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<mods:title id="DBBDF02416205C518D4D6CCCDA1DE227">The genus Coleobonzia declared synonymous with Neobonzia Smiley, 1992 (Bdelloidea: Cunaxidae: Coleoscirinae)</mods:title>
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<paragraph id="A75C0629592B454EE9AAFF5CFDF1FF0E" blockId="1.[151,536,220,245]" box="[151,536,220,245]" pageId="1" pageNumber="60">
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<emphasis id="9597DA3B592B454EE9AAFF5CFDF1FF0E" bold="true" box="[151,536,220,245]" pageId="1" pageNumber="60">
Genus
<taxonomicName id="60E37DAA592B454EE9DBFF5DFDF1FF0E" authority="Smiley, 1992" authorityName="Smiley" authorityYear="1992" box="[230,536,220,245]" class="Arachnida" family="Cunaxidae" genus="Neobonzia" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="9597DA3B592B454EE9DBFF5DFE6AFF0F" bold="true" box="[230,387,221,245]" italics="true" pageId="1" pageNumber="60">NEOBONZIA</emphasis>
<bibRefCitation id="C3727BD8592B454EE8B7FF5CFDF1FF0E" author="Smiley" box="[394,536,220,244]" pageId="1" pageNumber="60" refString="Smiley, R. L. (1992) The predatory mite family Cunaxidae (Acari) of the world with a new classification. Indira Publishing House, West Bloomfield, Michigan. 356 pp." type="book" year="1992">Smiley, 1992</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<treatmentCitationGroup id="87F32107592B454EE9AAFEA2FCF7FEC3" box="[151,798,290,313]" pageId="1" pageNumber="60">
<taxonomicName id="60E37DAA592B454EE9AAFEA2FDD2FEC3" authority="Den Heyer 1977: 178" authorityName="Den Heyer" authorityPageNumber="178" authorityYear="1977" box="[151,571,290,313]" class="Arachnida" family="Cunaxidae" genus="Pseudobonzia" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="9597DA3B592B454EE9AAFEA2FEC4FEC3" box="[151,301,290,313]" italics="true" pageId="1" pageNumber="60">Pseudobonzia</emphasis>
<treatmentCitation id="26422038592B454EE869FEA2FDD2FEC3" author="Den" box="[340,571,290,313]" page="178" pageId="1" pageNumber="60" year="1977">
<bibRefCitation id="C3727BD8592B454EE869FEA2FDE1FEC3" author="Den" box="[340,520,290,313]" pageId="1" pageNumber="60" refString="Den Heyer, J. (1977) Six new species of Pseudobonzia Smiley, 1975 (Prostigmata: Acari) from the Ethiopian Region. Journal of the entomological Society of southern Africa, 40 (2), 171 - 194." type="journal article" year="1977">Den Heyer 1977</bibRefCitation>
: 178
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194; 1980: 121124
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</paragraph>
<paragraph id="A75C0629592B454EE9AAFEC2FD20FEA3" blockId="1.[151,924,290,409]" box="[151,713,322,345]" pageId="1" pageNumber="60">
<treatmentCitationGroup id="87F32107592B454EE9AAFEC2FD20FEA3" box="[151,713,322,345]" pageId="1" pageNumber="60">
<taxonomicName id="60E37DAA592B454EE9AAFEC2FD72FEA3" authority="Den Heyer &amp; Castro 2008: 42" authorityName="Den Heyer &amp; Castro" authorityPageNumber="42" authorityYear="2008" box="[151,667,322,345]" class="Arachnida" family="Cunaxidae" genus="Coleobonzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="9597DA3B592B454EE9AAFEC2FEF7FEA3" box="[151,286,322,345]" italics="true" pageId="1" pageNumber="60">Coleobonzia</emphasis>
<treatmentCitation id="26422038592B454EE867FEC2FD72FEA3" author="Den" box="[346,667,322,345]" page="42" pageId="1" pageNumber="60" year="2008">
<bibRefCitation id="C3727BD8592B454EE867FEC2FD9CFEA3" author="Den" box="[346,629,322,345]" pageId="1" pageNumber="60" refString="Den Heyer, J. &amp; Castro, T. M. M. G de. (2008) A new cunaxid genus with descriptions of two new species from Brazil (Acari: Prostigmata: Bdelloidea: Cunaxidae). Zootaxa, 1731, 42 - 50." type="journal article" year="2008">Den Heyer &amp; Castro 2008</bibRefCitation>
: 42
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</taxonomicName>
50.
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</paragraph>
<paragraph id="A75C0629592B454DE9AAFEE2FC56F831" blockId="1.[151,924,290,409]" lastBlockId="2.[151,1436,1729,1995]" lastPageId="2" lastPageNumber="61" pageId="1" pageNumber="60">
<treatmentCitationGroup id="87F32107592B454DE9AAFEE2FC56F831" infered="true" lastPageId="2" lastPageNumber="61" pageId="1" pageNumber="60">
<taxonomicName id="60E37DAA592B454EE9AAFEE2FDFFFE83" authority="Smiley 1992: 331" authorityName="Smiley" authorityPageNumber="331" authorityYear="1992" box="[151,534,354,377]" class="Arachnida" family="Cunaxidae" genus="Neobonzia" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="9597DA3B592B454EE9AAFEE2FEE0FE83" box="[151,265,354,377]" italics="true" pageId="1" pageNumber="60">Neobonzia</emphasis>
<treatmentCitation id="26422038592B454EE864FEE2FDFFFE83" author="Smiley" box="[345,534,354,377]" page="331" pageId="1" pageNumber="60" year="1992">
<bibRefCitation id="C3727BD8592B454EE864FEE2FE0DFE83" author="Smiley" box="[345,484,354,377]" pageId="1" pageNumber="60" refString="Smiley, R. L. (1992) The predatory mite family Cunaxidae (Acari) of the world with a new classification. Indira Publishing House, West Bloomfield, Michigan. 356 pp." type="book" year="1992">Smiley 1992</bibRefCitation>
: 331
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</taxonomicName>
333;
<treatmentCitation id="26422038592B454EEB64FEE2FCBAFE83" author="Corpuz-Raros" box="[601,851,354,377]" page="87" pageId="1" pageNumber="60" year="2008">
<bibRefCitation id="C3727BD8592B454EEB64FEE2FCC6FE83" author="Corpuz-Raros" box="[601,815,354,377]" pageId="1" pageNumber="60" refString="Corpuz-Raros, L. A. (2008) Additional species of Cunaxidae and first report of the subfamily Neobonzinae (Cunaxidae, Acari) from the Philippines. Asia Life Sciences, 17 (1), 71 - 89." type="journal article" year="2008">Corpuz-Raros 2008</bibRefCitation>
: 87
</treatmentCitation>
89
<typeStatus id="7858B88B592B454EE9AAFE02FF25FE63" box="[151,204,386,409]" pageId="1" pageNumber="60">Type</typeStatus>
species:
<taxonomicName id="60E37DAA592B454EE868FE02FD40FE63" authority="Smiley, 1992" authorityName="Smiley" authorityYear="1992" box="[341,681,386,409]" class="Arachnida" family="Cunaxidae" genus="Neobonzia" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="moseri">
<emphasis id="9597DA3B592B454EE868FE02FDFEFE63" box="[341,535,386,409]" italics="true" pageId="1" pageNumber="60">Neobonzia moseri</emphasis>
<bibRefCitation id="C3727BD8592B454EEB20FE02FD40FE63" author="Smiley" box="[541,681,386,409]" pageId="1" pageNumber="60" refString="Smiley, R. L. (1992) The predatory mite family Cunaxidae (Acari) of the world with a new classification. Indira Publishing House, West Bloomfield, Michigan. 356 pp." type="book" year="1992">Smiley, 1992</bibRefCitation>
</taxonomicName>
(Original designation)
<emphasis id="9597DA3B5928454DE9AAF941FCEBF923" bold="true" box="[151,770,1729,1753]" pageId="2" pageNumber="61">
Redescription of the genus
<taxonomicName id="60E37DAA5928454DE8D9F941FD15F923" authority="Smiley, 1992" authorityName="Smiley" authorityYear="1992" box="[484,764,1729,1753]" class="Arachnida" family="Cunaxidae" genus="Neobonzia" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="9597DA3B5928454DE8D9F941FDB0F923" bold="true" box="[484,601,1729,1753]" italics="true" pageId="2" pageNumber="61">Neobonzia</emphasis>
<bibRefCitation id="C3727BD85928454DEB5BF941FD15F923" author="Smiley" box="[614,764,1729,1753]" pageId="2" pageNumber="61" refString="Smiley, R. L. (1992) The predatory mite family Cunaxidae (Acari) of the world with a new classification. Indira Publishing House, West Bloomfield, Michigan. 356 pp." type="book" year="1992">Smiley, 1992</bibRefCitation>
</taxonomicName>
.
</emphasis>
<taxonomicName id="60E37DAA5928454DEA33F942FC69F923" box="[782,896,1730,1753]" class="Arachnida" family="Cunaxidae" genus="Neobonzia" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="9597DA3B5928454DEA33F942FC69F923" box="[782,896,1730,1753]" italics="true" pageId="2" pageNumber="61">Neobonzia</emphasis>
</taxonomicName>
are soft-bodied and are often, but not always, yellowish-brown. The weakly-sclerotised dorsal propodosomal areas in females are without subcuticular reticulations. In males an additional weakly sclerotised dorsal hysterosomal area may be present or absent. The gnathosoma may be darker in colour due to somewhat heavier sclerotisation. Palps 5-jointed and extend by approximately half the length of the tibiotarsus past the entomalae. Basi- and telofemora bear dorsal simple tactile seta. Tibiotarsus usually S-shaped and proximally stouter than in
<taxonomicName id="60E37DAA5928454DE880F8EFFDBAF87C" box="[445,595,1903,1926]" class="Arachnida" family="Cunaxidae" genus="Pseudobonzia" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="9597DA3B5928454DE880F8EFFDBAF87C" box="[445,595,1903,1926]" italics="true" pageId="2" pageNumber="61">Pseudobonzia</emphasis>
</taxonomicName>
, with or without a tubercle and the (pen-)ultimate seta rather long. Setae f2 present in both sexes. Coxal plates I+II separated medially. Genital setae in both sexes either three or four pairs. Legs usually shorter than idiosoma. All leg femora divided by articulation facets.
</treatmentCitationGroup>
</paragraph>
<caption id="F39C56A1592B454EE9AAF845FB21F826" box="[151,1224,1989,2013]" httpUri="https://zenodo.org/record/203916/files/figure.png" pageId="1" pageNumber="60" targetBox="[185,1412,501,1950]" targetPageId="1">
<paragraph id="A75C0629592B454EE9AAF845FB21F826" blockId="1.[151,1224,1989,2013]" box="[151,1224,1989,2013]" pageId="1" pageNumber="60">
<emphasis id="9597DA3B592B454EE9AAF845FEABF827" bold="true" box="[151,322,1989,2013]" pageId="1" pageNumber="60">FIGURES 1, 2.</emphasis>
<taxonomicName id="60E37DAA592B454EE874F846FD49F826" authority="Smiley, 1992" authorityName="Smiley" authorityYear="1992" box="[329,672,1989,2013]" class="Arachnida" family="Cunaxidae" genus="Neobonzia" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="moseri">
<emphasis id="9597DA3B592B454EE874F846FDE2F827" box="[329,523,1990,2013]" italics="true" pageId="1" pageNumber="60">Neobonzia moseri</emphasis>
Smiley, 1992
</taxonomicName>
, dorsal aspect (1) and palp (2) (after Smiley 1992).
</paragraph>
</caption>
<caption id="F39C56A15928454DE9AAF9D9FBBAF969" httpUri="https://zenodo.org/record/203917/files/figure.png" pageId="2" pageNumber="61" targetBox="[157,1429,195,1600]" targetPageId="2">
<paragraph id="A75C06295928454DE9AAF9D9FBBAF969" blockId="2.[151,1436,1625,1683]" pageId="2" pageNumber="61">
<emphasis id="9597DA3B5928454DE9AAF9D9FEAFF98B" bold="true" box="[151,326,1625,1649]" pageId="2" pageNumber="61">FIGURES 38.</emphasis>
<taxonomicName id="60E37DAA5928454DE86CF9DAFD5AF98B" authority="Smiley, 1992" authorityName="Smiley" authorityYear="1992" box="[337,691,1626,1649]" class="Arachnida" family="Cunaxidae" genus="Neobonzia" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="moseri">
<emphasis id="9597DA3B5928454DE86CF9DAFDFFF98B" box="[337,534,1626,1649]" italics="true" pageId="2" pageNumber="61">Neobonzia moseri</emphasis>
Smiley, 1992
</taxonomicName>
. (3) Entire specimen; (4) Gnathosoma and leg I; (5) Idiosoma and sensillae; (6) Legs I and II; (7) Hysterosoma and leg IV; (8) Hysterosoma, legs III and IV.
</paragraph>
</caption>
<paragraph id="A75C06295929454CE9FAFF18FD75FEC3" blockId="3.[151,1436,152,383]" pageId="3" pageNumber="62">
The above redescription is based on all the species described by
<bibRefCitation id="C3727BD85929454CEA8EFF18FBB1FF55" author="Berlese" box="[947,1112,152,175]" pageId="3" pageNumber="62" refString="Berlese, A. (1910) Acari nuovi. Redia, 6 (2), 199 - 234." type="journal article" year="1910">Berlese (1910)</bibRefCitation>
,
<bibRefCitation id="C3727BD85929454CED54FF18FA7EFF55" author="Baker" box="[1129,1431,152,175]" pageId="3" pageNumber="62" refString="Baker, E. W. &amp; Hoffmann, Anita (1948) Acaros de la familia Cunaxidae. Anales de la Escuela Nacional Ciencias Biologicas (Mexico), 5 (3 - 4), 229 - 273." type="journal article" year="1948">
<collectingCountry id="DFF446B95929454CED54FF18FB40FF55" box="[1129,1193,152,175]" name="United States Pacific Island Wildlife Refuges" pageId="3" pageNumber="62">Baker</collectingCountry>
&amp; Hoffmann (1948)
</bibRefCitation>
,
<bibRefCitation id="C3727BD85929454CE9AAFF3AFEACFF2B" author="Chaudhri" box="[151,325,186,209]" pageId="3" pageNumber="62" refString="Chaudhri, W. M. (1977) Descriptions of the mites of the Family Cunaxidae (Acarina) from Pakistan. Pakistan Journal of Agricultural Science, 14 (2 - 3), 41 - 52." type="journal article" year="1977">Chaudhri (1977</bibRefCitation>
,
<bibRefCitation id="C3727BD85929454CE86EFF3AFE65FF2B" author="Chaudhri" box="[339,396,186,209]" pageId="3" pageNumber="62" refString="Chaudhri, W. M. (1980) Studies on the biosystematics and control of mites on fieldcrops, vegetables and fruit plants in Pakistan. Report on project # PK-ARS- 150. University of Agriculture, Faisalabad. 2 nd Annual report, 1 - 78." type="journal article" year="1980">1980</bibRefCitation>
),
<bibRefCitation id="C3727BD85929454CE89FFF3AFD8DFF2B" author="Den" box="[418,612,186,209]" pageId="3" pageNumber="62" refString="Den Heyer, J. (1977) Six new species of Pseudobonzia Smiley, 1975 (Prostigmata: Acari) from the Ethiopian Region. Journal of the entomological Society of southern Africa, 40 (2), 171 - 194." type="journal article" year="1977">Den Heyer (1977</bibRefCitation>
,
<bibRefCitation id="C3727BD85929454CEB4FFF3AFD42FF2B" author="Heyer" box="[626,683,186,209]" pageId="3" pageNumber="62" refString="Den Heyer, J. (1980) Six new species of the subfamily Coleoscirinae (Cunaxidae: Actinedida: Acarida). Phytophylactica, 12, 105 - 128." type="journal article" year="1980">1980</bibRefCitation>
),
<bibRefCitation id="C3727BD85929454CEBFCFF3AFC89FF2B" author="Smiley" box="[705,864,186,209]" pageId="3" pageNumber="62" refString="Smiley, R. L. (1992) The predatory mite family Cunaxidae (Acari) of the world with a new classification. Indira Publishing House, West Bloomfield, Michigan. 356 pp." type="book" year="1992">Smiley (1992)</bibRefCitation>
,
<bibRefCitation id="C3727BD85929454CEA52FF3AFBA8FF2B" author="Sergeyenko" box="[879,1089,186,209]" pageId="3" pageNumber="62" refString="Sergeyenko, A. L. (2005) A new species of the mite genus Pseudobonzia Smiley, 1975 (Acarina: Prostigmata: Cunaxidae) from Ukraine. Acarina, 13 (2), 159 - 163." type="journal article" year="2005">Sergeyenko (2005)</bibRefCitation>
,
<bibRefCitation id="C3727BD85929454CED73FF3AFADFFF2B" author="Corpuz-Raros" box="[1102,1334,186,209]" pageId="3" pageNumber="62" refString="Corpuz-Raros, L. A. (2008) Additional species of Cunaxidae and first report of the subfamily Neobonzinae (Cunaxidae, Acari) from the Philippines. Asia Life Sciences, 17 (1), 71 - 89." type="journal article" year="2008">Corpuz-Raros (2008)</bibRefCitation>
and
<bibRefCitation id="C3727BD85929454CEC52FF3AFE7FFF0E" pageId="3" pageNumber="62" refString="Den Heyer, J. &amp; Castro, T. M. M. G de. (2008) A new cunaxid genus with descriptions of two new species from Brazil (Acari: Prostigmata: Bdelloidea: Cunaxidae). Zootaxa, 1731, 42 - 50." type="journal article">Den Heyer &amp; Castro (2008)</bibRefCitation>
. Therefore the genus concept, when all described species are taken in consideration, should be robust. However, to compile a key to all species of
<taxonomicName id="60E37DAA5929454CEBF5FE80FCD3FEED" box="[712,826,256,279]" class="Arachnida" family="Cunaxidae" genus="Neobonzia" kingdom="Animalia" order="Prostigmata" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="9597DA3B5929454CEBF5FE80FCD3FEED" box="[712,826,256,279]" italics="true" pageId="3" pageNumber="62">Neobonzia</emphasis>
</taxonomicName>
from literature, i.e. without having seen specimens, is a case of guesswork; therefore no key is provided.
</paragraph>
<paragraph id="A75C06295929454CE9FAFEC5FB90FE85" blockId="3.[151,1436,152,383]" pageId="3" pageNumber="62">
One curiousity is that it seems strange that
<bibRefCitation id="C3727BD85929454CEB94FEC5FCAEFEA6" author="Smiley" box="[681,839,325,348]" pageId="3" pageNumber="62" refString="Smiley, R. L. (1992) The predatory mite family Cunaxidae (Acari) of the world with a new classification. Indira Publishing House, West Bloomfield, Michigan. 356 pp." type="book" year="1992">Smiley (1992)</bibRefCitation>
, having incorporated all the species described by
<bibRefCitation id="C3727BD85929454CEC52FEC5FEC9FE85" author="Den" pageId="3" pageNumber="62" refString="Den Heyer, J. (1977) Six new species of Pseudobonzia Smiley, 1975 (Prostigmata: Acari) from the Ethiopian Region. Journal of the entomological Society of southern Africa, 40 (2), 171 - 194." type="journal article" year="1977">Den Heyer (1977</bibRefCitation>
,
<bibRefCitation id="C3727BD85929454CE817FEE8FE8AFE85" author="Heyer" box="[298,355,360,383]" pageId="3" pageNumber="62" refString="Den Heyer, J. (1980) Six new species of the subfamily Coleoscirinae (Cunaxidae: Actinedida: Acarida). Phytophylactica, 12, 105 - 128." type="journal article" year="1980">1980</bibRefCitation>
), did not recognise the similarity of
<taxonomicName id="60E37DAA5929454CEBD8FEE8FCBEFE85" box="[741,855,360,383]" class="Arachnida" family="Cunaxidae" genus="Neobonzia" kingdom="Animalia" order="Prostigmata" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="9597DA3B5929454CEBD8FEE8FCBEFE85" box="[741,855,360,383]" italics="true" pageId="3" pageNumber="62">Neobonzia</emphasis>
</taxonomicName>
with that group of species.
</paragraph>
</subSubSection>
</treatment>
</document>