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<document id="141EDEB34C89C04D658184DD77F5F78B" ID-CLB-Dataset="30434" ID-DOI="10.11646/zootaxa.4399.4.10" ID-GBIF-Dataset="f5adc190-982e-4678-bfa8-5f53b66457aa" ID-ISSN="1175-5326" ID-Zenodo-Dep="1206812" ID-ZooBank="9268BDE4-5A5A-4A1D-9DD6-321BE70355B8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1521900264889" checkinUser="plazi" docAuthor="Sehnal, Richard" docDate="2018" docId="1619EF4B77275E2669FFF9BD9F2AC329" docLanguage="en" docName="zootaxa.4399.4.10.pdf" docOrigin="Zootaxa 4399 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eulepida mbala Sehnal 2018, new species" docType="treatment" docVersion="8" lastPageNumber="593" masterDocId="EA20973377275E246968FF819A52C660" masterDocTitle="Eulepida mbala, a new species from Zambia (Coleoptera: Scarabaeidae: Melolonthinae)" masterLastPageNumber="595" masterPageNumber="591" pageNumber="591" updateTime="1698533218777" updateUser="ExternalLinkService">
<mods:mods id="BB77BE9A0418C7AF64D5CD06B363000B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1C9FB9936F411A4C19BD323B1FEF5E59">
<mods:title id="19DAEA2572B09E379FFF1953FAEB3248">Eulepida mbala, a new species from Zambia (Coleoptera: Scarabaeidae: Melolonthinae)</mods:title>
</mods:titleInfo>
<mods:name id="71B341325260398A5789223AFF2B8C22" type="personal">
<mods:role id="536C1A02112A3D175EB5A00C4D02F2A1">
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</mods:role>
<mods:namePart id="ACE464F2F6AAE4C58B7364252A6E480D">Sehnal, Richard</mods:namePart>
</mods:name>
<mods:typeOfResource id="C9C69E06F78A22231A469A34086A2E03">text</mods:typeOfResource>
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<mods:title id="10F30F0D1C6D2DA1BCA46DBF3FF7EE2A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="919FF91EA45D44A67D59B9548DB030EB">
<mods:date id="783DA485F7BB10269A25CAED93A70EE1">2018</mods:date>
<mods:detail id="586FE5B5AA6F4D210CB694B9A6525149" type="pubDate">
<mods:number id="DB60826DE5424CB1D46067E051084450">2018-03-23</mods:number>
</mods:detail>
<mods:detail id="7AB028B9A423745B77187D8FE5E8CFEC" type="volume">
<mods:number id="03F98CA5823AA1ABF9A27469028A6843">4399</mods:number>
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<mods:number id="93EA9DFF1D2A71D04C5540EDC08FAB1B">4</mods:number>
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<mods:classification id="CA5E7B05E198A201FD604D6E373E5E67">journal article</mods:classification>
<mods:identifier id="B7969AF4775ADFA929A3C5BA0C224D9F" type="CLB-Dataset">30434</mods:identifier>
<mods:identifier id="91829AD4EDBAF576344C710BAE92C4C5" type="DOI">10.11646/zootaxa.4399.4.10</mods:identifier>
<mods:identifier id="4A596E08E89D593F2E091C101ED2F3CA" type="GBIF-Dataset">f5adc190-982e-4678-bfa8-5f53b66457aa</mods:identifier>
<mods:identifier id="493DBB7E0D2AB19C92D3DA96D0E3A0A0" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="1619EF4B77275E2669FFF9BD9F2AC329" ID-DOI="http://doi.org/10.5281/zenodo.4485086" ID-GBIF-Taxon="142524468" ID-Zenodo-Dep="4485086" LSID="urn:lsid:plazi:treatment:1619EF4B77275E2669FFF9BD9F2AC329" httpUri="http://treatment.plazi.org/id/1619EF4B77275E2669FFF9BD9F2AC329" lastPageId="2" lastPageNumber="593" pageId="0" pageNumber="591">
<subSubSection id="D6AA0DD677275E2469FFF9BD9803C036" box="[151,593,1596,1622]" pageId="0" pageNumber="591" type="nomenclature">
<paragraph id="9E0F5E5D77275E2469FFF9BD9803C036" blockId="0.[151,1436,1596,2035]" box="[151,593,1596,1622]" pageId="0" pageNumber="591">
<heading id="C547E93177275E2469FFF9BD9803C036" bold="true" box="[151,593,1596,1622]" fontSize="11" level="1" pageId="0" pageNumber="591" reason="1">
<emphasis id="ACC4824F77275E2469FFF9BD9803C036" bold="true" box="[151,593,1596,1622]" pageId="0" pageNumber="591">
<taxonomicName id="59B025DE77275E2469FFF9BD9BE1C036" authority="Sehnal" authorityName="Sehnal" authorityYear="2018" box="[151,435,1596,1622]" class="Insecta" family="Melolonthidae" genus="Eulepida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="591" phylum="Arthropoda" rank="species" species="mbala" status="sp. nov.">
<emphasis id="ACC4824F77275E2469FFF9BD9B04C036" bold="true" box="[151,342,1596,1622]" italics="true" pageId="0" pageNumber="591">Eulepida mbala</emphasis>
Sehnal
</taxonomicName>
,
<taxonomicNameLabel id="B7F73F3477275E2468A9F9BD9803C036" box="[449,593,1596,1622]" pageId="0" pageNumber="591" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D6AA0DD677275E246B30F9BD997AC036" box="[600,808,1596,1622]" pageId="0" pageNumber="591" type="description">
<paragraph id="9E0F5E5D77275E246B30F9BD997AC036" blockId="0.[151,1436,1596,2035]" box="[600,808,1596,1622]" pageId="0" pageNumber="591">
<heading id="C547E93177275E246B30F9BD997AC036" bold="true" box="[600,808,1596,1622]" fontSize="11" level="1" pageId="0" pageNumber="591" reason="1">
<emphasis id="ACC4824F77275E246B30F9BD997AC036" bold="true" box="[600,808,1596,1622]" pageId="0" pageNumber="591">
(
<figureCitation id="068B42D877275E246B08F9BD98BFC036" box="[608,749,1596,1622]" captionStart="FIGURES 1" captionStartId="1.[151,258,1914,1936]" captionTargetBox="[171,1426,470,1853]" captionTargetId="figure@1.[151,1436,437,1891]" captionTargetPageId="1" captionText="FIGURES 1A–F. Eulepida mbala new species, holotype (male) and paratype No. 1 (female). Habitus, dorsal view: A, holotype; B, paratype. Pygidium, lateral view, male (upper) and female (lower). C, Eulepida mbala new species; D, Eulepida anatina Brenske, 1896; E, Eulepida tschindeanaPéringuey, 1904;F, Eulepida werneri Lacroix,2010.Not to scale." httpUri="https://zenodo.org/record/1206814/files/figure.png" pageId="0" pageNumber="591">Figs. 1A–C</figureCitation>
,
<figureCitation id="068B42D877275E246B93F9BD9972C036" box="[763,800,1596,1622]" captionStart="FIGURES 2" captionStartId="3.[151,258,1307,1329]" captionTargetBox="[180,1410,211,1262]" captionTargetId="figure@3.[151,1436,191,1285]" captionTargetPageId="3" captionText="FIGURES 2A–D. Aedeagus of species group II of Eulepida, dorsal (left) and lateral (right) views. A, Eulepida mbala new species, holotype; B, Eulepida anatina Brenske, 1896; C, Eulepida tschindeana Péringuey, 1904; D, Eulepida werneri Lacroix, 2010. Not to scale." httpUri="https://zenodo.org/record/1206816/files/figure.png" pageId="0" pageNumber="591">2A</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D6AA0DD677275E2469FFF9DF9EA1C0BD" pageId="0" pageNumber="591" type="materials_examined">
<paragraph id="9E0F5E5D77275E2469FFF9DF996AC015" blockId="0.[151,1436,1596,2035]" box="[151,824,1630,1653]" pageId="0" pageNumber="591">
<materialsCitation id="2ED8540077275E2469FFF9DF996AC015" ID-GBIF-Occurrence="1836204545" box="[151,824,1630,1653]" country="Zambia" location="Mbala." pageId="0" pageNumber="591" specimenCount="1" stateProvince="Northern Province" typeStatus="holotype">
<emphasis id="ACC4824F77275E2469FFF9DF9B7CC015" bold="true" box="[151,302,1630,1653]" pageId="0" pageNumber="591">
<typeStatus id="410BE0FF77275E2469FFF9DF9A9DC015" box="[151,207,1630,1653]" pageId="0" pageNumber="591">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="E6A71ECD77275E24685CF9DF9BDAC015" box="[308,392,1630,1653]" name="Zambia" pageId="0" pageNumber="591">Zambia</collectingCountry>
,
<collectingRegion id="5C7490BF77275E2468FBF9DF9809C015" box="[403,603,1630,1653]" country="Zambia" name="Northern" pageId="0" pageNumber="591">Northern Province</collectingRegion>
,
<quantity id="5948F3B877275E246B0DF9DF98FBC015" box="[613,681,1630,1653]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="0" pageNumber="591" unit="km" value="15.0">15 km</quantity>
W of
<location id="9B6F088677275E246B84F9DF996AC015" LSID="urn:lsid:plazi:treatment:1619EF4B77275E2669FFF9BD9F2AC329:9B6F088677275E246B84F9DF996AC015" box="[748,824,1630,1653]" country="Zambia" name="Mbala." pageId="0" pageNumber="591" stateProvince="Northern Province">Mbala.</location>
</materialsCitation>
</paragraph>
<paragraph id="9E0F5E5D77275E2469AFF9009EA1C0BD" blockId="0.[151,1436,1596,2035]" pageId="0" pageNumber="591">
<materialsCitation id="2ED8540077275E2469AFF9009964C0DB" ID-GBIF-Occurrence="1836204546" collectingDate="2009-12-15" collectorName="Michal Bednarik" country="Zambia" location="Tanganyika" pageId="0" pageNumber="591" specimenCount="4" specimenCount-female="1" specimenCount-male="1" stateProvince="Northern" typeStatus="holotype">
<emphasis id="ACC4824F77275E2469AFF9009B22C0F8" bold="true" box="[199,368,1665,1688]" pageId="0" pageNumber="591">
<typeStatus id="410BE0FF77275E2469AFF9009AADC0F8" box="[199,255,1665,1688]" pageId="0" pageNumber="591">Type</typeStatus>
material.
</emphasis>
<typeStatus id="410BE0FF77275E246811F9009B8EC0F8" box="[377,476,1665,1688]" pageId="0" pageNumber="591" type="holotype">Holotype</typeStatus>
(male) and
<specimenCount id="88B695D477275E246B0DF90098B0C0F8" box="[613,738,1665,1688]" pageId="0" pageNumber="591" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
(
<specimenCount id="88B695D477275E246B9FF9009912C0F8" box="[759,832,1665,1688]" pageId="0" pageNumber="591" type="male">1 male</specimenCount>
and
<specimenCount id="88B695D477275E246A12F900998DC0F8" box="[890,991,1665,1688]" pageId="0" pageNumber="591" type="female">1 female</specimenCount>
): “
<collectingCountry id="E6A71ECD77275E246A95F9009E39C0F8" box="[1021,1131,1665,1688]" name="Zambia" pageId="0" pageNumber="591">ZAMBIA</collectingCountry>
,
<collectingRegion id="5C7490BF77275E246D11F9009E8BC0F8" box="[1145,1241,1665,1688]" country="Zambia" name="Northern" pageId="0" pageNumber="591">Northern</collectingRegion>
prov. |
<locationDeviation id="8FBD477F77275E246C5AF9009A89C0DB" pageId="0" pageNumber="591">
<quantity id="5948F3B877275E246C5AF9009F2AC0F8" box="[1330,1400,1665,1688]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="0" pageNumber="591" unit="km" value="15.0">15 km</quantity>
W Mbala
</locationDeviation>
- lake |
<location id="9B6F088677275E246847F9259BFCC0DB" LSID="urn:lsid:plazi:treatment:1619EF4B77275E2669FFF9BD9F2AC329:9B6F088677275E246847F9259BFCC0DB" box="[303,430,1700,1723]" country="Zambia" name="Tanganyika" pageId="0" pageNumber="591" stateProvince="Northern">Tanganyika</location>
,
<collectingDate id="FA4A817577275E2468D0F9259814C0DB" box="[440,582,1700,1723]" pageId="0" pageNumber="591" value="2009-12-15">
<date id="EA0E789D77275E2468D0F9259814C0DB" box="[440,582,1700,1723]" pageId="0" pageNumber="591" value="2009-12-15">15. XII. 2009</date>
</collectingDate>
|
<collectorName id="33453B8B77275E246B30F925995EC0DB" box="[600,780,1700,1723]" pageId="0" pageNumber="591">Michal Bednařík</collectorName>
leg
</materialsCitation>
.
<materialsCitation id="2ED8540077275E246A57F9259EBDC0BD" ID-GBIF-Occurrence="1836204547" country="Czech Republic" county="Velenice" elevation="1500" location="Narodni" municipality="Richard Sehnal" pageId="0" pageNumber="591" specimenCount="3" stateProvince="Praha" typeStatus="holotype">
<elevation id="159DB96E77275E246A57F92599DDC0DB" box="[831,911,1700,1723]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="0" pageNumber="591" unit="m" value="1500.0">
<quantity id="5948F3B877275E246A57F92599DDC0DB" box="[831,911,1700,1723]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="0" pageNumber="591" unit="m" value="1500.0">1500 m</quantity>
</elevation>
[printed]”.
