295 lines
26 KiB
XML
295 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.122.33032" ID-GBIF-Dataset="f497907c-ee54-41cd-b666-7292c766b9c3" ID-PMC="PMC6548746" ID-Pensoft-Pub="1314-2003-122-29" ID-Pensoft-UUID="FFB7FFA5CF25AD00FFBDFFEBFFFCFFDD" ID-PubMed="31182907" ID-Zenodo-Dep="3238737" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-2003-122-29" ModsDocOrigin="PhytoKeys 122" ModsDocTitle="Páramo Calamagrostis s.l. (Poaceae): An updated list and key to the species known or likely to occur in páramos of NW South America and southern Central America including two new species, one new variety and five new records for Colombia" checkinTime="1559110536903" checkinUser="pensoft" docAuthor="Sylvester, Steven P., Soreng, Robert J., Bravo-Pedraza, William J., Cuta-Alarcon, Lia E., Giraldo-Canas, Diego, Aguilar-Cano, Jose & Peterson, Paul M." docDate="2019" docId="761B5CC2302CA1E78803B3D234DDD61E" docLanguage="en" docName="PhytoKeys 122: 29-78" docOrigin="PhytoKeys 122" docSource="http://dx.doi.org/10.3897/phytokeys.122.33032" docTitle="Deschampsia podophora Saarela, PhytoKeys 87: 90. 2017. Deyeuxia podophora (Pilg.) Sodiro, Rev. Col. Nac. Vicente Rocafuerte 11: 79. 1930. Calamagrostis podophora Pilg., Bot. Jahrb. Syst. 42 (1): 66. 1908" docType="treatment" docVersion="6" id="FFB7FFA5CF25AD00FFBDFFEBFFFCFFDD" lastPageNumber="29" masterDocId="FFB7FFA5CF25AD00FFBDFFEBFFFCFFDD" masterDocTitle="Paramo Calamagrostis s. l. (Poaceae): An updated list and key to the species known or likely to occur in paramos of NW South America and southern Central America including two new species, one new variety and five new records for Colombia" masterLastPageNumber="78" masterPageNumber="29" pageNumber="29" updateTime="1668139302904" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Paramo Calamagrostis s. l. (Poaceae): An updated list and key to the species known or likely to occur in paramos of NW South America and southern Central America including two new species, one new variety and five new records for Colombia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sylvester, Steven P.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Soreng, Robert J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bravo-Pedraza, William J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cuta-Alarcon, Lia E.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Giraldo-Canas, Diego</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Aguilar-Cano, Jose</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Peterson, Paul M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>122</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>29</mods:start>
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<mods:end>78</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.122.33032</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.122.33032</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-122-29</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFB7FFA5CF25AD00FFBDFFEBFFFCFFDD</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3238737</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="157294009" LSID="urn:lsid:plazi:treatment:761B5CC2302CA1E78803B3D234DDD61E" httpUri="http://treatment.plazi.org/id/761B5CC2302CA1E78803B3D234DDD61E" lastPageNumber="29" pageId="0" pageNumber="29">
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<subSubSection pageId="0" pageNumber="29" type="nomenclature">
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<paragraph pageId="0" pageNumber="29">
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<taxonomicName LSID="6C6A8A00-6598-5ED5-A050-32827968A02B" authority="(Pilg.) Saarela, PhytoKeys 87: 90. 2017. Deyeuxia podophora (Pilg.) Sodiro, Rev. Col. Nac. Vicente Rocafuerte 11: 79. 1930. Calamagrostis podophora Pilg., Bot. Jahrb. Syst. 42 (1): 66. 1908." class="Liliopsida" family="Poaceae" genus="Deschampsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Deschampsia podophora" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="podophora">Deschampsia podophora (Pilg.) Saarela, PhytoKeys 87: 90. 2017. Deyeuxia podophora (Pilg.) Sodiro, Rev. Col. Nac. Vicente Rocafuerte 11: 79. 1930. Calamagrostis podophora Pilg., Bot. Jahrb. Syst. 42 (1): 66. 1908.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="materials_examined">
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<paragraph pageId="0" pageNumber="29">Type.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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PERU.
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<normalizedToken originalValue="Junín">Junin</normalizedToken>
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: Berge Westlich von Huacapistana [Prov. Tarma, in montibus prope Huacapistana ad occid. in stepposis], 3500 m alt., 18 Jan. 1903, A. Weberbauer 2231 (lectotype, designated by
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<bibRefCitation author="Vega, AS" journalOrPublisher="Botanica" pageId="0" pageNumber="29" pagination="31 - 35" refId="B44" refString="Vega, AS, Rugolo de Agrasar, ZE, 2013. Lectotypifications in taxa of the genera Calamagrostis, Deyeuxia, and Digitaria (Poaceae). Gayana. Botanica 70 (1): 31 - 35" title="Lectotypifications in taxa of the genera Calamagrostis, Deyeuxia, and Digitaria (Poaceae). Gayana." volume="70" year="2013">
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Vega and
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<normalizedToken originalValue="Rúgolo">Rugolo</normalizedToken>
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de Agrasar (2013
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</bibRefCitation>
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: 28): BAA (BAA00000767 [image!]) fragm. ex B; isolectotype: US (US00149282!)).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="description">
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<paragraph pageId="0" pageNumber="29">Description.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" pageId="0" pageNumber="29">Tufted perennial</emphasis>
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with vertical rhizomes forming short solitary tufts to medium-sized tussocks, with leaf blades mostly basal with inflorescences usually greatly exerted from basal foliage or both basal and cauline with some cauline blades often surpassing the inflorescence.
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<emphasis bold="true" pageId="0" pageNumber="29">Tillers</emphasis>
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intravaginal.
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<emphasis bold="true" pageId="0" pageNumber="29">Culms</emphasis>
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20
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<normalizedToken originalValue="–75(–">-75(-</normalizedToken>
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110) cm tall, to 3 mm wide, erect, striate, nodes and internodes terete, smooth and lustrous; nodes hidden in the sheaths with no nodes exposed at flowering; uppermost internodes 20-32.5 cm long, as long or longer than the sheath.
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<emphasis bold="true" pageId="0" pageNumber="29">Sheaths</emphasis>
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striate;
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<emphasis bold="true" pageId="0" pageNumber="29">flag leaf sheaths</emphasis>
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22-38 cm long;
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<emphasis bold="true" pageId="0" pageNumber="29">upper culm sheaths</emphasis>
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lax, glabrous and smooth;
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<emphasis bold="true" pageId="0" pageNumber="29">basal leaf sheaths</emphasis>
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4.5-20 cm long, longer than the internodes, glabrous and smooth.
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<emphasis bold="true" pageId="0" pageNumber="29">Ligules</emphasis>
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not stipulate;
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<emphasis bold="true" pageId="0" pageNumber="29">upper culm ligules</emphasis>
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7.5-22 mm long, strongly decurrent with the sheaths, long acuminate, membranous to slightly coriaceous, without notable lateral keels, apices entire, erose or narrowly bifid, sometimes fimbriate, abaxial surface smooth;
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<emphasis bold="true" pageId="0" pageNumber="29">ligules of innovations</emphasis>
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4
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<normalizedToken originalValue="–15(–">-15(-</normalizedToken>
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20) mm long, slightly to strongly and broadly decurrent with the sheaths, long acuminate, membranous to slightly coriaceous, lateral keels sometimes notable, apices entire or a narrow bifid point, sometimes slightly erose, abaxial surface smooth or sometimes slightly scabrous at the apex.
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<emphasis bold="true" pageId="0" pageNumber="29">Leaf blades</emphasis>
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(2.5
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<normalizedToken originalValue="–)5–">-)5-</normalizedToken>
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22 cm long, 0.6-7 mm wide when opened out, flat, conduplicate or involute/convolute and filiform and cylindrical to subelliptical in outline, sometimes opening out to become flat at their apices, straight and erect to slightly curved, glabrous, isomorphic or more or less dimorphic, when dimorphic those of the innovations filiform and cylindrical to subelliptic in outline while those of the upper flowering culm are usually wider and flat, conduplicate or convolute towards the apices, abaxially smooth, adaxially smooth or lightly scaberulous along the veins, sometimes becoming densely scabrous towards the apex, edges smooth or slightly scaberulous, veins usually pronounced, numerous and tightly packed, apex obtuse to slightly pungent;
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<emphasis bold="true" pageId="0" pageNumber="29">flag leaf blades</emphasis>
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2.9-6 cm long, 2-7 mm wide when opened out.
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<emphasis bold="true" pageId="0" pageNumber="29">Panicles</emphasis>
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10-25 cm long, 3-8 cm wide, open to slightly condensed, oval, greenish-purple with spikelets tending to be laxly glomerate on the distal half of the inflorescence branches with the proximal half usually lacking spikelets, largely exerted to moderately included in the uppermost sheath and/or blade;
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<emphasis bold="true" pageId="0" pageNumber="29">main panicle axis</emphasis>
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terete to slightly compressed, usually with a narrow groove running down both sides, glabrous, smooth to lightly scaberulous, internodes tending to be long, lower internode 3-11.5 cm long;
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<emphasis bold="true" pageId="0" pageNumber="29">panicle branches</emphasis>
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1.5-8 cm long, bearing 10 to over 50 spikelets per branch, flexuous, spreading, pendulous or divergent at a 45° angle to slightly ascending, verticillate in clusters of 2 or 3, terete or slightly grooved, glabrous, almost smooth to scabrous;
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<emphasis bold="true" pageId="0" pageNumber="29">pedicels</emphasis>
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0.5-2.5 mm long, usually shorter than the spikelets, glabrous, lightly to densely scabrous.
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<emphasis bold="true" pageId="0" pageNumber="29">Spikelets</emphasis>
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1-flowered, sometimes with a rudimentary floret at the apex of the rachilla that appears like a slightly broader section of the rachilla covered in sparse diminute hairs, not strongly laterally compressed, disarticulating above the glumes with the florets disarticulating from the apex of the extended rachilla internode, this remaining attached to the glumes.
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<emphasis bold="true" pageId="0" pageNumber="29">Glumes</emphasis>
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3.5-5.5 mm long, subequal, the lower glume 0.2-0.6 mm shorter than the upper glume, lanceolate, membranous, purplish-green, lustrous, smooth or sometimes lightly scabrous throughout the keel of the upper glume;
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<emphasis bold="true" pageId="0" pageNumber="29">lower glumes</emphasis>
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1-veined, apex acuminate or bidentate, less frequently finely denticulate or erose;
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<emphasis bold="true" pageId="0" pageNumber="29">upper glumes</emphasis>
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3-veined, lateral veins either short <
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<normalizedToken originalValue="½">1/2</normalizedToken>
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length of glume or reaching from
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<normalizedToken originalValue="½">1/2</normalizedToken>
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to 2/3 the length of the glume, apex usually acuminate, sometimes finely denticulate.
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<emphasis bold="true" pageId="0" pageNumber="29">Floret</emphasis>
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stipitate, much shorter than the glumes, never passing the apex of the lower glume.
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<emphasis bold="true" pageId="0" pageNumber="29">Lowermost rachilla internode</emphasis>
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0.4-0.7 mm long, prolonged between the glumes and the floret, often slightly geniculate at its apex and bent in a ca. 30°-45° angle, slightly dilated at its apex, usually glabrous, less often with a few long hairs ca. 0.7 mm long emerging from it, smooth.
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<emphasis bold="true" pageId="0" pageNumber="29">Lemmas</emphasis>
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2.4-3.5 mm long, of the same consistency as the glumes, light green with purple tinges towards apex, becoming golden at maturity, glabrous, smooth with the keel apex rarely scaberulous, apex truncate and denticulate, usually with 4 clearly distinguished teeth, 0.3-0.5 mm long and erose between the teeth, 5-veined, veins not evident; awns 1.5-4 mm long, sometimes absent, inserted in the middle or lower third of the lemma, usually as long as the lemma or passing the glume apex by as much as 1.5 mm.
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<emphasis bold="true" pageId="0" pageNumber="29">Paleas</emphasis>
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0.5-0.8 mm shorter than the lemma, of the same consistency and colour, keels sparsely scabrous and notable, apex bidentate or 4-dentate.
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<emphasis bold="true" pageId="0" pageNumber="29">Callus</emphasis>
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rounded, short, with a basal tuft of hairs 0.5-2.2 mm long, reaching from 1/3 the length of the lemma to almost the lemma apex.
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<emphasis bold="true" pageId="0" pageNumber="29">Rachilla</emphasis>
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1.5-2.5 mm long, reaching from 2/3 to 4/5 the length of the lemma, with copious short to medium-sized hairs 0.5-1.4 mm long, the hairs reaching from 4/5 to sometimes passing the apex of the lemma, apex of rachilla sometimes clavate.
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<emphasis bold="true" pageId="0" pageNumber="29">Lodicules</emphasis>
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2, ca. 0.5 mm long, membranous, acute.
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<emphasis bold="true" pageId="0" pageNumber="29">Stamens</emphasis>
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3, anthers (0.7
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<normalizedToken originalValue="–)1.2–">-)1.2-</normalizedToken>
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1.9 mm long.
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<emphasis bold="true" pageId="0" pageNumber="29">Ovary</emphasis>
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ca. 0.5 mm long, small, styles 2, stigmas plumose, short.
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<emphasis bold="true" pageId="0" pageNumber="29">Caryopses</emphasis>
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1-1.4 mm long, dorsally slightly gibbose, surcus not noticeable, embryo short, hilum basal, oval;
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<emphasis bold="true" pageId="0" pageNumber="29">endosperm</emphasis>
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dry.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="distribution">
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<paragraph pageId="0" pageNumber="29">Distribution and ecology.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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Colombia, Ecuador, Peru, Venezuela. The ecology of this species is distinct compared to many
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<taxonomicName class="Liliopsida" family="Gramineae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="29">Calamagrostis</emphasis>
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</taxonomicName>
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s.l. species as it is usually found in very damp, swamp-like conditions by the side of high-elevation lakes or watercourses in Andean
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<normalizedToken originalValue="páramo">paramo</normalizedToken>
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or jalca vegetation, less often in humid open
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<normalizedToken originalValue="páramo">paramo</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="materials_examined">
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<paragraph pageId="0" pageNumber="29">Other specimens examined.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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COLOMBIA.
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<emphasis bold="true" pageId="0" pageNumber="29">
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<normalizedToken originalValue="Boyacá">Boyaca</normalizedToken>
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</emphasis>
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: Below Las Playas de Ritacuba in the Cocuy mountains above Guican, on the banks of a fast-flowing stream coming off the snow fields and passing through thinly vegetated moraine covered country, 4100 m alt., 24 Jun. 1984, J.R.I. Wood 4457 (K).
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<emphasis bold="true" pageId="0" pageNumber="29">Cauca</emphasis>
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: Purace National Park, Laguna de San Rafael, in open boggy
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<normalizedToken originalValue="páramo">paramo</normalizedToken>
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in the lake basin, particularly in banks by ditches, 3300 m alt., 6 Apr. 1985, J.R.I. Wood 4803 (K); Volcan Purace, above Pilimbala, frequent in bog pools in high
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<normalizedToken originalValue="páramo">paramo</normalizedToken>
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, 3700-4000 m alt., 5 Apr. 1985, J.R.I. Wood 4787 (K).
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</paragraph>
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<paragraph pageId="0" pageNumber="29">
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ECUADOR.
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<emphasis bold="true" pageId="0" pageNumber="29">Pichincha</emphasis>
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: road Olmedo-Laguna San Marcos, W of the pass, 0°5'N; 78°1-2'W, 3600 m alt., 10 Jul. 1980, B.
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<normalizedToken originalValue="Øllgaard">Ollgaard</normalizedToken>
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et al. 34406 (K); Along road to Refugio, Volcan Cayembe,
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<normalizedToken originalValue="páramo">paramo</normalizedToken>
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and swamp,
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<geoCoordinate degrees="00" direction="south" minutes="04" orientation="latitude" precision="925" value="-0.06666667">00°04'S</geoCoordinate>
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;
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<geoCoordinate degrees="77" direction="west" minutes="54" orientation="longitude" precision="925" value="-77.9">77°54'W</geoCoordinate>
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, 4300 m alt., 2 Mar. 1988, S. Renvoize 70510 (K).
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<emphasis bold="true" pageId="0" pageNumber="29">Napo</emphasis>
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: Eastern Cordillera, Llanganati Mountains, by Lake Aucacocha, on stream sides in bog, forming large tussocks up to 20 cm across, 3750 m alt., 16 Aug. 1969, P.J. Edwards 127 (K); Eastern Cordillera, Llanganati Mountains, by Lake Aucacocha, growing in the wettest area of the bog, in clusters of tufts, sheath bases submerged in peat, 3700 m alt., Aug. 1969, P.J. Edwards 62 (K).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="notes">
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<paragraph pageId="0" pageNumber="29">Notes.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Deschampsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Deschampsia podophora" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="podophora">
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<emphasis italics="true" pageId="0" pageNumber="29">Deschampsia podophora</emphasis>
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</taxonomicName>
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has been traditionally treated as belonging to
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<taxonomicName authority="subsect. Stylagrostis" class="Liliopsida" family="Gramineae" genus="Calamagrostis" higherTaxonomySource="CoL" infraspecific-rank="subsect." kingdom="Plantae" lsidName="Calamagrostis" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="genus" subsection="Stylagrostis">Calamagrostis subsect. Stylagrostis</taxonomicName>
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due to the presence of an extended rachilla internode between the glumes and the floret.
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Deschampsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Deschampsia podophora" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="podophora">
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<emphasis italics="true" pageId="0" pageNumber="29">Deschampsia podophora</emphasis>
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</taxonomicName>
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is closely related to
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<taxonomicName lsidName="D. parodiana" pageId="0" pageNumber="29" rank="species" species="parodiana">
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<emphasis italics="true" pageId="0" pageNumber="29">D. parodiana</emphasis>
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</taxonomicName>
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(=
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<taxonomicName authorityName="Hitchc" authorityYear="1927" baseAuthorityName="Kunth" class="Liliopsida" family="Gramineae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis ligulata" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="ligulata">
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<emphasis italics="true" pageId="0" pageNumber="29">Calamagrostis ligulata</emphasis>
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</taxonomicName>
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) and has been placed as a synonym of this in previous works (
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<bibRefCitation author="Escalona, FD" journalOrPublisher="Contributions from the United States National Herbarium" pageId="0" pageNumber="29" refId="B6" refString="Escalona, FD, 1988a. Systematics of CalamagrostissectionDeyeuxia, subsection Stylagrostis, (Poaceae: Pooideae). PhD. Thesis, Iowa State University, U.S.A." title="Systematics of CalamagrostissectionDeyeuxia, subsection Stylagrostis, (Poaceae: Pooideae). PhD. Thesis, Iowa State University, U. S. A." year="1988 a">Escalona 1988a</bibRefCitation>
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,
|
||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.13488" author="Escalona, FD" journalOrPublisher="Phytologia" pageId="0" pageNumber="29" pagination="337 - 347" refId="B7" refString="Escalona, FD, 1988b. Stylagrostis a new subsection of genus Calamagrostis (Poaceae: Pooideae) and three new species from Colombia, Ecuador and Venezuela. Phytologia 65 (5): 337 - 347, DOI: https://doi.org/10.5962/bhl.part.13488" title="Stylagrostis a new subsection of genus Calamagrostis (Poaceae: Pooideae) and three new species from Colombia, Ecuador and Venezuela." url="https://doi.org/10.5962/bhl.part.13488" volume="65" year="1988 b">1988b</bibRefCitation>
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;
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||
<bibRefCitation author="Tovar, O" journalOrPublisher="Ruizia" pageId="0" pageNumber="29" pagination="1 - 480" refId="B43" refString="Tovar, O, 1993. Las Gramineas (Poaceae) del Peru. Ruizia 13: 1 - 480" title="Las Gramineas (Poaceae) del Peru." volume="13" year="1993">Tovar 1993</bibRefCitation>
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||
;
|
||
<bibRefCitation author="Luteyn, JL" journalOrPublisher="Memoirs of the New York Botanical Garden" pageId="0" pageNumber="29" refId="B20" refString="Luteyn, JL, 1999. Paramos, a checklist of plant diversity, geographical distribution, and botanical literature. Memoirs of the New York Botanical Garden 84: viii-xv, 1-278." title="Paramos, a checklist of plant diversity, geographical distribution, and botanical literature. Memoirs of the New York Botanical Garden 84: viii-xv, 1 - 278." year="1999">Luteyn 1999</bibRefCitation>
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;
|
||
<bibRefCitation author="Bono, G" editor="Bono, G" journalOrPublisher="Universita degle Studi di Firenze, Firenze" pageId="0" pageNumber="29" pagination="127 - 154" refId="B1" refString="Bono, G, 2010. Poaceae. In: Bono, G, Ed., Catalogo de la Flora y Vegetacion de los Valles de la Vertiente Occidental de los Andes de la Cordillera de Merida. Universita degle Studi di Firenze, Firenze: 127 - 154" title="Poaceae." volumeTitle="Catalogo de la Flora y Vegetacion de los Valles de la Vertiente Occidental de los Andes de la Cordillera de Merida." year="2010">Bono 2010</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Briceno, B" editor="Morillo, GN" journalOrPublisher="2. Centro Editorial Litoramma C. A., Merida" pageId="0" pageNumber="29" pagination="599 - 711" refId="B2" refString="Briceno, B, 2010. Capitulo XXI Familia Poaceae (R.Br.) Barnhart (Gramineae). In: Morillo, GN, Briceno, B, Silva, JF, Eds., Botanica y Ecologia de las Monocotiledoneas de los Paramos en Venezuela, vol. 2. Centro Editorial Litoramma C. A., Merida: 599 - 711" title="Capitulo XXI Familia Poaceae (R. Br.) Barnhart (Gramineae)." volumeTitle="Botanica y Ecologia de las Monocotiledoneas de los Paramos en Venezuela, vol." year="2010">
|
||
<normalizedToken originalValue="Briceño">Briceno</normalizedToken>
|
||
2010
|
||
</bibRefCitation>
|
||
). The principal differentiating characters that separate
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Deschampsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Deschampsia parodiana" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="parodiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="29">Deschampsia parodiana</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName lsidName="D. podophora" pageId="0" pageNumber="29" rank="species" species="podophora">
|
||
<emphasis italics="true" pageId="0" pageNumber="29">D. podophora</emphasis>
|
||
</taxonomicName>
|
||
are the smaller anthers, 0.4-0.5 mm long (vs. (0.7
|
||
<normalizedToken originalValue="–)1.2–1.5(–">-)1.2-1.5(-</normalizedToken>
|
||
1.9) mm long in
|
||
<taxonomicName lsidName="D. podophora" pageId="0" pageNumber="29" rank="species" species="podophora">
|
||
<emphasis italics="true" pageId="0" pageNumber="29">D. podophora</emphasis>
|
||
</taxonomicName>
|
||
) and the shorter rachilla, 1-1.2 mm long, that is sparsely pilose with hairs not usually reaching the apex of the palea (vs. (1.2
|
||
<normalizedToken originalValue="–)1.4–">-)1.4-</normalizedToken>
|
||
2.5 mm long, with copious hairs that usually surpass the lemma in
|
||
<taxonomicName lsidName="D. podophora" pageId="0" pageNumber="29" rank="species" species="podophora">
|
||
<emphasis italics="true" pageId="0" pageNumber="29">D. podophora</emphasis>
|
||
</taxonomicName>
|
||
). Characters of shape and density of the inflorescence mentioned in
|
||
<bibRefCitation DOI="https://doi.org/10.2307/3391886" author="Laegaard, S" journalOrPublisher="Novon" pageId="0" pageNumber="29" pagination="23 - 30" refId="B17" refString="Laegaard, S, 1998. New species and names in Ecuadorian grasses (Poaceae). Novon 8 (1): 23 - 30, DOI: https://doi.org/10.2307/3391886" title="New species and names in Ecuadorian grasses (Poaceae)." url="https://doi.org/10.2307/3391886" volume="8" year="1998">Laegaard (1998</bibRefCitation>
|
||
: 27) were found to be not good for differentiating the two species. While
|
||
<taxonomicName lsidName="D. parodiana" pageId="0" pageNumber="29" rank="species" species="parodiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="29">D. parodiana</emphasis>
|
||
</taxonomicName>
|
||
only has a lax open inflorescence with long pendant branches and spikelets glomerate on the distal part of the branches,
|
||
<taxonomicName lsidName="D. podophora" pageId="0" pageNumber="29" rank="species" species="podophora">
|
||
<emphasis italics="true" pageId="0" pageNumber="29">D. podophora</emphasis>
|
||
</taxonomicName>
|
||
exhibits both denser semi-spikelike inflorescences with inflorescence branches having spikelets from the base as well as open inflorescences with pendulous branches and spikelets glomerate on the distal part of the branches like that of
|
||
<taxonomicName lsidName="D. parodiana" pageId="0" pageNumber="29" rank="species" species="parodiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="29">D. parodiana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="29">
|
||
Some specimens from Colombia were found to be generally larger than those from Ecuador, in terms of the number and length of the culms with mainly cauline leaf blades that were longer and wider. These specimens had a habit appearance similar to var.
|
||
<taxonomicName lsidName="var. mutica" pageId="0" pageNumber="29" rank="variety" species="parodiana" variety="mutica">mutica</taxonomicName>
|
||
being larger tussock-forming plants with multiple culms and inflorescences held within sheaths.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |