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<document id="2B8CAE2585DDC624A2BE7E09F9331536" ID-DOI="10.1080/00222933.2018.1444804" ID-ISSN="1464-5262" ID-Zenodo-Dep="4747420" ID-ZooBank="72474144-5D65-48BE-9021-0490952AAA3C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620671357015" checkinUser="carolina" docAuthor="Kim, Sora, Bae, Yang-Seop &amp; Lee, Seunghwan" docDate="2018" docId="2B183D78FF9AC16874CD415991A44036" docLanguage="en" docName="JNATHIST.52.9-10.607-620.pdf" docOrigin="Journal of Natural History 52 (9 - 10)" docSource="http://dx.doi.org/10.1080/00222933.2018.1444804" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Promalactis petasumella Kim &amp; Bae &amp; Lee 2018, sp. nov." docType="treatment" docUuid="5E5276B9-0F6E-4D69-A1E6-6C71FBCE7AD4" docUuidSource="ZooBank" docVersion="4" lastPageNumber="619" masterDocId="D7214500FF91C165756147379002426E" masterDocTitle="Genus Promalactis Meyrick (Lepidoptera: Oecophoridae) from Cambodia, Part II: five new species, checklist and taxonomic key for the species in Cambodia" masterLastPageNumber="620" masterPageNumber="607" pageNumber="617" updateTime="1698914663135" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D1B8FB57FF195627CB92AEFCB5EDED65">Genus Promalactis Meyrick (Lepidoptera: Oecophoridae) from Cambodia, Part II: five new species, checklist and taxonomic key for the species in Cambodia</mods:title>
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<mods:namePart id="1980060851F5FCDE91ECD5257D8F6B7A">Kim, Sora</mods:namePart>
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<mods:namePart id="24FFD2190543F971D57D995D78DB1E3C">Bae, Yang-Seop</mods:namePart>
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<mods:namePart id="641239991B7C705B5E35867078D642CE">Lee, Seunghwan</mods:namePart>
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<mods:title id="03458C4E50626898582EA7DD89DA3FF6">Journal of Natural History</mods:title>
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<mods:date id="F7368B868AB1C3E0A29CA913C8D0101D">2018</mods:date>
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<mods:number id="D25DB4760B99733AD8D7F8B435D4F4D4">2018-03-15</mods:number>
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<mods:number id="BC495CA8D5C8C684403DD37C08D17302">52</mods:number>
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<mods:number id="37E2E488A84683FB411CEB7A6B794894">9 - 10</mods:number>
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<mods:identifier id="625A60FAB4A7DE601DA1B47514FDB6C5" type="ISSN">1464-5262</mods:identifier>
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<subSubSection id="EBABDFE5FF9AC16E74CD4159937A44E6" box="[428,888,1646,1672]" pageId="11" pageNumber="617" type="nomenclature">
<paragraph id="A30E8C6EFF9AC16E74CD4159937A44E6" blockId="11.[428,888,1646,1707]" box="[428,888,1646,1672]" pageId="11" pageNumber="617">
<heading id="F8463B02FF9AC16E74CD4159937A44E6" box="[428,888,1646,1672]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="617" reason="2">
<taxonomicName id="64B1F7EDFF9AC16E74CD4159931044E6" authority="Kim" authorityName="Kim &amp; Bae &amp; Lee" authorityYear="2018" box="[428,786,1646,1672]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="617" phylum="Arthropoda" rank="species" species="petasumella" status="sp. nov.">
<emphasis id="91C5507CFF9AC16E74CD415992D944E6" bold="true" box="[428,731,1646,1672]" italics="true" pageId="11" pageNumber="617">Promalactis petasumella</emphasis>
Kim
</taxonomicName>
<taxonomicNameLabel id="8AF6ED07FF9AC16E767A4158937A44E6" box="[795,888,1647,1672]" pageId="11" pageNumber="617" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EBABDFE5FF9AC16E74B141A5935144C5" box="[464,851,1682,1707]" pageId="11" pageNumber="617" type="description">
<paragraph id="A30E8C6EFF9AC16E74B141A5935144C5" blockId="11.[428,888,1646,1707]" box="[464,851,1682,1707]" pageId="11" pageNumber="617">
(
<figureCitation id="3B8A90EBFF9AC16E74B941A5924644C5" box="[472,580,1682,1707]" captionStart="Figure 1" captionStartId="4.[160,225,1677,1700]" captionTargetBox="[190,1120,152,1632]" captionTargetId="figure-1@4.[190,1125,142,1646]" captionTargetPageId="4" captionText="Figure 1. Adult of Promalactis. Promalactis ceratiscus sp. nov.: (a) head; (b) wing pattern. P. flavidius sp. nov.: (c) head; (d) wing pattern. P. latifasciata sp. nov.: (e) head; (f) wing pattern. P. mecodigita sp. nov.: (g) head; (h) wing pattern. P. petasumella sp. nov.: (i) head; (j) wing pattern." figureDoi="http://doi.org/10.5281/zenodo.4747422" httpUri="https://zenodo.org/record/4747422/files/figure.png" pageId="11" pageNumber="617">Figures 1</figureCitation>
(i), 1(j), 2(mo), 3(eg))
</paragraph>
</subSubSection>
<subSubSection id="EBABDFE5FF9DC16975C147BA93BC433A" pageId="12" pageNumber="618" type="diagnosis">
<paragraph id="A30E8C6EFF9DC16975C147BA911642C8" blockId="12.[160,1156,141,340]" box="[160,276,141,166]" pageId="12" pageNumber="618">
<emphasis id="91C5507CFF9DC16975C147BA911642C8" bold="true" box="[160,276,141,166]" italics="true" pageId="12" pageNumber="618">Diagnosis</emphasis>
</paragraph>
<paragraph id="A30E8C6EFF9DC16975C1478693BC433A" blockId="12.[160,1156,141,340]" pageId="12" pageNumber="618">
This species is similar to
<taxonomicName id="64B1F7EDFF9DC16974D24787923542A7" authority="Kim &amp; Bae &amp; Lee, 2018" authorityName="Kim &amp; Bae &amp; Lee" authorityYear="2018" box="[435,567,176,201]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="618" phylum="Arthropoda" rank="species" species="ceratiscus" status="sp. nov.">
<emphasis id="91C5507CFF9DC16974D24787923542A7" box="[435,567,176,201]" italics="true" pageId="12" pageNumber="618">P. ceratiscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8AF6ED07FF9DC169775F4786929742A7" box="[574,661,177,201]" pageId="12" pageNumber="618" rank="species">sp. nov.</taxonomicNameLabel>
in the forewing pattern, but it can be easily distinguished from the latter by the antemedial band oblique toward wing base and the presence of an apical patch. The male genitalia are similar to those of
<taxonomicName id="64B1F7EDFF9DC16976B547C294584360" authority="Kim &amp; Bae &amp; Lee, 2018" authorityName="Kim &amp; Bae &amp; Lee" authorityYear="2018" box="[980,1114,245,270]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="618" phylum="Arthropoda" rank="species" species="ceratiscus" status="sp. nov.">
<emphasis id="91C5507CFF9DC16976B547C294584360" box="[980,1114,245,270]" italics="true" pageId="12" pageNumber="618">P. ceratiscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8AF6ED07FF9DC169710547C190D1435F" pageId="12" pageNumber="618" rank="species">sp. nov.</taxonomicNameLabel>
, by sharing large and bifurcate lobes of juxta, but differs from this species by having the lateral margin of lobe of the juxta bearing a projection at middle.
</paragraph>
</subSubSection>
<subSubSection id="EBABDFE5FF9DC16975C146A2944344F8" pageId="12" pageNumber="618" type="description">
<paragraph id="A30E8C6EFF9DC16975C146A2912443C0" blockId="12.[160,1156,405,985]" box="[160,294,405,430]" pageId="12" pageNumber="618">
<emphasis id="91C5507CFF9DC16975C146A2912443C0" bold="true" box="[160,294,405,430]" italics="true" pageId="12" pageNumber="618">Description</emphasis>
</paragraph>
<paragraph id="A30E8C6EFF9DC16975C1468E920341B7" blockId="12.[160,1156,405,985]" pageId="12" pageNumber="618">
Adult (
<figureCitation id="3B8A90EBFF9DC1697591468E914D43BF" box="[240,335,441,465]" captionStart="Figure 1" captionStartId="4.[160,225,1677,1700]" captionTargetBox="[190,1120,152,1632]" captionTargetId="figure-1@4.[190,1125,142,1646]" captionTargetPageId="4" captionText="Figure 1. Adult of Promalactis. Promalactis ceratiscus sp. nov.: (a) head; (b) wing pattern. P. flavidius sp. nov.: (c) head; (d) wing pattern. P. latifasciata sp. nov.: (e) head; (f) wing pattern. P. mecodigita sp. nov.: (g) head; (h) wing pattern. P. petasumella sp. nov.: (i) head; (j) wing pattern." figureDoi="http://doi.org/10.5281/zenodo.4747422" httpUri="https://zenodo.org/record/4747422/files/figure.png" pageId="12" pageNumber="618">Figure 1</figureCitation>
(i), 1(j)). Head: Frons blackish dark brown; vertex white basally, tinged with pale greyish brown; occiput blackish dark brown. Antenna: Scape white entirely, 1/2 length of diameter of eye; fragellum dark brown and white alternately dorsally. Labial palpus: 2nd palpomere pale greyish dark brown; 3rd palpomere blackish dark brown, slightly shorter than 2nd palpomere. Thorax: Thorax and tegula blackish dark brown, latter tinged with brown. Wing expanse 8.0
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. Forewing ground colour brownish yellow; four bands all white, edged with fuscous scales: subbasal band relatively narrow, slightly arched toward wing base, reaching to costal margin; antemedial band slightly oblique toward wing base, not reaching costal margin; medial band oblique toward apex, 1/2 length of antemedial band, connected to postmedial band; postmedial band oblique toward wing base, as long as medial band; two white patches, edged with fuscous scales: costal patch at 1/2, darker near costa, reaching beyond half across wing, semi-ovate; apical patch semi-ovate, occupied from sub-apical costa to tornus followed by termen; fringes greyish dark brown tinged with yellow near apex. Hind wing sub-lanceolate; ground colour pale greyish brown; fringes greyish brown.
</paragraph>
<paragraph id="A30E8C6EFF9DC16975C1432D94534727" blockId="12.[160,1156,1050,1353]" pageId="12" pageNumber="618">
<emphasis id="91C5507CFF9DC16975C1432D9223465A" bold="true" box="[160,545,1050,1076]" pageId="12" pageNumber="618">
<emphasis id="91C5507CFF9DC16975C1432D914C465D" bold="true" box="[160,334,1050,1075]" italics="true" pageId="12" pageNumber="618">Male genitalia</emphasis>
(
<figureCitation id="3B8A90EBFF9DC1697402432D91CE465A" box="[355,460,1050,1076]" captionStart="Figure 2" captionStartId="5.[160,225,1535,1558]" captionTargetBox="[166,1150,144,1501]" captionTargetId="figure-1@5.[163,1153,142,1504]" captionTargetPageId="5" captionText="Figure 2. Male genitalia of Promalactis. (a-c) Promalactis ceratiscus sp. nov.; (d-f) P. flavidius sp. nov.; (g-i) P. latifasciata sp. nov.; (j-l) P. mecodigita sp. nov.; (m-o) P. petasumella sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4747424" httpUri="https://zenodo.org/record/4747424/files/figure.png" pageId="12" pageNumber="618">Figure 2</figureCitation>
(mo)).
</emphasis>
Uncus large, broadly rounded apically, incised apically, forming two lateral lobes. Gnathos wide at base, up-turned at 1/2, gradually narrowed to apex, short, tongue shaped, slightly longer than uncus. Valva with costal margin slightly concave before middle; cucullus at 3/5 of costal margin, wide at base, straight from base to 1/2, gradually narrowed from 1/2 to sub-apex, apex spine-shaped; sacculus elongate, finger-shaped, setose from 4/5 of costal margin to apex; juxta large, bifurcate medially forming two large lobes, as long as valva: the lobe with lateral margin covered by dense setose from middle to apex, bearing projection in middle. Saccus large, triangular. Phallus slightly bent at middle, as long as valva. Cornutus absent.
</paragraph>
<paragraph id="A30E8C6EFF9DC16975C142BD944344F8" blockId="12.[160,1156,1418,1686]" pageId="12" pageNumber="618">
<emphasis id="91C5507CFF9DC16975C142BD923447CD" bold="true" box="[160,566,1418,1444]" pageId="12" pageNumber="618">
<emphasis id="91C5507CFF9DC16975C142BD916947CD" bold="true" box="[160,363,1418,1443]" italics="true" pageId="12" pageNumber="618">Female genitalia</emphasis>
(
<figureCitation id="3B8A90EBFF9DC16974E142BD91E847CA" box="[384,490,1418,1444]" captionStart="Figure 3" captionStartId="10.[160,225,1556,1579]" captionTargetBox="[169,1151,142,1517]" captionTargetId="figure-1@10.[163,1152,142,1525]" captionTargetPageId="10" captionText="Figure 3. Female genitalia of Promalactis. (a-d) Promalactis flavidius sp. nov.; (e-g) P. petasumella sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4747426" httpUri="https://zenodo.org/record/4747426/files/figure.png" pageId="12" pageNumber="618">Figure 3</figureCitation>
(eg)).
</emphasis>
Papillae anales setose. Apophyses posteriores more than 1.5 times longer than apophyses anteriores. Apophyses anteriores as long as papilla anales. Lamella postvaginalis large, sclerotized, incised caudal-medially, hat-shaped, bearing small thumb-shaped projection sub-apically. Lamella antevaginalis large, sclerotized, both sides of antrum forming projection elongated. Antrum broad cylindrical-shaped, rather straight to ductus bursae, as long as 3/5 of apophyses posteriores. Seventh sternum bearing sclerotized, long-oval projection anteriorly. Ductus bursae wider, slightly longer than antrum. Corpus bursae ovate, bearing a signum; signum bearing several different sized spines.
</paragraph>
</subSubSection>
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<emphasis id="91C5507CFF9CC16875C147BA910942C8" bold="true" box="[160,267,141,166]" italics="true" pageId="13" pageNumber="619">
<typeStatus id="7C0A32CCFF9CC16875C147BA910942C8" box="[160,267,141,166]" pageId="13" pageNumber="619" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="A30E8C6EFF9CC16875C14786943E435F" blockId="13.[160,1156,141,305]" pageId="13" pageNumber="619">
<materialsCitation id="13D98633FF9CC16875C14786943E435F" collectingDate="2012-02-20" collectingDateMax="2014-12-23" collectingDateMin="2012-02-20" collectorName="Bae &amp; S Kim &amp; S Kim." country="Cambodia" county="Male" elevation="644" location="Bae" municipality="Sam Kos" pageId="13" pageNumber="619" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Pursat" typeStatus="paratype">
<collectingCounty id="4A6FF4E2FF9CC16875C1478690DA42A7" box="[160,216,177,201]" pageId="13" pageNumber="619">Male</collectingCounty>
,
<collectingCountry id="DBA6CCFEFF9CC16875854786915542A7" box="[228,343,177,201]" name="Cambodia" pageId="13" pageNumber="619">Cambodia</collectingCountry>
,
<collectingRegion id="6175428CFF9CC1687402478691EA42A7" box="[355,488,177,201]" country="Cambodia" name="Pursat" pageId="13" pageNumber="619">Pursat Prov.</collectingRegion>
<collectingMunicipality id="436A1614FF9CC16874914786925142A7" box="[496,595,177,201]" pageId="13" pageNumber="619">Sam Kos</collectingMunicipality>
,
<date id="D70FAAAEFF9CC168773E4786932742A7" box="[607,805,177,201]" pageId="13" pageNumber="619" value="2012-02-20">
<collectingDate id="C74B5346FF9CC168773E4786932742A7" box="[607,805,177,201]" pageId="13" pageNumber="619" value="2012-02-20">20 February 2012</collectingDate>
</date>
,
<location id="A66EDAB5FF9CC16876534786935842A7" LSID="urn:lsid:plazi:treatment:2B183D78FF9AC16874CD415991A44036:A66EDAB5FF9CC16876534786935842A7" box="[818,858,177,201]" country="Cambodia" county="Male" municipality="Sam Kos" name="Bae" pageId="13" pageNumber="619" stateProvince="Pursat">Bae</location>
<emphasis id="91C5507CFF9CC16876024787939142A7" box="[867,915,176,201]" italics="true" pageId="13" pageNumber="619">et al</emphasis>
., gen. slide no. 9599.
<emphasis id="91C5507CFF9CC16875C147E4910A4282" box="[160,264,211,236]" italics="true" pageId="13" pageNumber="619">
<typeStatus id="7C0A32CCFF9CC16875C147E4910A4282" box="[160,264,211,236]" pageId="13" pageNumber="619" type="paratype">Paratypes</typeStatus>
</emphasis>
. One male and
<specimenCount id="B5B747E7FF9CC16874D347E3922D4282" box="[434,559,212,236]" pageId="13" pageNumber="619" type="female">one female</specimenCount>
,
<collectingCountry id="DBA6CCFEFF9CC168775647E392A84282" box="[567,682,212,236]" name="Cambodia" pageId="13" pageNumber="619">Cambodia</collectingCountry>
,
<collectingRegion id="6175428CFF9CC16877D247E393374282" box="[691,821,212,236]" country="Cambodia" name="Pursat" pageId="13" pageNumber="619">Pursat Prov.</collectingRegion>
<location id="A66EDAB5FF9CC168765847E3939B4282" LSID="urn:lsid:plazi:treatment:2B183D78FF9AC16874CD415991A44036:A66EDAB5FF9CC168765847E3939B4282" box="[825,921,212,236]" country="Cambodia" county="Male" municipality="Sam Kos" name="Sam Kos" pageId="13" pageNumber="619" stateProvince="Pursat">Sam Kos</location>
,
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,
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<collectingDate id="C74B5346FF9CC168769147E390DA4360" pageId="13" pageNumber="619" value="2014-12-23">23 December 2014</collectingDate>
</date>
,
<collectorName id="0E44E9B8FF9CC168758247C191094360" box="[227,267,246,270]" pageId="13" pageNumber="619">Bae</collectorName>
<emphasis id="91C5507CFF9CC168747547C291414360" box="[276,323,245,270]" italics="true" pageId="13" pageNumber="619">et al</emphasis>
., gen. slide no. 9606(M)/
<collectorName id="0E44E9B8FF9CC168773547C1929A4360" box="[596,664,246,270]" pageId="13" pageNumber="619">S Kim</collectorName>
;
<specimenCount id="B5B747E7FF9CC16877C547C1930C4360" box="[676,782,246,270]" pageId="13" pageNumber="619" type="male">one male</specimenCount>
,
<specimenCount id="B5B747E7FF9CC168767B47C193A04360" box="[794,930,246,270]" pageId="13" pageNumber="619" type="female">two females</specimenCount>
and one ex(missing abdomen)., same locality,
<date id="D70FAAAEFF9CC16874A0462E9287435F" box="[449,645,281,305]" pageId="13" pageNumber="619" value="2012-02-20">
<collectingDate id="C74B5346FF9CC16874A0462E9287435F" box="[449,645,281,305]" pageId="13" pageNumber="619" value="2012-02-20">20 February 2012</collectingDate>
</date>
,
<collectorName id="0E44E9B8FF9CC16877EE462E92B5435F" box="[655,695,281,305]" pageId="13" pageNumber="619">Bae</collectorName>
<emphasis id="91C5507CFF9CC16877DE462F92EC435F" box="[703,750,280,305]" italics="true" pageId="13" pageNumber="619">et al</emphasis>
., gen. slide no. 9605(F)/
<collectorName id="0E44E9B8FF9CC1687694462E943E435F" box="[1013,1084,281,305]" pageId="13" pageNumber="619">S Kim.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="EBABDFE5FF9CC16875C1465192464389" pageId="13" pageNumber="619" type="etymology">
<paragraph id="A30E8C6EFF9CC16875C14651911F4311" blockId="13.[160,1155,358,487]" box="[160,285,358,383]" pageId="13" pageNumber="619">
<emphasis id="91C5507CFF9CC16875C14651911F4311" bold="true" box="[160,285,358,383]" italics="true" pageId="13" pageNumber="619">Etymology</emphasis>
</paragraph>
<paragraph id="A30E8C6EFF9CC16875C146BD92464389" blockId="13.[160,1155,358,487]" pageId="13" pageNumber="619">
The species epithet,
<taxonomicName id="64B1F7EDFF9CC16874F246BE921543CC" authorityName="Kim &amp; Bae &amp; Lee" authorityYear="2018" box="[403,535,393,418]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="619" phylum="Arthropoda" rank="species" species="petasumella">
<emphasis id="91C5507CFF9CC16874F246BE921543CC" box="[403,535,393,418]" italics="true" pageId="13" pageNumber="619">petasumella</emphasis>
</taxonomicName>
, is derived from the Latin,
<emphasis id="91C5507CFF9CC168763F46BE93C643CC" box="[862,964,393,418]" italics="true" pageId="13" pageNumber="619">petasum-</emphasis>
(=hat) and -
<emphasis id="91C5507CFF9CC168713C46BE948143CC" box="[1117,1155,393,418]" italics="true" pageId="13" pageNumber="619">ella</emphasis>
(=Latin diminutive suffix), referring to the hat-shape of the lamella antevaginalis and postvaginalis of the female genitalia.
</paragraph>
</subSubSection>
<subSubSection id="EBABDFE5FF9CC16875C1452B91A44036" box="[160,422,540,600]" pageId="13" pageNumber="619" type="distribution">
<paragraph id="A30E8C6EFF9CC16875C1452B912E405B" blockId="13.[160,422,540,600]" box="[160,300,540,565]" pageId="13" pageNumber="619">
<emphasis id="91C5507CFF9CC16875C1452B912E405B" bold="true" box="[160,300,540,565]" italics="true" pageId="13" pageNumber="619">Distribution</emphasis>
</paragraph>
<paragraph id="A30E8C6EFF9CC16875C1457791A44036" blockId="13.[160,422,540,600]" box="[160,422,576,600]" pageId="13" pageNumber="619">
<collectingCountry id="DBA6CCFEFF9CC16875C1457791114036" box="[160,275,576,600]" name="Cambodia" pageId="13" pageNumber="619">Cambodia</collectingCountry>
(West: new).
</paragraph>
</subSubSection>
</treatment>
</document>