE6BF993" collectingDate="2011-05-05" collectionCode="T" collectorName="Louros, W &amp; Szederjesi &amp; Zs. &amp; The &amp; Soil Zoology Collections &amp; Hungarian Natural History Museum &amp; Budapest" country="Greece" county="Material" elevation="130" latitude="39.2171" location="Ano Kotsanopoulo" longLatPrecision="1" longitude="20.713717" municipality="Preveza" pageId="5" pageNumber="67" specimenCount="1" specimenCount-female="1" stateProvince="Ipeiros" typeStatus="holotype">
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examined —
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. Female. Collected from
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<emphasis id="CDBD300A35086E1DFED8FB71FD98FB5F" box="[320,532,1251,1275]" italics="true" pageId="5" pageNumber="67">Oryctes nasicornis</emphasis>
(
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)
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,
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,
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,
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peripheral unit,
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, garden of a cafe bar along the road towards
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of the village,
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a.s.l.
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,
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,
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. KontschAEn, J., MurAEnyi, D.,
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,
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. and UjvAEri,
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coll.
<typeStatus id="207252BA35086E1DFF07FA24FE9FFA6A" box="[159,275,1462,1486]" pageId="5" pageNumber="67" type="paratype">Paratypes</typeStatus>
<specimenCount id="E9CF279135086E1DFE82FA24FE26FA69" box="[282,426,1461,1485]" pageId="5" pageNumber="67" type="female">two females</specimenCount>
, locality, date and host same as in
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.
<collectorName id="523C89CE35086E1DFEC4FA4AFE04FA54" box="[348,392,1496,1520]" pageId="5" pageNumber="67">The</collectorName>
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was deposited in the
<collectorName id="523C89CE35086E1DFF07FA69FE4DF9B7" box="[159,449,1531,1555]" pageId="5" pageNumber="67">Soil Zoology Collections</collectorName>
of the
<collectorName id="523C89CE35086E1DFD8DFA69FEE6F992" pageId="5" pageNumber="67">Hungarian Natural History Museum</collectorName>
,
<collectorName id="523C89CE35086E1DFEEDF98DFE6BF993" box="[373,487,1567,1591]" pageId="5" pageNumber="67">Budapest</collectorName>
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.
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<paragraph id="FF76EC1835086E1DFF5DF9DBFBA0FB83" blockId="5.[159,762,845,1948]" lastBlockId="5.[826,1429,653,1063]" pageId="5" pageNumber="67">
Description — Female. Dorsum (
<figureCitation id="67F2F09D35086E1DFDCDF9DBFD41F9C5" box="[597,717,1609,1633]" captionStart="FIGURE" captionStartId="4.[159,171,1746,1765]" captionTargetBox="[282,1301,459,1693]" captionTargetId="figure-4@4.[265,1323,449,1715]" captionTargetPageId="4" captionText="FIGURE 1: Neopodocinum longisetum n. sp. holotype, female: A Body in dorsal view; B Body in ventral view; C Ventral view of gnathosoma and palp; D Epistome; E Chelicera." figureDoi="http://doi.org/10.5281/zenodo.5404292" httpUri="https://zenodo.org/record/5404292/files/figure.png" pageId="5" pageNumber="67">Figure 1A</figureCitation>
) — Dorsal shield oblong, with length 994 1090 µm and width 628 705 µm at level of coxae II (n=3), micropunctation on surface without ornamentation. Number of setae increased, bearing more than 30 pairs of dorsal setae and one unpaired seta. Setae j4-j6, z5-z6, Jx, J2, J5, Z5 minute, needle-like, other dorsal setae longer to very long; j1 plumose distally, twice the length of adjacent z1; setae j3, z2 and z4 especially long and distally pilose. Marginally bears neotrichous setae which are very long, slightly bent and pilose in their distal half. Length of dorsal setae: j1 = 83 96; j2 = 122 128; j3, z2, z4 and s4 = 224 288; j4 and z5 = 32 38; j5, j6, z6 and Jx = 7 13; z1 = 45 58; s2 = 77 109; s5-s6 = 340 372; J2 and Z5 = 26 38; J5 = 19 26; Z1, Z2 and Z4 = 71 83; S1 = 58 83; S2, S4 and S5 = 391 481. One median unpaired seta present (Jx), posterior to j6. Dorsal shield with 11 pairs of pores (5 pairs of lyrifissures [ids6, idZ1, idZ2, idS3, idJ4] and 6 pairs of gland pores) and 8 pairs of microspicules (ms). Peritreme not reaching the bases of setae z1.
</paragraph>
<paragraph id="FF76EC1835086E1DFCF8FBDAFC44F935" blockId="5.[826,1429,1096,1681]" pageId="5" pageNumber="67">
Venter (
<figureCitation id="67F2F09D35086E1DFC27FBDAFBBAFBC4" box="[959,1078,1096,1120]" captionStart="FIGURE" captionStartId="4.[159,171,1746,1765]" captionTargetBox="[282,1301,459,1693]" captionTargetId="figure-4@4.[265,1323,449,1715]" captionTargetPageId="4" captionText="FIGURE 1: Neopodocinum longisetum n. sp. holotype, female: A Body in dorsal view; B Body in ventral view; C Ventral view of gnathosoma and palp; D Epistome; E Chelicera." figureDoi="http://doi.org/10.5281/zenodo.5404292" httpUri="https://zenodo.org/record/5404292/files/figure.png" pageId="5" pageNumber="67">Figure 1B</figureCitation>
) — Sternal shield 103 109 long and 353 365 wide at level of coxae II. Bearing 3 pairs of needle-like setae and two pairs of lyrifissures. Posterior margin of sternal shield with a pair of clearly visible spur-like structure. Measurements of setae: St1 = 109 115, St2 = 103 115 and St3 = 90 96. Cuticule with a slight punctation. Metasternal setae 58 71 long, inserted on small metasternal platelets. Genital shield with straight posterior margin, 308 340 long and 244 269 wide. Length of setae on epigynal shield 83 109. Anal shield egg-shaped, with length 166 173 and width 128 135. Para-anal setae 38 58 long, needle-like, post-anal seta shorter, 32 38 long. Cribrum developed with pores on lateral margin of anal shield. Opisthogaster bearing more than 60 pairs of slightly pilose setae.
</paragraph>
<paragraph id="FF76EC1835086E1EFCF8F920FEC2FDC3" blockId="5.[826,1429,1714,1949]" lastBlockId="6.[159,762,311,1548]" lastPageId="6" lastPageNumber="68" pageId="5" pageNumber="67">
Gnathosoma (
<figureCitation id="67F2F09D35086E1DFB92F920FB08F96E" box="[1034,1156,1714,1738]" captionStart="FIGURE" captionStartId="4.[159,171,1746,1765]" captionTargetBox="[282,1301,459,1693]" captionTargetId="figure-4@4.[265,1323,449,1715]" captionTargetPageId="4" captionText="FIGURE 1: Neopodocinum longisetum n. sp. holotype, female: A Body in dorsal view; B Body in ventral view; C Ventral view of gnathosoma and palp; D Epistome; E Chelicera." figureDoi="http://doi.org/10.5281/zenodo.5404292" httpUri="https://zenodo.org/record/5404292/files/figure.png" pageId="5" pageNumber="67">Figure 1C</figureCitation>
) — Gnathosoma well developed. Deutosternal groove with 5 rows of denticles, 3 pairs of hypostomal setae and one pair of palpcoxal seta present, all setae needle-like. Internal posterior hypostomal seta (h2) longer than others. Measurements of hypostomal setae: h1 = 71 77, h2 = 103 135, h3 = 64 and capitulate seta = 71 96. Tectum (
<figureCitation id="67F2F09D350B6E1EFE11FEA5FD8DFEEB" box="[393,513,311,335]" captionStart="FIGURE" captionStartId="4.[159,171,1746,1765]" captionTargetBox="[282,1301,459,1693]" captionTargetId="figure-4@4.[265,1323,449,1715]" captionTargetPageId="4" captionText="FIGURE 1: Neopodocinum longisetum n. sp. holotype, female: A Body in dorsal view; B Body in ventral view; C Ventral view of gnathosoma and palp; D Epistome; E Chelicera." figureDoi="http://doi.org/10.5281/zenodo.5404292" httpUri="https://zenodo.org/record/5404292/files/figure.png" pageId="6" pageNumber="68">Figure 1D</figureCitation>
) unipartite with serration, length 115 135. Cheliceral measurements: fixed digit: 90, moveable digit 115 128. Fixed digit with simple dorsal seta, one median big tooth, distal smaller tooth, pilus dentilis and hooked terminally (
<figureCitation id="67F2F09D350B6E1EFF3EFE74FE97FE5A" box="[166,283,486,510]" captionStart="FIGURE" captionStartId="4.[159,171,1746,1765]" captionTargetBox="[282,1301,459,1693]" captionTargetId="figure-4@4.[265,1323,449,1715]" captionTargetPageId="4" captionText="FIGURE 1: Neopodocinum longisetum n. sp. holotype, female: A Body in dorsal view; B Body in ventral view; C Ventral view of gnathosoma and palp; D Epistome; E Chelicera." figureDoi="http://doi.org/10.5281/zenodo.5404292" httpUri="https://zenodo.org/record/5404292/files/figure.png" pageId="6" pageNumber="68">Figure 1E</figureCitation>
). Moveable digit with a bidentate tooth, small teeth and terminal hook. Arthrodial brush densely pilose. Length of fixed digit 90, moveable digit 115 128.
</paragraph>
<paragraph id="FF76EC18350B6E1EFF5DFD13FDFAFC2A" blockId="6.[159,762,311,1548]" pageId="6" pageNumber="68">
Legs — Tarsus I without ambulacrum, terminates distally in several small setae and one long seta, anterolaterally with a tridentate hook. Tarsi II- IV with well-developed ambulacra and claws. Most of the leg setae simple. Genu IV with 6 simple and 1 pilose setae. Leg chaetotaxy typical for the genus (
<bibRefCitation id="9B5891E9350B6E1EFF3FFCC1FEBBFCCF" author="Krantz G. W." box="[167,311,851,875]" pageId="6" pageNumber="68" pagination="139 - 226" refId="ref5516" refString="Krantz G. W. 1965 - A review of the genus Neopodocinum Oudemans, 1902 (Acarina: Macrochelidae) - Acarologia, 7 (2): 139 - 226." type="journal article" year="1965">Krantz 1965</bibRefCitation>
). Leg lengths: leg I 724 795, leg II 660 720, leg III 756 788, leg IV 833 1147.
</paragraph>
<paragraph id="FF76EC18350B6E1EFF5DFC35FD2DFBCA" blockId="6.[159,762,311,1548]" pageId="6" pageNumber="68">
Habitat —
<taxonomicName id="38C9979B350B6E1EFEDFFC3AFE6EFC1B" authority="Ács &amp; Sutak &amp; Kontschan, 2016" authorityName="Ács &amp; Sutak &amp; Kontschan" authorityYear="2016" box="[327,482,935,959]" class="Arachnida" family="Macrochelidae" genus="Neopodocinum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="68" phylum="Arthropoda" rank="species" species="longisetum" status="sp. nov.">
<emphasis id="CDBD300A350B6E1EFEDFFC3AFE6EFC1B" box="[327,482,935,959]" italics="true" pageId="6" pageNumber="68">N. longisetum</emphasis>
</taxonomicName>
<emphasis id="CDBD300A350B6E1EFE72FC35FDA3FC1B" bold="true" box="[490,559,935,959]" pageId="6" pageNumber="68">
<taxonomicNameLabel id="D68E8D71350B6E1EFE72FC35FDA3FC1B" box="[490,559,935,959]" pageId="6" pageNumber="68" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
was collected as phoretic on
<taxonomicName id="38C9979B350B6E1EFEB5FC58FDF0FC46" authority="(Linnaeus)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[301,636,970,994]" class="Magnoliopsida" family="Solanaceae" genus="Oryctes" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="68" phylum="Tracheophyta" rank="species" species="nasicornis">
<emphasis id="CDBD300A350B6E1EFEB5FC58FE7BFC46" box="[301,503,970,994]" italics="true" pageId="6" pageNumber="68">Oryctes nasicornis</emphasis>
(Linnaeus)
</taxonomicName>
in
<collectingCountry id="87DEAC88350B6E1EFD3BFC58FD7BFC46" box="[675,759,970,994]" name="Greece" pageId="6" pageNumber="68">Greece</collectingCountry>
. Phoresy on beetles is a common phenomenon in the family
<taxonomicName id="38C9979B350B6E1EFE83FB82FE46FB8C" authorityName="Vitzthum" authorityYear="1930" box="[283,458,1040,1064]" class="Arachnida" family="Macrochelidae" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="68" phylum="Arthropoda" rank="family">Macrochelidae</taxonomicName>
, enabling the mites to disperse easily and thus have the chance to find more suitable substrates. (
<bibRefCitation id="9B5891E9350B6E1EFE17FBC4FD9DFBCA" author="Costa M." box="[399,529,1110,1134]" pageId="6" pageNumber="68" pagination="411 - 428" refId="ref4606" refString="Costa M. 1969 - The association between mesostigmatic mites and coprid beetles - Acarologia, 11: 411 - 428." type="journal article" year="1969">Costa 1969</bibRefCitation>
,
<bibRefCitation id="9B5891E9350B6E1EFD83FBC4FD16FBCA" author="Binns E. S." box="[539,666,1110,1134]" pageId="6" pageNumber="68" pagination="571 - 620" refId="ref4356" refString="Binns E. S. 1982 - Phoresy as migration - some functional aspects of phoresy in mites - Biol. Rev., 57: 571 - 620. doi: 10.1111 / j. 1469 - 185 X. 1982. tb 00374. x" type="journal article" year="1982">Binns 1982</bibRefCitation>
)
</paragraph>
<paragraph id="FF76EC18350B6E1EFF5DFB15FDE3FB41" blockId="6.[159,762,311,1548]" pageId="6" pageNumber="68">
Etymology — The specific name is taken from the Latin &quot;
<taxonomicName id="38C9979B350B6E1EFEABFB38FE27FB66" authorityName="Ács &amp; Sutak &amp; Kontschan" authorityYear="2016" box="[307,427,1194,1218]" class="Arachnida" family="Macrochelidae" genus="Neopodocinum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="68" phylum="Arthropoda" rank="species" species="longisetum">
<emphasis id="CDBD300A350B6E1EFEABFB38FE27FB66" box="[307,427,1194,1218]" italics="true" pageId="6" pageNumber="68">longisetum</emphasis>
</taxonomicName>
&quot; and refers to the long marginal and submarginal dorsal setae.
</paragraph>
<paragraph id="FF76EC18350B6E1EFF5DFB6DFDFFF9A8" blockId="6.[159,762,311,1548]" pageId="6" pageNumber="68">
Notes —
<taxonomicName id="38C9979B350B6E1EFEA1FB6DFE5BFAB2" authority="Ács &amp; Sutak &amp; Kontschan, 2016" authorityName="Ács &amp; Sutak &amp; Kontschan" authorityYear="2016" box="[313,471,1278,1302]" class="Arachnida" family="Macrochelidae" genus="Neopodocinum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="68" phylum="Arthropoda" rank="species" species="longisetum" status="sp. nov.">
<emphasis id="CDBD300A350B6E1EFEA1FB6DFE5BFAB2" box="[313,471,1278,1302]" italics="true" pageId="6" pageNumber="68">N. longisetum</emphasis>
</taxonomicName>
<emphasis id="CDBD300A350B6E1EFE79FB6DFDA3FAB3" bold="true" box="[481,559,1279,1303]" pageId="6" pageNumber="68">
<taxonomicNameLabel id="D68E8D71350B6E1EFE79FB6DFDA3FAB3" box="[481,559,1279,1303]" pageId="6" pageNumber="68" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="38C9979B350B6E1EFD46FB6DFE7AFA9E" authority="(Berlese, 1908)" baseAuthorityName="Berlese" baseAuthorityYear="1908" class="Arachnida" family="Macrochelidae" genus="Neopodocinum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="68" phylum="Arthropoda" rank="species" species="caputmedusae">
<emphasis id="CDBD300A350B6E1EFD46FB6DFEBAFA9D" italics="true" pageId="6" pageNumber="68">N. caputmedusae</emphasis>
(
<bibRefCitation id="9B5891E9350B6E1EFED3FAB3FE62FA9E" author="Berlese A." box="[331,494,1313,1338]" pageId="6" pageNumber="68" pagination="1 - 15" refId="ref4334" refString="Berlese A. 1908 - Elenco di genere e specie nuove di Acari - Redia, 5: 1 - 15." type="journal article" year="1908">Berlese, 1908</bibRefCitation>
)
</taxonomicName>
, differences between them are shown in the
<tableCitation id="B24BD9A3350B6E1EFE3CFAD6FE79FAF9" box="[420,501,1348,1373]" captionStart="TABLE" captionStartId="5.[211,224,308,327]" captionText="TABLE 1: Most important differences between Neopodocinum longisetum n. sp. and Neopodocinum caputmedusae (Berlese, 1908)" pageId="6" pageNumber="68">Table 1</tableCitation>
. Differences also were confirmed between
<taxonomicName id="38C9979B350B6E1EFE10FAFAFDAEFA24" authorityName="Ács &amp; Sutak &amp; Kontschan" authorityYear="2016" box="[392,546,1384,1408]" class="Arachnida" family="Macrochelidae" genus="Neopodocinum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="68" phylum="Arthropoda" rank="species" species="longisetum">
<emphasis id="CDBD300A350B6E1EFE10FAFAFDAEFA24" box="[392,546,1384,1408]" italics="true" pageId="6" pageNumber="68">N. longisetum</emphasis>
</taxonomicName>
and the other earlier described species of
<taxonomicName id="38C9979B350B6E1EFE56FA19FDE3FA07" authorityName="Oudemans" authorityYear="1902" box="[462,623,1419,1443]" class="Arachnida" family="Macrochelidae" genus="Neopodocinum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="CDBD300A350B6E1EFE56FA19FDE3FA07" box="[462,623,1419,1443]" italics="true" pageId="6" pageNumber="68">Neopodocinum</emphasis>
</taxonomicName>
(
<bibRefCitation id="9B5891E9350B6E1EFD1AFA19FF5BFA62" author="Bregetova N. G." pageId="6" pageNumber="68" pagination="176 - 179" refId="ref4403" refString="Bregetova N. G. 1958 - A new species of Gamasid mite of the genus Megalolaelaps (Parasitiformes, Gamasoidea) - Paraz. Sbornik, 18: 176 - 179." type="journal article" year="1958">Bregetova 1958</bibRefCitation>
;
<bibRefCitation id="9B5891E9350B6E1EFF7CFA3CFEEAFA62" author="Costa M." box="[228,358,1454,1478]" pageId="6" pageNumber="68" pagination="63 - 86" refId="ref4562" refString="Costa M. 1965 - Neopodocinum caputmedusae comb., nov., a polymorphic mesostigmatic mite associated with Copris hispanus (L.) (Coleoptera: Scarabaeidae) - Isr. J. Zool., 14: 63 - 86." type="journal article" year="1965">
<collectingRegion id="3D0D22FA350B6E1EFF7CFA3CFEABFA62" box="[228,295,1454,1478]" country="New Zealand" name="West Coast" pageId="6" pageNumber="68">Costa</collectingRegion>
1965
</bibRefCitation>
;
<bibRefCitation id="9B5891E9350B6E1EFEEBFA3CFD88FA62" author="Krantz G. W." box="[371,516,1454,1478]" pageId="6" pageNumber="68" pagination="139 - 226" refId="ref5516" refString="Krantz G. W. 1965 - A review of the genus Neopodocinum Oudemans, 1902 (Acarina: Macrochelidae) - Acarologia, 7 (2): 139 - 226." type="journal article" year="1965">Krantz 1965</bibRefCitation>
,
<bibRefCitation id="9B5891E9350B6E1EFD89FA3CFF5BFA4D" author="Hartini S. &amp; Takaku G." pageId="6" pageNumber="68" pagination="47 - 65" refId="ref5157" refString="Hartini S., Takaku G. 2003 - Mites of the macrochelid genus Neopodocinum (Arachnida: Acari: Gamasida: Macrochelidae) associated with dung beetles in West Java, Indonesia - Species Diversity, 8: 47 - 65." type="journal article" year="2003">Hartini and Takaku 2003</bibRefCitation>
;
<bibRefCitation id="9B5891E9350B6E1EFF76FA43FDA7FA4D" author="Hartini S. &amp; Takaku G." box="[238,555,1489,1513]" pageId="6" pageNumber="68" pagination="77 - 89" refId="ref5200" refString="Hartini S., Takaku G. 2004 - Neopodocinum Mites (Arachnida: Acari: Macrochelidae) in Kalimantan - Species Diversity, 9: 77 - 89." type="journal article" year="2004">Hartini and Takaku 2004</bibRefCitation>
;
<bibRefCitation id="9B5891E9350B6E1EFDDAFA43FD7AFA4D" author="Iavorschi V." box="[578,758,1489,1513]" pageId="6" pageNumber="68" pagination="99 - 107" refId="ref5268" refString="Iavorschi V. 1975 - Neopodocinum ilincae, nouvelle espLce de Macrochelidae (Acarina - Anactinotrichida) de Roumanie - Trav. Inst. SpOol. Emile Racovitza, 14: 99 - 107." type="journal article" year="1975">Iavorschi 1975</bibRefCitation>
;
<bibRefCitation id="9B5891E9350B6E1EFF07FA66FEBBF9A8" author="Moraza M. L." box="[159,311,1524,1548]" pageId="6" pageNumber="68" pagination="261 - 269" refId="ref6077" refString="Moraza M. L. 2004 - The phoretic genus Neopodocinum (Oudemans, 1902) in the Iberian Peninsula (Acari: Mesostigmata: Macrochelidae) - Rev. Iber. Aracnol., 10: 261 - 269." type="journal article" year="2004">Moraza 2004</bibRefCitation>
;
<bibRefCitation id="9B5891E9350B6E1EFEDAFA66FDEBF9A8" author="Takaku G. &amp; Hartini S." box="[322,615,1524,1548]" pageId="6" pageNumber="68" pagination="323 - 345" refId="ref6805" refString="Takaku G., Hartini S. 2001 - Macrochelid Mites (Arachnida: Acari: Macrochelidae: Glyptholaspis, Macrocheles, Neopodocinum) associated with Dung Beetles in Bali, Indonesia - Species Diversity, 6: 323 - 345." type="journal article" year="2001">Takaku and Hartini 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>