377 lines
50 KiB
XML
377 lines
50 KiB
XML
<document id="A4C46BD489A6261A2F82BA74CFDE3C8C" ID-DOI="10.5281/zenodo.203260" ID-GBIF-Dataset="9ffb9edc-7879-4d55-932d-f50fe7be77ea" ID-ISSN="1175-5326" ID-Zenodo-Dep="203260" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460294051109" checkinUser="plazi" docAuthor="Schmelz, Rüdiger M., Collado, Rut & Römbke, Jörg" docDate="2011" docId="520EAA7DD648450DFF408067FB77FAB2" docLanguage="en" docName="zt02838p029.pdf" docOrigin="Zootaxa 2838" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Guaranidrilus cingulatus Schmelz, Collado & Römbke, 2011, sp. nov." docType="treatment" docVersion="9" lastPageNumber="10" masterDocId="AE37D205D64F4504FFD78220FFC2FF82" masterDocTitle="Mata Atlântica enchytraeids (Paraná, Brazil): A new genus, Xetadrilus gen. nov., with three new species, and four new species of Guaranidrilus Černosvitov (Enchytraeidae, Oligochaeta)" masterLastPageNumber="29" masterPageNumber="1" pageNumber="8" updateTime="1698251872398" updateUser="plazi">
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<mods:mods id="D8B9A854BEB35834A0B0AAA0B84AA319" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5DC87F82958BDE0BBCFF00F1CB8B30D3">
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<mods:title id="C99C70B25482357FB52C7EC6DF10B5DE">Mata Atlântica enchytraeids (Paraná, Brazil): A new genus, Xetadrilus gen. nov., with three new species, and four new species of Guaranidrilus Černosvitov (Enchytraeidae, Oligochaeta)</mods:title>
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</mods:titleInfo>
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<mods:name id="EE4C38F5EB200A834F879C2477EB7792" type="personal">
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<mods:role id="D0D92F493D74A6ED0391B022947EBFD4">
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<mods:roleTerm id="2285BBC0D62403646516948E3E8B1240">Author</mods:roleTerm>
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<mods:namePart id="DBA4CD2F1053845F8D0A0F789F7E9E17">Schmelz, Rüdiger M.</mods:namePart>
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<mods:roleTerm id="5B9252C4403D062A71F259E0E9B75EB6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="192809BC3E9E314D956741EE20D7962E">Collado, Rut</mods:namePart>
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</mods:name>
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<mods:name id="DA3BBB3A2E2C8B9760D8C2690A5E1A94" type="personal">
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<mods:roleTerm id="018C979986B2AC06BA36B5F0357ACCDD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="703E9606E0EC78FCAC4015E95BFC5F84">Römbke, Jörg</mods:namePart>
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<mods:typeOfResource id="ED02D0D71D4013C9BA767D380F8F4B20">text</mods:typeOfResource>
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<mods:title id="DF96AACF285463E31A84620BDAD12A83">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F4BC7908D70BF8A128E900F7B364E4DD">
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<mods:date id="FA86FA750FC47E7CF23142AF20ED1EE7">2011</mods:date>
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<mods:detail id="20212B9C455CF6C04A2631DAC465ECD6" type="volume">
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<mods:number id="894B2CC2B4E10266BA83D1C7444532E8">2838</mods:number>
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<mods:classification id="3AC6603FD1FA61F0E370C458B55CA4FD">journal article</mods:classification>
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<mods:identifier id="E5D9D285DCE480FA0AA24E7E4282AF0C" type="DOI">10.5281/zenodo.203260</mods:identifier>
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<mods:identifier id="62209E0EF35096B1B8B6CD098C777A69" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E995EE352FD2BB1999D78CAF673B4BE1" type="Zenodo-Dep">203260</mods:identifier>
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<treatment id="520EAA7DD648450DFF408067FB77FAB2" ID-DOI="http://doi.org/10.5281/zenodo.5611823" ID-GBIF-Taxon="119399035" ID-Zenodo-Dep="5611823" LSID="urn:lsid:plazi:treatment:520EAA7DD648450DFF408067FB77FAB2" httpUri="http://treatment.plazi.org/id/520EAA7DD648450DFF408067FB77FAB2" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
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<subSubSection id="92BD48E0D6484503FF408067FEDCFD00" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="DA181B6BD6484503FF408067FDEDFDE3" blockId="7.[151,559,583,642]" box="[151,559,583,609]" pageId="7" pageNumber="8">
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<heading id="8150AC07D6484503FF408067FDEDFDE3" bold="true" box="[151,559,583,609]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
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<emphasis id="E8D3C779D6484503FF408067FDEDFDE3" bold="true" box="[151,559,583,609]" pageId="7" pageNumber="8">
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<taxonomicName id="1DA760E8D6484503FF408067FE08FDE3" ID-CoL="852NR" box="[151,458,583,609]" class="Clitellata" family="Enchytraeidae" genus="Guaranidrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="cingulatus" status="sp. nov.">
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<emphasis id="E8D3C779D6484503FF408067FE08FDE3" bold="true" box="[151,458,583,609]" italics="true" pageId="7" pageNumber="8">Guaranidrilus cingulatus</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="F3E07A02D6484503FE068067FDEDFDE3" box="[465,559,583,609]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="DA181B6BD6484503FF40804AFEDCFD00" blockId="7.[151,559,583,642]" box="[151,286,618,642]" pageId="7" pageNumber="8">
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(
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<figureCitation id="429C07EED6484503FF48804AFF25FD00" box="[159,231,618,642]" captionStart="FIGURE 3" captionStartId="8.[151,250,1559,1582]" captionTargetBox="[221,1358,196,1530]" captionTargetId="figure@8.[220,1366,193,1538]" captionTargetPageId="8" captionText="FIGURE 3. Guaranidrilus cingulatus sp. nov. A. Segments IV – IX, lateral view. B. Anterior (left) and caudal (right) chaeta. C. Segments IV – VIII, dorsal to dorso-lateral view. D. Spermatheca. E. Clitellum dorsally. F. Preclitellar nephridium. G. Postclitellar nephridium. H. Male efferent apparatus. Male pore in ventral view. I. Male pore in lateral view, same specimen as in H. All drawings from whole mounts." httpUri="https://zenodo.org/record/203263/files/figure.png" pageId="7" pageNumber="8">Figs 3</figureCitation>
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,
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<figureCitation id="429C07EED6484503FF24804AFEC2FD00" box="[243,256,618,642]" captionStart="FIGURE 8" captionStartId="24.[151,250,1826,1849]" captionTargetBox="[342,1223,190,1783]" captionTargetId="figure@24.[342,1250,174,1802]" captionTargetPageId="24" captionText="FIGURE 8. Schematic representation of taxonomic differences of species described here. The following traits are shown: Genus level: Chaetal pattern, shape of epidermal gland cells (eg, only shown in a few anterior segments, except in E), presence / absence of prostomial ganglion (g) in front of brain (br), prostomial papillae (p), and frontal recess in prostomial epithelium (r). Species level: Distribution of epidermal gland cells, pattern of secondary ventral pharyngeal gland lobes (pg), presence / absence of oesophageal appendages (oa) and intestinal diverticula (id), extension of the latter, distribution of preclitellar nephridia (ne). A. Guaranidrilus andreolii sp. nov., B. G. m a rq u e s i sp. nov. C. G. cingulatus sp. nov. D. G. h o e f e r i sp. nov. E. Xetadrilus maacki sp. nov. F. X. aphanus sp. nov. G. X. fabryi sp. nov." httpUri="https://zenodo.org/record/203268/files/figure.png" pageId="7" pageNumber="8">8</figureCitation>
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C)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="92BD48E0D6484503FF408093FA9FFAAC" pageId="7" pageNumber="8" type="materials_examined">
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<paragraph id="DA181B6BD6484503FF408093FC57FD6C" blockId="7.[151,1437,690,1939]" pageId="7" pageNumber="8">
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<emphasis id="E8D3C779D6484503FF408093FEC6FD49" bold="true" box="[151,260,691,715]" pageId="7" pageNumber="8">
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<typeStatus id="051CA5C9D6484503FF408093FEC6FD49" box="[151,260,691,715]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
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</emphasis>
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||
.
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||
<collectionCode id="BCB683AED6484503FEC48092FEB2FD48" box="[275,368,690,714]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="8">MZUSP</collectionCode>
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1225, adult specimen, stained whole mount,
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<taxonomicName id="1DA760E8D6484503FCA58092FB9CFD48" authority="Cachoeira" authorityName="Cachoeira" box="[882,1118,690,714]" class="Insecta" family="Pseudococcidae" genus="Antonina" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Antonina, Cachoeira</taxonomicName>
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||
, medium old secondary forest on Cambisol [site 17],
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||
<date id="AE193DABD6484503FE6A80F7FDEFFD6C" box="[445,557,726,750]" pageId="7" pageNumber="8" value="2004-03">Mar 2004</date>
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||
, leg. B. Förster, R. M. Schmelz.
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||
</paragraph>
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||
<paragraph id="DA181B6BD6484503FF1280DBFC97FC90" blockId="7.[151,1437,690,1939]" box="[197,853,762,787]" pageId="7" pageNumber="8">
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<emphasis id="E8D3C779D6484503FF1280DBFEFFFC91" bold="true" box="[197,317,763,787]" pageId="7" pageNumber="8">
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<typeStatus id="051CA5C9D6484503FF1280DBFEFFFC91" box="[197,317,763,787]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
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</emphasis>
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.
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||
<collectionCode id="BCB683AED6484503FE9C80DAFE6AFC90" box="[331,424,762,786]" pageId="7" pageNumber="8">MZSUP</collectionCode>
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,
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<specimenCount id="CCA1D0E2D6484503FE6480DAFD8FFC90" box="[435,589,762,786]" pageId="7" pageNumber="8" type="generic">14 specimens</specimenCount>
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, stained whole mounts:
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||
</paragraph>
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<paragraph id="DA181B6BD6484503FF12813EFCCDFCD8" blockId="7.[151,1437,690,1939]" pageId="7" pageNumber="8">
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<collectionCode id="BCB683AED6484503FF12813EFEE0FCB4" box="[197,290,798,822]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="8">MZUSP</collectionCode>
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1226,
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||
<specimenCount id="CCA1D0E2D6484503FEA6813EFDC2FCB4" box="[369,512,798,822]" pageId="7" pageNumber="8" type="generic">6 specimens</specimenCount>
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,
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||
<taxonomicName id="1DA760E8D6484503FDDB813EFD3AFCB4" authority="Cachoeira" authorityName="Cachoeira" box="[524,760,798,822]" class="Insecta" family="Pseudococcidae" genus="Antonina" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Antonina, Cachoeira</taxonomicName>
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,
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<geoCoordinate id="BF937DACD6484503FCD3813EFCBFFCB4" box="[772,893,798,822]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="15" value="-25.333055">25°19'59"S</geoCoordinate>
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–
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<geoCoordinate id="BF937DACD6484503FC5A813EFBD9FCB4" box="[909,1051,798,822]" direction="west" orientation="longitude" pageId="7" pageNumber="8" precision="15" value="-48.700558">48°42'02"W</geoCoordinate>
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||
, young secondary forest on Gleysol [site 25],
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||
<date id="AE193DABD6484503FEFE8163FE5CFCD8" box="[297,414,834,859]" pageId="7" pageNumber="8" value="2003-05">May 2003</date>
|
||
, leg. J. Römbke, R. M. Schmelz.
|
||
</paragraph>
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||
<paragraph id="DA181B6BD6484503FF128146FCC7FCFC" blockId="7.[151,1437,690,1939]" box="[197,773,870,894]" pageId="7" pageNumber="8">
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<collectionCode id="BCB683AED6484503FF128146FEE0FCFC" box="[197,290,870,894]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="8">MZUSP</collectionCode>
|
||
1227,
|
||
<specimenCount id="CCA1D0E2D6484503FEB88147FE3EFCFC" box="[367,508,870,894]" pageId="7" pageNumber="8" type="generic">7 specimens</specimenCount>
|
||
, same data as
|
||
<typeStatus id="051CA5C9D6484503FD4C8146FCC3FCFC" box="[667,769,870,894]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
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.
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||
</paragraph>
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||
<paragraph id="DA181B6BD6484503FF1281AAFA45FC44" blockId="7.[151,1437,690,1939]" pageId="7" pageNumber="8">
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<collectionCode id="BCB683AED6484503FF1281AAFEE0FC20" box="[197,290,906,930]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="8">MZUSP</collectionCode>
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||
1228,
|
||
<specimenCount id="CCA1D0E2D6484503FEAB81AAFDCDFC20" box="[380,527,906,930]" pageId="7" pageNumber="8" type="generic">3 specimens</specimenCount>
|
||
,
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||
<taxonomicName id="1DA760E8D6484503FDF781AAFCD0FC20" authority="Cachoeira" authorityName="Cachoeira" box="[544,786,906,930]" class="Insecta" family="Pseudococcidae" genus="Antonina" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Antonina, Cachoeira</taxonomicName>
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||
, 25°19'04"–41''S, 48°40'36"–41'51"W,
|
||
<quantity id="1D5FB68ED6484503FB0B81AAFA91FC20" box="[1244,1363,906,930]" metricMagnitude="1" metricUnit="m" metricValue="7.0" metricValueMax="12.0" metricValueMin="2.0" pageId="7" pageNumber="8" unit="m" value="70.0" valueMax="120.0" valueMin="20.0">20–120 m</quantity>
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a.s.l., young and medium old secondary forests on Gleysol [sites 12, 24, 30],
|
||
<date id="AE193DABD6484503FC61818EFBE2FC44" box="[950,1056,942,966]" pageId="7" pageNumber="8" value="2004-10">Oct 2004</date>
|
||
, leg. B. Förster, R. M. Schmelz.
|
||
</paragraph>
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||
<paragraph id="DA181B6BD6484503FF1281F2FC00FB8C" blockId="7.[151,1437,690,1939]" pageId="7" pageNumber="8">
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<collectionCode id="BCB683AED6484503FF1281F2FEE0FC68" box="[197,290,978,1002]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="8">MZUSP</collectionCode>
|
||
1229,
|
||
<specimenCount id="CCA1D0E2D6484503FEA781F2FE30FC68" box="[368,498,978,1002]" pageId="7" pageNumber="8" type="generic">1 specimen</specimenCount>
|
||
, Curitiba outskirts,
|
||
<geoCoordinate id="BF937DACD6484503FD0781F2FC92FC68" box="[720,848,978,1002]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="15" value="-25.320278">25°19'13''S</geoCoordinate>
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,
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||
<geoCoordinate id="BF937DACD6484503FC8B81F2FC27FC68" box="[860,997,978,1002]" direction="west" orientation="longitude" pageId="7" pageNumber="8" precision="15" value="-49.10611">49°06'22''W</geoCoordinate>
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,
|
||
<quantity id="1D5FB68ED6484503FC2681F2FBFAFC68" box="[1009,1080,978,1002]" metricMagnitude="2" metricUnit="m" metricValue="9.57" pageId="7" pageNumber="8" unit="m" value="957.0">957 m</quantity>
|
||
a.s.l.,
|
||
<taxonomicName id="1DA760E8D6484503FB5581F3FB37FC69" box="[1154,1269,979,1003]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="genus">
|
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<emphasis id="E8D3C779D6484503FB5581F3FB37FC69" box="[1154,1269,979,1003]" italics="true" pageId="7" pageNumber="8">Araucaria</emphasis>
|
||
</taxonomicName>
|
||
forest on Podzol, property of R. Marques,
|
||
<date id="AE193DABD6484503FE0B81D6FD84FB8C" box="[476,582,1014,1038]" pageId="7" pageNumber="8" value="2004-10">Oct 2004</date>
|
||
, leg. R. Marques, R. M. Schmelz.
|
||
</paragraph>
|
||
<paragraph id="DA181B6BD6484503FF12863BFB69FBB0" blockId="7.[151,1437,690,1939]" box="[197,1195,1050,1074]" pageId="7" pageNumber="8">
|
||
<collectionCode id="BCB683AED6484503FF12863BFECCFBB0" box="[197,270,1051,1074]" pageId="7" pageNumber="8">UFPR</collectionCode>
|
||
,
|
||
<specimenCount id="CCA1D0E2D6484503FECE863AFE67FBB0" box="[281,421,1050,1074]" pageId="7" pageNumber="8" type="generic">6 specimens</specimenCount>
|
||
, 3 stained whole mounts, 3 ethanol-preserved, Guarequeçaba, Itaqui:
|
||
</paragraph>
|
||
<paragraph id="DA181B6BD6484503FF10861FFD29FBF8" blockId="7.[151,1437,690,1939]" pageId="7" pageNumber="8">
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||
<collectionCode id="BCB683AED6484503FF10861FFECFFBD4" box="[199,269,1087,1110]" pageId="7" pageNumber="8">UFPR</collectionCode>
|
||
OL-10,
|
||
<specimenCount id="CCA1D0E2D6484503FEB9861EFE38FBD4" box="[366,506,1086,1110]" pageId="7" pageNumber="8" type="generic">2 specimens</specimenCount>
|
||
,
|
||
<geoCoordinate id="BF937DACD6484503FDD1861EFD5EFBD4" box="[518,668,1086,1110]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="-25.245916">25°14'45.3''S</geoCoordinate>
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||
,
|
||
<geoCoordinate id="BF937DACD6484503FD7F861EFC84FBD4" box="[680,838,1086,1110]" direction="west" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="-48.50539">48°30'19.4''W</geoCoordinate>
|
||
,
|
||
<quantity id="1D5FB68ED6484503FC85861EFC49FBD4" box="[850,907,1086,1110]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="7" pageNumber="8" unit="m" value="27.0">27 m</quantity>
|
||
a.s.l., medium old secondary forest on Cambisol [site 38],
|
||
<date id="AE193DABD6484503FEFE8642FE57FBF8" box="[297,405,1122,1146]" pageId="7" pageNumber="8" value="2007-09">Sep 2007</date>
|
||
, leg. P. Heine, R. M. Schmelz.
|
||
</paragraph>
|
||
<paragraph id="DA181B6BD6484503FF1286A7FD9BFB40" blockId="7.[151,1437,690,1939]" pageId="7" pageNumber="8">
|
||
<collectionCode id="BCB683AED6484503FF1286A7FEC9FB1C" box="[197,267,1159,1182]" pageId="7" pageNumber="8">UFPR</collectionCode>
|
||
OL-11,
|
||
<specimenCount id="CCA1D0E2D6484503FEBA86A6FE32FB1C" box="[365,496,1158,1182]" pageId="7" pageNumber="8" type="generic">1 specimen</specimenCount>
|
||
,
|
||
<geoCoordinate id="BF937DACD6484503FE2A86A6FD50FB1C" box="[509,658,1158,1182]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="-25.308832">25°18'31.8''S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="BF937DACD6484503FD4886A6FCFFFB1C" box="[671,829,1158,1182]" direction="west" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="-48.46511">48°27'54,4''W</geoCoordinate>
|
||
,
|
||
<quantity id="1D5FB68ED6484503FC9D86A6FC47FB1C" box="[842,901,1158,1182]" metricMagnitude="1" metricUnit="m" metricValue="9.3" pageId="7" pageNumber="8" unit="m" value="93.0">93 m</quantity>
|
||
a.s.l., old-growth forest on Cambisol [site 40],
|
||
<date id="AE193DABD6484503FF40868AFEC6FB40" box="[151,260,1194,1218]" pageId="7" pageNumber="8" value="2007-09">Sep 2007</date>
|
||
. leg. P. Heine, R. M. Schmelz.
|
||
</paragraph>
|
||
<paragraph id="DA181B6BD6484503FF1286EFFC77FA88" blockId="7.[151,1437,690,1939]" pageId="7" pageNumber="8">
|
||
<collectionCode id="BCB683AED6484503FF1286EFFEC9FB64" box="[197,267,1231,1254]" pageId="7" pageNumber="8">UFPR</collectionCode>
|
||
OL-12:
|
||
<specimenCount id="CCA1D0E2D6484503FEBB86EEFE38FB64" box="[364,506,1230,1254]" pageId="7" pageNumber="8" type="generic">3 specimens</specimenCount>
|
||
, ethanol-preserved,
|
||
<geoCoordinate id="BF937DACD6484503FD0B86EEFCB3FB64" box="[732,881,1230,1254]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="-25.247528">25°14'51,1''S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="BF937DACD6484503FCAA86EEFBD9FB64" box="[893,1051,1230,1254]" direction="west" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="-48.492054">48°29'31.4''W</geoCoordinate>
|
||
,
|
||
<quantity id="1D5FB68ED6484503FBF086EEFB90FB64" box="[1063,1106,1230,1254]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="8" unit="m" value="8.0">8 m</quantity>
|
||
a.s.l., medium old secondary forest on Cambisol [site 39],
|
||
<date id="AE193DABD6484503FE0A86D3FD87FA88" box="[477,581,1266,1290]" pageId="7" pageNumber="8" value="2008-01">Jan 2008</date>
|
||
, leg. J. Römbke, R. M. Schmelz.
|
||
</paragraph>
|
||
<paragraph id="DA181B6BD6484503FF108736FA9FFAAC" blockId="7.[151,1437,690,1939]" box="[199,1373,1302,1327]" pageId="7" pageNumber="8">
|
||
<emphasis id="E8D3C779D6484503FF108736FE78FAAD" bold="true" box="[199,442,1302,1327]" pageId="7" pageNumber="8">Additional material.</emphasis>
|
||
Eight specimens from Cachoeira and Itaqui sites, examined in vivo, not preserved.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92BD48E0D6484503FF10871BFF22FAF5" pageId="7" pageNumber="8" type="etymology">
|
||
<paragraph id="DA181B6BD6484503FF10871BFF22FAF5" blockId="7.[151,1437,690,1939]" pageId="7" pageNumber="8">
|
||
<emphasis id="E8D3C779D6484503FF10871BFE8CFAD1" bold="true" box="[199,334,1339,1363]" pageId="7" pageNumber="8">Etymology.</emphasis>
|
||
'cingulatus' (lat.) = girded; named in reference to the girdle-shaped clitellum, so far unique in the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92BD48E0D648450DFF1087A2FAA9FD06" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="description">
|
||
<paragraph id="DA181B6BD6484503FF1087A2FB59F95C" blockId="7.[151,1437,690,1939]" pageId="7" pageNumber="8">
|
||
<emphasis id="E8D3C779D6484503FF1087A2FDE8FA19" bold="true" box="[199,554,1410,1435]" pageId="7" pageNumber="8">Description. Body dimensions</emphasis>
|
||
. Small species, living adults ca.
|
||
<quantity id="1D5FB68ED6484503FC4087A2FBCBFA18" box="[919,1033,1410,1434]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="5.0" metricValueMin="3.5" pageId="7" pageNumber="8" unit="mm" value="4.25" valueMax="5.0" valueMin="3.5">3.5–5 mm</quantity>
|
||
long and
|
||
<quantity id="1D5FB68ED6484503FBA087A2FACEFA18" box="[1143,1292,1410,1434]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="7" pageNumber="8" unit="mm" value="0.125" valueMax="0.15" valueMin="0.1">0.1–0.15 mm</quantity>
|
||
wide, whole mounts ca.
|
||
<quantity id="1D5FB68ED6484503FECC8786FE4CFA3C" box="[283,398,1446,1470]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="7" pageNumber="8" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.5–4 mm</quantity>
|
||
long and
|
||
<quantity id="1D5FB68ED6484503FDD58786FD5BFA3C" box="[514,665,1446,1470]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="7" pageNumber="8" unit="mm" value="0.175" valueMax="0.2" valueMin="0.15">0.15–0.2 mm</quantity>
|
||
wide.
|
||
<emphasis id="E8D3C779D6484503FD3F8786FC74FA3D" bold="true" box="[744,950,1446,1471]" pageId="7" pageNumber="8">Segment number</emphasis>
|
||
26–32.
|
||
<emphasis id="E8D3C779D6484503FBC38786FBB4FA3D" bold="true" box="[1044,1142,1446,1471]" pageId="7" pageNumber="8">Chaetae</emphasis>
|
||
(
|
||
<figureCitation id="429C07EED6484503FB508786FB08FA3C" box="[1159,1226,1446,1470]" captionStart="FIGURE 3" captionStartId="8.[151,250,1559,1582]" captionTargetBox="[221,1358,196,1530]" captionTargetId="figure@8.[220,1366,193,1538]" captionTargetPageId="8" captionText="FIGURE 3. Guaranidrilus cingulatus sp. nov. A. Segments IV – IX, lateral view. B. Anterior (left) and caudal (right) chaeta. C. Segments IV – VIII, dorsal to dorso-lateral view. D. Spermatheca. E. Clitellum dorsally. F. Preclitellar nephridium. G. Postclitellar nephridium. H. Male efferent apparatus. Male pore in ventral view. I. Male pore in lateral view, same specimen as in H. All drawings from whole mounts." httpUri="https://zenodo.org/record/203263/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
|
||
B) two per bundle throughout, absent at XII, straight distally. In caudal segments chaetae 1.5–
|
||
<date id="AE193DABD6484503FC3A87EAFC39FA60" box="[1005,1019,1482,1506]" pageId="7" pageNumber="8">2</date>
|
||
x as large as largest anterior chaetae, equally large ventrally and laterally (ca. 60 µm long and 4–5 µm wide); in anterior segments laterals smaller than ventrals (30 µm vs. 40 µm).
|
||
<emphasis id="E8D3C779D6484503FE358432FD28F9A9" bold="true" box="[482,746,1554,1579]" pageId="7" pageNumber="8">Epidermal gland cells</emphasis>
|
||
invisible in vivo; in well-preserved whole mounts two rows distinguishable of inconspicuous, transversely elongate cells in dorsal half of anterior segments.
|
||
<emphasis id="E8D3C779D6484503FB3A8416FA9FF9CD" bold="true" box="[1261,1373,1590,1615]" pageId="7" pageNumber="8">Clitellum</emphasis>
|
||
(
|
||
<figureCitation id="429C07EED6484503FAB98416FF66F9F0" captionStart="FIGURE 3" captionStartId="8.[151,250,1559,1582]" captionTargetBox="[221,1358,196,1530]" captionTargetId="figure@8.[220,1366,193,1538]" captionTargetPageId="8" captionText="FIGURE 3. Guaranidrilus cingulatus sp. nov. A. Segments IV – IX, lateral view. B. Anterior (left) and caudal (right) chaeta. C. Segments IV – VIII, dorsal to dorso-lateral view. D. Spermatheca. E. Clitellum dorsally. F. Preclitellar nephridium. G. Postclitellar nephridium. H. Male efferent apparatus. Male pore in ventral view. I. Male pore in lateral view, same specimen as in H. All drawings from whole mounts." httpUri="https://zenodo.org/record/203263/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
|
||
E) well-developed, perfectly girdle-shaped, cells in ca. 34 transverse rows ventrally, dorsally rows indefinite when fully developed, here hyalocytes and granulocytes alternating, granulocytes smaller than hyalocytes and more numerous; ventrally only granulocytes; granulocytes almost square-shaped, giving a 'pavement'-like aspect.
|
||
<emphasis id="E8D3C779D6484503FF4084E7FED1F95D" bold="true" box="[151,275,1735,1759]" pageId="7" pageNumber="8">Head pore</emphasis>
|
||
on prostomium mid-dorsally, prostomial inner papillae inconspicuous or absent.
|
||
</paragraph>
|
||
<paragraph id="DA181B6BD648450DFF1084CBFACFFE9E" blockId="7.[151,1437,690,1939]" lastBlockId="9.[151,1437,151,1328]" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
|
||
<emphasis id="E8D3C779D6484503FF1084CBFEFCF881" bold="true" box="[199,318,1771,1795]" pageId="7" pageNumber="8">Body wall</emphasis>
|
||
thick in vivo, 5–10 µm thick in whole mounts, longitudinal muscle layer well-developed, especially in adults.
|
||
<emphasis id="E8D3C779D6484503FED3852EFE98F8A5" bold="true" box="[260,346,1806,1831]" pageId="7" pageNumber="8">Cuticle</emphasis>
|
||
distinguishable, ca. 1.5 µm thick.
|
||
<emphasis id="E8D3C779D6484503FD0E852EFCDEF8A5" bold="true" box="[729,796,1806,1831]" pageId="7" pageNumber="8">
|
||
<collectingCountry id="A2B05BFBD6484503FD0E852EFCDEF8A5" box="[729,796,1806,1831]" name="Spain" pageId="7" pageNumber="8">Septa</collectingCountry>
|
||
</emphasis>
|
||
slightly thickened in preclitellar segments, from 4/5 to 5/ 6 or 8/9.
|
||
<emphasis id="E8D3C779D6484503FF2B8512FE82F8C9" bold="true" box="[252,320,1842,1867]" pageId="7" pageNumber="8">Brain</emphasis>
|
||
concave posteriorly, twice as long as wide (length ca. 70 µm, fix), prostomial ganglia very small or absent. Ventral nerve cord with suboesophageal ganglion in II–IV and segmental ganglia from V on.
|
||
<emphasis id="E8D3C779D6484503FAD38576FEE1F811" bold="true" pageId="7" pageNumber="8">Oesophageal appendages</emphasis>
|
||
(
|
||
<figureCitation id="429C07EED6484503FEEE855AFEB8F810" box="[313,378,1914,1938]" captionStart="FIGURE 3" captionStartId="8.[151,250,1559,1582]" captionTargetBox="[221,1358,196,1530]" captionTargetId="figure@8.[220,1366,193,1538]" captionTargetPageId="8" captionText="FIGURE 3. Guaranidrilus cingulatus sp. nov. A. Segments IV – IX, lateral view. B. Anterior (left) and caudal (right) chaeta. C. Segments IV – VIII, dorsal to dorso-lateral view. D. Spermatheca. E. Clitellum dorsally. F. Preclitellar nephridium. G. Postclitellar nephridium. H. Male efferent apparatus. Male pore in ventral view. I. Male pore in lateral view, same specimen as in H. All drawings from whole mounts." httpUri="https://zenodo.org/record/203263/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
|
||
A,C) in anterior of VI, paired, spherical in dorsal view, sessile, side by side dorsally on gut wall, not larger than secondary ventral pharyngeal gland lobes.
|
||
<emphasis id="E8D3C779D647450CFCB18496FB87F94C" bold="true" box="[870,1093,1718,1742]" pageId="8" pageNumber="9">Pharyngeal glands</emphasis>
|
||
(
|
||
<figureCitation id="429C07EED647450CFB828496FB5AF94C" box="[1109,1176,1718,1742]" captionStart="FIGURE 3" captionStartId="8.[151,250,1559,1582]" captionTargetBox="[221,1358,196,1530]" captionTargetId="figure@8.[220,1366,193,1538]" captionTargetPageId="8" captionText="FIGURE 3. Guaranidrilus cingulatus sp. nov. A. Segments IV – IX, lateral view. B. Anterior (left) and caudal (right) chaeta. C. Segments IV – VIII, dorsal to dorso-lateral view. D. Spermatheca. E. Clitellum dorsally. F. Preclitellar nephridium. G. Postclitellar nephridium. H. Male efferent apparatus. Male pore in ventral view. I. Male pore in lateral view, same specimen as in H. All drawings from whole mounts." httpUri="https://zenodo.org/record/203263/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
|
||
A,C) dorsally unpaired in IV, V, VI; primary and secondary ventral lobes in V and VI. Primary ventral lobes of VI largest.
|
||
<emphasis id="E8D3C779D647450CFAFD84F9FEDAF894" bold="true" pageId="8" pageNumber="9">Intestinal diverticula</emphasis>
|
||
(
|
||
<figureCitation id="429C07EED647450CFEFE84DEFEAFF894" box="[297,365,1790,1814]" captionStart="FIGURE 3" captionStartId="8.[151,250,1559,1582]" captionTargetBox="[221,1358,196,1530]" captionTargetId="figure@8.[220,1366,193,1538]" captionTargetPageId="8" captionText="FIGURE 3. Guaranidrilus cingulatus sp. nov. A. Segments IV – IX, lateral view. B. Anterior (left) and caudal (right) chaeta. C. Segments IV – VIII, dorsal to dorso-lateral view. D. Spermatheca. E. Clitellum dorsally. F. Preclitellar nephridium. G. Postclitellar nephridium. H. Male efferent apparatus. Male pore in ventral view. I. Male pore in lateral view, same specimen as in H. All drawings from whole mounts." httpUri="https://zenodo.org/record/203263/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
|
||
A,C) paired, lateral to intestine, confined to VII, occupying entire segment length, rarely less (
|
||
<figureCitation id="429C07EED647450CFF488502FF20F8B8" box="[159,226,1826,1850]" captionStart="FIGURE 3" captionStartId="8.[151,250,1559,1582]" captionTargetBox="[221,1358,196,1530]" captionTargetId="figure@8.[220,1366,193,1538]" captionTargetPageId="8" captionText="FIGURE 3. Guaranidrilus cingulatus sp. nov. A. Segments IV – IX, lateral view. B. Anterior (left) and caudal (right) chaeta. C. Segments IV – VIII, dorsal to dorso-lateral view. D. Spermatheca. E. Clitellum dorsally. F. Preclitellar nephridium. G. Postclitellar nephridium. H. Male efferent apparatus. Male pore in ventral view. I. Male pore in lateral view, same specimen as in H. All drawings from whole mounts." httpUri="https://zenodo.org/record/203263/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
|
||
A), separate from each other; each with large anterior and small stump-like posterior projection, the latter situated dorso-laterally, extending slightly into VIII in few specimens (
|
||
<figureCitation id="429C07EED647450CFC108566FBCEF8DC" box="[967,1036,1862,1886]" captionStart="FIGURE 3" captionStartId="8.[151,250,1559,1582]" captionTargetBox="[221,1358,196,1530]" captionTargetId="figure@8.[220,1366,193,1538]" captionTargetPageId="8" captionText="FIGURE 3. Guaranidrilus cingulatus sp. nov. A. Segments IV – IX, lateral view. B. Anterior (left) and caudal (right) chaeta. C. Segments IV – VIII, dorsal to dorso-lateral view. D. Spermatheca. E. Clitellum dorsally. F. Preclitellar nephridium. G. Postclitellar nephridium. H. Male efferent apparatus. Male pore in ventral view. I. Male pore in lateral view, same specimen as in H. All drawings from whole mounts." httpUri="https://zenodo.org/record/203263/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
|
||
C); connections of diverticula with intestine in posterior of VII; outer and inner surface often irregularly lobed or wavy; walls ca. 7–10 µm thick, no radial striation.
|
||
<emphasis id="E8D3C779D647450CFE9E85AEFDEDF824" bold="true" box="[329,559,1934,1958]" pageId="8" pageNumber="9">Dorsal blood vessel</emphasis>
|
||
from
|
||
<date id="AE193DABD647450CFDA385AFFD72F824" box="[628,688,1934,1958]" bridgedPair="-" pageId="8" pageNumber="9">X – 1</date>
|
||
/
|
||
<date id="AE193DABD647450CFD6D85AEFCC2F824" box="[698,768,1934,1958]" pageId="8" pageNumber="9">2 XII.</date>
|
||
<emphasis id="E8D3C779D647450CFCD085ADFBC3F824" bold="true" box="[775,1025,1933,1958]" pageId="8" pageNumber="9">Preclitellar nephrida</emphasis>
|
||
(
|
||
<figureCitation id="429C07EED647450CFBC685AEFB91F824" box="[1041,1107,1934,1958]" captionStart="FIGURE 3" captionStartId="8.[151,250,1559,1582]" captionTargetBox="[221,1358,196,1530]" captionTargetId="figure@8.[220,1366,193,1538]" captionTargetPageId="8" captionText="FIGURE 3. Guaranidrilus cingulatus sp. nov. A. Segments IV – IX, lateral view. B. Anterior (left) and caudal (right) chaeta. C. Segments IV – VIII, dorsal to dorso-lateral view. D. Spermatheca. E. Clitellum dorsally. F. Preclitellar nephridium. G. Postclitellar nephridium. H. Male efferent apparatus. Male pore in ventral view. I. Male pore in lateral view, same specimen as in H. All drawings from whole mounts." httpUri="https://zenodo.org/record/203263/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
|
||
F) 3 pairs, at 6/7 – 8/9, occasionally absent at either 6/7 or 8/9. Anteseptale slightly longer than wide, with ca. 1/3 of entire nephridial body, postseptale slightly bulging dorsad, with dorsal vesicle, gradually merging into short efferent duct, terminal vesicle absent; ante- and postseptale separated by a slight constriction at level of septa.
|
||
<emphasis id="E8D3C779D646450DFBC882B7FAE9FF32" bold="true" box="[1055,1323,151,176]" pageId="9" pageNumber="10">Postclitellar nephridia</emphasis>
|
||
(
|
||
<figureCitation id="429C07EED646450DFAEC82B7FABFFF2D" box="[1339,1405,151,175]" captionStart="FIGURE 3" captionStartId="8.[151,250,1559,1582]" captionTargetBox="[221,1358,196,1530]" captionTargetId="figure@8.[220,1366,193,1538]" captionTargetPageId="8" captionText="FIGURE 3. Guaranidrilus cingulatus sp. nov. A. Segments IV – IX, lateral view. B. Anterior (left) and caudal (right) chaeta. C. Segments IV – VIII, dorsal to dorso-lateral view. D. Spermatheca. E. Clitellum dorsally. F. Preclitellar nephridium. G. Postclitellar nephridium. H. Male efferent apparatus. Male pore in ventral view. I. Male pore in lateral view, same specimen as in H. All drawings from whole mounts." httpUri="https://zenodo.org/record/203263/files/figure.png" pageId="9" pageNumber="10">Fig. 3</figureCitation>
|
||
G) present in less than half of all positions; shape as in preclitellar nephridia, but more elongate.
|
||
<emphasis id="E8D3C779D646450DFB20829BFA5EFF56" bold="true" box="[1271,1436,187,212]" pageId="9" pageNumber="10">Coelomocytes</emphasis>
|
||
slightly longer than wide, 17–28 µm long, with very fine wavy or groovy texture, no granula distinguished; cell margins smooth, refractile in oblique position; cells abundant but not obscuring other inner structures.
|
||
</paragraph>
|
||
<caption id="8ED84BE3D647450CFF408437FE05F90B" httpUri="https://zenodo.org/record/203263/files/figure.png" pageId="8" pageNumber="9" targetBox="[221,1358,196,1530]" targetPageId="8">
|
||
<paragraph id="DA181B6BD647450CFF408437FE05F90B" blockId="8.[151,1437,1559,1673]" pageId="8" pageNumber="9">
|
||
<emphasis id="E8D3C779D647450CFF408437FED0F9AE" bold="true" box="[151,274,1559,1582]" pageId="8" pageNumber="9">FIGURE 3.</emphasis>
|
||
<taxonomicName id="1DA760E8D647450CFECF8437FDDAF9AF" box="[280,536,1559,1581]" class="Clitellata" family="Enchytraeidae" genus="Guaranidrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="cingulatus" status="sp. nov.">
|
||
<emphasis id="E8D3C779D647450CFECF8437FDDAF9AF" box="[280,536,1559,1581]" italics="true" pageId="8" pageNumber="9">Guaranidrilus cingulatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E8D3C779D647450CFDC88438FDADF9AF" bold="true" box="[543,623,1560,1581]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="F3E07A02D647450CFDC88438FDADF9AF" box="[543,623,1560,1581]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. Segments IV–IX, lateral view. B. Anterior (left) and caudal (right) chaeta. C. Segments IV–VIII, dorsal to dorso-lateral view. D. Spermatheca. E. Clitellum dorsally. F. Preclitellar nephridium. G. Postclitellar nephridium. H. Male efferent apparatus. Male pore in ventral view. I. Male pore in lateral view, same specimen as in H. All drawings from whole mounts.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DA181B6BD646450DFF108307FD69FDBE" blockId="9.[151,1437,151,1328]" pageId="9" pageNumber="10">
|
||
<emphasis id="E8D3C779D646450DFF108307FEBEFEC2" bold="true" box="[199,380,295,320]" pageId="9" pageNumber="10">Seminal vesicle</emphasis>
|
||
absent, free cysts in XI sparse.
|
||
<emphasis id="E8D3C779D646450DFD368307FCBCFEC2" bold="true" box="[737,894,295,320]" pageId="9" pageNumber="10">Spermatozoa</emphasis>
|
||
ca. 47 µm long, heads 18 µm long.
|
||
<emphasis id="E8D3C779D646450DFAC38307FF79FEE6" bold="true" pageId="9" pageNumber="10">Sperm funnel</emphasis>
|
||
(
|
||
<figureCitation id="429C07EED646450DFF1C836CFECCFEE6" box="[203,270,332,356]" captionStart="FIGURE 3" captionStartId="8.[151,250,1559,1582]" captionTargetBox="[221,1358,196,1530]" captionTargetId="figure@8.[220,1366,193,1538]" captionTargetPageId="8" captionText="FIGURE 3. Guaranidrilus cingulatus sp. nov. A. Segments IV – IX, lateral view. B. Anterior (left) and caudal (right) chaeta. C. Segments IV – VIII, dorsal to dorso-lateral view. D. Spermatheca. E. Clitellum dorsally. F. Preclitellar nephridium. G. Postclitellar nephridium. H. Male efferent apparatus. Male pore in ventral view. I. Male pore in lateral view, same specimen as in H. All drawings from whole mounts." httpUri="https://zenodo.org/record/203263/files/figure.png" pageId="9" pageNumber="10">Fig. 3</figureCitation>
|
||
E,H) small, length less than 1/4 body diameter, 1.2–1.5x as long as wide, barrel-shaped, flattened. Vas deferens short with few loops, diameter 8 µm proximally, 5 µm distally.
|
||
<emphasis id="E8D3C779D646450DFC1C8350FB1CFE0A" bold="true" box="[971,1246,368,392]" pageId="9" pageNumber="10">Male copulatory organ</emphasis>
|
||
(
|
||
<figureCitation id="429C07EED646450DFB39834FFAF3FE05" box="[1262,1329,367,391]" captionStart="FIGURE 3" captionStartId="8.[151,250,1559,1582]" captionTargetBox="[221,1358,196,1530]" captionTargetId="figure@8.[220,1366,193,1538]" captionTargetPageId="8" captionText="FIGURE 3. Guaranidrilus cingulatus sp. nov. A. Segments IV – IX, lateral view. B. Anterior (left) and caudal (right) chaeta. C. Segments IV – VIII, dorsal to dorso-lateral view. D. Spermatheca. E. Clitellum dorsally. F. Preclitellar nephridium. G. Postclitellar nephridium. H. Male efferent apparatus. Male pore in ventral view. I. Male pore in lateral view, same specimen as in H. All drawings from whole mounts." httpUri="https://zenodo.org/record/203263/files/figure.png" pageId="9" pageNumber="10">Fig. 3</figureCitation>
|
||
H,I) minute, as if absent, male glands small, not thicker than clitellum. Male pores on body surface, surrounded by male glands, no bursa. Transverse copulatory body muscles sparse, not seen in vivo.
|
||
<emphasis id="E8D3C779D646450DFBFB8397FB39FE52" bold="true" box="[1068,1275,439,464]" pageId="9" pageNumber="10">Accessory glands</emphasis>
|
||
absent.
|
||
<emphasis id="E8D3C779D646450DFA8B8397FF3EFE76" bold="true" pageId="9" pageNumber="10">Spermatheca</emphasis>
|
||
(
|
||
<figureCitation id="429C07EED646450DFEDC83FCFE8FFE76" box="[267,333,476,500]" captionStart="FIGURE 3" captionStartId="8.[151,250,1559,1582]" captionTargetBox="[221,1358,196,1530]" captionTargetId="figure@8.[220,1366,193,1538]" captionTargetPageId="8" captionText="FIGURE 3. Guaranidrilus cingulatus sp. nov. A. Segments IV – IX, lateral view. B. Anterior (left) and caudal (right) chaeta. C. Segments IV – VIII, dorsal to dorso-lateral view. D. Spermatheca. E. Clitellum dorsally. F. Preclitellar nephridium. G. Postclitellar nephridium. H. Male efferent apparatus. Male pore in ventral view. I. Male pore in lateral view, same specimen as in H. All drawings from whole mounts." httpUri="https://zenodo.org/record/203263/files/figure.png" pageId="9" pageNumber="10">Fig. 3</figureCitation>
|
||
C,D) simple, extending into V or VI, about as long as body diameter or slightly longer. Ectal duct not longer than 15 µm, with cuticle-lined canal only in distal half; distal part of ampulla ca. 15 µm wide, often without sperm; sperm present more proximally.
|
||
</paragraph>
|
||
<paragraph id="DA181B6BD646450DFF108067FAA9FD06" blockId="9.[151,1437,151,1328]" pageId="9" pageNumber="10">
|
||
<emphasis id="E8D3C779D646450DFF108067FEE8FDE2" bold="true" box="[199,298,583,608]" pageId="9" pageNumber="10">Habitat.</emphasis>
|
||
<taxonomicName id="1DA760E8D646450DFEE58068FE04FDE2" box="[306,454,584,608]" class="Clitellata" family="Enchytraeidae" genus="Guaranidrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="cingulatus">
|
||
<emphasis id="E8D3C779D646450DFEE58068FE04FDE2" box="[306,454,584,608]" italics="true" pageId="9" pageNumber="10">G. cingulatus</emphasis>
|
||
</taxonomicName>
|
||
was found in young to medium secondary forests and old-growth forest. It was absent in pastures and at agroforestry sites, and almost absent at recently abandoned pastures (
|
||
<specimenCount id="CCA1D0E2D646450DFBBC804CFB2AFD06" box="[1131,1256,620,644]" pageId="9" pageNumber="10" type="generic">1 specimen</specimenCount>
|
||
identified).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92BD48E0D646450DFF1080B0FB77FAB2" pageId="9" pageNumber="10" type="discussion">
|
||
<paragraph id="DA181B6BD646450DFF1080B0FA9CFCFD" blockId="9.[151,1437,151,1328]" pageId="9" pageNumber="10">
|
||
<emphasis id="E8D3C779D646450DFF1080B0FEF9FD2A" bold="true" box="[199,315,656,680]" pageId="9" pageNumber="10">Remarks.</emphasis>
|
||
<taxonomicName id="1DA760E8D646450DFE9280B0FE1EFD2A" box="[325,476,656,680]" class="Clitellata" family="Enchytraeidae" genus="Guaranidrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="cingulatus">
|
||
<emphasis id="E8D3C779D646450DFE9280B0FE1EFD2A" box="[325,476,656,680]" italics="true" pageId="9" pageNumber="10">G. cingulatus</emphasis>
|
||
</taxonomicName>
|
||
is conspicuous (1) by the girdle-shaped clitellum, well-developed on all sides and quite prominent in living and preserved specimens. All other reproductive structures are small and inconspicuous. Further species-specific traits, diagnostic in combination: (2) epidermal gland cells appear absent; (3) cuticle conspicuous, longitudinal muscle layer well-developed, body wall often thick; (4) terminal (= caudal) chaetae up to twice as large as anterior chaetae; (5) oesophageal appendages in anterior of VI; (6) intestinal diverticula in VII, with posterior extension and often irregularly lobed; (7) three pairs of preclitellar nephridia, from 6/7 to 8/9. Traits 2–7 suffice to identify the species; they were used in the routine identification of animals, juveniles included.
|
||
</paragraph>
|
||
<paragraph id="DA181B6BD646450DFF1281ADFC7AFB46" blockId="9.[151,1437,151,1328]" pageId="9" pageNumber="10">
|
||
Most similar to
|
||
<taxonomicName id="1DA760E8D646450DFEA181ADFDCBFC26" box="[374,521,908,932]" class="Clitellata" family="Enchytraeidae" genus="Guaranidrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="cingulatus">
|
||
<emphasis id="E8D3C779D646450DFEA181ADFDCBFC26" box="[374,521,908,932]" italics="true" pageId="9" pageNumber="10">G. cingulatus</emphasis>
|
||
</taxonomicName>
|
||
is the North-American
|
||
<taxonomicName id="1DA760E8D646450DFCC681ADFB6AFC26" authority="Coates & Diaz, 1988" authorityName="Coates & Diaz" authorityYear="1988" box="[785,1192,908,932]" class="Clitellata" family="Enchytraeidae" genus="Guaranidrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="oregonensis">
|
||
<emphasis id="E8D3C779D646450DFCC681ADFC77FC26" box="[785,949,909,932]" italics="true" pageId="9" pageNumber="10">G. oregonensis</emphasis>
|
||
<bibRefCitation id="BE36669AD646450DFC6B81ACFB6AFC26" author="Coates" box="[956,1192,908,932]" pageId="9" pageNumber="10" refString="Coates, K. A. & Diaz, R. J. (1988) Description of Guaranidrilus oregonensis (Oligochaeta: Enchytraeidae) from North America, with additional comments on the genus. Proceedings of the Biological Society of Washington, 101, 773 - 783." type="journal article" year="1988">Coates & Diaz, 1988</bibRefCitation>
|
||
</taxonomicName>
|
||
, described from fixed material. The species is similar in body length (
|
||
<quantity id="1D5FB68ED646450DFD65818FFCE8FC45" box="[690,810,943,967]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="9" pageNumber="10" unit="mm" value="3.75" valueMax="4.5" valueMin="3.0">3–4.5 mm</quantity>
|
||
), segment number (26–34), chaetae (enlarged in caudal segments), epidermal gland cells (few rows), origin of dorsal blood vessel (X–XII), position of preclitellar nephridia (3 pairs, 6/7–8/9), and sperm-related structures (small).
|
||
<taxonomicName id="1DA760E8D646450DFCA381D8FBDBFB92" box="[884,1049,1016,1040]" class="Clitellata" family="Enchytraeidae" genus="Guaranidrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="oregonensis">
|
||
<emphasis id="E8D3C779D646450DFCA381D8FBDBFB92" box="[884,1049,1016,1040]" italics="true" pageId="9" pageNumber="10">G. oregonensis</emphasis>
|
||
</taxonomicName>
|
||
may also have a girdle-shaped clitellum, but the trait is described ambiguously and requires reinvestigation: "Clitellum ... in transverse rows ventrolaterally; reduced but complete ventrally between male pores" (ibd.: 775). The species differs in the absence of oesophageal appendages (present in
|
||
<taxonomicName id="1DA760E8D646450DFDFA8645FD02FBFE" box="[557,704,1124,1148]" class="Clitellata" family="Enchytraeidae" genus="Guaranidrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="cingulatus">
|
||
<emphasis id="E8D3C779D646450DFDFA8645FD02FBFE" box="[557,704,1124,1148]" italics="true" pageId="9" pageNumber="10">G. cingulatus</emphasis>
|
||
</taxonomicName>
|
||
) and in the extension over 2 segments of the intestinal diverticula (1 segment in
|
||
<taxonomicName id="1DA760E8D646450DFE9386A8FE19FB22" box="[324,475,1160,1184]" class="Clitellata" family="Enchytraeidae" genus="Guaranidrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="cingulatus">
|
||
<emphasis id="E8D3C779D646450DFE9386A8FE19FB22" box="[324,475,1160,1184]" italics="true" pageId="9" pageNumber="10">G. cingulatus</emphasis>
|
||
</taxonomicName>
|
||
). Specimens had been sampled from freshwater marsh soils in the estuary of the
|
||
<collectingCountry id="A2B05BFBD646450DFF40868CFEC7FB46" box="[151,261,1196,1220]" name="United States of America" pageId="9" pageNumber="10">Columbia</collectingCountry>
|
||
river (Oregon,
|
||
<collectingCountry id="A2B05BFBD646450DFE64868CFDC0FB46" box="[435,514,1196,1220]" name="United States of America" pageId="9" pageNumber="10">U.S.A.</collectingCountry>
|
||
), see
|
||
<bibRefCitation id="BE36669AD646450DFD94868CFCFBFB46" author="Coates" box="[579,825,1196,1220]" pageId="9" pageNumber="10" refString="Coates, K. A. & Diaz, R. J. (1988) Description of Guaranidrilus oregonensis (Oligochaeta: Enchytraeidae) from North America, with additional comments on the genus. Proceedings of the Biological Society of Washington, 101, 773 - 783." type="journal article" year="1988">Coates & Diaz (1988)</bibRefCitation>
|
||
for details.
|
||
</paragraph>
|
||
<paragraph id="DA181B6BD646450DFF1286F0FB77FAB2" blockId="9.[151,1437,151,1328]" pageId="9" pageNumber="10">
|
||
Further similar species are
|
||
<taxonomicName id="1DA760E8D646450DFE2F86F0FCF5FB65" authority="Cernosvitov, 1937" authorityName="Cernosvitov" authorityYear="1937" box="[504,823,1226,1256]" class="Clitellata" family="Enchytraeidae" genus="Guaranidrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="rarus">
|
||
<emphasis id="E8D3C779D646450DFE2F86F0FD96FB6A" box="[504,596,1232,1256]" italics="true" pageId="9" pageNumber="10">G. rarus</emphasis>
|
||
<bibRefCitation id="BE36669AD646450DFD8A86EAFCF5FB65" author="Cernosvitov" box="[605,823,1226,1255]" pageId="9" pageNumber="10" refString="Cernosvitov, L. (1937 a) Notes sur les Oligochaeta (Naididees et Enchytraeidees) de l'Argentine. Anales del Museo Argentino de Ciencias Naturales, Buenos Aires, 39, 135 - 157." type="journal article" year="1937" yearSuffix="a">Černosvitov, 1937a</bibRefCitation>
|
||
</taxonomicName>
|
||
emend.
|
||
<bibRefCitation id="BE36669AD646450DFC4E86EFFB57FB65" author="Coates" box="[921,1173,1231,1255]" pageId="9" pageNumber="10" refString="Coates, K. A. & Diaz, R. J. (1988) Description of Guaranidrilus oregonensis (Oligochaeta: Enchytraeidae) from North America, with additional comments on the genus. Proceedings of the Biological Society of Washington, 101, 773 - 783." type="journal article" year="1988">Coates & Diaz (1988)</bibRefCitation>
|
||
, and
|
||
<emphasis id="E8D3C779D646450DFB0486F0FA89FB6A" box="[1235,1355,1232,1256]" italics="true" pageId="9" pageNumber="10">G. s a w a y a i</emphasis>
|
||
<bibRefCitation id="BE36669AD646450DFA8086EFFF10FA8E" author="Righi" pageId="9" pageNumber="10" refString="Righi, G. (1973) Sobre tres especies Brasileiras de Enchytraeidae (Oligochaeta). Boletim de Zoologia e Biologia Marina, Sao Paulo, 30, 469 - 482." type="journal article" year="1973">Righi, 1973</bibRefCitation>
|
||
. Both species, which may be synonymous (
|
||
<bibRefCitation id="BE36669AD646450DFD6286D4FC5EFA8E" author="Coates" box="[693,924,1268,1292]" pageId="9" pageNumber="10" refString="Coates, K. A. & Diaz, R. J. (1988) Description of Guaranidrilus oregonensis (Oligochaeta: Enchytraeidae) from North America, with additional comments on the genus. Proceedings of the Biological Society of Washington, 101, 773 - 783." type="journal article" year="1988">Coates & Diaz 1988</bibRefCitation>
|
||
), differ from
|
||
<taxonomicName id="1DA760E8D646450DFBE786D5FB01FA8E" box="[1072,1219,1268,1292]" class="Clitellata" family="Enchytraeidae" genus="Guaranidrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enchytraeida" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="cingulatus">
|
||
<emphasis id="E8D3C779D646450DFBE786D5FB01FA8E" box="[1072,1219,1268,1292]" italics="true" pageId="9" pageNumber="10">G. cingulatus</emphasis>
|
||
</taxonomicName>
|
||
in having a saddleshaped clitellum. Other differences are inconclusive and require a reinvestigation of the
|
||
<typeStatus id="051CA5C9D646450DFBA58738FB73FAB2" box="[1138,1201,1304,1328]" pageId="9" pageNumber="10">types</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |