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<document id="1C7F406E5317FE410925B85634CDE180" ID-DOI="10.11646/zootaxa.3981.4.1" ID-GBIF-Dataset="15c3bf52-4481-4a75-8643-7c919bdde6c4" ID-ISSN="1175-5326" ID-Zenodo-Dep="235846" ID-ZooBank="EF71D83B-17B4-49CA-826E-D3A8E7979750" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461248924452" checkinUser="plazi" docAuthor="Zwick, Peter" docDate="2015" docId="5C5BE52CFF8BBC662CB5FE979174094F" docLanguage="en" docName="zt03981p490.pdf" docOrigin="Zootaxa 3981 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptocyphon furcalonga Zwick, 2015, n. sp." docType="treatment" docVersion="8" lastPageNumber="480" masterDocId="A0629D54FF91BC7B2C22FFE99334080F" masterDocTitle="Australian Marsh Beetles (Coleoptera: Scirtidae). 8. The new genera Cygnocyphon, Eximiocyphon, Paracyphon, Leptocyphon, Tectocyphon, and additions to Contacyphon de Gozis, Nanocyphon Zwick and Eurycyphon Watts" masterLastPageNumber="490" masterPageNumber="451" pageNumber="477" updateTime="1698607092210" updateUser="plazi">
<mods:mods id="B2CB859C7D1F9B4F7DBE37E7BE08B679" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="AAC3862B17652100A427920651DBFCD8">Australian Marsh Beetles (Coleoptera: Scirtidae). 8. The new genera Cygnocyphon, Eximiocyphon, Paracyphon, Leptocyphon, Tectocyphon, and additions to Contacyphon de Gozis, Nanocyphon Zwick and Eurycyphon Watts</mods:title>
</mods:titleInfo>
<mods:name id="4E00078BACA99E31D90AEDB691747948" type="personal">
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<mods:namePart id="BB90340AF704053F0519E488E6A8A4FD">Zwick, Peter</mods:namePart>
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<mods:title id="EEA0C7B9CC1F5C67046477C58124723E">Zootaxa</mods:title>
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<mods:date id="7C06D3CBFFEA7AC3F1EBDC5B2CEA2AB0">2015</mods:date>
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<mods:number id="BF07A21A3D8658DDB990C63E66EE93BC">3981</mods:number>
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<mods:classification id="DE267A7B9A25CB728ADDD8031F01401D">journal article</mods:classification>
<mods:identifier id="B64FFEFDAF6B437408DEDD224162DF41" type="DOI">10.11646/zootaxa.3981.4.1</mods:identifier>
<mods:identifier id="A9D660D82EF55789236BDD6F2626AD80" type="GBIF-Dataset">15c3bf52-4481-4a75-8643-7c919bdde6c4</mods:identifier>
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<treatment id="5C5BE52CFF8BBC662CB5FE979174094F" ID-DOI="http://doi.org/10.5281/zenodo.6110578" ID-GBIF-Taxon="119635595" ID-Zenodo-Dep="6110578" LSID="urn:lsid:plazi:treatment:5C5BE52CFF8BBC662CB5FE979174094F" httpUri="http://treatment.plazi.org/id/5C5BE52CFF8BBC662CB5FE979174094F" lastPageId="29" lastPageNumber="480" pageId="26" pageNumber="477">
<subSubSection id="9CE807B1FF8BBC612CB5FE97921A09B5" pageId="26" pageNumber="477" type="nomenclature">
<paragraph id="D44D543AFF8BBC612CB5FE9791250997" blockId="26.[151,529,382,442]" box="[151,529,382,408]" pageId="26" pageNumber="477">
<heading id="8F05E356FF8BBC612CB5FE9791250997" bold="true" box="[151,529,382,408]" fontSize="11" level="1" pageId="26" pageNumber="477" reason="1">
<emphasis id="E6868828FF8BBC612CB5FE9791250997" bold="true" box="[151,529,382,408]" pageId="26" pageNumber="477">
<emphasis id="E6868828FF8BBC612CB5FE9792F30997" bold="true" box="[151,455,382,408]" italics="true" pageId="26" pageNumber="477">
<taxonomicName id="13F22FB9FF8BBC612CB5FE9792F50997" box="[151,449,382,408]" class="Insecta" family="Scirtidae" genus="Leptocyphon" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="furcalonga" status="sp. nov.">Leptocyphon furcalonga</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="FDB53553FF8BBC612DECFE9791250997" box="[462,529,382,408]" pageId="26" pageNumber="477" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D44D543AFF8BBC612CB5FE48921A09B5" blockId="26.[151,529,382,442]" box="[151,302,417,442]" pageId="26" pageNumber="477">
(
<figureCitation id="4CC948BFFF8BBC612CBDFE48921209B5" box="[159,294,417,442]" captionStart-0="FIGURES 76 – 79" captionStart-1="FIGURES 80 – 88" captionStartId-0="26.[151,264,1961,1984]" captionStartId-1="27.[151,264,1910,1933]" captionTargetBox-0="[150,1436,1070,1933]" captionTargetBox-1="[194,1393,193,1857]" captionTargetId-0="figure@26.[150,1436,1070,1940]" captionTargetId-1="figure@27.[182,1393,193,1859]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionText-0="FIGURES 76 – 79. Leptocyphon furcalonga, n. gen., n. sp.: 76, 77, habitus dorsal and lateral; 78, forebody, oblique frontodorsal view; 79, cleared head, ventral. Not to scale." captionText-1="FIGURES 80 – 88. Leptocyphon furcalonga, n. gen., n. sp.: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T 8; 83, S 8; 84, T 9 and S 9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S 8 shown in black; 88, prehensor. 80 – 84 and 85, 86 and 88 to the same scales, respectively. aT 8, apodemes of T 8; ba, bacula of ovipositor; lc, laterocervicalia." httpUri-0="https://zenodo.org/record/235858/files/figure.png" httpUri-1="https://zenodo.org/record/235859/files/figure.png" pageId="26" pageNumber="477">Figs. 76–87</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9CE807B1FF8BBC672CB5FE0091F80A6F" lastPageId="28" lastPageNumber="479" pageId="26" pageNumber="477" type="materials_examined">
<paragraph id="D44D543AFF8BBC612CB5FE00964F0A61" blockId="26.[151,1437,489,1018]" pageId="26" pageNumber="477">
<emphasis id="E6868828FF8BBC612CB5FE0092720A0D" bold="true" box="[151,326,489,514]" pageId="26" pageNumber="477">
<typeStatus id="0B49EA98FF8BBC612CB5FE0093E60A0D" box="[151,210,489,514]" pageId="26" pageNumber="477">Type</typeStatus>
material.
</emphasis>
<specimenCount id="C2F49FB3FF8BBC612D6CFE0092450A0D" box="[334,369,489,514]" pageId="26" pageNumber="477" type="generic">
1
<emphasis id="E6868828FF8BBC612D7EFE0092450A0D" bold="true" box="[348,369,489,514]" pageId="26" pageNumber="477">♂</emphasis>
</specimenCount>
<typeStatus id="0B49EA98FF8BBC612D58FE0092D60A0D" box="[378,482,489,514]" pageId="26" pageNumber="477" type="holotype">holotype</typeStatus>
,
<specimenCount id="C2F49FB3FF8BBC612DD2FE0091270A0D" box="[496,531,489,514]" pageId="26" pageNumber="477" type="generic">
6
<emphasis id="E6868828FF8BBC612DDCFE0091270A0D" bold="true" box="[510,531,489,514]" pageId="26" pageNumber="477">♂</emphasis>
</specimenCount>
<typeStatus id="0B49EA98FF8BBC612E39FE0091A10A0D" box="[539,661,489,514]" pageId="26" pageNumber="477" type="paratype">paratypes</typeStatus>
:
<quantity id="130AF9DFFF8BBC612E81FE0091D70A0D" box="[675,739,489,514]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="477" unit="km" value="20.0">20km</quantity>
NE Cervantes [
<geoCoordinate id="B1C632FDFF8BBC612FBAFE00973C0A0D" box="[920,1032,489,514]" direction="south" orientation="latitude" pageId="26" pageNumber="477" precision="5" value="-30.5022">30.5022S</geoCoordinate>
,
<geoCoordinate id="B1C632FDFF8BBC612837FE0097A70A0D" box="[1045,1171,489,514]" direction="east" orientation="longitude" pageId="26" pageNumber="477" precision="5" value="115.0681">115.0681E</geoCoordinate>
],
<date id="A04C72FAFF8BBC612884FE00966B0A0E" box="[1190,1375,489,513]" pageId="26" pageNumber="477" value="1987-08-20" valueMax="1987-08-23" valueMin="1987-08-20">20-23 Aug.1987</date>
<emphasis id="E6868828FF8BBC61294AFE0096A60A0D" bold="true" box="[1384,1426,489,514]" pageId="26" pageNumber="477">WA</emphasis>
, C.Reid ex Acacia (
<collectionCode id="B2E3CCFFFF8BBC612D4DFDE7928C0A29" LSID="urn:lsid:biocol.org:col:32981" box="[367,440,526,550]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="26" pageNumber="477">ANIC</collectionCode>
; the entirely cleared
<typeStatus id="0B49EA98FF8BBC612E85FDE4903D0A29" box="[679,777,525,550]" pageId="26" pageNumber="477" type="holotype">holotype</typeStatus>
was dissected and mounted in Euparal, on several pinned slides; 1 PT in
<collectionCode id="B2E3CCFFFF8BBC612D69FDD892440A46" box="[331,368,561,585]" pageId="26" pageNumber="477">PZ</collectionCode>
);
<specimenCount id="C2F49FB3FF8BBC612DA4FDD8929D0A45" box="[390,425,561,586]" pageId="26" pageNumber="477" type="generic">
1
<emphasis id="E6868828FF8BBC612DB6FDD8929D0A45" bold="true" box="[404,425,561,586]" pageId="26" pageNumber="477">♂</emphasis>
</specimenCount>
<typeStatus id="0B49EA98FF8BBC612D91FDD891160A45" box="[435,546,561,586]" pageId="26" pageNumber="477" type="paratype">paratype</typeStatus>
:
<geoCoordinate id="B1C632FDFF8BBC612E10FDD891B50A46" box="[562,641,561,585]" direction="south" orientation="latitude" pageId="26" pageNumber="477" precision="555" value="-33.48">33.48S</geoCoordinate>
<geoCoordinate id="B1C632FDFF8BBC612EA8FDD891DF0A45" box="[650,747,561,586]" direction="east" orientation="longitude" pageId="26" pageNumber="477" precision="555" value="115.17">115.17E</geoCoordinate>
, Creek
<quantity id="130AF9DFFF8BBC612F67FDD890430A45" box="[837,887,561,586]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="26" pageNumber="477" unit="km" value="8.0">8km</quantity>
N of Osmington, WA,
<date id="A04C72FAFF8BBC6128A8FDD897DC0A45" box="[1162,1256,561,586]" pageId="26" pageNumber="477" value="1969-11-15">15.xi.69</date>
, E. Britton. 1♀
<emphasis id="E6868828FF8BBC612CB5FDBC92380A61" bold="true" box="[151,268,597,622]" pageId="26" pageNumber="477">
<typeStatus id="0B49EA98FF8BBC612CB5FDBC92330A61" box="[151,263,597,622]" pageId="26" pageNumber="477" type="paratype">paratype</typeStatus>
:
</emphasis>
<geoCoordinate id="B1C632FDFF8BBC612D30FDBF92540A61" box="[274,352,598,622]" direction="south" orientation="latitude" pageId="26" pageNumber="477" precision="555" value="-33.39">33.39S</geoCoordinate>
<geoCoordinate id="B1C632FDFF8BBC612D4AFDBC92FC0A61" box="[360,456,597,622]" direction="east" orientation="longitude" pageId="26" pageNumber="477" precision="555" value="115.01">115.01E</geoCoordinate>
,
<quantity id="130AF9DFFF8BBC612DF1FDBC91310A61" box="[467,517,597,622]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="477" unit="km" value="1.0">1km</quantity>
S of Yallingup WA
<date id="A04C72FAFF8BBC612EC5FDBF90520A61" box="[743,870,598,622]" pageId="26" pageNumber="477" value="1981-10-04">4 Oct.1981</date>
, I.D.Naumann, J.C.Cardale ex ethanol (
<collectionCode id="B2E3CCFFFF8BBC612907FDBF96440A61" LSID="urn:lsid:biocol.org:col:32981" box="[1317,1392,598,622]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="26" pageNumber="477">ANIC</collectionCode>
).
</paragraph>
<paragraph id="D44D543AFF8BBC612CE5FD90919F0B2E" blockId="26.[151,1437,489,1018]" pageId="26" pageNumber="477">
<emphasis id="E6868828FF8BBC612CE5FD9092120A9D" bold="true" box="[199,294,633,658]" pageId="26" pageNumber="477">Habitus</emphasis>
. BL
<quantity id="130AF9DFFF8BBC612D4BFD9392CC0A9E" box="[361,504,634,658]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.2" metricValueMin="2.4" pageId="26" pageNumber="477" unit="mm" value="2.8" valueMax="3.2" valueMin="2.4">2.4–3.2 mm</quantity>
, BL/BW ~1.8. Body elongate, the almost parallel elytra widest a little behind midlength of elytra which are more than five times longer than the transverse pronotum. Most specimens dark brown to almost black, some (
<figureCitation id="4CC948BFFF8BBC612DDAFD2891B60AD5" box="[504,642,705,730]" captionStart="FIGURES 76 – 79" captionStartId="26.[151,264,1961,1984]" captionTargetBox="[150,1436,1070,1933]" captionTargetId="figure@26.[150,1436,1070,1940]" captionTargetPageId="26" captionText="FIGURES 76 – 79. Leptocyphon furcalonga, n. gen., n. sp.: 76, 77, habitus dorsal and lateral; 78, forebody, oblique frontodorsal view; 79, cleared head, ventral. Not to scale." httpUri="https://zenodo.org/record/235858/files/figure.png" pageId="26" pageNumber="477">Figs. 76–78</figureCitation>
) with ochre pronotum and elytra. The semi-erect pilosity is greyish. Elytra with very fine dense punctation. The punctures on head and especially pronotum are much larger, granular, almost pearl-shaped, barely one diameter apart.
</paragraph>
<paragraph id="D44D543AFF8BBC612CE5FCC493C80BF5" blockId="26.[151,1437,489,1018]" pageId="26" pageNumber="477">
Head (
<figureCitation id="4CC948BFFF8BBC612D30FCC492510B49" box="[274,357,813,838]" captionStart="FIGURES 76 – 79" captionStartId="26.[151,264,1961,1984]" captionTargetBox="[150,1436,1070,1933]" captionTargetId="figure@26.[150,1436,1070,1940]" captionTargetPageId="26" captionText="FIGURES 76 – 79. Leptocyphon furcalonga, n. gen., n. sp.: 76, 77, habitus dorsal and lateral; 78, forebody, oblique frontodorsal view; 79, cleared head, ventral. Not to scale." httpUri="https://zenodo.org/record/235858/files/figure.png" pageId="26" pageNumber="477">Fig. 79</figureCitation>
) about 1.25 times wider across the eyes than long from occiput to front of frontoclypeus. Eyes distinctly projecting. Antenna long and slender, scape ovoid, without anterior edge, pedicel smaller than scape, ovoid. Antennomere 3 thinner than pedicel but as long as it, conical. Distal antennomeres about twice as long as wide at apex. Right mandible with a small tooth, left without tooth. Maxillary palpus unmodified, terminal segment apically narrowed, tip blunt. Terminal segment of labial palpus bean-shaped, inserted terminally on penultimate segment.
</paragraph>
<caption id="808D04B2FF8BBC612CB5F84091AA0FD1" httpUri="https://zenodo.org/record/235858/files/figure.png" pageId="26" pageNumber="477" targetBox="[150,1436,1070,1933]" targetPageId="26">
<paragraph id="D44D543AFF8BBC612CB5F84091AA0FD1" blockId="26.[151,1435,1961,2014]" pageId="26" pageNumber="477">
<emphasis id="E6868828FF8BBC612CB5F840927B0FB1" bold="true" box="[151,335,1961,1984]" pageId="26" pageNumber="477">FIGURES 76–79</emphasis>
.
<emphasis id="E6868828FF8BBC612D7CF840916A0FB0" box="[350,606,1961,1983]" italics="true" pageId="26" pageNumber="477">
<taxonomicName id="13F22FB9FF8BBC612D7CF840916F0FB0" box="[350,603,1961,1983]" class="Insecta" family="Scirtidae" genus="Leptocyphon" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="furcalonga" status="gen. nov.">Leptocyphon furcalonga</taxonomicName>
,
</emphasis>
<emphasis id="E6868828FF8BBC612E45F84391850FB0" bold="true" box="[615,689,1962,1983]" pageId="26" pageNumber="477">
<taxonomicNameLabel id="FDB53553FF8BBC612E45F84391850FB0" box="[615,689,1962,1983]" pageId="26" pageNumber="477" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="E6868828FF8BBC612E9DF84391CF0FB0" bold="true" box="[703,763,1962,1983]" pageId="26" pageNumber="477">n. sp.</emphasis>
: 76, 77, habitus dorsal and lateral; 78, forebody, oblique frontodorsal view; 79, cleared head, ventral. Not to scale.
</paragraph>
</caption>
<caption id="808D04B2FF8ABC602CB5F89F97950FE6" httpUri="https://zenodo.org/record/235859/files/figure.png" pageId="27" pageNumber="478" targetBox="[194,1393,193,1857]" targetPageId="27">
<paragraph id="D44D543AFF8ABC602CB5F89F97950FE6" blockId="27.[151,1436,1910,2025]" pageId="27" pageNumber="478">
<emphasis id="E6868828FF8ABC602CB5F89F92790F83" bold="true" box="[151,333,1910,1933]" pageId="27" pageNumber="478">FIGURES 80–88</emphasis>
.
<emphasis id="E6868828FF8ABC602D78F89E916C0F82" box="[346,600,1911,1933]" italics="true" pageId="27" pageNumber="478">
<taxonomicName id="13F22FB9FF8ABC602D78F89E91600F82" box="[346,596,1911,1933]" class="Insecta" family="Scirtidae" genus="Leptocyphon" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="furcalonga" status="gen. nov.">Leptocyphon furcalonga</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="FDB53553FF8ABC602E7CF89E91920F83" box="[606,678,1911,1932]" pageId="27" pageNumber="478" rank="genus">
<emphasis id="E6868828FF8ABC602E7CF89E91920F83" bold="true" box="[606,678,1911,1932]" pageId="27" pageNumber="478">n. gen.</emphasis>
</taxonomicNameLabel>
,
<emphasis id="E6868828FF8ABC602E90F89E91D80F83" bold="true" box="[690,748,1911,1932]" pageId="27" pageNumber="478">n. sp.</emphasis>
: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T8; 83, S8; 84, T9 and S9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S8 shown in black; 88, prehensor. 80–84 and 85, 86 and 88 to the same scales, respectively. aT8, apodemes of T8; ba, bacula of ovipositor; lc, laterocervicalia.
</paragraph>
</caption>
<paragraph id="D44D543AFF8DBC672CE5FF7E929F094F" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="479">The pronotum is transverse, twice as wide at base as its length along midline. The rear margin projects towards the scutellum, the rear angles are about rectangular, the curved sides converge towards the blunt front angles which do not project. The notum is a little domed, side margin in lateral view straight. A pair of small paramedian pits at the rear pronotal margin. In the whole mount of a cleared specimen they resemble minute punctures. Legs ambulatory, unmodified.</paragraph>
<paragraph id="D44D543AFF8DBC672CE5FEA591F80A6F" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="479">
Underside: The subgenal ridge runs on the lower face of the head, far from the eye. In front it meets a short branch coming from the end of the gular suture, together they enter the side edge of the oral cavity (
<figureCitation id="4CC948BFFF8DBC672925FE8696680987" box="[1287,1372,367,392]" captionStart="FIGURES 76 – 79" captionStartId="26.[151,264,1961,1984]" captionTargetBox="[150,1436,1070,1933]" captionTargetId="figure@26.[150,1436,1070,1940]" captionTargetPageId="26" captionText="FIGURES 76 – 79. Leptocyphon furcalonga, n. gen., n. sp.: 76, 77, habitus dorsal and lateral; 78, forebody, oblique frontodorsal view; 79, cleared head, ventral. Not to scale." httpUri="https://zenodo.org/record/235858/files/figure.png" pageId="28" pageNumber="479">Fig. 79</figureCitation>
). The short and thin, blade-like prosternal process projects barely back beyond the coxae, its tip is insignificantly widened, with single hairs (
<figureCitation id="4CC948BFFF8DBC672DC7FE5E910B09DF" box="[485,575,439,464]" captionStart="FIGURES 80 – 88" captionStartId="27.[151,264,1910,1933]" captionTargetBox="[194,1393,193,1857]" captionTargetId="figure@27.[182,1393,193,1859]" captionTargetPageId="27" captionText="FIGURES 80 – 88. Leptocyphon furcalonga, n. gen., n. sp.: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T 8; 83, S 8; 84, T 9 and S 9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S 8 shown in black; 88, prehensor. 80 – 84 and 85, 86 and 88 to the same scales, respectively. aT 8, apodemes of T 8; ba, bacula of ovipositor; lc, laterocervicalia." httpUri="https://zenodo.org/record/235859/files/figure.png" pageId="28" pageNumber="479">Fig. 80</figureCitation>
). The mesoventral groove is poorly circumscribed and shallow. It is an anteriorly open small triangle (
<figureCitation id="4CC948BFFF8DBC672E28FE35915709FB" box="[522,611,476,501]" captionStart="FIGURES 80 – 88" captionStartId="27.[151,264,1910,1933]" captionTargetBox="[194,1393,193,1857]" captionTargetId="figure@27.[182,1393,193,1859]" captionTargetPageId="27" captionText="FIGURES 80 – 88. Leptocyphon furcalonga, n. gen., n. sp.: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T 8; 83, S 8; 84, T 9 and S 9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S 8 shown in black; 88, prehensor. 80 – 84 and 85, 86 and 88 to the same scales, respectively. aT 8, apodemes of T 8; ba, bacula of ovipositor; lc, laterocervicalia." httpUri="https://zenodo.org/record/235859/files/figure.png" pageId="28" pageNumber="479">Fig. 81</figureCitation>
), no vertical carinae in front of it on vertical front of mesoventrite. Mesoventral process narrower than an antennomere but separating the coxae completely. Its finely incised end meets the front of the metaventrite, the metaventral process is barely developed. The discrimen extends over ¾ of the metaventrite. Abdominal sternites unmodified.
</paragraph>
</subSubSection>
<subSubSection id="9CE807B1FF8DBC662CE5FD8590E108F7" lastPageId="29" lastPageNumber="480" pageId="28" pageNumber="479" type="description">
<paragraph id="D44D543AFF8DBC672CE5FD8592840B53" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="479">
<emphasis id="E6868828FF8DBC672CE5FD8592300A8A" bold="true" box="[199,260,620,645]" pageId="28" pageNumber="479">Male</emphasis>
. T8 with pentagonal plate, apodemes short, straight, basally connected by fine rods delimiting a deep backward incision into the plate. Most of the plate is rough, densely covered with microtrichia, the interspersed true setae are barely noticeable. Microtrichia and socketed setae along the caudal margin are of similar length. S8 large, sclerite Y-shaped, with very long unpaired branch, the weakly sclerotized plate reduced to lateral plaques with few setae (
<figureCitation id="4CC948BFFF8DBC672CC1FD1592410B1B" box="[227,373,764,789]" captionStart="FIGURES 80 – 88" captionStartId="27.[151,264,1910,1933]" captionTargetBox="[194,1393,193,1857]" captionTargetId="figure@27.[182,1393,193,1859]" captionTargetPageId="27" captionText="FIGURES 80 – 88. Leptocyphon furcalonga, n. gen., n. sp.: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T 8; 83, S 8; 84, T 9 and S 9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S 8 shown in black; 88, prehensor. 80 – 84 and 85, 86 and 88 to the same scales, respectively. aT 8, apodemes of T 8; ba, bacula of ovipositor; lc, laterocervicalia." httpUri="https://zenodo.org/record/235859/files/figure.png" pageId="28" pageNumber="479">Figs. 82, 83</figureCitation>
). Apodemes of T9 well developed, basally connected by an angular sclerite. The plate is membranous, bare and short. S9 is short, two indistinct sclerotized stripes support the semicircular plate which is caudally pilose (
<figureCitation id="4CC948BFFF8DBC672D6DFCAD92970B53" box="[335,419,836,861]" captionStart="FIGURES 80 – 88" captionStartId="27.[151,264,1910,1933]" captionTargetBox="[194,1393,193,1857]" captionTargetId="figure@27.[182,1393,193,1859]" captionTargetPageId="27" captionText="FIGURES 80 – 88. Leptocyphon furcalonga, n. gen., n. sp.: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T 8; 83, S 8; 84, T 9 and S 9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S 8 shown in black; 88, prehensor. 80 – 84 and 85, 86 and 88 to the same scales, respectively. aT 8, apodemes of T 8; ba, bacula of ovipositor; lc, laterocervicalia." httpUri="https://zenodo.org/record/235859/files/figure.png" pageId="28" pageNumber="479">Fig. 84</figureCitation>
).
</paragraph>
<paragraph id="D44D543AFF8DBC672CE5FC8E978C0BE2" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="479">
The slender penis is very long (
<figureCitation id="4CC948BFFF8DBC672E0EFC8E91B40B8F" box="[556,640,871,896]" captionStart="FIGURES 80 – 88" captionStartId="27.[151,264,1910,1933]" captionTargetBox="[194,1393,193,1857]" captionTargetId="figure@27.[182,1393,193,1859]" captionTargetPageId="27" captionText="FIGURES 80 – 88. Leptocyphon furcalonga, n. gen., n. sp.: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T 8; 83, S 8; 84, T 9 and S 9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S 8 shown in black; 88, prehensor. 80 – 84 and 85, 86 and 88 to the same scales, respectively. aT 8, apodemes of T 8; ba, bacula of ovipositor; lc, laterocervicalia." httpUri="https://zenodo.org/record/235859/files/figure.png" pageId="28" pageNumber="479">Fig. 85</figureCitation>
). The calyx-shaped pala supports long flat rod-like parameroids. They are narrow, apparent width varies with position. Their apices are short, cap-like, with sensory pores. The trigonium is much longer than the parameroids, very slender in its basal half, then gently widened and deeply divided into two narrow parallel branches. Each branch bears apically a fine membranous process or flagellum.
</paragraph>
<paragraph id="D44D543AFF8DBC672CE5FC1196B40C72" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="479">
Base of tegmen (
<figureCitation id="4CC948BFFF8DBC672DACFC1E92D70C1F" box="[398,483,1015,1040]" captionStart="FIGURES 80 – 88" captionStartId="27.[151,264,1910,1933]" captionTargetBox="[194,1393,193,1857]" captionTargetId="figure@27.[182,1393,193,1859]" captionTargetPageId="27" captionText="FIGURES 80 – 88. Leptocyphon furcalonga, n. gen., n. sp.: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T 8; 83, S 8; 84, T 9 and S 9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S 8 shown in black; 88, prehensor. 80 – 84 and 85, 86 and 88 to the same scales, respectively. aT 8, apodemes of T 8; ba, bacula of ovipositor; lc, laterocervicalia." httpUri="https://zenodo.org/record/235859/files/figure.png" pageId="28" pageNumber="479">Fig. 86</figureCitation>
) spade-shaped, with strongly sclerotized frame and membranous centre. From the membranous centre rises a pair of long paramedian weakly sclerotized blades with sensory pores in distal half and a few sensory pegs near the rounded tip. From the sclerotized sides of the tegmen originate very long soft processes tapering to a long acute tip. There is a dense rough pelt of pointed scales (detail in
<figureCitation id="4CC948BFFF8DBC672816FB8D97B30C72" box="[1076,1159,1124,1149]" captionStart="FIGURES 80 – 88" captionStartId="27.[151,264,1910,1933]" captionTargetBox="[194,1393,193,1857]" captionTargetId="figure@27.[182,1393,193,1859]" captionTargetPageId="27" captionText="FIGURES 80 – 88. Leptocyphon furcalonga, n. gen., n. sp.: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T 8; 83, S 8; 84, T 9 and S 9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S 8 shown in black; 88, prehensor. 80 – 84 and 85, 86 and 88 to the same scales, respectively. aT 8, apodemes of T 8; ba, bacula of ovipositor; lc, laterocervicalia." httpUri="https://zenodo.org/record/235859/files/figure.png" pageId="28" pageNumber="479">Fig. 86</figureCitation>
), except basolaterally.
</paragraph>
<paragraph id="D44D543AFF8DBC672CE5FB60975E0CCA" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="479">In specimens with everted genitalia, the fork of the penis is directed dorsad, the median paramere blades are wrapped around the dorsal part of the penis. The pelted acute processes diverge widely.</paragraph>
<paragraph id="D44D543AFF8DBC672CE5FB2697F80E07" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="479">
<emphasis id="E6868828FF8DBC672CE5FB26922A0CE7" bold="true" box="[199,286,1231,1256]" pageId="28" pageNumber="479">Female</emphasis>
(
<figureCitation id="4CC948BFFF8DBC672D13FB2692F50CE7" box="[305,449,1231,1256]" captionStart="FIGURES 80 – 88" captionStartId="27.[151,264,1910,1933]" captionTargetBox="[194,1393,193,1857]" captionTargetId="figure@27.[182,1393,193,1859]" captionTargetPageId="27" captionText="FIGURES 80 – 88. Leptocyphon furcalonga, n. gen., n. sp.: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T 8; 83, S 8; 84, T 9 and S 9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S 8 shown in black; 88, prehensor. 80 – 84 and 85, 86 and 88 to the same scales, respectively. aT 8, apodemes of T 8; ba, bacula of ovipositor; lc, laterocervicalia." httpUri="https://zenodo.org/record/235859/files/figure.png" pageId="28" pageNumber="479">Figs. 87, 88</figureCitation>
). S7 with slightly angular apex, other sternites unmodified. Ovipositor modified, the bacula are
<emphasis id="E6868828FF8DBC672D32FB1C92060D03" box="[272,306,1269,1292]" italics="true" pageId="28" pageNumber="479">via</emphasis>
a plate rigidly connected with the gonocoxites, the usual articulation is missing. The gonocoxites are slender triangles with sclerotised barb-like basolateral angles. A deep notch separates them from an elongate weak sclerite from which the long bacula extend forward. The stylites are wart-like, with one sensillum near the base and another at the tip. T8 large, soft, shape not recognizable, apodemes long. S8 is a triangular cone with microtrichia to which two short anteriorly widely separate apodemes (black in
<figureCitation id="4CC948BFFF8DBC672F59FA6D90FF0D93" box="[891,971,1412,1437]" captionStart="FIGURES 80 – 88" captionStartId="27.[151,264,1910,1933]" captionTargetBox="[194,1393,193,1857]" captionTargetId="figure@27.[182,1393,193,1859]" captionTargetPageId="27" captionText="FIGURES 80 – 88. Leptocyphon furcalonga, n. gen., n. sp.: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T 8; 83, S 8; 84, T 9 and S 9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S 8 shown in black; 88, prehensor. 80 – 84 and 85, 86 and 88 to the same scales, respectively. aT 8, apodemes of T 8; ba, bacula of ovipositor; lc, laterocervicalia." httpUri="https://zenodo.org/record/235859/files/figure.png" pageId="28" pageNumber="479">Fig. 87</figureCitation>
) attach. Prehensor (
<figureCitation id="4CC948BFFF8DBC672889FA6D97C90D93" box="[1195,1277,1412,1437]" captionStart="FIGURES 80 – 88" captionStartId="27.[151,264,1910,1933]" captionTargetBox="[194,1393,193,1857]" captionTargetId="figure@27.[182,1393,193,1859]" captionTargetPageId="27" captionText="FIGURES 80 – 88. Leptocyphon furcalonga, n. gen., n. sp.: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T 8; 83, S 8; 84, T 9 and S 9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S 8 shown in black; 88, prehensor. 80 – 84 and 85, 86 and 88 to the same scales, respectively. aT 8, apodemes of T 8; ba, bacula of ovipositor; lc, laterocervicalia." httpUri="https://zenodo.org/record/235859/files/figure.png" pageId="28" pageNumber="479">Fig. 88</figureCitation>
) composed of a transverse sclerite bar which is laterally connected to cuticular folds densely covered with microtrichia. The bar has a short median extension. Two narrowly separated triangular plates with strong teeth along the sides and finer ones along the wide caudal edge lie on the opposite side of the gonoduct. Bursella not observed.
</paragraph>
<paragraph id="D44D543AFF8DBC672CE5F9FD926E0EEF" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="479">
<emphasis id="E6868828FF8DBC672CE5F9FD923E0E22" bold="true" box="[199,266,1556,1581]" pageId="28" pageNumber="479">Notes</emphasis>
.The minute pronotal pits suggested this might be an exceptionally small species of
<emphasis id="E6868828FF8DBC6728D5F9FD96A90E22" box="[1271,1437,1556,1581]" italics="true" pageId="28" pageNumber="479">
<taxonomicName id="13F22FB9FF8DBC6728D5F9FD96AC0E22" box="[1271,1432,1556,1581]" class="Insecta" family="Scirtidae" genus="Heterocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="genus">Heterocyphon</taxonomicName>
.
</emphasis>
However, there are profound differences, e.g., several mandibular teeth in
<taxonomicName id="13F22FB9FF8DBC67286BF9DE96AC0E5F" authority="Watts 2009" authorityName="Watts" authorityYear="2009" box="[1097,1432,1591,1616]" class="Insecta" family="Scirtidae" genus="Heterocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="genus">
<emphasis id="E6868828FF8DBC67286BF9DE97D20E5F" box="[1097,1254,1591,1616]" italics="true" pageId="28" pageNumber="479">Heterocyphon</emphasis>
(
<bibRefCitation id="B06329CBFF8DBC672922F9D196A50E5F" author="Watts" box="[1280,1425,1592,1616]" pageId="28" pageNumber="479" refString="Watts, C. H. S. (2009) Revision of Heterocyphon Armstrong, Peneveronatus Armstrong and Accolabass gen. nov. (Scirtidae: Coleoptera). Transactions of the Royal Society of South Australia, 133 (1), 108 - 149." type="journal article" year="2009">Watts 2009</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="13F22FB9FF8DBC672CB5F9B592010E7A" box="[151,309,1628,1653]" class="Insecta" family="Scirtidae" genus="Heterocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="genus">
<emphasis id="E6868828FF8DBC672CB5F9B592010E7A" box="[151,309,1628,1653]" italics="true" pageId="28" pageNumber="479">Heterocyphon</emphasis>
</taxonomicName>
species (also the single western species,
<taxonomicName id="13F22FB9FF8DBC672F2EF9B490CB0E7B" authority="Watts" authorityName="Watts" box="[780,1023,1628,1653]" class="Insecta" family="Scirtidae" genus="Heterocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="E6868828FF8DBC672F2EF9B490830E7A" box="[780,951,1628,1653]" italics="true" pageId="28" pageNumber="479">H. occidentalis</emphasis>
Watts
</taxonomicName>
which I examined in ANIC) have a large lanceolate prosternal process, the long tip resting in an elongate U-shaped mesoventral groove. In side view the prosternal process appears bent because there is a stepwise decrease from the thick front part to the flat pilose posterior portion.
</paragraph>
<paragraph id="D44D543AFF8DBC672CE5F905900D0FB7" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="479">
Paired punctures (also called &quot;small depressed areas&quot; (
<bibRefCitation id="B06329CBFF8DBC672F12F90590DB0F0A" author="Armstrong" box="[816,1007,1772,1797]" pageId="28" pageNumber="479" refString="Armstrong, J. W. T. (1953) On Australian Helodidae (Coleoptera). 1. Descriptions of new genera and species. 1. Proceedings of the Linnean Society of New South Wales, 78, 19 - 32." type="journal article" year="1953">Armstrong 1953</bibRefCitation>
), &quot;pits, indentations&quot; (
<bibRefCitation id="B06329CBFF8DBC6728C9F90596440F0B" author="Watts" box="[1259,1392,1772,1796]" pageId="28" pageNumber="479" refString="Watts, C. H. S. (2009) Revision of Heterocyphon Armstrong, Peneveronatus Armstrong and Accolabass gen. nov. (Scirtidae: Coleoptera). Transactions of the Royal Society of South Australia, 133 (1), 108 - 149." type="journal article" year="2009">Watts 2009</bibRefCitation>
), or &quot;Porenpunkt&quot; (Klausnitzer 2013) at the base of the pronotum mark the point where the posterior and lateral carinae of the pronotum meet with the medial edge of the pronotal hypomeron on the underside. This detail is variably expressed in
<taxonomicName id="13F22FB9FF8DBC672D09F8BE91510F7F" authority="Watts 2009" authorityName="Watts" authorityYear="2009" box="[299,613,1879,1904]" class="Insecta" family="Scirtidae" genus="Heterocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="genus">
<emphasis id="E6868828FF8DBC672D09F8BE92FD0F7F" box="[299,457,1879,1904]" italics="true" pageId="28" pageNumber="479">Heterocyphon</emphasis>
(
<bibRefCitation id="B06329CBFF8DBC672DFAF8B1916A0F7F" author="Watts" box="[472,606,1880,1904]" pageId="28" pageNumber="479" refString="Watts, C. H. S. (2009) Revision of Heterocyphon Armstrong, Peneveronatus Armstrong and Accolabass gen. nov. (Scirtidae: Coleoptera). Transactions of the Royal Society of South Australia, 133 (1), 108 - 149." type="journal article" year="2009">Watts 2009</bibRefCitation>
)
</taxonomicName>
. It received not much attention in the past but was also mentioned in the Australian
<taxonomicName id="13F22FB9FF8DBC672D35F89491690F9B" authority="Watts 2009" authorityName="Watts" authorityYear="2009" box="[279,605,1916,1940]" class="Insecta" family="Scirtidae" genus="Peneveronatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="genus">
<emphasis id="E6868828FF8DBC672D35F89492880F9B" box="[279,444,1917,1940]" italics="true" pageId="28" pageNumber="479">Peneveronatus</emphasis>
(
<bibRefCitation id="B06329CBFF8DBC672DEDF89591620F9B" author="Watts" box="[463,598,1916,1940]" pageId="28" pageNumber="479" refString="Watts, C. H. S. (2009) Revision of Heterocyphon Armstrong, Peneveronatus Armstrong and Accolabass gen. nov. (Scirtidae: Coleoptera). Transactions of the Royal Society of South Australia, 133 (1), 108 - 149." type="journal article" year="2009">Watts 2009</bibRefCitation>
)
</taxonomicName>
and in the northern hemisphere genera
<emphasis id="E6868828FF8DBC67281AF89597910F9A" box="[1080,1189,1916,1941]" italics="true" pageId="28" pageNumber="479">Nyholmia</emphasis>
Klausnitzer,
<emphasis id="E6868828FF8DBC672961F89496A80F9B" box="[1347,1436,1917,1940]" italics="true" pageId="28" pageNumber="479">Exneria</emphasis>
Klausnitzer, and
<emphasis id="E6868828FF8DBC672D76F84992E40FB6" box="[340,464,1952,1977]" italics="true" pageId="28" pageNumber="479">Yoshitomia</emphasis>
Klausnitzer (Klausnitzer 2013).
</paragraph>
<paragraph id="D44D543AFF8DBC672CE5F82D96A80FD2" blockId="28.[151,1437,151,2013]" box="[199,1436,1988,2013]" pageId="28" pageNumber="479">
The female of
<taxonomicName id="13F22FB9FF8DBC672D4CF82C91330FD2" box="[366,519,1988,2013]" class="Insecta" family="Scirtidae" genus="Leptocyphon" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="furcalonga">
<emphasis id="E6868828FF8DBC672D4CF82C91330FD2" box="[366,519,1988,2013]" italics="true" pageId="28" pageNumber="479">L. furcalonga</emphasis>
</taxonomicName>
lacks hair patches on S4 described in females of
<taxonomicName id="13F22FB9FF8DBC672819F82D97EC0FD2" box="[1083,1240,1988,2013]" class="Insecta" family="Scirtidae" genus="Heterocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="genus">
<emphasis id="E6868828FF8DBC672819F82D97EC0FD2" box="[1083,1240,1988,2013]" italics="true" pageId="28" pageNumber="479">Heterocyphon</emphasis>
</taxonomicName>
which (from fig.
</paragraph>
<paragraph id="D44D543AFF8CBC662CB5FF7E90E108F7" blockId="29.[151,1436,151,320]" pageId="29" pageNumber="480">
<quantity id="130AF9DFFF8CBC662CB5FF7E93E308BF" box="[151,215,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="29" pageNumber="480" unit="in" value="16.0">16 in</quantity>
<bibRefCitation id="B06329CBFF8CBC662CC6FF71925808BF" author="Watts" box="[228,364,152,176]" pageId="29" pageNumber="480" refString="Watts, C. H. S. (2009) Revision of Heterocyphon Armstrong, Peneveronatus Armstrong and Accolabass gen. nov. (Scirtidae: Coleoptera). Transactions of the Royal Society of South Australia, 133 (1), 108 - 149." type="journal article" year="2009">Watts 2009</bibRefCitation>
) have articulated gonocoxites with an obliquely inserted subterminal stylite. There is no information on prehensors in
<taxonomicName id="13F22FB9FF8CBC662DC7FF5591B608DA" box="[485,642,188,213]" class="Insecta" family="Scirtidae" genus="Heterocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="genus">
<emphasis id="E6868828FF8CBC662DC7FF5591B608DA" box="[485,642,188,213]" italics="true" pageId="29" pageNumber="480">Heterocyphon</emphasis>
</taxonomicName>
.The rigid connection of the bacula to the sharply pointed gonocoxites with a miniature gonostyle suggests that eggs are laid into some substrate.
</paragraph>
</subSubSection>
<subSubSection id="9CE807B1FF8CBC662CE5FEED9174094F" pageId="29" pageNumber="480" type="etymology">
<paragraph id="D44D543AFF8CBC662CE5FEED9174094F" blockId="29.[151,1436,151,320]" pageId="29" pageNumber="480">
<emphasis id="E6868828FF8CBC662CE5FEED92720912" bold="true" box="[199,326,260,285]" pageId="29" pageNumber="480">Etymology</emphasis>
. The name is a noun in apposition composed of two Latin words meaning &quot;the long fork&quot;, describing the shape of the trigonium.
</paragraph>
</subSubSection>
</treatment>
</document>
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