treatments-xml/data/7E/42/17/7E421778FF8FFFB1FF10FCD1A6485FEA.xml
2024-06-21 12:41:40 +02:00

220 lines
27 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="B3135F6CF2E4D153A7D9FC417CF61EF2" ID-CLB-Dataset="21199" ID-DOI="10.11646/zootaxa.4615.1.10" ID-GBIF-Dataset="806f3a8f-219d-48ff-9b05-8ff10546940a" ID-ISSN="1175-5326" ID-Zenodo-Dep="3995294" ID-ZooBank="70175096-AA78-4B74-BF38-44804F8FAD59" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1598056528139" checkinUser="felipe" docAuthor="Benetti, Cesar J., Short, Andrew E. Z. &amp; Michat, Mariano C." docDate="2019" docId="7E421778FF8FFFB1FF10FCD1A6485FEA" docLanguage="en" docName="zootaxa.4615.1.10.pdf" docOrigin="Zootaxa 4615 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hamadiana Benetti &amp; Short &amp; Michat 2019, gen. n." docType="treatment" docUuid="15244304-1BD6-4054-B414-CB8216E4F5C7" docUuidSource="ZooBank" docVersion="7" lastPageNumber="177" masterDocId="827B6F00FF8EFFB0FF87FFF0A2645971" masterDocTitle="Hamadiana chapadensis, a new genus and species of diving beetle from Brazil (Coleoptera, Dytiscidae, Laccophilinae, Laccophilini)" masterLastPageNumber="184" masterPageNumber="176" pageNumber="177" updateTime="1698845169827" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="19F9C96FA542A5019CDA7B0E019D2194" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B0FD6EA9877F535EB5C4E3E785C6A444">
<mods:title id="0484010C49E34750CA5256BA041188EF">Hamadiana chapadensis, a new genus and species of diving beetle from Brazil (Coleoptera, Dytiscidae, Laccophilinae, Laccophilini)</mods:title>
</mods:titleInfo>
<mods:name id="DAA81F7CC636B2433DC306CA3AA90961" type="personal">
<mods:role id="752C047218A77C70606DC0C259CE96E6">
<mods:roleTerm id="231AC18E48800610EF3FB2B6FF00F992">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9AEEBC0A4473654018AC6D069C77A889">Benetti, Cesar J.</mods:namePart>
<mods:affiliation id="67F922A329207C2B30425F00D3169DE5">Coordenação de Biodiversidade, Programa de Pós-Graduação em Entomologia (PPGEnt), Instituto Nacional de Pesquisas da Amazônia (INPA), Av. André Araújo 2936, CEP 69067 - 375, Manaus, AM, Brazil. E-mail: cjbenetti @ gmail. com</mods:affiliation>
</mods:name>
<mods:name id="AAB20BAE93BA8BFAB412FB8106DB8FEC" type="personal">
<mods:role id="19C32E6F1055AF1714FA246AFDD1CEB2">
<mods:roleTerm id="A53B4BAE0CDC04C7D03FA4A0159A77FA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E20816C5535E2A0BAEBE5D159778FFBB">Short, Andrew E. Z.</mods:namePart>
<mods:affiliation id="1D8985E507C1DC1FCA2F62807259FB35">Department of Ecology &amp; Evolutionary Biology, Division of Entomology, Biology Institute, University of Kansas, Lawrence, United States. E-mail: aezshort @ ku. edu</mods:affiliation>
</mods:name>
<mods:name id="1E8EA541B1D6A8B1A128BC48BCB8521E" type="personal">
<mods:role id="C23F45408662599A3DA2275A79EEB19E">
<mods:roleTerm id="5F3463C6BA1E7627AD848B95A035B661">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CFE11279D9A49C11293882A4144DCB15">Michat, Mariano C.</mods:namePart>
<mods:affiliation id="27B0C3B12D2459C95C428BC11A3AD869">Laboratory of Entomology, IBBEA, CONICET-UBA., DBBE-FCEN, University of Buenos Aires, Argentina.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="27D3EF0AC4FC62FD34C3ED55A80A2F8A">text</mods:typeOfResource>
<mods:relatedItem id="C8C73E970C193864DEF4857F47C0582C" type="host">
<mods:titleInfo id="6D8FB5C3FD329BD3D1900A807542AB13">
<mods:title id="959C82C957D3698A103A7C0CA567E220">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="22716BD5723DAFD508CEC678631506E1">
<mods:date id="85F408D385201DE1B0ACD8D72F7068AB">2019</mods:date>
<mods:detail id="6E7B3ACD8E1D22E4C6F7AEF11222832F" type="pubDate">
<mods:number id="C58292042627567A60C4102106B6FE9B">2019-06-12</mods:number>
</mods:detail>
<mods:detail id="16239399E8B30353243F041EB4013237" type="volume">
<mods:number id="B0330FEA48BDED1DD7C73E9D1AA9D291">4615</mods:number>
</mods:detail>
<mods:detail id="1D1FFA88F09FAAB629F802DC27B8B420" type="issue">
<mods:number id="BB7574D809C02DA0D7B040155848CB77">1</mods:number>
</mods:detail>
<mods:extent id="7FD922CBB5685AFF05696D866C49A68E" unit="page">
<mods:start id="EAE78BBC3BFDF4C5142AF5E2B3561F06">176</mods:start>
<mods:end id="B28B9B2D2888D9C185DA8284A5155FC6">184</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="1FD0CC0B9CBCE8EB15292B75F1E9858E">journal article</mods:classification>
<mods:identifier id="7BB202EE0CD7A4C49229BD55B52D72D5" type="CLB-Dataset">21199</mods:identifier>
<mods:identifier id="8E3B0A3DA584E09889A5776E2FF6C491" type="DOI">10.11646/zootaxa.4615.1.10</mods:identifier>
<mods:identifier id="C4AF271FE802FF2EEF7515CF8F05D68C" type="GBIF-Dataset">806f3a8f-219d-48ff-9b05-8ff10546940a</mods:identifier>
<mods:identifier id="41F934D329821994BA6CD4772AF9D457" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A3DA28692D7D74AE2DF476EA08A16998" type="Zenodo-Dep">3995294</mods:identifier>
<mods:identifier id="ABC840272CCAC5120EAB919F3F5860E1" type="ZooBank">70175096-AA78-4B74-BF38-44804F8FAD59</mods:identifier>
</mods:mods>
<treatment id="7E421778FF8FFFB1FF10FCD1A6485FEA" ID-DOI="http://doi.org/10.5281/zenodo.4324071" ID-GBIF-Taxon="167292781" ID-Zenodo-Dep="4324071" LSID="urn:lsid:zoobank.org:act:15244304-1BD6-4054-B414-CB8216E4F5C7" httpUri="http://treatment.plazi.org/id/7E421778FF8FFFB1FF10FCD1A6485FEA" lastPageNumber="177" pageId="1" pageNumber="177">
<subSubSection id="BEF1F5E5FF8FFFB1FF10FCD1A3E45A4A" box="[151,384,801,828]" pageId="1" pageNumber="177" type="nomenclature">
<paragraph id="F654A66EFF8FFFB1FF10FCD1A3E45A4A" blockId="1.[151,951,801,899]" box="[151,384,801,828]" pageId="1" pageNumber="177">
<heading id="AD1C1102FF8FFFB1FF10FCD1A3E45A4A" bold="true" box="[151,384,801,828]" fontSize="11" level="1" pageId="1" pageNumber="177" reason="1">
<emphasis id="C49F7A7CFF8FFFB1FF10FCD1A3E45A4A" bold="true" box="[151,384,801,828]" pageId="1" pageNumber="177">
<taxonomicName id="31EBDDEDFF8FFFB1FF10FCD1A3425A4D" ID-CoL="8LZLD" authority="Benetti &amp; Short &amp; Michat, 2019" authorityName="Benetti &amp; Short &amp; Michat" authorityYear="2019" box="[151,294,801,828]" class="Insecta" family="Dytiscidae" genus="Hamadiana" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="C49F7A7CFF8FFFB1FF10FCD1A3425A4D" bold="true" box="[151,294,801,828]" italics="true" pageId="1" pageNumber="177">Hamadiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFACC707FF8FFFB1FEA9FCD1A3E45A4A" box="[302,384,801,827]" pageId="1" pageNumber="177" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BEF1F5E5FF8FFFB1FF10FCB5A1D35AF2" pageId="1" pageNumber="177" type="description">
<paragraph id="F654A66EFF8FFFB1FF10FCB5A37D5A2E" blockId="1.[151,951,801,899]" box="[151,281,836,863]" pageId="1" pageNumber="177">
(
<figureCitation id="6ED0BAEBFF8FFFB1FF18FCB5A3745A2E" box="[159,272,836,863]" captionStart-0="FIGURES 12" captionStart-1="FIGURES 37" captionStart-2="FIGURE 811" captionStart-3="FIGURES 1215" captionStartId-0="4.[151,264,1536,1561]" captionStartId-1="5.[151,264,1803,1828]" captionStartId-2="6.[151,250,1220,1245]" captionStartId-3="6.[151,264,1972,1997]" captionTargetBox-0="[151,1435,538,1448]" captionTargetBox-1="[151,1435,181,1732]" captionTargetBox-2="[184,1320,233,1174]" captionTargetBox-3="[163,1335,1312,1937]" captionTargetId-0="figure@4.[151,1435,538,1448]" captionTargetId-1="figure@5.[151,1435,181,1732]" captionTargetId-2="figure@6.[169,1418,174,1209]" captionTargetId-3="figure@6.[151,1435,1296,1960]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="6" captionText-0="FIGURES 12. Hamadiana chapadensis sp. n.: 1) habitus, dorsal view; 2) habitus, ventral view." captionText-1="FIGURES 37. Hamadiana chapadensis sp. n.: 3) posterior pronotal angle; 4) prosternal process; 5) metacoxa, arrow indicating stridulatory file; 6) abdomen and right hind-leg, arrows indicating simple apical tibial spurs and medial marginal protrusion on abdominal ventrite 3; 7) left metatarsus." captionText-2="FIGURE 811. Hamadiana chapadensis sp. n., ventral view: 8) prosternal process; 9) metacoxal processes; 10) last abdominal ventrite; 11) left metatarsus." captionText-3="FIGURES 1215. Hamadiana chapadensis sp. n.: 12) median lobe, dorsal view; 13) median lobe, left lateral view; 14) left paramere, left lateral view; 15) right paramere, left lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.3995296" figureDoi-1="http://doi.org/10.5281/zenodo.3995298" figureDoi-2="http://doi.org/10.5281/zenodo.3995300" figureDoi-3="http://doi.org/10.5281/zenodo.3995302" httpUri-0="https://zenodo.org/record/3995296/files/figure.png" httpUri-1="https://zenodo.org/record/3995298/files/figure.png" httpUri-2="https://zenodo.org/record/3995300/files/figure.png" httpUri-3="https://zenodo.org/record/3995302/files/figure.png" pageId="1" pageNumber="177">Figs 115</figureCitation>
)
</paragraph>
<paragraph id="F654A66EFF8FFFB1FF10FC98A1D35AF2" blockId="1.[151,951,801,899]" box="[151,951,872,899]" pageId="1" pageNumber="177">
<uri id="827AAA6CFF8FFFB1FF10FC98A1D35AF2" box="[151,951,872,899]" pageId="1" pageNumber="177">
urn:lsid:zoobank.org:act:
<uuid id="824D9CBBFF8FFFB1FE2CFC98A1D35AF2" box="[427,951,872,899]" pageId="1" pageNumber="177">15244304-1BD6-4054-B414-CB8216E4F5C7</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="BEF1F5E5FF8FFFB1FF10FC41A1C45ABA" box="[151,928,944,971]" pageId="1" pageNumber="177" type="type_taxon">
<paragraph id="F654A66EFF8FFFB1FF10FC41A1C45ABA" blockId="1.[151,928,944,971]" box="[151,928,944,971]" pageId="1" pageNumber="177">
<emphasis id="C49F7A7CFF8FFFB1FF10FC41A3565ABA" bold="true" box="[151,306,944,971]" pageId="1" pageNumber="177">
<typeStatus id="295018CCFF8FFFB1FF10FC41A2B65ABA" box="[151,210,945,971]" pageId="1" pageNumber="177">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="31EBDDEDFF8FFFB1FEBEFC41A02A5ABA" ID-CoL="8LZ6Q" authority="Benetti &amp; Short &amp; Michat, 2019" authorityName="Benetti &amp; Short &amp; Michat" authorityYear="2019" box="[313,590,945,971]" class="Insecta" family="Dytiscidae" genus="Hamadiana" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="species" species="chapadensis" status="sp. nov.">
<emphasis id="C49F7A7CFF8FFFB1FEBEFC41A02A5ABA" box="[313,590,945,971]" italics="true" pageId="1" pageNumber="177">Hamadiana chapadensis</emphasis>
</taxonomicName>
<emphasis id="C49F7A7CFF8FFFB1FDD1FC41A0F15ABA" bold="true" box="[598,661,945,971]" pageId="1" pageNumber="177">
<taxonomicNameLabel id="DFACC707FF8FFFB1FDD1FC41A0F15ABA" box="[598,661,945,971]" pageId="1" pageNumber="177" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by present designation.
</paragraph>
</subSubSection>
<subSubSection id="BEF1F5E5FF8FFFB1FF10FC08A6EE5DB6" pageId="1" pageNumber="177" type="description">
<paragraph id="F654A66EFF8FFFB1FF10FC08A6EE5DB6" blockId="1.[151,1437,1016,1691]" pageId="1" pageNumber="177">
<emphasis id="C49F7A7CFF8FFFB1FF10FC08A34D5D62" bold="true" box="[151,297,1016,1043]" pageId="1" pageNumber="177">Description.</emphasis>
The new genus is unique among
<taxonomicName id="31EBDDEDFF8FFFB1FD26FC09A14B5D62" baseAuthorityName="Miller &amp; Bergsten" baseAuthorityYear="2016" box="[673,815,1017,1043]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
in having the hind margin of metacoxal process deeply incised and medially slightly protruded backwards (
<figureCitation id="6ED0BAEBFF8FFFB1FD67FBEDA1415D46" box="[736,805,1053,1079]" captionStart="FIGURE 811" captionStartId="6.[151,250,1220,1245]" captionTargetBox="[184,1320,233,1174]" captionTargetId="figure@6.[169,1418,174,1209]" captionTargetPageId="6" captionText="FIGURE 811. Hamadiana chapadensis sp. n., ventral view: 8) prosternal process; 9) metacoxal processes; 10) last abdominal ventrite; 11) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995300" httpUri="https://zenodo.org/record/3995300/files/figure.png" pageId="1" pageNumber="177">Fig. 9</figureCitation>
). Other diagnostic characters are: antennomeres simple, not expanded; metacoxal lines not straight, converging anteriad in posterior 3/4 of their length, slightly diverging anteriad in anterior 1/4 (
<figureCitation id="6ED0BAEBFF8FFFB1FE2EFB95A38A5D0E" box="[425,494,1125,1151]" captionStart="FIGURE 811" captionStartId="6.[151,250,1220,1245]" captionTargetBox="[184,1320,233,1174]" captionTargetId="figure@6.[169,1418,174,1209]" captionTargetPageId="6" captionText="FIGURE 811. Hamadiana chapadensis sp. n., ventral view: 8) prosternal process; 9) metacoxal processes; 10) last abdominal ventrite; 11) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995300" httpUri="https://zenodo.org/record/3995300/files/figure.png" pageId="1" pageNumber="177">Fig. 9</figureCitation>
); metatibia with two simple apical spurs (
<figureCitation id="6ED0BAEBFF8FFFB1FC4FFB95A6695DF1" box="[968,1037,1125,1152]" captionStart="FIGURES 37" captionStartId="5.[151,264,1803,1828]" captionTargetBox="[151,1435,181,1732]" captionTargetId="figure@5.[151,1435,181,1732]" captionTargetPageId="5" captionText="FIGURES 37. Hamadiana chapadensis sp. n.: 3) posterior pronotal angle; 4) prosternal process; 5) metacoxa, arrow indicating stridulatory file; 6) abdomen and right hind-leg, arrows indicating simple apical tibial spurs and medial marginal protrusion on abdominal ventrite 3; 7) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995298" httpUri="https://zenodo.org/record/3995298/files/figure.png" pageId="1" pageNumber="177">Fig. 6</figureCitation>
). Moreover, the last metatarsomere is bilobed, with inner lobe long and acute and outer lobe about half shorter; the single metatarsal claw is curved, apically rounded and short, about as long as the outer lobe of last metatarsomere (
<figureCitation id="6ED0BAEBFF8FFFB1FBACFB5DA6185DB6" box="[1067,1148,1197,1223]" captionStart="FIGURE 811" captionStartId="6.[151,250,1220,1245]" captionTargetBox="[184,1320,233,1174]" captionTargetId="figure@6.[169,1418,174,1209]" captionTargetPageId="6" captionText="FIGURE 811. Hamadiana chapadensis sp. n., ventral view: 8) prosternal process; 9) metacoxal processes; 10) last abdominal ventrite; 11) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995300" httpUri="https://zenodo.org/record/3995300/files/figure.png" pageId="1" pageNumber="177">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BEF1F5E5FF8FFFB1FF40FB21A6595C7E" pageId="1" pageNumber="177" type="etymology">
<paragraph id="F654A66EFF8FFFB1FF40FB21A6595C7E" blockId="1.[151,1437,1016,1691]" pageId="1" pageNumber="177">
<emphasis id="C49F7A7CFF8FFFB1FF40FB21A32A5D9A" bold="true" box="[199,334,1233,1259]" pageId="1" pageNumber="177">Etymology.</emphasis>
This genus is named after Neusa Hamada, INPA, in recognition of her significant contribution to the knowledge of aquatic insects in Amazonia. The gender of the name is feminine.
</paragraph>
</subSubSection>
<subSubSection id="BEF1F5E5FF8FFFB1FF40FAE8A6485FEA" pageId="1" pageNumber="177" type="discussion">
<paragraph id="F654A66EFF8FFFB1FF40FAE8A6485FEA" blockId="1.[151,1437,1016,1691]" pageId="1" pageNumber="177">
<emphasis id="C49F7A7CFF8FFFB1FF40FAE8A3285C42" bold="true" box="[199,332,1304,1331]" pageId="1" pageNumber="177">Discussion.</emphasis>
Among the Neotropical genera of
<taxonomicName id="31EBDDEDFF8FFFB1FD5CFAE9A10F5C42" baseAuthorityName="Miller &amp; Bergsten" baseAuthorityYear="2016" box="[731,875,1305,1331]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
, the new genus is easily differentiated from
<taxonomicName id="31EBDDEDFF8FFFB1FAEEFAEAA2925C26" authorityName="Leach" authorityYear="1815" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="C49F7A7CFF8FFFB1FAEEFAEAA2925C26" italics="true" pageId="1" pageNumber="177">Laccophilus</emphasis>
</taxonomicName>
and
<taxonomicName id="31EBDDEDFF8FFFB1FEA9FACDA3C05C26" authorityName="Steiner" authorityYear="1981" box="[302,420,1341,1367]" class="Insecta" family="Dytiscidae" genus="Napodytes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="C49F7A7CFF8FFFB1FEA9FACDA3C05C26" box="[302,420,1341,1367]" italics="true" pageId="1" pageNumber="177">Napodytes</emphasis>
</taxonomicName>
in having two simple apical spurs on metatibiae (single apical spur in
<taxonomicName id="31EBDDEDFF8FFFB1FB43FACDA75E5C26" authorityName="Steiner" authorityYear="1981" box="[1220,1338,1341,1367]" class="Insecta" family="Dytiscidae" genus="Napodytes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="C49F7A7CFF8FFFB1FB43FACDA75E5C26" box="[1220,1338,1341,1367]" italics="true" pageId="1" pageNumber="177">Napodytes</emphasis>
</taxonomicName>
and two apically bifid spurs in
<taxonomicName id="31EBDDEDFF8FFFB1FE0BFA91A0715C0A" authorityName="Leach" authorityYear="1815" box="[396,533,1377,1403]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="C49F7A7CFF8FFFB1FE0BFA91A0715C0A" box="[396,533,1377,1403]" italics="true" pageId="1" pageNumber="177">Laccophilus</emphasis>
</taxonomicName>
). This character is shared with
<taxonomicName id="31EBDDEDFF8FFFB1FCEBFA91A18E5C0A" authorityName="Regimbart" authorityYear="1895" box="[876,1002,1377,1403]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="C49F7A7CFF8FFFB1FCEBFA91A18E5C0A" box="[876,1002,1377,1403]" italics="true" pageId="1" pageNumber="177">Laccodytes</emphasis>
</taxonomicName>
and
<taxonomicName id="31EBDDEDFF8FFFB1FB99FA92A6C85C0A" authorityName="Toledo &amp; Michat" authorityYear="2015" box="[1054,1196,1378,1403]" class="Insecta" family="Dytiscidae" genus="Laccomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="C49F7A7CFF8FFFB1FB99FA92A6C85C0A" box="[1054,1196,1378,1403]" italics="true" pageId="1" pageNumber="177">Laccomimus</emphasis>
</taxonomicName>
, but
<taxonomicName id="31EBDDEDFF8FFFB1FB66FA91A7015C0A" authority="Benetti &amp; Short &amp; Michat, 2019" authorityName="Benetti &amp; Short &amp; Michat" authorityYear="2019" box="[1249,1381,1377,1403]" class="Insecta" family="Dytiscidae" genus="Hamadiana" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="C49F7A7CFF8FFFB1FB66FA91A7015C0A" box="[1249,1381,1377,1403]" italics="true" pageId="1" pageNumber="177">Hamadiana</emphasis>
</taxonomicName>
<emphasis id="C49F7A7CFF8FFFB1FAECFA91A2CA5CEE" bold="true" pageId="1" pageNumber="177">
<taxonomicNameLabel id="DFACC707FF8FFFB1FAECFA91A2CA5CEE" pageId="1" pageNumber="177" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
can be differentiated from these two genera by the shape of metacoxal processes, which are unique in having the hind margin strongly incised and medially protruded backwards (
<figureCitation id="6ED0BAEBFF8FFFB1FCFCFA59A1A45CB2" box="[891,960,1449,1475]" captionStart="FIGURE 811" captionStartId="6.[151,250,1220,1245]" captionTargetBox="[184,1320,233,1174]" captionTargetId="figure@6.[169,1418,174,1209]" captionTargetPageId="6" captionText="FIGURE 811. Hamadiana chapadensis sp. n., ventral view: 8) prosternal process; 9) metacoxal processes; 10) last abdominal ventrite; 11) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995300" httpUri="https://zenodo.org/record/3995300/files/figure.png" pageId="1" pageNumber="177">Fig. 9</figureCitation>
). From
<taxonomicName id="31EBDDEDFF8FFFB1FB9CFA59A6FD5CB2" authorityName="Regimbart" authorityYear="1895" box="[1051,1177,1449,1475]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="C49F7A7CFF8FFFB1FB9CFA59A6FD5CB2" box="[1051,1177,1449,1475]" italics="true" pageId="1" pageNumber="177">Laccodytes</emphasis>
</taxonomicName>
, the new genus can be distinguished by the metacoxal lines converging anteriad in posterior 3/4 of their length, slightly diverging anteriad in anterior 1/4 (straight in
<taxonomicName id="31EBDDEDFF8FFFB1FE40FA01A0215F7A" authorityName="Regimbart" authorityYear="1895" box="[455,581,1521,1547]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="C49F7A7CFF8FFFB1FE40FA01A0215F7A" box="[455,581,1521,1547]" italics="true" pageId="1" pageNumber="177">Laccodytes</emphasis>
</taxonomicName>
) and by the larger body (more than
<quantity id="31130B8BFF8FFFB1FC60FA01A64E5F7A" box="[999,1066,1521,1547]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="177" unit="mm" value="3.0">3 mm</quantity>
in length in
<taxonomicName id="31EBDDEDFF8FFFB1FB38FA01A7275F7A" authority="Benetti &amp; Short &amp; Michat, 2019" authorityName="Benetti &amp; Short &amp; Michat" authorityYear="2019" box="[1215,1347,1521,1547]" class="Insecta" family="Dytiscidae" genus="Hamadiana" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="C49F7A7CFF8FFFB1FB38FA01A7275F7A" box="[1215,1347,1521,1547]" italics="true" pageId="1" pageNumber="177">Hamadiana</emphasis>
</taxonomicName>
<emphasis id="C49F7A7CFF8FFFB1FACBFA01A7F95F7A" bold="true" box="[1356,1437,1521,1547]" pageId="1" pageNumber="177">
<taxonomicNameLabel id="DFACC707FF8FFFB1FACBFA01A7F95F7A" box="[1356,1437,1521,1547]" pageId="1" pageNumber="177" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
and less than
<quantity id="31130B8BFF8FFFB1FEB5F9E5A3ED5F5E" box="[306,393,1557,1583]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="177" unit="mm" value="2.4">2.4 mm</quantity>
in
<taxonomicName id="31EBDDEDFF8FFFB1FE29F9E5A0485F5E" authorityName="Regimbart" authorityYear="1895" box="[430,556,1557,1583]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="C49F7A7CFF8FFFB1FE29F9E5A0485F5E" box="[430,556,1557,1583]" italics="true" pageId="1" pageNumber="177">Laccodytes</emphasis>
</taxonomicName>
). From
<taxonomicName id="31EBDDEDFF8FFFB1FD0DF9E6A17C5F5E" authorityName="Toledo &amp; Michat" authorityYear="2015" box="[650,792,1558,1583]" class="Insecta" family="Dytiscidae" genus="Laccomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="C49F7A7CFF8FFFB1FD0DF9E6A17C5F5E" box="[650,792,1558,1583]" italics="true" pageId="1" pageNumber="177">Laccomimus</emphasis>
</taxonomicName>
,
<taxonomicName id="31EBDDEDFF8FFFB1FCA1F9E5A1CE5F5E" authority="Benetti &amp; Short &amp; Michat, 2019" authorityName="Benetti &amp; Short &amp; Michat" authorityYear="2019" box="[806,938,1557,1583]" class="Insecta" family="Dytiscidae" genus="Hamadiana" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="C49F7A7CFF8FFFB1FCA1F9E5A1CE5F5E" box="[806,938,1557,1583]" italics="true" pageId="1" pageNumber="177">Hamadiana</emphasis>
</taxonomicName>
<emphasis id="C49F7A7CFF8FFFB1FC35F9E5A6675F5E" bold="true" box="[946,1027,1557,1583]" pageId="1" pageNumber="177">
<taxonomicNameLabel id="DFACC707FF8FFFB1FC35F9E5A6675F5E" box="[946,1027,1557,1583]" pageId="1" pageNumber="177" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
differs also in having the prosternal process elongate and apically acute (relatively short and apically rounded in
<taxonomicName id="31EBDDEDFF8FFFB1FC7FF9CAA6E25F22" authorityName="Toledo &amp; Michat" authorityYear="2015" box="[1016,1158,1594,1619]" class="Insecta" family="Dytiscidae" genus="Laccomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="C49F7A7CFF8FFFB1FC7FF9CAA6E25F22" box="[1016,1158,1594,1619]" italics="true" pageId="1" pageNumber="177">Laccomimus</emphasis>
</taxonomicName>
), in the mesotibial spurs not longer than mesotarsomeres 12 (longer than mesotarsomeres
<quantity id="31130B8BFF8FFFB1FCFFF9ADA1DA5F06" box="[888,958,1629,1655]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="1" pageNumber="177" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">14 in</quantity>
<taxonomicName id="31EBDDEDFF8FFFB1FC43F9AEA6365F06" authorityName="Toledo &amp; Michat" authorityYear="2015" box="[964,1106,1630,1655]" class="Insecta" family="Dytiscidae" genus="Laccomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="C49F7A7CFF8FFFB1FC43F9AEA6365F06" box="[964,1106,1630,1655]" italics="true" pageId="1" pageNumber="177">Laccomimus</emphasis>
</taxonomicName>
) and in the larger body (more than
<quantity id="31130B8BFF8FFFB1FF49F971A36B5FEA" box="[206,271,1665,1691]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="177" unit="mm" value="3.0">3 mm</quantity>
in length in
<taxonomicName id="31EBDDEDFF8FFFB1FE1BF971A0445FEA" authority="Benetti &amp; Short &amp; Michat, 2019" authorityName="Benetti &amp; Short &amp; Michat" authorityYear="2019" box="[412,544,1665,1691]" class="Insecta" family="Dytiscidae" genus="Hamadiana" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="C49F7A7CFF8FFFB1FE1BF971A0445FEA" box="[412,544,1665,1691]" italics="true" pageId="1" pageNumber="177">Hamadiana</emphasis>
</taxonomicName>
<emphasis id="C49F7A7CFF8FFFB1FDAFF971A0135FEA" bold="true" box="[552,631,1665,1691]" pageId="1" pageNumber="177">
<taxonomicNameLabel id="DFACC707FF8FFFB1FDAFF971A0135FEA" box="[552,631,1665,1691]" pageId="1" pageNumber="177" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
and less than
<quantity id="31130B8BFF8FFFB1FC92F970A10F5FEA" box="[789,875,1664,1691]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="177" unit="mm" value="2.5">2.5 mm</quantity>
in
<taxonomicName id="31EBDDEDFF8FFFB1FC09F972A6785FEA" authorityName="Toledo &amp; Michat" authorityYear="2015" box="[910,1052,1666,1691]" class="Insecta" family="Dytiscidae" genus="Laccomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="C49F7A7CFF8FFFB1FC09F972A6785FEA" box="[910,1052,1666,1691]" italics="true" pageId="1" pageNumber="177">Laccomimus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>