<typeStatus id="410BE0FF77275E246D64F9259E13C0DB" box="[1036,1089,1700,1723]" pageId="0" pageNumber="591">Type</typeStatus>
depository:
<typeStatus id="410BE0FF77275E246DADF9259F70C0DB" box="[1221,1314,1700,1723]" pageId="0" pageNumber="591" type="holotype">holotype</typeStatus>
in
<location id="9B6F088677275E246C2CF9259FC9C0DB" LSID="urn:lsid:plazi:treatment:1619EF4B77275E2669FFF9BD9F2AC329:9B6F088677275E246C2CF9259FC9C0DB" box="[1348,1435,1700,1723]" country="Czech Republic" county="Velenice" municipality="Richard Sehnal" name="Narodni" pageId="0" pageNumber="591" stateProvince="Praha">Národní</location>
muzeum
<collectingRegion id="5C7490BF77275E246991F9479B6AC0BD" box="[249,312,1734,1757]" country="Czech Republic" name="Praha" pageId="0" pageNumber="591">Praha</collectingRegion>
,
<collectingCountry id="E6A71ECD77275E24682BF9479BBDC0BD" box="[323,495,1734,1757]" name="Czech Republic" pageId="0" pageNumber="591">Czech Republic</collectingCountry>
, both
<typeStatus id="410BE0FF77275E246B58F94798C4C0BD" box="[560,662,1734,1757]" pageId="0" pageNumber="591" type="paratype">paratypes</typeStatus>
in
<collectingMunicipality id="7E6BC42777275E246BD0F9479909C0BD" box="[696,859,1734,1757]" pageId="0" pageNumber="591">Richard Sehnal</collectingMunicipality>
collection,
<collectingCounty id="776E26D177275E246AB1F9479E6AC0BD" box="[985,1080,1734,1757]" pageId="0" pageNumber="591">Velenice</collectingCounty>
,
<collectingCountry id="E6A71ECD77275E246D2AF9479EBDC0BD" box="[1090,1263,1734,1757]" name="Czech Republic" pageId="0" pageNumber="591">Czech Republic</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D6AA0DD677275E2669AFF9689941C237" lastPageId="2" lastPageNumber="593" pageId="0" pageNumber="591" type="description">
<paragraph id="9E0F5E5D77275E2469AFF9689EF0C1CD" blockId="0.[151,1436,1596,2035]" pageId="0" pageNumber="591">
<emphasis id="ACC4824F77275E2469AFF968987BC160" bold="true" box="[199,553,1769,1792]" pageId="0" pageNumber="591">
Description of
<typeStatus id="410BE0FF77275E246818F9689B80C160" box="[368,466,1769,1792]" pageId="0" pageNumber="591" type="holotype">holotype</typeStatus>
(male).
</emphasis>
Body length
<quantity id="5948F3B877275E246BD4F968994CC160" box="[700,798,1769,1792]" metricMagnitude="-2" metricUnit="m" metricValue="2.23" pageId="0" pageNumber="591" unit="mm" value="22.3">22.3 mm</quantity>
, elongate, almost parallel-sided, gently convex. Dorsal and ventral surface weakly shiny, head blackish brown, pronotum and elytra chestnut brown, pronotum with darker disk, pronotal and elytral sides darker, macrosetation almost white (
<figureCitation id="068B42D877275E246A2BF8AF99E9C125" box="[835,955,1838,1861]" captionStart="FIGURES 1" captionStartId="1.[151,258,1914,1936]" captionTargetBox="[171,1426,470,1853]" captionTargetId="figure@1.[151,1436,437,1891]" captionTargetPageId="1" captionText="FIGURES 1A–F. Eulepida mbala new species, holotype (male) and paratype No. 1 (female). Habitus, dorsal view: A, holotype; B, paratype. Pygidium, lateral view, male (upper) and female (lower). C, Eulepida mbala new species; D, Eulepida anatina Brenske, 1896; E, Eulepida tschindeanaPéringuey, 1904;F, Eulepida werneri Lacroix,2010.Not to scale." httpUri="https://zenodo.org/record/1206814/files/figure.png" pageId="0" pageNumber="591">Fig. 1A–B</figureCitation>
). Dorsal surface of head covered with erect macrosetae, pronotum, scutellum and elytra covered with narrow, medium long, recumbent macrosetae; ventral surface of thorax and abdomen with narrow, pale yellow, short and long, recumbent macrosetae and with sparse, yellow, long, erect macrosetae. Head appendages and legs covered with pale yellow, medium long macrosetae.
</paragraph>
<paragraph id="9E0F5E5D77275E2569AFF8389B0FC71F" blockId="0.[151,1436,1596,2035]" lastBlockId="1.[151,1436,152,383]" lastPageId="1" lastPageNumber="592" pageId="0" pageNumber="591">
Head with labrum transverse, deeply bilobed; lobes rounded, irregularly and weakly punctate; lobes covered with long, erect macrosetae. Head including clypeus densely wrinkled and coarsely punctate; each puncture with long, erect macroseta. Macrosetae
<date id="EA0E789D77265E2568CEFF199B92C6CF" box="[422,448,152,175]" pageId="1" pageNumber="592">4x</date>
longer than puncture diameter. Clypeus sparsely, evenly, moderately punctate with macrosetae, as on head. Without frontoclypeal carina. Frons densely punctate with macrosetae, as on head. Occipital carina poorly indicated. Occiput sparsely, evenly, moderately punctate. Canthus narrow, short, glabrous. Eye large, distinctly extending beyond canthus. Antenna with 10 antennomeres; club with three antennomeres, almost straight, 1.7x longer than antennal shaft (antennomeres 1–7 combined). Antennomeres 1–7 with sparse long macrosetae; club with sparse short macrosetae. Terminal maxillary palpomere expanded near midlength, club-shaped, shorter than palpomeres 2 and 3 combined.
</paragraph>
<caption id="CACF0ED577265E2569FFF8FB9E6EC1AB" httpUri="https://zenodo.org/record/1206814/files/figure.png" pageId="1" pageNumber="592" startId="1.[151,258,1914,1936]" targetBox="[171,1426,470,1853]" targetPageId="1">
<paragraph id="9E0F5E5D77265E2569FFF8FB9E6EC1AB" blockId="1.[151,1436,1914,1995]" pageId="1" pageNumber="592">
<emphasis id="ACC4824F77265E2569FFF8FB9B16C1F0" bold="true" box="[151,324,1914,1936]" pageId="1" pageNumber="592">FIGURES 1A–F.</emphasis>
<taxonomicName id="59B025DE77265E256823F8FB9BB7C1F0" authority="Sehnal, 2018" authorityName="Sehnal" authorityYear="2018" box="[331,485,1914,1936]" class="Insecta" family="Melolonthidae" genus="Eulepida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="mbala" status="sp. nov.">
<emphasis id="ACC4824F77265E256823F8FB9BB7C1F0" box="[331,485,1914,1936]" italics="true" pageId="1" pageNumber="592">Eulepida mbala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7F73F3477265E256884F8FB9830C1F0" box="[492,610,1914,1936]" pageId="1" pageNumber="592" rank="species">
<emphasis id="ACC4824F77265E256884F8FB9830C1F0" bold="true" box="[492,610,1914,1936]" pageId="1" pageNumber="592">new species</emphasis>
</taxonomicNameLabel>
, holotype (male) and paratype No. 1 (female). Habitus, dorsal view: A, holotype; B, paratype. Pygidium, lateral view, male (upper) and female (lower). C,
<taxonomicName id="59B025DE77265E256A26F81699BBC1CD" authority="Sehnal, 2018" authorityName="Sehnal" authorityYear="2018" box="[846,1001,1943,1965]" class="Insecta" family="Melolonthidae" genus="Eulepida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="mbala" status="sp. nov.">
<emphasis id="ACC4824F77265E256A26F81699BBC1CD" box="[846,1001,1943,1965]" italics="true" pageId="1" pageNumber="592">Eulepida mbala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7F73F3477265E256A99F8199E3BC1CE" box="[1009,1129,1944,1966]" pageId="1" pageNumber="592" rank="species">
<emphasis id="ACC4824F77265E256A99F8199E3BC1CE" bold="true" box="[1009,1129,1944,1966]" pageId="1" pageNumber="592">new species</emphasis>
</taxonomicNameLabel>
; D,
<taxonomicName id="59B025DE77265E256DFFF8169A98C1AB" authority="Brenske, 1896" authorityName="Brenske" authorityYear="1896" class="Insecta" family="Melolonthidae" genus="Eulepida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="anatina">
<emphasis id="ACC4824F77265E256DFFF8169F6DC1CD" box="[1175,1343,1943,1965]" italics="true" pageId="1" pageNumber="592">Eulepida anatina</emphasis>
Brenske, 1896
</taxonomicName>
; E,
<taxonomicName id="59B025DE77265E256986F8349831C1AB" authority="Peringuey, 1904" authorityName="Peringuey" authorityYear="1904" box="[238,611,1973,1995]" class="Insecta" family="Melolonthidae" genus="Eulepida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="tschindeana">
<emphasis id="ACC4824F77265E256986F8349BEDC1AB" box="[238,447,1973,1995]" italics="true" pageId="1" pageNumber="592">Eulepida tschindeana</emphasis>
Péringuey, 1904
</taxonomicName>
; F,
<taxonomicName id="59B025DE77265E256BECF83499E8C1AB" authority="Lacroix, 2010" authorityName="Lacroix" authorityYear="2010" box="[644,954,1973,1995]" class="Insecta" family="Melolonthidae" genus="Eulepida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="werneri">
<emphasis id="ACC4824F77265E256BECF8349979C1AB" box="[644,811,1973,1995]" italics="true" pageId="1" pageNumber="592">Eulepida werneri</emphasis>
<bibRefCitation id="FA2123AC77265E256A59F83499E8C1AB" author="Lacroix, M." bookContentInfo="277 pp." box="[817,954,1973,1995]" journalOrPublisher="Editions Marc Lacroix, Paris" pageId="1" pageNumber="592" refId="ref2395" refString="Lacroix, M. (2010) Melolonthinae afrotropicaux (Scarabaeoidea, Melolonthidae). Genera et catalogue commente. Editions Marc Lacroix, Paris, 277 pp., 192 pls." title="Melolonthinae afrotropicaux (Scarabaeoidea, Melolonthidae). Genera et catalogue commente" type="book" year="2010">Lacroix, 2010</bibRefCitation>
</taxonomicName>
. Not to scale.
</paragraph>
</caption>
<paragraph id="9E0F5E5D77255E2669AFFF199FD2C777" blockId="2.[151,1436,152,1353]" pageId="2" pageNumber="593">
Pronotum transverse, widest at base, anterolateral wrinkles present. Lateral margins crenulate, with short macrosetose. Base bordered, medially gently arched toward elytra, with smooth margin paralleled by a row of fine punctures and sharp lateral angles. Disk with deep punctures separated by 1–2 puncture diameters and macrosetae
<date id="EA0E789D77255E266CE9FF5C9FC9C694" box="[1409,1435,221,244]" pageId="2" pageNumber="593">3x</date>
longer than puncture diameters. Punctation of remaining surface similar, macrosetae yellowish white, semirecumbent.
</paragraph>
<paragraph id="9E0F5E5D77255E2669AFFEA39ECBC73C" blockId="2.[151,1436,152,1353]" pageId="2" pageNumber="593">
Scutellum large, almost equilaterally triangular, with sides and apex rounded; punctation irregular, each puncture bearing narrow, white, almost recumbent scale-like macroseta
<date id="EA0E789D77255E266A59FEC4991EC73C" box="[817,844,325,348]" pageId="2" pageNumber="593">3x</date>
longer than puncture diameter.
</paragraph>
<paragraph id="9E0F5E5D77255E2669AFFEE99E17C469" blockId="2.[151,1436,152,1353]" pageId="2" pageNumber="593">Elytra weakly convex, widest near rounded apex; apical angle of elytron approximately rectangular. Striae absent. Humeral umbone present, weakly swollen. Surface not microsculptured, matt, with shallow, coarse punctures arranged in evenly spaced rows separated by 2 puncture diameters, and interspersed with fine punctures separated by 1–3 puncture diameters. Each puncture bears a narrow, yellowish-white, weakly recumbent macroseta, laterally some macrosetae become scale-like. Epipleuron distinct, complete, wide, membranous, laterally glabrous.</paragraph>
<paragraph id="9E0F5E5D77255E2669AFFD949910C44C" blockId="2.[151,1436,152,1353]" box="[199,834,533,556]" pageId="2" pageNumber="593">Macropterous species, wings fully developed in both sexes.</paragraph>
<paragraph id="9E0F5E5D77255E2669AFFDB99899C4D7" blockId="2.[151,1436,152,1353]" pageId="2" pageNumber="593">Legs with femora narrow, shiny, irregularly punctate, macrosetaceous. Protibia narrow, distinctly bidentate; inner terminal spur at level of proximal tooth. Mesotibia and metatibia slightly expanded distally, with two macrosetiferous longitudinal carinae. Metatibial apical carina vague. Upper terminal spine of metatibia slightly longer than lower spine, apically pointed. Claws bifid, with ventrobasal teeth.</paragraph>
<paragraph id="9E0F5E5D77255E2669AFFD4399FEC4B9" blockId="2.[151,1436,152,1353]" box="[199,940,706,729]" pageId="2" pageNumber="593">Ventrum covered by narrow, recumbent, yellowish-white macrosetae.</paragraph>
<paragraph id="9E0F5E5D77255E2669AFFD649B54C521" blockId="2.[151,1436,152,1353]" pageId="2" pageNumber="593">
Pygidium slightly longer than wide, convex, completely bordered, apically broadly rounded and with a pronounced elongate cuticular invagination; evenly covered by deep macrosetose punctures, macrosetae
<date id="EA0E789D77255E266DFBFC899EFFC57F" box="[1171,1197,776,799]" pageId="2" pageNumber="593">3x</date>
longer than puncture diameters.
</paragraph>
<paragraph id="9E0F5E5D77255E2669AFFCCC9E84C5E7" blockId="2.[151,1436,152,1353]" pageId="2" pageNumber="593">
Male genitalia (
<figureCitation id="068B42D877255E266807FCCC9B82C504" box="[367,464,845,868]" captionStart="FIGURES 2" captionStartId="3.[151,258,1307,1329]" captionTargetBox="[180,1410,211,1262]" captionTargetId="figure@3.[151,1436,191,1285]" captionTargetPageId="3" captionText="FIGURES 2A–D. Aedeagus of species group II of Eulepida, dorsal (left) and lateral (right) views. A, Eulepida mbala new species, holotype; B, Eulepida anatina Brenske, 1896; C, Eulepida tschindeana Péringuey, 1904; D, Eulepida werneri Lacroix, 2010. Not to scale." httpUri="https://zenodo.org/record/1206816/files/figure.png" pageId="2" pageNumber="593">Figs. 2A</figureCitation>
). Parameres symmetrical, longer than phallobase, evenly curved in ventral aspect, rounded with double hump in lateral aspect, apically curved and expanded in dorsal aspect; macrosetae absent.
</paragraph>
<paragraph id="9E0F5E5D77255E2669AFFC139FC6C5C9" blockId="2.[151,1436,152,1353]" box="[199,1428,914,937]" pageId="2" pageNumber="593">
<emphasis id="ACC4824F77255E2669AFFC139BFEC5C9" bold="true" box="[199,428,914,937]" pageId="2" pageNumber="593">Variability in males.</emphasis>
The only known male
<typeStatus id="410BE0FF77255E266BCAFC1398ACC5C9" box="[674,766,914,937]" pageId="2" pageNumber="593" type="paratype">paratype</typeStatus>
is slightly smaller,
<quantity id="5948F3B877255E266AA7FC139E7CC5C9" box="[975,1070,914,937]" metricMagnitude="-2" metricUnit="m" metricValue="2.2199999999999998" pageId="2" pageNumber="593" unit="mm" value="22.2">22.2 mm</quantity>
long. Colouration as in
<typeStatus id="410BE0FF77255E266C47FC139FC2C5C9" box="[1327,1424,914,937]" pageId="2" pageNumber="593" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="9E0F5E5D77255E2669AFFC349B68C254" blockId="2.[151,1436,152,1353]" pageId="2" pageNumber="593">
<emphasis id="ACC4824F77255E2669AFFC349BFAC5AC" bold="true" box="[199,424,949,972]" pageId="2" pageNumber="593">Sexual dimorphism</emphasis>
. Female differs from male in the following respects: body length 23.0 mm; antennal club shorter; border of clypeus evenly rounded (
<figureCitation id="068B42D877255E266B1AFC599895C58F" box="[626,711,984,1007]" captionStart="FIGURES 1" captionStartId="1.[151,258,1914,1936]" captionTargetBox="[171,1426,470,1853]" captionTargetId="figure@1.[151,1436,437,1891]" captionTargetPageId="1" captionText="FIGURES 1A–F. Eulepida mbala new species, holotype (male) and paratype No. 1 (female). Habitus, dorsal view: A, holotype; B, paratype. Pygidium, lateral view, male (upper) and female (lower). C, Eulepida mbala new species; D, Eulepida anatina Brenske, 1896; E, Eulepida tschindeanaPéringuey, 1904;F, Eulepida werneri Lacroix,2010.Not to scale." httpUri="https://zenodo.org/record/1206814/files/figure.png" pageId="2" pageNumber="593">Fig. 1B</figureCitation>
); protibial teeth wider and longer; elytra widest behind midlength (
<figureCitation id="068B42D877255E2669F7FC7B9AA0C271" box="[159,242,1018,1041]" captionStart="FIGURES 1" captionStartId="1.[151,258,1914,1936]" captionTargetBox="[171,1426,470,1853]" captionTargetId="figure@1.[151,1436,437,1891]" captionTargetPageId="1" captionText="FIGURES 1A–F. Eulepida mbala new species, holotype (male) and paratype No. 1 (female). Habitus, dorsal view: A, holotype; B, paratype. Pygidium, lateral view, male (upper) and female (lower). C, Eulepida mbala new species; D, Eulepida anatina Brenske, 1896; E, Eulepida tschindeanaPéringuey, 1904;F, Eulepida werneri Lacroix,2010.Not to scale." httpUri="https://zenodo.org/record/1206814/files/figure.png" pageId="2" pageNumber="593">Fig. 1B</figureCitation>
); pygidium simple, wider, shorter, concave (
<figureCitation id="068B42D877255E266BB8FC7B9971C271" box="[720,803,1018,1041]" captionStart="FIGURES 1" captionStartId="1.[151,258,1914,1936]" captionTargetBox="[171,1426,470,1853]" captionTargetId="figure@1.[151,1436,437,1891]" captionTargetPageId="1" captionText="FIGURES 1A–F. Eulepida mbala new species, holotype (male) and paratype No. 1 (female). Habitus, dorsal view: A, holotype; B, paratype. Pygidium, lateral view, male (upper) and female (lower). C, Eulepida mbala new species; D, Eulepida anatina Brenske, 1896; E, Eulepida tschindeanaPéringuey, 1904;F, Eulepida werneri Lacroix,2010.Not to scale." httpUri="https://zenodo.org/record/1206814/files/figure.png" pageId="2" pageNumber="593">Fig. 1C</figureCitation>
); metatibia more strongly expanded distally; tarsomeres of all legs shorter.
</paragraph>
<paragraph id="9E0F5E5D77255E2669AFFBC19941C237" blockId="2.[151,1436,152,1353]" box="[199,787,1088,1111]" pageId="2" pageNumber="593">
<emphasis id="ACC4824F77255E2669AFFBC19B88C237" bold="true" box="[199,474,1088,1111]" pageId="2" pageNumber="593">Geographic distribution</emphasis>
. Zambia, Northern Province.
</paragraph>
</subSubSection>
<subSubSection id="D6AA0DD677255E2669AFFBE39ABBC347" pageId="2" pageNumber="593" type="diagnosis">
<paragraph id="9E0F5E5D77255E2669AFFBE39ABBC347" blockId="2.[151,1436,152,1353]" pageId="2" pageNumber="593">
<emphasis id="ACC4824F77255E2669AFFBE39BE9C219" bold="true" box="[199,443,1122,1145]" pageId="2" pageNumber="593">Differential diagnosis</emphasis>
.
<taxonomicName id="59B025DE77255E2668A1FBE39821C219" authority="Sehnal, 2018" authorityName="Sehnal" authorityYear="2018" box="[457,627,1122,1145]" class="Insecta" family="Melolonthidae" genus="Eulepida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="mbala" status="sp. nov.">
<emphasis id="ACC4824F77255E2668A1FBE39821C219" box="[457,627,1122,1145]" italics="true" pageId="2" pageNumber="593">Eulepida mbala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7F73F3477255E266B13FBE398A9C219" box="[635,763,1122,1145]" pageId="2" pageNumber="593" rank="species">new species</taxonomicNameLabel>
can be differentiated from all other species of the genus using the following diagnostic characters in males: protibia bidentate; antennal club distinctly longer than antennal shaft; pygidium narrow, longer than wide, with a pronounced apical invagination; scales on pronotum shorter and wider than on elytra; macrosetae on elytra narrow, with interspersed isolated scale-like macrosetae; aedeagus evenly curved in ventral aspect, rounded with double hump in lateral aspect, apically curved and expanded in dorsal aspect; body length &lt;
<quantity id="5948F3B877255E2669FFFA919AB7C347" box="[151,229,1296,1319]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="593" unit="mm" value="23.0">23 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="D6AA0DD677255E2669AFFAB39F2AC329" box="[199,1400,1330,1353]" pageId="2" pageNumber="593" type="etymology">
<paragraph id="9E0F5E5D77255E2669AFFAB39F2AC329" blockId="2.[151,1436,152,1353]" box="[199,1400,1330,1353]" pageId="2" pageNumber="593">
<emphasis id="ACC4824F77255E2669AFFAB39B12C329" bold="true" box="[199,320,1330,1353]" pageId="2" pageNumber="593">Etymology</emphasis>
. Derived from the name of the
<typeStatus id="410BE0FF77255E266BF8FAB398ECC329" box="[656,702,1330,1353]" pageId="2" pageNumber="593">type</typeStatus>
locality Mbala (
<collectingRegion id="5C7490BF77255E266A05FAB39E65C329" box="[877,1079,1330,1353]" country="Zambia" name="Northern" pageId="2" pageNumber="593">Northern Province</collectingRegion>
,
<collectingCountry id="E6A71ECD77255E266D2AFAB39ECAC329" box="[1090,1176,1330,1353]" name="Zambia" pageId="2" pageNumber="593">Zambia</collectingCountry>
); noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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