192 lines
19 KiB
XML
192 lines
19 KiB
XML
<document id="0EB9181EF6F90591182EC4624F1D0E27" ID-DOI="10.11646/zootaxa.1100.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5051728" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625110131895" checkinUser="felipe" docAuthor="Prena, Jens" docDate="2005" docId="4540A14CCF709E20B436DD57FCF733D9" docLanguage="en" docName="zt01100p126.pdf" docOrigin="Zootaxa 1100 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1100.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Embates oculifer Prena 2005, sp. n." docType="treatment" docUuid="22427C55-420C-486A-A8B7-435C333C3A90" docUuidSource="ZooBank" docVersion="6" lastPageNumber="151" masterDocId="B979D934CF3D9E6EB53ED853FFA1377F" masterDocTitle="The Middle American species of Embates Chevrolat (Coleoptera: Curculionidae: Baridinae)" masterLastPageNumber="151" masterPageNumber="1" pageNumber="1" updateTime="1698981629170" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="B5A59F4D0F0E2564A6DC892D1DD5B34C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="58781953CA7B217840EB6B798BD9F40D">
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<mods:title id="90A815BC56415A374F578EAB2F9B7DA8">The Middle American species of Embates Chevrolat (Coleoptera: Curculionidae: Baridinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="6ADC96036D122FECB7487873D7EA20FC" type="personal">
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<mods:role id="1EB6CD3282841CA6315E3E1558533832">
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<mods:roleTerm id="47D115E0FD15B7AE8EB5673CC5975E5B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A3BD3791F0D3F87121FFED9097A719FD">Prena, Jens</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="480572B06DB536D82663C25E9BE187BB">text</mods:typeOfResource>
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<mods:titleInfo id="95A7A41762FD4A156DB46ED7FFC4E312">
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<mods:title id="C9319A57F48E816CE10C982179644482">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="17DC3FF74FA9532170A5D2CCA889286C">
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<mods:date id="E8DC9D4321273F5E493A8D25E4C490BE">2005</mods:date>
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<mods:detail id="63E7207EF1CDF786113158280C292225" type="pubDate">
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<mods:number id="D591A076CEB89BF16F7FEB919CB7D5F1">2005-12-19</mods:number>
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</mods:detail>
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<mods:detail id="8902E0D18B5110A4129C0C1C30CC0EA6" type="volume">
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<mods:number id="FB4F478B2326796DD67E3992B0AC92A4">1100</mods:number>
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</mods:detail>
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<mods:detail id="CBF621637347E5F4DBBA5C7D10502CA9" type="issue">
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<mods:number id="F55F09421C01F681BD3805B35DEA360E">1</mods:number>
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<mods:extent id="A756C4E3DD08CA9D9529340D3AF11AFC" unit="page">
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<mods:start id="94CF2F222778DAD72D20FEF79699FD02">1</mods:start>
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<mods:end id="A58096DEF235773D8880F6ABA5FBF4CC">151</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="C3204F47169A8463F3A38554A36817D8">
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<mods:url id="E9430CF99C7CB901E56C1E5AD3975372">https://biotaxa.org/Zootaxa/article/view/zootaxa.1100.1.1</mods:url>
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</mods:location>
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<mods:classification id="B633115365EB0E7D4C229396ACBA625A">journal article</mods:classification>
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<mods:identifier id="7A4113D59C3E4EFC51CA7A3AC0A39EC6" type="DOI">10.11646/zootaxa.1100.1.1</mods:identifier>
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<mods:identifier id="2A5F4F821952ABB93F6223A1387636BC" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="AA071891C9040EC6E9AC505C45AC54F0" type="Zenodo-Dep">5051728</mods:identifier>
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<mods:identifier id="97C1E4E0A8566B84FD6F6E20E9592492" type="ZooBank">7C1F1264-5F23-4557-BFC2-4D015289CF7E</mods:identifier>
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</mods:mods>
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<treatment id="4540A14CCF709E20B436DD57FCF733D9" ID-DOI="http://doi.org/10.5281/zenodo.5054449" ID-Zenodo-Dep="5054449" LSID="urn:lsid:zoobank.org:act:22427C55-420C-486A-A8B7-435C333C3A90" httpUri="http://treatment.plazi.org/id/4540A14CCF709E20B436DD57FCF733D9" lastPageId="78" pageId="77">
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<subSubSection id="85F343D1CF709E23B436DD57FD3C3261" box="[264,669,1284,1310]" pageId="77" type="nomenclature">
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<paragraph id="CD56105ACF709E23B436DD57FD3C3261" blockId="77.[264,669,1284,1345]" box="[264,669,1284,1310]" pageId="77">
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<heading id="961EA736CF709E23B436DD57FD3C3261" bold="true" box="[264,669,1284,1310]" fontSize="11" level="1" pageId="77" reason="1">
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||
<emphasis id="FF9DCC48CF709E23B436DD57FD3C3261" bold="true" box="[264,669,1284,1310]" pageId="77">
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<emphasis id="FF9DCC48CF709E23B436DD57FE8D3261" bold="true" box="[264,300,1284,1310]" italics="true" pageId="77">41.</emphasis>
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<taxonomicName id="0AE96BD9CF709E23B40ADD57FDF23261" ID-CoL="6F7FD" authority="Prena" authorityName="Prena" authorityYear="2005" box="[308,595,1284,1310]" class="Insecta" family="Curculionidae" genus="Embates" kingdom="Animalia" order="Coleoptera" pageId="77" phylum="Arthropoda" rank="species" species="oculifer" status="sp. nov.">
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<emphasis id="FF9DCC48CF709E23B40ADD57FDA03261" bold="true" box="[308,513,1284,1310]" italics="true" pageId="77">Embates oculifer</emphasis>
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Prena
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</taxonomicName>
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<taxonomicNameLabel id="E4AE7133CF709E23B765DD57FD3C3261" box="[603,669,1284,1310]" pageId="77" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="85F343D1CF709E20B436DD74FCF733D9" lastPageId="78" pageId="77" type="description">
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<paragraph id="CD56105ACF709E23B436DD74FE16323E" blockId="77.[264,669,1284,1345]" box="[264,439,1319,1345]" pageId="77">(Fig. 151, 258)</paragraph>
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<paragraph id="CD56105ACF709E23B436DD24FEC8329E" blockId="77.[264,1323,1399,1905]" pageId="77">
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<materialsCitation id="7D811A07CF709E23B436DD24FEC8329E" collectingDate="2000-03-29" collectionCode="INBC" collectorName="Prena" country="Costa Rica" elevation="1350" latitude="9.4375" location="Chucuyo" municipality="Cerro" pageId="77" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
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<typeStatus id="1252AEF8CF709E23B436DD24FED432EE" box="[264,373,1399,1425]" pageId="77" type="holotype">Holotype</typeStatus>
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male (dissected),
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<collectingCountry id="B5FE50CACF709E23B772DD24FD6E32EE" box="[588,719,1399,1425]" name="Costa Rica" pageId="77">Costa Rica</collectingCountry>
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, labeled: “
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<collectingCountry id="B5FE50CACF709E23B675DD24FC7032EE" box="[843,977,1399,1425]" name="Costa Rica" pageId="77">Costa Rica</collectingCountry>
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,
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<collectingRegion id="0F2DDEB8CF709E23B6E0DD24FBC932EE" box="[990,1128,1399,1425]" country="Costa Rica" name="San Jose" pageId="77">SAN JOSÉ</collectingRegion>
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: 12/ km NE San Isidro,
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<collectingMunicipality id="2D328A20CF709E23B463DDCCFE3E32C6" box="[349,415,1439,1465]" pageId="77">Cerro</collectingMunicipality>
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/
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<location id="C8364681CF709E23B48EDDCCFDBC32C6" LSID="urn:lsid:plazi:treatment:4540A14CCF709E20B436DD57FCF733D9:C8364681CF709E23B48EDDCCFDBC32C6" box="[432,541,1439,1465]" country="Costa Rica" latitude="9.4375" municipality="Cerro" name="Chucuyo" pageId="77" stateProvince="San Jose">Chucuyo</location>
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,
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<geoCoordinate id="A8DD769DCF709E23B715DDCCFD1E32C6" box="[555,703,1439,1465]" degrees="9" direction="north" minutes="26" orientation="latitude" pageId="77" precision="15" seconds="15" value="9.4375">9° 26' 15'' N</geoCoordinate>
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83°/ 36' 55'' W,
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<quantity id="0A11BDBFCF709E23B6B6DDCCFC4532C6" box="[904,996,1439,1465]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="77" unit="m" value="1350.0">
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<elevation id="46C4F769CF709E23B6B6DDCCFC4532C6" box="[904,996,1439,1465]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="77" unit="m" value="1350.0">1350 m</elevation>
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</quantity>
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,/ 28/
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<date id="B957369ACF709E23B11EDDCCFB3832C6" box="[1056,1177,1439,1465]" pageId="77" value="2000-03-29">
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<collectingDate id="A913CF72CF709E23B11EDDCCFB3832C6" box="[1056,1177,1439,1465]" pageId="77" value="2000-03-29">29.3.2000</collectingDate>
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</date>
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, leg.
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<collectorName id="601C758CCF709E23B1E5DDCCFA8032C6" box="[1243,1313,1439,1465]" pageId="77">Prena</collectorName>
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” (
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<collectionCode id="ABF8889FCF709E23B42EDD94FEFD329E" box="[272,348,1479,1505]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="77">INBC</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="CD56105ACF709E23B406DDBCFA843176" blockId="77.[264,1323,1399,1905]" box="[312,1317,1519,1545]" pageId="77">
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<materialsCitation id="7D811A07CF709E23B406DDBCFA803176" box="[312,1313,1519,1545]" collectionCode="CWOB" country="Costa Rica" location="Costa Rica" pageId="77" specimenCode="INBC 2, JPPC 7" specimenCount="10" specimenCount-female="2" specimenCount-male="8" typeStatus="paratype">
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<typeStatus id="1252AEF8CF709E23B406DDBCFE0B3176" box="[312,426,1519,1545]" pageId="77" type="paratype">Paratypes</typeStatus>
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10 (
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<specimenCount id="DBEFDBD3CF709E23B4E1DDBCFD9D3176" box="[479,572,1519,1545]" pageId="77" type="male">8 males</specimenCount>
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,
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<specimenCount id="DBEFDBD3CF709E23B776DDBCFD1C3176" box="[584,701,1519,1545]" pageId="77" type="female">2 females</specimenCount>
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),
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<collectingCountry id="B5FE50CACF709E23B7EFDDBCFCF23176" box="[721,851,1519,1545]" name="Costa Rica" pageId="77">Costa Rica</collectingCountry>
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, same label (
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<collectionCode id="ABF8889FCF709E23B6D5DDBCFBE63176" box="[1003,1095,1519,1545]" pageId="77">CWOB</collectionCode>
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,
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<specimenCode id="9D4FB821CF709E23B16ADDBCFB103176" box="[1108,1201,1519,1545]" collectionCode="INBC" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="77">INBC 2</specimenCode>
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,
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<specimenCode id="9D4FB821CF709E23B180DDBCFAB73176" box="[1214,1302,1519,1545]" collectionCode="JPPC" pageId="77">JPPC 7</specimenCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="CD56105ACF709E20B406DE44FB913509" blockId="77.[264,1323,1399,1905]" lastBlockId="78.[264,1323,284,1190]" lastPageId="78" pageId="77">
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Description. Habitus: total length
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<quantity id="0A11BDBFCF709E23B7DDDE44FCD6314E" box="[739,887,1559,1585]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" metricValueMax="4.7" metricValueMin="3.7" pageId="77" unit="mm" value="4.2" valueMax="4.7" valueMin="3.7">3.7–4.7 mm</quantity>
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(m=4.1, n=11). Color: integument rufouscastaneous, venter piceous; basic vestiture moderately dense, cupreous, dorsolateral pronotal vitta of cupreous scales broad, illdefined, scales velvety black in pentagonal elytral macula (Fig. 151); venter with white scales on prosternum and along flank. Head: frontal fovea absent, rostrum rather slender, curved, subcylindrical (as Fig. 144), apex slightly expanded laterally, subcostate dorsomedially, basolateral margin slightly edged, length of rostrum
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<specimenCount id="DBEFDBD3CF709E23B4DFDF5BFDA93065" box="[481,520,1800,1818]" pageId="77" type="male">♂♂</specimenCount>
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1.11–1.27 x (m=1.18, n=9),
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<specimenCount id="DBEFDBD3CF709E23B65DDF5BFC273065" box="[867,902,1800,1818]" pageId="77" type="female">♀♀</specimenCount>
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1.18–1.24 x (n=2) pronotal length, length of anteantennal portion
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<specimenCount id="DBEFDBD3CF709E23B746DF63FD3E303D" box="[632,671,1840,1858]" pageId="77" type="male">♂♂</specimenCount>
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0.42–0.45 x (m=0.43, n=9),
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<specimenCount id="DBEFDBD3CF709E23B6CFDF63FBB4303D" box="[1009,1045,1840,1858]" pageId="77" type="female">♀♀</specimenCount>
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0.47–0.49 x (n=2) total rostral length, dorsal margin of antennal scrobe reaching rostral base before eye; length of funicular segments 1 and 2 subequal, club oblong ovate. Pronotum: length 0.95–1.01 x (m=0.98, n=11) maximum width, sides subparallel to slightly rounded in basal half, anterior portion tubulate; disk densely punctate, intervals granulose, subcostate dorsomedially. Elytra: length 1.83–1.92 x (m=1.88, n=11) width at humeri, width 1.32–1.39 x (m=1.35, n=11) maximum pronotal width, sides subparallel in basal half, apices rounded conjointly, preapical callus developed, striae fine, punctures indistinct, interstriae flat, 9 costate, 7 subcostate near subapical callus. Legs: slender, tibia nearly straight and parallelsided, ventral margin with distal cluster of cupreous hairs, tarsal claws flat and approximate at base. Male: aedeagus as
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<taxonomicName id="0AE96BD9CF739E20B717DA0FFD1E350A" authorityName="Prena" authorityYear="2005" box="[553,703,604,629]" class="Insecta" family="Curculionidae" genus="Embates" kingdom="Animalia" order="Coleoptera" pageId="78" phylum="Arthropoda" rank="species" species="euchasma">
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<emphasis id="FF9DCC48CF739E20B717DA0FFD1E350A" box="[553,703,604,629]" italics="true" pageId="78">E. euchasma</emphasis>
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</taxonomicName>
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, with apex round (as Fig. 147).
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</paragraph>
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<paragraph id="CD56105ACF739E20B406DAD7FEDA3591" blockId="78.[264,1323,284,1190]" pageId="78">
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Plant association. Species of
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<taxonomicName id="0AE96BD9CF739E20B7B2DAD7FD6C35E2" box="[652,717,644,669]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="78" phylum="Tracheophyta" rank="genus">
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<emphasis id="FF9DCC48CF739E20B7B2DAD7FD6C35E2" box="[652,717,644,669]" italics="true" pageId="78">Piper</emphasis>
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</taxonomicName>
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with the shootapex emerging from within the leafbase at flowering nodes:
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<taxonomicName id="0AE96BD9CF739E20B705DAFFFD1B35BA" box="[571,698,684,709]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="78" phylum="Tracheophyta" rank="species" species="maxonii">
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<emphasis id="FF9DCC48CF739E20B705DAFFFD1B35BA" box="[571,698,684,709]" italics="true" pageId="78">P. maxonii</emphasis>
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</taxonomicName>
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(Prena 6),
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<taxonomicName id="0AE96BD9CF739E20B679DAFFFC7635BA" box="[839,983,684,709]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="78" phylum="Tracheophyta" rank="species" species="imperiale">
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<emphasis id="FF9DCC48CF739E20B679DAFFFC7635BA" box="[839,983,684,709]" italics="true" pageId="78">P. imperiale</emphasis>
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</taxonomicName>
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(Prena 4),
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<taxonomicName id="0AE96BD9CF739E20B15ADAFFFA8A35BA" box="[1124,1323,684,709]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="78" phylum="Tracheophyta" rank="species" species="fimbriculatum">
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<emphasis id="FF9DCC48CF739E20B15ADAFFFA8A35BA" box="[1124,1323,684,709]" italics="true" pageId="78">P. fimbriculatum</emphasis>
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</taxonomicName>
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(Prena 1).
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</paragraph>
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<paragraph id="CD56105ACF739E20B406DAAFFB673469" blockId="78.[264,1323,284,1190]" box="[312,1222,764,790]" pageId="78">
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Distribution.
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<collectingCountry id="B5FE50CACF739E20B4EADAAFFDF73469" box="[468,598,764,790]" name="Costa Rica" pageId="78">Costa Rica</collectingCountry>
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, Pacific side, one location near San Isidro (Fig. 258).
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</paragraph>
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<paragraph id="CD56105ACF739E20B406DB77FD9A3419" blockId="78.[264,1323,284,1190]" pageId="78">Specific epithet. The name is a Latin compound adjective derived from oculus (spot, stigma) and fero (to bear).</paragraph>
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<paragraph id="CD56105ACF739E20B406DB27FCF733D9" blockId="78.[264,1323,284,1190]" pageId="78">
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Discussion. The meristic data indicate that
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<taxonomicName id="0AE96BD9CF739E20B67CDB27FC6034F2" authorityName="Prena" authorityYear="2005" box="[834,961,884,909]" class="Insecta" family="Curculionidae" genus="Embates" kingdom="Animalia" order="Coleoptera" pageId="78" phylum="Arthropoda" rank="species" species="oculifer">
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<emphasis id="FF9DCC48CF739E20B67CDB27FC6034F2" box="[834,961,884,909]" italics="true" pageId="78">E. oculifer</emphasis>
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</taxonomicName>
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is related more closely to the largersized
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<taxonomicName id="0AE96BD9CF739E20B4A6DBCFFDED34CA" authorityName="Prena" authorityYear="2005" box="[408,588,924,949]" class="Insecta" family="Curculionidae" genus="Embates" kingdom="Animalia" order="Coleoptera" pageId="78" phylum="Arthropoda" rank="species" species="subulirostris">
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<emphasis id="FF9DCC48CF739E20B4A6DBCFFDED34CA" box="[408,588,924,949]" italics="true" pageId="78">E. subulirostris</emphasis>
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</taxonomicName>
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rather than to the similarly small
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<taxonomicName id="0AE96BD9CF739E20B6E8DBCFFBFA34CA" authorityName="Prena" authorityYear="2005" box="[982,1115,924,949]" class="Insecta" family="Curculionidae" genus="Embates" kingdom="Animalia" order="Coleoptera" pageId="78" phylum="Arthropoda" rank="species" species="cordiger">
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<emphasis id="FF9DCC48CF739E20B6E8DBCFFBFA34CA" box="[982,1115,924,949]" italics="true" pageId="78">E. cordiger</emphasis>
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</taxonomicName>
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from the Atlantic side. Both smallsized species differ significantly (p<0.001) in the length of the rostrum, antennal insertion, elytral proportion and total length. However,
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<taxonomicName id="0AE96BD9CF739E20B132DBBFFB2B337A" authorityName="Prena" authorityYear="2005" box="[1036,1162,1004,1029]" class="Insecta" family="Curculionidae" genus="Embates" kingdom="Animalia" order="Coleoptera" pageId="78" phylum="Arthropoda" rank="species" species="oculifer">
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<emphasis id="FF9DCC48CF739E20B132DBBFFB2B337A" box="[1036,1162,1004,1029]" italics="true" pageId="78">E. oculifer</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="0AE96BD9CF739E20B1F9DBBFFEC53352" authorityName="Prena" authorityYear="2005" class="Insecta" family="Curculionidae" genus="Embates" kingdom="Animalia" order="Coleoptera" pageId="78" phylum="Arthropoda" rank="species" species="subulirostris">
|
||
<emphasis id="FF9DCC48CF739E20B1F9DBBFFEC53352" italics="true" pageId="78">E. subulirostris</emphasis>
|
||
</taxonomicName>
|
||
differ in total length only (p<0.001). I infer that
|
||
<taxonomicName id="0AE96BD9CF739E20B6A4DC47FBEC3352" authorityName="Prena" authorityYear="2005" box="[922,1101,1044,1069]" class="Insecta" family="Curculionidae" genus="Embates" kingdom="Animalia" order="Coleoptera" pageId="78" phylum="Arthropoda" rank="species" species="subulirostris">
|
||
<emphasis id="FF9DCC48CF739E20B6A4DC47FBEC3352" box="[922,1101,1044,1069]" italics="true" pageId="78">E. subulirostris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="0AE96BD9CF739E20B1BBDC47FAA13352" authorityName="Prena" authorityYear="2005" box="[1157,1280,1044,1069]" class="Insecta" family="Curculionidae" genus="Embates" kingdom="Animalia" order="Coleoptera" pageId="78" phylum="Arthropoda" rank="species" species="oculifer">
|
||
<emphasis id="FF9DCC48CF739E20B1BBDC47FAA13352" box="[1157,1280,1044,1069]" italics="true" pageId="78">E. oculifer</emphasis>
|
||
</taxonomicName>
|
||
are Pacific relict populations that became isolated geographically by climatic changes in the past. It should be noted, that male
|
||
<taxonomicName id="0AE96BD9CF739E20B7A4DC37FCB73302" authorityName="Prena" authorityYear="2005" box="[666,790,1124,1149]" class="Insecta" family="Curculionidae" genus="Embates" kingdom="Animalia" order="Coleoptera" pageId="78" phylum="Arthropoda" rank="species" species="oculifer">
|
||
<emphasis id="FF9DCC48CF739E20B7A4DC37FCB73302" box="[666,790,1124,1149]" italics="true" pageId="78">E. oculifer</emphasis>
|
||
</taxonomicName>
|
||
have medially depressed ventrites, while this character state is unreliable for sexing
|
||
<taxonomicName id="0AE96BD9CF739E20B7F2DCDFFCEE33DA" authorityName="Prena" authorityYear="2005" box="[716,847,1164,1189]" class="Insecta" family="Curculionidae" genus="Embates" kingdom="Animalia" order="Coleoptera" pageId="78" phylum="Arthropoda" rank="species" species="cordiger">
|
||
<emphasis id="FF9DCC48CF739E20B7F2DCDFFCEE33DA" box="[716,847,1164,1189]" italics="true" pageId="78">E. cordiger</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |