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<document id="19AF9222EBA57A5DDCA1D298AEBEDA04" ID-DOI="10.11646/zootaxa.3995.1.21" ID-GBIF-Dataset="810c1097-eff4-4262-acd6-1f76d05f1f83" ID-ISSN="1175-5326" ID-Zenodo-Dep="236857" ID-ZooBank="1D5CD052-9754-493C-836D-6A69756747D9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461278201543" checkinUser="plazi" docAuthor="Oliver, P. Graham" docDate="2015" docId="3956C47FFFE4E32AFF44F8D4FE78FEB2" docLanguage="en" docName="zt03995p263.pdf" docOrigin="Zootaxa 3995 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mendicula ferruginosa Forbes 1844" docType="treatment" docVersion="8" lastPageNumber="261" masterDocId="C56FBC07FFE3E323FFD3FFE5FF80FFD7" masterDocTitle="Deep-water Thyasiridae (Mollusca: Bivalvia) from the Oman Margin, Arabian Sea, new species and examples of endemism and cosmopolitanism" masterLastPageNumber="263" masterPageNumber="252" pageNumber="259" updateTime="1698613566852" updateUser="plazi">
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<mods:title id="0E0B9AC1CCF414A70A3C39F0C10483EE">Deep-water Thyasiridae (Mollusca: Bivalvia) from the Oman Margin, Arabian Sea, new species and examples of endemism and cosmopolitanism</mods:title>
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<mods:namePart id="BC8DA2A9952FF682B67767CC4C5CDEB3">Oliver, P. Graham</mods:namePart>
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<mods:date id="729D22AAD646A5353241E52D6758BF8C">2015</mods:date>
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<mods:number id="952C721C784C3F855B1A4FE0BA02F531">3995</mods:number>
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<mods:identifier id="BDDCC91B5CF30C9896B884E739F3AE0E" type="DOI">10.11646/zootaxa.3995.1.21</mods:identifier>
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<treatment id="3956C47FFFE4E32AFF44F8D4FE78FEB2" ID-DOI="http://doi.org/10.5281/zenodo.6098591" ID-GBIF-Taxon="119640192" ID-Zenodo-Dep="6098591" LSID="urn:lsid:plazi:treatment:3956C47FFFE4E32AFF44F8D4FE78FEB2" httpUri="http://treatment.plazi.org/id/3956C47FFFE4E32AFF44F8D4FE78FEB2" lastPageId="9" lastPageNumber="261" pageId="7" pageNumber="259">
<subSubSection id="F9E526E2FFE4E324FF44F8D4FE8EF8BB" pageId="7" pageNumber="259" type="nomenclature">
<paragraph id="B1407569FFE4E324FF44F8D4FD06F89C" blockId="7.[151,646,1841,1901]" box="[151,646,1841,1867]" pageId="7" pageNumber="259">
<heading id="EA08C205FFE4E324FF44F8D4FD06F89C" bold="true" box="[151,646,1841,1867]" fontSize="11" level="1" pageId="7" pageNumber="259" reason="1">
<taxonomicName id="76FF0EEAFFE4E324FF44F8D4FD06F89C" authority="Forbes, 1844" authorityName="Forbes" authorityYear="1844" box="[151,646,1841,1867]" class="Bivalvia" family="Thyasiridae" genus="Mendicula" kingdom="Animalia" order="Lucinoida" pageId="7" pageNumber="259" phylum="Mollusca" rank="species" species="ferruginosa">
<emphasis id="838BA97BFFE4E324FF44F8D4FD06F89C" bold="true" box="[151,646,1841,1867]" pageId="7" pageNumber="259">
<emphasis id="838BA97BFFE4E324FF44F8D4FE99F89C" bold="true" box="[151,281,1841,1867]" italics="true" pageId="7" pageNumber="259">Mendicula</emphasis>
aff.
<emphasis id="838BA97BFFE4E324FE82F8D4FE5EF89C" bold="true" box="[337,478,1841,1867]" italics="true" pageId="7" pageNumber="259">ferruginosa</emphasis>
<bibRefCitation id="D56E0898FFE4E324FE35F8D4FD06F89C" author="Forbes" box="[486,646,1841,1867]" pageId="7" pageNumber="259" refString="Forbes, E. (1844) Report on the Mollusca and Radiata of the Aegean Sea, and on their distribution, considered as bearing on geology. Reports of the British Association for the Advancement of Science, 1843, 130 - 193." type="journal article" year="1844">Forbes, 1844</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="B1407569FFE4E324FF44F8B1FE8EF8BB" blockId="7.[151,646,1841,1901]" box="[151,270,1876,1901]" pageId="7" pageNumber="259">
<figureCitation id="29C469ECFFE4E324FF44F8B1FF5BF8BB" box="[151,219,1876,1901]" captionStart="FIGURE 4. A" captionStartId="8.[151,250,1893,1915]" captionTargetBox="[151,1436,373,1871]" captionTargetId="figure@8.[151,1436,373,1872]" captionTargetPageId="8" captionText="FIGURE 4. A, Axinulus aff. croulinensis, Oman margin; B, Axinulus croulinensis Jeffreys, Loch Fyne, 40 m, Scotland, National Museum Wales; C, Axinulus undescribed, Gulf of Mexico, National Museum Wales; D, A. brevis Verrill &amp; Bush, Rockall Trough, 1595 m, National Museum Wales; E – F, Mendicula aff. ferruginosa, Oman margin; G – H, Mendicula ferruginosa Forbes, Rockall Trough, 1595 m, National Museum Wales; I, Leptaxinus indusarium Oliver &amp; Levin, holotype, National Museum Wales, NMWZ. 2005.34.1." httpUri="https://zenodo.org/record/236861/files/figure.png" pageId="7" pageNumber="259">Figs 4</figureCitation>
F–G
</paragraph>
</subSubSection>
<subSubSection id="F9E526E2FFE4E324FF44F879FB9AF80E" pageId="7" pageNumber="259" type="materials_examined">
<paragraph id="B1407569FFE4E324FF44F879FB9AF80E" blockId="7.[151,1436,1948,2009]" pageId="7" pageNumber="259">
<emphasis id="838BA97BFFE4E324FF44F879FE86F862" bold="true" box="[151,262,1948,1973]" pageId="7" pageNumber="259">Material.</emphasis>
Five specimens, Off Ras Madrakah, southern
<collectingCountry id="C9E835F9FFE4E324FCEFF879FC03F863" box="[828,899,1948,1972]" name="Oman" pageId="7" pageNumber="259">Oman</collectingCountry>
, Discovery cruise 211 st. 12719#1,
<geoCoordinate id="D4CB13AEFFE4E324FAE8F879FA1CF863" box="[1339,1436,1948,1972]" direction="north" orientation="latitude" pageId="7" pageNumber="259" precision="925" value="19.133333">19°08´N</geoCoordinate>
<geoCoordinate id="D4CB13AEFFE4E324FF44F825FF77F80E" box="[151,247,1984,2009]" direction="east" orientation="longitude" pageId="7" pageNumber="259" precision="925" value="58.65">58°39´E</geoCoordinate>
,
<quantity id="7607D88CFFE4E324FED0F825FED3F80E" box="[259,339,1984,2009]" metricMagnitude="3" metricUnit="m" metricValue="3.15" pageId="7" pageNumber="259" unit="m" value="3150.0">3150m</quantity>
, collected P.G. Oliver,
<date id="C54153A9FFE4E324FD82F825FC91F80F" box="[593,785,1984,2009]" pageId="7" pageNumber="259" value="1994-10-31">31 October 1994</date>
.
<collectionCode id="D7EEEDACFFE4E324FCC8F824FCE7F80F" box="[795,871,1985,2008]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="259">NMW</collectionCode>
.Z. 1995.009.19.
</paragraph>
</subSubSection>
<subSubSection id="F9E526E2FFEBE32BFF14FF72FD07FECA" pageId="8" pageNumber="260" type="description">
<paragraph id="B1407569FFEBE32BFF14FF72FD07FECA" blockId="8.[151,1437,151,285]" pageId="8" pageNumber="260">
<emphasis id="838BA97BFFEBE32BFF14FF72FED9FF67" bold="true" box="[199,345,151,176]" pageId="8" pageNumber="260">Description.</emphasis>
To
<quantity id="7607D88CFFEBE32BFE58FF7DFE50FF78" box="[395,464,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="260" unit="mm" value="2.0">2 mm</quantity>
in length. Equivalve. Equilateral prosogyrous beaks. Subcircular, posterior margin more narrowly rounded than anterior. Posterior area weakly demarcated by a radial angulation; submarginal sulcus narrow; lunule a slight depression. Hinge with an anterior thickening that in the right valve most developed. Exteriorly with a heavy ferruginous coating.
</paragraph>
</subSubSection>
<caption id="E58025E1FFEBE32BFF44F880FDE0F821" httpUri="https://zenodo.org/record/236861/files/figure.png" pageId="8" pageNumber="260" targetBox="[151,1436,373,1871]" targetPageId="8">
<paragraph id="B1407569FFEBE32BFF44F880FDE0F821" blockId="8.[151,1436,1893,2038]" pageId="8" pageNumber="260">
<emphasis id="838BA97BFFEBE32BFF44F880FE97F8AD" bold="true" box="[151,279,1893,1915]" pageId="8" pageNumber="260">FIGURE 4.</emphasis>
A,
<taxonomicName id="76FF0EEAFFEBE32BFE95F880FDD0F8AC" box="[326,592,1893,1915]" class="Bivalvia" family="Thyasiridae" genus="Axinulus" kingdom="Animalia" order="Lucinoida" pageId="8" pageNumber="260" phylum="Mollusca" rank="species" species="croulinensis">
<emphasis id="838BA97BFFEBE32BFE95F880FDD0F8AC" box="[326,592,1893,1915]" italics="true" pageId="8" pageNumber="260">Axinulus aff. croulinensis</emphasis>
</taxonomicName>
, Oman margin; B,
<taxonomicName id="76FF0EEAFFEBE32BFCF5F880FB65F8AC" authority="Jeffreys, Loch Fyne" authorityName="Jeffreys, Loch Fyne" box="[806,1253,1893,1915]" class="Bivalvia" family="Thyasiridae" genus="Axinulus" kingdom="Animalia" order="Lucinoida" pageId="8" pageNumber="260" phylum="Mollusca" rank="species" species="croulinensis">
<emphasis id="838BA97BFFEBE32BFCF5F880FB85F8AC" box="[806,1029,1893,1915]" italics="true" pageId="8" pageNumber="260">Axinulus croulinensis</emphasis>
Jeffreys, Loch Fyne
</taxonomicName>
, 40 m, Scotland, National Museum Wales; C,
<taxonomicName id="76FF0EEAFFEBE32BFE15F861FD9FF84D" box="[454,543,1924,1946]" class="Bivalvia" family="Thyasiridae" genus="Axinulus" kingdom="Animalia" order="Lucinoida" pageId="8" pageNumber="260" phylum="Mollusca" rank="genus">
<emphasis id="838BA97BFFEBE32BFE15F861FD9FF84D" box="[454,543,1924,1946]" italics="true" pageId="8" pageNumber="260">Axinulus</emphasis>
</taxonomicName>
undescribed, Gulf of Mexico, National Museum Wales; D,
<taxonomicName id="76FF0EEAFFEBE32BFB45F860FE04F86E" authority="Verrill &amp; Bush, Rockall Trough, 1595" authorityName="Verrill &amp; Bush, Rockall Trough" authorityYear="1595" class="Bivalvia" family="Thyasiridae" genus="Cryptodon" higherTaxonomySource="CoL" kingdom="Animalia" order="Lucinida" pageId="8" pageNumber="260" phylum="Mollusca" rank="species" species="brevis">
<emphasis id="838BA97BFFEBE32BFB45F860FB72F84D" box="[1174,1266,1924,1946]" italics="true" pageId="8" pageNumber="260">A. brevis</emphasis>
Verrill &amp; Bush, Rockall Trough, 1595
</taxonomicName>
m, National Museum Wales; E–F,
<taxonomicName id="76FF0EEAFFEBE32BFCC2F846FBAFF86E" box="[785,1071,1955,1977]" class="Bivalvia" family="Thyasiridae" genus="Mendicula" kingdom="Animalia" order="Lucinoida" pageId="8" pageNumber="260" phylum="Mollusca" rank="species" species="ferruginosa">
<emphasis id="838BA97BFFEBE32BFCC2F846FBAFF86E" box="[785,1071,1955,1977]" italics="true" pageId="8" pageNumber="260">Mendicula aff. ferruginosa</emphasis>
</taxonomicName>
, Oman margin; G–H,
<taxonomicName id="76FF0EEAFFEBE32BFAE3F846FDCCF800" authority="Forbes, Rockall Trough, 1595" authorityName="Forbes, Rockall Trough" authorityYear="1595" class="Bivalvia" family="Thyasiridae" genus="Mendicula" kingdom="Animalia" order="Lucinoida" pageId="8" pageNumber="260" phylum="Mollusca" rank="species" species="ferruginosa">
<emphasis id="838BA97BFFEBE32BFAE3F846FE8DF800" italics="true" pageId="8" pageNumber="260">Mendicula ferruginosa</emphasis>
Forbes, Rockall Trough, 1595
</taxonomicName>
m, National Museum Wales; I,
<taxonomicName id="76FF0EEAFFEBE32BFC48F827FAB0F800" authority="Oliver &amp; Levin" authorityName="Oliver &amp; Levin" box="[923,1328,1985,2007]" class="Bivalvia" family="Thyasiridae" genus="Leptaxinus" kingdom="Animalia" order="Lucinoida" pageId="8" pageNumber="260" phylum="Mollusca" rank="species" species="indusarium">
<emphasis id="838BA97BFFEBE32BFC48F827FB05F800" box="[923,1157,1985,2007]" italics="true" pageId="8" pageNumber="260">Leptaxinus indusarium</emphasis>
Oliver &amp; Levin
</taxonomicName>
, holotype, National Museum Wales, NMWZ.2005.34.1.
</paragraph>
</caption>
<subSubSection id="F9E526E2FFEAE32AFF14FF72FE78FEB2" pageId="9" pageNumber="261" type="discussion">
<paragraph id="B1407569FFEAE32AFF14FF72FE78FEB2" blockId="9.[151,1436,151,357]" pageId="9" pageNumber="261">
<emphasis id="838BA97BFFEAE32AFF14FF72FEBBFF67" bold="true" box="[199,315,151,176]" pageId="9" pageNumber="261">Remarks.</emphasis>
These shells are inseparable from the deep water examples of
<taxonomicName id="76FF0EEAFFEAE32AFC2FFF7CFB25FF67" box="[1020,1189,152,176]" class="Bivalvia" family="Thyasiridae" genus="Mendicula" kingdom="Animalia" order="Lucinoida" pageId="9" pageNumber="261" phylum="Mollusca" rank="species" species="ferruginosa">
<emphasis id="838BA97BFFEAE32AFC2FFF7CFB25FF67" box="[1020,1189,152,176]" italics="true" pageId="9" pageNumber="261">M. ferruginosa</emphasis>
</taxonomicName>
in the Atlantic Ocean (
<figureCitation id="29C469ECFFEAE32AFF4CFF59FF67FF03" box="[159,231,188,213]" captionStart="FIGURE 4. A" captionStartId="8.[151,250,1893,1915]" captionTargetBox="[151,1436,373,1871]" captionTargetId="figure@8.[151,1436,373,1872]" captionTargetPageId="8" captionText="FIGURE 4. A, Axinulus aff. croulinensis, Oman margin; B, Axinulus croulinensis Jeffreys, Loch Fyne, 40 m, Scotland, National Museum Wales; C, Axinulus undescribed, Gulf of Mexico, National Museum Wales; D, A. brevis Verrill &amp; Bush, Rockall Trough, 1595 m, National Museum Wales; E – F, Mendicula aff. ferruginosa, Oman margin; G – H, Mendicula ferruginosa Forbes, Rockall Trough, 1595 m, National Museum Wales; I, Leptaxinus indusarium Oliver &amp; Levin, holotype, National Museum Wales, NMWZ. 2005.34.1." httpUri="https://zenodo.org/record/236861/files/figure.png" pageId="9" pageNumber="261">Figs 4</figureCitation>
H–I) and the eastern Pacific Ocean (Coan
<emphasis id="838BA97BFFEAE32AFD1DFF58FC8FFF02" box="[718,783,188,213]" italics="true" pageId="9" pageNumber="261">et al.,</emphasis>
2000). They are thinner and less heavily coated than the shelf examples. Very similar taxa are
<taxonomicName id="76FF0EEAFFEAE32AFD9AFF04FD63FF2F" box="[585,739,225,248]" class="Bivalvia" family="Thyasiridae" genus="Mendicula" kingdom="Animalia" order="Lucinoida" pageId="9" pageNumber="261" phylum="Mollusca" rank="species" species="memorata">
<emphasis id="838BA97BFFEAE32AFD9AFF04FD63FF2F" box="[585,739,225,248]" italics="true" pageId="9" pageNumber="261">M. memorata</emphasis>
</taxonomicName>
from SE
<collectingCountry id="C9E835F9FFEAE32AFC86FF3AFC3FFF2F" box="[853,959,223,248]" name="Australia" pageId="9" pageNumber="261">Australia</collectingCountry>
,
<taxonomicName id="76FF0EEAFFEAE32AFC1FFF04FAB1FF2F" authority="Zelaya, 2010" authorityName="Zelaya" authorityYear="2010" box="[972,1329,223,248]" class="Bivalvia" family="Thyasiridae" genus="Mendicula" kingdom="Animalia" order="Lucinoida" pageId="9" pageNumber="261" phylum="Mollusca" rank="species" species="sudamericana">
<emphasis id="838BA97BFFEAE32AFC1FFF04FB13FF2F" box="[972,1171,223,248]" italics="true" pageId="9" pageNumber="261">M. sudamericana</emphasis>
<bibRefCitation id="D56E0898FFEAE32AFB4FFF3AFAB1FF2F" author="Zelaya" box="[1180,1329,223,248]" pageId="9" pageNumber="261" refString="Zelaya, D. G. (2010) New species of Thyasira, Mendicula, and Axinulus (Bivalvia, Thyasiroidea) from Sub-Antarctic and Antarctic waters. Polar Biology, 33, 607 - 616." type="journal article" year="2010">Zelaya, 2010</bibRefCitation>
</taxonomicName>
from the southern Ocean and
<taxonomicName id="76FF0EEAFFEAE32AFEACFEE1FC27FECB" authority="Keuning &amp; Schander, 2010" authorityName="Keuning &amp; Schander" authorityYear="2010" box="[383,935,260,285]" class="Bivalvia" family="Thyasiridae" genus="Thyasira" kingdom="Animalia" order="Lucinoida" pageId="9" pageNumber="261" phylum="Mollusca" rank="species" species="ockelmanni">
<emphasis id="838BA97BFFEAE32AFEACFEE1FDEBFECA" box="[383,619,260,285]" italics="true" pageId="9" pageNumber="261">Thyasira ockelmanni</emphasis>
<bibRefCitation id="D56E0898FFEAE32AFDA0FEE1FC27FECB" author="Keuning" box="[627,935,260,285]" pageId="9" pageNumber="261" refString="Keuning, R. &amp; Schander, C. (2010) Thyasira ockelmanni (Mollusca: Bivalvia: Veneroidea), A new species of Thyasiridae from the Norwegian Sea. Fauna norvegica, 30, 21 - 24." type="journal article" year="2010">Keuning &amp; Schander, 2010</bibRefCitation>
</taxonomicName>
from the abyssal Norwegian Sea. Variation seen in
<taxonomicName id="76FF0EEAFFEAE32AFF3EFECCFE15FE97" box="[237,405,296,320]" class="Bivalvia" family="Thyasiridae" genus="Mendicula" kingdom="Animalia" order="Lucinoida" pageId="9" pageNumber="261" phylum="Mollusca" rank="species" species="ferruginosa">
<emphasis id="838BA97BFFEAE32AFF3EFECCFE15FE97" box="[237,405,296,320]" italics="true" pageId="9" pageNumber="261">M. ferruginosa</emphasis>
</taxonomicName>
from the NE Atlantic (see
<bibRefCitation id="D56E0898FFEAE32AFD15FEC2FC15FE97" author="Oliver" box="[710,917,295,320]" pageId="9" pageNumber="261" refString="Oliver, P. G., Holmes, A. M., Killeen, I. J. &amp; Turner, J. A. (2010) Marine Bivalve Shells of the British Isles (Mollusca: Bivalvia). Amgueddfa Cymru - National Museum Wales. Available from: http: // naturalhistory. museumwales. ac. uk / britishbivalves (accessed 28 May 2015)" type="journal article" year="2010">
Oliver
<emphasis id="838BA97BFFEAE32AFCC7FECCFCCDFE97" box="[788,845,295,320]" italics="true" pageId="9" pageNumber="261">et al.</emphasis>
, 2010
</bibRefCitation>
) makes definitive recognition of these species from shell characters uncertain.
</paragraph>
</subSubSection>
</treatment>
</document>
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