139 lines
9.9 KiB
XML
139 lines
9.9 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.113.1353" ID-GBIF-Dataset="953ba201-d189-4d18-b7cb-dfceb46900a3" ID-PMC="PMC3187627" ID-Pensoft-Pub="1313-2970-113-39" ID-PubMed="21976992" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-113-39" ModsDocOrigin="ZooKeys 113" ModsDocTitle="The dazed and confused identity of Agassiz’s land tortoise, Gopherus agassizii (Testudines, Testudinidae) with the description of a new species, and its consequences for conservation" checkinTime="1451250228616" checkinUser="pensoft" docAuthor="Murphy, Robert W., Berry, Kristin H., Edwards, Taylor, Leviton, Alan E., Lathrop, Amy & Riedle, J. Daren" docDate="2011" docId="6E53861C67BFC80390CBF48B0B2984EF" docLanguage="en" docName="ZooKeys 113: 39-71" docOrigin="ZooKeys 113" docSource="http://dx.doi.org/10.3897/zookeys.113.1353" docTitle="Gopherus agassizii Cooper 1861" docType="treatment" docVersion="3" lastPageNumber="49" masterDocId="FFFB155FD763FF9D2C30FF8882076558" masterDocTitle="The dazed and confused identity of Agassiz's land tortoise, Gopherus agassizii (Testudines, Testudinidae) with the description of a new species, and its consequences for conservation" masterLastPageNumber="71" masterPageNumber="39" pageNumber="48" updateTime="1668151780330" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The dazed and confused identity of Agassiz's land tortoise, Gopherus agassizii (Testudines, Testudinidae) with the description of a new species, and its consequences for conservation</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Murphy, Robert W.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Berry, Kristin H.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Edwards, Taylor</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Leviton, Alan E.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lathrop, Amy</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Riedle, J. Daren</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>113</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>39</mods:start>
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<mods:end>71</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.113.1353</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.113.1353</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-113-39</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152030064" LSID="urn:lsid:plazi:treatment:6E53861C67BFC80390CBF48B0B2984EF" httpUri="http://treatment.plazi.org/id/6E53861C67BFC80390CBF48B0B2984EF" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48">
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<subSubSection pageId="9" pageNumber="48" type="nomenclature">
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<paragraph pageId="9" pageNumber="48">
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<normalizedToken originalValue="Agassiz’s">Agassiz's</normalizedToken>
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Desert Tortoise
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<taxonomicName LSID="http://species-id.net/wiki/Gopherus_agassizii" authority="Cooper, 1861" authorityName="Cooper" authorityYear="1861" class="Reptilia" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus agassizii" order="Testudines" pageId="9" pageNumber="48" phylum="Chordata" rank="species" species="agassizii">Gopherus agassizii (Cooper, 1861)</taxonomicName>
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Figs 48
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48" type="lectotype">
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<paragraph pageId="9" pageNumber="48">Lectotype.</paragraph>
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<paragraph lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48">
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USNM (National Museum of Natural History, Smithsonian Institution) 7888; terra typica restricta: California, San Bernardino County; Mountains of California, near Fort Mojave; Soda Valley (very approximately
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="35.1">35°6'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-116.1">116°6'W</geoCoordinate>
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). We restrict the type locality to that published by
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<bibRefCitation author="Cooper, JG" journalOrPublisher="1)" pageId="21" pageNumber="60" pagination="118 - 123" title="New Californian animals. Proceedings of the California Academy of Sciences (ser." volume="2" year="1861">Cooper (1861)</bibRefCitation>
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and the parchment tag
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<pageBreakToken pageId="10" pageNumber="49" start="start">associated</pageBreakToken>
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with the specimen. References to other localities, including Solado Valley, an apparent synonym of Soda Valley, should be considered to be in error.
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</paragraph>
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<paragraph pageId="10" pageNumber="49">
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The evolutionary species concept (
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<bibRefCitation author="Simpson, GG" journalOrPublisher="Columbia University Press, New York" pageId="21" pageNumber="60" title="Principles of Animal Taxonomy" year="1961">Simpson 1961</bibRefCitation>
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;
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<bibRefCitation author="Wiley, EO" journalOrPublisher="Systematic Biology" pageId="21" pageNumber="60" pagination="17 - 26" title="The evolutionary species concept reconsidered" volume="27" year="1978">Wiley 1978</bibRefCitation>
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) suggests that the Sonoran population of the desert tortoise should be recognized as a new taxon.
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<bibRefCitation author="Frost, DR" journalOrPublisher="Herpetologica" pageId="21" pageNumber="60" pagination="87 - 104" title="Species in concept and practice: herpetological applications." volume="46" year="1990">Frost and Hillis (1990)</bibRefCitation>
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effectively argue that subspecies should not be recognized for continuously distributed species; we agree. Given these two observations, at least two species of desert tortoise should be recognized. The DNA sequence data exclude application of the available name
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<taxonomicName class="Reptilia" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus lepidocephalus" order="Testudines" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="lepidocephalus">Gopherus lepidocephalus</taxonomicName>
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for the Sonoran Desert population of
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<taxonomicName class="Reptilia" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus" order="Testudines" pageId="10" pageNumber="49" phylum="Chordata" rank="genus">Gopherus</taxonomicName>
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that occurs west and south of the Colorado River and they confirm that the lectotype of
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<taxonomicName class="Reptilia" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus agassizii" order="Testudines" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="agassizii">Gopherus agassizii</taxonomicName>
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is from the Mojave Desert, and not Arizona. Because no names are available for the tortoise population occurring in the Sonoran Desert south and east of the Colorado River, we describe it as a new species.
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</paragraph>
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<caption pageId="10" pageNumber="49">
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<paragraph pageId="10" pageNumber="49">
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Figure 4. Dorsal view of the lectotype of
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<taxonomicName class="Reptilia" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus agassizii" order="Testudines" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="agassizii">Gopherus agassizii</taxonomicName>
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, USNM 7888. Black bar is 3 cm.
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</paragraph>
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</caption>
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<caption pageId="10" pageNumber="49">
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<paragraph pageId="10" pageNumber="49">
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Figure 5. Ventral view of the holotype of
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<taxonomicName class="Reptilia" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus agassizii" order="Testudines" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="agassizii">Gopherus agassizii</taxonomicName>
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, USNM 7888. Black bar is 3 cm.
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</paragraph>
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</caption>
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<caption pageId="10" pageNumber="49">
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<paragraph pageId="10" pageNumber="49">
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Figure 6. Anterior view of the holotype of
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<taxonomicName class="Reptilia" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus agassizii" order="Testudines" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="agassizii">Gopherus agassizii</taxonomicName>
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, USNM 7888. Black bar is 3 cm.
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</paragraph>
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</caption>
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<caption pageId="10" pageNumber="49">
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<paragraph pageId="10" pageNumber="49">
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Figure 7. Posterior view of the holotype of
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<taxonomicName class="Reptilia" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus agassizii" order="Testudines" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="agassizii">Gopherus agassizii</taxonomicName>
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, USNM 7888.
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</paragraph>
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</caption>
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<caption pageId="10" pageNumber="49">
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<paragraph pageId="10" pageNumber="49">
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Figure 8. Left, lateral view of the holotype of
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<taxonomicName class="Reptilia" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus agassizii" order="Testudines" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="agassizii">Gopherus agassizii</taxonomicName>
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, USNM 7888.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |