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<document id="D659E3B8ECE8B49636A0AA98311C00C6" ID-DOI="10.1649/1035.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="10111177" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630267768663" checkinUser="valdenar" docAuthor="Chandler, Donald S. &amp; Lewis, Julian J." docDate="2008" docId="03F3C44DFFB9552CFEFB6E75FC2B5229" docLanguage="en" docName="1035.1.pdf" docOrigin="The Coleopterists Bulletin 62 (1)" docSource="http://dx.doi.org/10.1649/1035.1" docStyle="DocumentStyle:76939A7658C5F80CF4BB6818E728552A.2:TheColeopBull.2003-2009.journal_article.1cover" docStyleId="76939A7658C5F80CF4BB6818E728552A" docStyleName="TheColeopBull.2003-2009.journal_article.1cover" docStyleVersion="2" docTitle="Batrisodes (Babnormodes) hairstoni Park 1947" docType="treatment" docVersion="3" lastPageNumber="117" masterDocId="FFCABC35FFBB5528FFA56A4FFFA2507B" masterDocTitle="Notes on the Cave-Associated Pselaphinae (Staphylinidae) of Indiana" masterLastPageNumber="119" masterPageNumber="114" pageNumber="115" updateTime="1699683566605" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="2BE670D05D669524E23D720051EF4576" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4968B2C6BEAB8705A8D97753F5F4FCDF">Notes on the Cave-Associated Pselaphinae (Staphylinidae) of Indiana</mods:title>
</mods:titleInfo>
<mods:name id="3A54E9D407F9ADCDCA2A79937BB7E9C4" type="personal">
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</mods:role>
<mods:namePart id="6E52C93E4AB26A013BFF64A5049D64FB">Chandler, Donald S.</mods:namePart>
</mods:name>
<mods:name id="06915992A605720E04F81924CE12CB2D" type="personal">
<mods:role id="B9322B200C34037F7C20820989EE83AF">
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</mods:role>
<mods:namePart id="2814CBED1E0C1918871EFEA2A11F7F72">Lewis, Julian J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="6CA6D1C12B52084CCC0E0070B0DE1979">text</mods:typeOfResource>
<mods:relatedItem id="AEC44054F9182150EAE3C67C29D3F576" type="host">
<mods:titleInfo id="DE733E579044AEC3647EEE129192AC84">
<mods:title id="E6D3AC71FF5C26466D86ABD6A4F6A981">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="39A296E2EA0BB376D57B810F4A26DE81">
<mods:date id="79314B540F4AA02436DE2E3DEBE199BB">2008</mods:date>
<mods:detail id="0692926876AFF3FC9044823725EC0C9E" type="pubDate">
<mods:number id="6FF625DEB5CA00F9C07392472E79E735">2008-03-31</mods:number>
</mods:detail>
<mods:detail id="7989FD7A468A155EC1FB3D0534C6E108" type="volume">
<mods:number id="C14875ED380C3710ECD896E0015C2820">62</mods:number>
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<mods:detail id="CC8DA6709B609F88F711049980B275DB" type="issue">
<mods:number id="A46672DFEE18AAB8E76B55288509F8B8">1</mods:number>
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<mods:start id="81DB846AA7237C7BAD7317E7EF11DFE6">114</mods:start>
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<mods:url id="7124C26B162DDB9FC5426804A529B276">http://dx.doi.org/10.1649/1035.1</mods:url>
</mods:location>
<mods:classification id="6B849B335961F643CB1247FC22FF30B8">journal article</mods:classification>
<mods:identifier id="F43703D9CEB3AAE01BD750885623E33A" type="DOI">10.1649/1035.1</mods:identifier>
<mods:identifier id="40EA47B988B15301A06C245764423F09" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="F954BD22E1E2EE47378B190009ABD955" type="Zenodo-Dep">10111177</mods:identifier>
</mods:mods>
<treatment id="03F3C44DFFB9552CFEFB6E75FC2B5229" LSID="urn:lsid:plazi:treatment:03F3C44DFFB9552CFEFB6E75FC2B5229" httpUri="http://treatment.plazi.org/id/03F3C44DFFB9552CFEFB6E75FC2B5229" lastPageId="4" lastPageNumber="117" pageId="2" pageNumber="115">
<subSubSection id="C34026D0FFB9552AFEFB6E75FC9C542B" box="[350,830,1082,1104]" pageId="2" pageNumber="115" type="nomenclature">
<paragraph id="8BE5755BFFB9552AFEFB6E75FC9C542B" blockId="2.[350,830,1082,1131]" box="[350,830,1082,1104]" pageId="2" pageNumber="115">
<heading id="D0ADC237FFB9552AFEFB6E75FC9C542B" box="[350,830,1082,1104]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="115" reason="2">
<taxonomicName id="4C5A0ED8FFB9552AFEFB6E75FC9C542B" ID-CoL="L2M2" authority="Park 1947" authorityName="Park" authorityYear="1947" box="[350,830,1082,1104]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="115" phylum="Arthropoda" rank="species" species="hairstoni" subGenus="Babnormodes">
<emphasis id="B92EA949FFB9552AFEFB6E75FC9C542B" bold="true" box="[350,830,1082,1104]" pageId="2" pageNumber="115">
<emphasis id="B92EA949FFB9552AFEFB6E75FD6E542B" bold="true" box="[350,716,1082,1104]" italics="true" pageId="2" pageNumber="115">Batrisodes (Babnormodes) hairstoni</emphasis>
Park 1947
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34026D0FFB9552AFDB26E1AFD275410" box="[535,645,1109,1131]" pageId="2" pageNumber="115" type="description">
<paragraph id="8BE5755BFFB9552AFDB26E1AFD275410" blockId="2.[350,830,1082,1131]" box="[535,645,1109,1131]" pageId="2" pageNumber="115">
(
<figureCitation id="136169DEFFB9552AFDBB6E1AFDDF5410" box="[542,637,1109,1131]" captionStart="Figs" captionStartId="2.[196,235,632,651]" captionTargetBox="[209,973,194,608]" captionTargetId="figure-341@2.[207,973,194,609]" captionTargetPageId="2" captionText="Figs. 1–3. Batrisodes hairstoni. 1. Lateral view of left antenna with ventral view of apical three segments. 2. Dorsal view of head. 3. Dorsal view of aedeagus. Line equals 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10111179" httpUri="https://zenodo.org/record/10111179/files/figure.png" pageId="2" pageNumber="115">Figs. 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34026D0FFB9552AFF0A6ECEFE235579" pageId="2" pageNumber="115" type="reference_group">
<paragraph id="8BE5755BFFB9552AFF0A6ECEFC4F54C9" blockId="2.[175,1005,1153,1282]" pageId="2" pageNumber="115">
<taxonomicName id="4C5A0ED8FFB9552AFF0A6ECEFDB654EC" ID-CoL="L2M2" authority="Park 1947: 101" authorityName="Park" authorityPageNumber="101" authorityYear="1947" box="[175,532,1153,1175]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="115" phylum="Arthropoda" rank="species" species="hairstoni">
<emphasis id="B92EA949FFB9552AFF0A6ECEFED454ED" box="[175,374,1153,1174]" italics="true" pageId="2" pageNumber="115">Batrisodes hairstoni</emphasis>
<materialsCitation id="3B327F06FFB9552AFEDA6ECEFDB654EC" box="[383,532,1153,1175]" collectionCode="VI, FMNH" county="Clark Co." location="Uno" municipality="Clark Co. State Park" pageId="2" pageNumber="115" specimenCount="1" stateProvince="Indiana">Park 1947:101</materialsCitation>
</taxonomicName>
, pl. VI, fig. 5.
<typeStatus id="54E1CBF9FFB9552AFD166ECEFD4454EC" box="[691,742,1153,1175]" pageId="2" pageNumber="115">Type</typeStatus>
locality:
<collectingMunicipality id="6B81EF21FFB9552AFCEC6ECEFE8054C9" pageId="2" pageNumber="115">Clark Co. State Park</collectingMunicipality>
, near
<location id="8E852380FFB9552AFEF86ED3FE2954C9" LSID="urn:lsid:plazi:treatment:03F3C44DFFB9552CFEFB6E75FC2B5229:8E852380FFB9552AFEF86ED3FE2954C9" box="[349,395,1180,1202]" county="Clark Co." municipality="Clark Co. State Park" name="Uno" pageId="2" pageNumber="115" stateProvince="Indiana">Uno</location>
[misspelling for Union],
<collectingCounty id="62840DD7FFB9552AFD216ED3FD4854C9" box="[644,746,1180,1202]" pageId="2" pageNumber="115">Clark Co.</collectingCounty>
,
<collectingRegion id="499EBBB9FFB9552AFD566ED3FCE354C9" box="[755,833,1180,1202]" country="United States of America" name="Indiana" pageId="2" pageNumber="115">Indiana</collectingRegion>
(FMNH).
<location id="8E852380FFB9552AFC176ED3FC4854C9" LSID="urn:lsid:plazi:treatment:03F3C44DFFB9552CFEFB6E75FC2B5229:8E852380FFB9552AFC176ED3FC4854C9" box="[946,1002,1180,1202]" county="Clark Co." municipality="Clark Co. State Park" name="Male" pageId="2" pageNumber="115" stateProvince="Indiana">Male</location>
.
</paragraph>
<paragraph id="8BE5755BFFB9552AFF0A6EF8FC4E5493" blockId="2.[175,1005,1153,1282]" pageId="2" pageNumber="115">
<treatmentCitationGroup id="AB4A5275FFB9552AFF0A6EF8FC4E5493" pageId="2" pageNumber="115">
<treatmentCitation id="0AFB534AFFB9552AFF0A6EF8FD1E54B6" author="Park" box="[175,700,1207,1229]" page="74" pageId="2" pageNumber="115" year="1960">
<taxonomicName id="4C5A0ED8FFB9552AFF0A6EF8FD1E54B6" ID-CoL="8W79S" authority="Park 1960: 74" authorityName="Park" authorityPageNumber="74" authorityYear="1960" box="[175,700,1207,1229]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="115" phylum="Arthropoda" rank="species" species="krekeleri" subGenus="Babmormodes">
<emphasis id="B92EA949FFB9552AFF0A6EF8FEB554B7" box="[175,279,1207,1228]" italics="true" pageId="2" pageNumber="115">Batrisodes</emphasis>
(
<emphasis id="B92EA949FFB9552AFE8E6EF8FE1A54B7" box="[299,440,1207,1228]" italics="true" pageId="2" pageNumber="115">Babmormodes</emphasis>
)
<emphasis id="B92EA949FFB9552AFE6E6EF8FD8154B7" box="[459,547,1207,1228]" italics="true" pageId="2" pageNumber="115">krekeleri</emphasis>
<materialsCitation id="3B327F06FFB9552AFD8B6EF8FD1E54B6" box="[558,700,1207,1229]" collectionCode="FMNH, NEW" county="Clark Co." location="Cave Spring" pageId="2" pageNumber="115" specimenCount="1" stateProvince="Washington">
<bibRefCitation id="EFCB08AAFFB9552AFD8B6EF8FD1E54B6" author="Park" box="[558,700,1207,1229]" pageId="2" pageNumber="115" pagination="66 - 104" refId="ref3481" refString="Park, O. 1960. Cavernicolous pselaphid beetles of the United States. American Midland Naturalist 64 (1): 66 - 104." type="journal article" year="1960">Park 1960:74</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54E1CBF9FFB9552AFD6C6EF8FD5F54B6" box="[713,765,1207,1229]" pageId="2" pageNumber="115">Type</typeStatus>
locality:
<location id="8E852380FFB9552AFCC16EF8FC4A54B6" LSID="urn:lsid:plazi:treatment:03F3C44DFFB9552CFEFB6E75FC2B5229:8E852380FFB9552AFCC16EF8FC4A54B6" box="[868,1000,1207,1229]" county="Clark Co." name="Cave Spring" pageId="2" pageNumber="115" stateProvince="Washington">Cave Spring</location>
,
<locationDeviation id="9A576C79FFB9552AFF4A6E9DFE3E5493" box="[239,412,1234,1256]" location="New" pageId="2" pageNumber="115">
<quantity id="4CA2D8BEFFB9552AFF4A6E9DFE8D5493" box="[239,303,1234,1256]" metricMagnitude="3" metricUnit="m" metricValue="4.02336" pageId="2" pageNumber="115" unit="mi" value="2.5">2.5 mi</quantity>
SSW New
</locationDeviation>
<collectingRegion id="499EBBB9FFB9552AFE076E9DFD835493" box="[418,545,1234,1256]" country="United States of America" name="Washington" pageId="2" pageNumber="115">Washington</collectingRegion>
,
<collectingCounty id="62840DD7FFB9552AFD8F6E9DFD335493" box="[554,657,1234,1256]" pageId="2" pageNumber="115">Clark Co.</collectingCounty>
,
<collectingRegion id="499EBBB9FFB9552AFD3E6E9DFD485493" box="[667,746,1234,1256]" country="United States of America" name="Indiana" pageId="2" pageNumber="115">Indiana</collectingRegion>
(FMNH). Female.
<emphasis id="B92EA949FFB9552AFC176E9DFC4E5493" bold="true" box="[946,1004,1234,1256]" pageId="2" pageNumber="115">NEW</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE5755BFFB9552AFF4A6EA3FE235579" blockId="2.[175,1005,1153,1282]" box="[239,385,1260,1282]" pageId="2" pageNumber="115">
<emphasis id="B92EA949FFB9552AFF4A6EA3FE235579" bold="true" box="[239,385,1260,1282]" pageId="2" pageNumber="115">SYNONYMY</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34026D0FFB9552BFF626F52FDA253B5" lastPageId="3" lastPageNumber="116" pageId="2" pageNumber="115" type="description">
<paragraph id="8BE5755BFFB9552AFF626F52FC4F5659" blockId="2.[175,1005,1309,1570]" pageId="2" pageNumber="115">
<emphasis id="B92EA949FFB9552AFF626F52FE065548" bold="true" box="[199,420,1309,1331]" pageId="2" pageNumber="115">Specimens examined.</emphasis>
<materialsCitation id="3B327F06FFB9552AFE166F52FC4E5548" box="[435,1004,1309,1331]" collectionCode="Synonymy" county="Clark Co." location="Cave Spring" pageId="2" pageNumber="115" specimenCount="1" stateProvince="Indiana">
<collectingRegion id="499EBBB9FFB9552AFE166F52FDA75548" box="[435,517,1309,1331]" country="United States of America" name="Indiana" pageId="2" pageNumber="115">Indiana</collectingRegion>
:
<emphasis id="B92EA949FFB9552AFDB36F52FD205548" bold="true" box="[534,642,1309,1331]" pageId="2" pageNumber="115">
<collectingCounty id="62840DD7FFB9552AFDB36F52FD205548" box="[534,642,1309,1331]" pageId="2" pageNumber="115">Clark Co.</collectingCounty>
</emphasis>
,
<location id="8E852380FFB9552AFD336F52FCBE5548" LSID="urn:lsid:plazi:treatment:03F3C44DFFB9552CFEFB6E75FC2B5229:8E852380FFB9552AFD336F52FCBE5548" box="[662,796,1309,1331]" county="Clark Co." name="Cave Spring" pageId="2" pageNumber="115" stateProvince="Indiana">Cave Spring</location>
,
<locationDeviation id="9A576C79FFB9552AFC8A6F52FC4E5548" box="[815,1004,1309,1331]" location="New" pageId="2" pageNumber="115">
<quantity id="4CA2D8BEFFB9552AFC8A6F52FCCD5548" box="[815,879,1309,1331]" metricMagnitude="3" metricUnit="m" metricValue="4.02336" pageId="2" pageNumber="115" unit="mi" value="2.5">2.5 mi</quantity>
SSW New
</locationDeviation>
</materialsCitation>
<materialsCitation id="3B327F06FFB9552AFF0A6F77FC4B5659" collectingDate="1947-05-01" collectingDateMax="2006-08-26" collectingDateMin="1947-05-01" collectionCode="FMNH" collectorName="C. H. Krekeler &amp; N. G. Hairston &amp; Julian Lewis &amp; James Lewis &amp; Salisa Lewis &amp; Cave &amp; Pierre Paquin &amp; Salisa Rafail" county="Crawford Co." location="Hoton Canyon Cave" municipality="Clark State Forest" pageId="2" pageNumber="115" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Washington" typeStatus="holotype">
<collectingRegion id="499EBBB9FFB9552AFF0A6F77FE8F5535" box="[175,301,1336,1358]" country="United States of America" name="Washington" pageId="2" pageNumber="115">Washington</collectingRegion>
,
<date id="FFE4539BFFB9552AFE9F6F77FE165535" box="[314,436,1336,1358]" pageId="2" pageNumber="115" value="1959-08-03">
<collectingDate id="EFA0AA73FFB9552AFE9F6F77FE165535" box="[314,436,1336,1358]" pageId="2" pageNumber="115" value="1959-08-03">VIII-3-1959</collectingDate>
</date>
,
<collectorName id="26AF108DFFB9552AFE646F77FDF15535" box="[449,595,1336,1358]" pageId="2" pageNumber="115">C.H. Krekeler</collectorName>
(
<collectionCode id="ED4BED9EFFB9552AFDC16F77FD145535" box="[612,694,1336,1358]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="115" type="Museum">FMNH</collectionCode>
,
<typeStatus id="54E1CBF9FFB9552AFD666F77FCBE5535" box="[707,796,1336,1358]" pageId="2" pageNumber="115" type="holotype">holotype</typeStatus>
<taxonomicName id="4C5A0ED8FFB9552AFC806F77FC395536" authorityName="Park" authorityYear="1960" box="[805,923,1336,1357]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="115" phylum="Arthropoda" rank="species" species="krekeleri">
<emphasis id="B92EA949FFB9552AFC806F77FC395536" box="[805,923,1336,1357]" italics="true" pageId="2" pageNumber="115">B. krekeleri</emphasis>
</taxonomicName>
);
<collectingMunicipality id="6B81EF21FFB9552AFC166F77FE8E5513" pageId="2" pageNumber="115">Clark State Forest</collectingMunicipality>
, near Uno [Union],
<date id="FFE4539BFFB9552AFE506F1DFDF15513" box="[501,595,1362,1384]" pageId="2" pageNumber="115" value="1947-05-01">
<collectingDate id="EFA0AA73FFB9552AFE506F1DFDF15513" box="[501,595,1362,1384]" pageId="2" pageNumber="115" value="1947-05-01">V-1-1947</collectingDate>
</date>
,
<collectorName id="26AF108DFFB9552AFDF96F1DFD515513" box="[604,755,1362,1384]" pageId="2" pageNumber="115">N.G. Hairston</collectorName>
, ex stomach
<emphasis id="B92EA949FFB9552AFCD76F1CFC4F5513" box="[882,1005,1363,1384]" italics="true" pageId="2" pageNumber="115">
<taxonomicName id="4C5A0ED8FFB9552AFCD76F1CFC745513" authorityName="Tschudi" authorityYear="1838" box="[882,982,1363,1384]" class="Amphibia" family="Plethodontidae" genus="Plethodon" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="115" phylum="Chordata" rank="genus">Plethodon</taxonomicName>
c.
</emphasis>
<taxonomicName id="4C5A0ED8FFB9552AFF0A6F22FEF555F8" authority="(Green)" baseAuthorityName="Green" baseAuthorityYear="1818" box="[175,343,1389,1411]" class="Amphibia" family="Plethodontidae" genus="Plethodon" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="115" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="B92EA949FFB9552AFF0A6F22FF5D55F9" box="[175,255,1389,1410]" italics="true" pageId="2" pageNumber="115">cinereus</emphasis>
(Green)
</taxonomicName>
(
<collectionCode id="ED4BED9EFFB9552AFECD6F22FE1855F8" box="[360,442,1389,1411]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="115" type="Museum">FMNH</collectionCode>
,
<typeStatus id="54E1CBF9FFB9552AFE626F22FD8255F8" box="[455,544,1389,1411]" pageId="2" pageNumber="115" type="holotype">holotype</typeStatus>
<taxonomicName id="4C5A0ED8FFB9552AFD8C6F22FD3D55F9" authorityName="Park" authorityYear="1947" box="[553,671,1389,1410]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="115" phylum="Arthropoda" rank="species" species="hairstoni">
<emphasis id="B92EA949FFB9552AFD8C6F22FD3D55F9" box="[553,671,1389,1410]" italics="true" pageId="2" pageNumber="115">B. hairstoni</emphasis>
</taxonomicName>
).
<emphasis id="B92EA949FFB9552AFD1D6F22FCE655F8" bold="true" box="[696,836,1389,1411]" pageId="2" pageNumber="115">
<collectingCounty id="62840DD7FFB9552AFD1D6F22FCE655F8" box="[696,836,1389,1411]" pageId="2" pageNumber="115">Crawford Co.</collectingCounty>
</emphasis>
,
<specimenCount id="9D5CBED2FFB9552AFCF66F22FC3E55F8" box="[851,924,1389,1411]" pageId="2" pageNumber="115" type="male">1 male</specimenCount>
,
<location id="8E852380FFB9552AFC0C6F22FE9D55E6" LSID="urn:lsid:plazi:treatment:03F3C44DFFB9552CFEFB6E75FC2B5229:8E852380FFB9552AFC0C6F22FE9D55E6" county="Crawford Co." municipality="Clark State Forest" name="Hoton Canyon Cave" pageId="2" pageNumber="115" stateProvince="Washington">Hoton Canyon Cave</location>
,
<locationDeviation id="9A576C79FFB9552AFEE86FC8FD8C55E6" box="[333,558,1415,1437]" location="Marengo" pageId="2" pageNumber="115">
<quantity id="4CA2D8BEFFB9552AFEE86FC8FE2F55E6" box="[333,397,1415,1437]" metricMagnitude="3" metricUnit="m" metricValue="4.02336" pageId="2" pageNumber="115" unit="mi" value="2.5">2.5 mi</quantity>
ESE Marengo
</locationDeviation>
,
<date id="FFE4539BFFB9552AFD9E6FC8FD0555E6" box="[571,679,1415,1437]" pageId="2" pageNumber="115" value="1996-10-26">
<collectingDate id="EFA0AA73FFB9552AFD9E6FC8FD0555E6" box="[571,679,1415,1437]" pageId="2" pageNumber="115" value="1996-10-26">X-26-1996</collectingDate>
</date>
,
<collectorName id="26AF108DFFB9552AFD106FC8FC9855E6" box="[693,826,1415,1437]" pageId="2" pageNumber="115">Julian Lewis</collectorName>
,
<collectorName id="26AF108DFFB9552AFCE26FC8FC6F55E6" box="[839,973,1415,1437]" pageId="2" pageNumber="115">James Lewis</collectorName>
, &amp;
<location id="8E852380FFB9552AFF0A6FEDFE0955C3" LSID="urn:lsid:plazi:treatment:03F3C44DFFB9552CFEFB6E75FC2B5229:8E852380FFB9552AFF0A6FEDFE0955C3" box="[175,427,1442,1464]" county="Crawford Co." municipality="Clark State Forest" name="Victor Lewis. Greene Co." pageId="2" pageNumber="115" stateProvince="Washington">
Victor Lewis.
<emphasis id="B92EA949FFB9552AFE9E6FEDFE0955C3" bold="true" box="[315,427,1442,1464]" pageId="2" pageNumber="115">Greene Co.</emphasis>
</location>
,
<specimenCount id="9D5CBED2FFB9552AFE136FEDFDAF55C3" box="[438,525,1442,1464]" pageId="2" pageNumber="115" type="female">1 female</specimenCount>
,
<location id="8E852380FFB9552AFDB36FEDFD0B55C3" LSID="urn:lsid:plazi:treatment:03F3C44DFFB9552CFEFB6E75FC2B5229:8E852380FFB9552AFDB36FEDFD0B55C3" box="[534,681,1442,1464]" county="Crawford Co." municipality="Clark State Forest" name="Ashcraft Cave" pageId="2" pageNumber="115" stateProvince="Washington">Ashcraft Cave</location>
,
<locationDeviation id="9A576C79FFB9552AFD176FEDFCFB55C3" box="[690,857,1442,1464]" location="Koleen" pageId="2" pageNumber="115">
<quantity id="4CA2D8BEFFB9552AFD176FEDFD4255C3" box="[690,736,1442,1464]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="2" pageNumber="115" unit="mi" value="2.0">2 mi</quantity>
NE Koleen
</locationDeviation>
,
<date id="FFE4539BFFB9552AFCC66FEDFC4B55C3" box="[867,1001,1442,1464]" pageId="2" pageNumber="115" value="2006-08-20">
<collectingDate id="EFA0AA73FFB9552AFCC66FEDFC4B55C3" box="[867,1001,1442,1464]" pageId="2" pageNumber="115" value="2006-08-20">VIII-20-2006</collectingDate>
</date>
,
<collectorName id="26AF108DFFB9552AFF0A6FF2FE8F55A8" box="[175,301,1469,1491]" pageId="2" pageNumber="115">Julian Lewis</collectorName>
&amp;
<collectorName id="26AF108DFFB9552AFEEC6FF2FE6555A8" box="[329,455,1469,1491]" pageId="2" pageNumber="115">Salisa Lewis</collectorName>
, entrance room under stone.
<emphasis id="B92EA949FFB9552AFD4D6FF2FCD255A8" bold="true" box="[744,880,1469,1491]" pageId="2" pageNumber="115">
<location id="8E852380FFB9552AFD4D6FF2FCD255A8" LSID="urn:lsid:plazi:treatment:03F3C44DFFB9552CFEFB6E75FC2B5229:8E852380FFB9552AFD4D6FF2FCD255A8" box="[744,880,1469,1491]" county="Crawford Co." municipality="Clark State Forest" name="Lawrence Co." pageId="2" pageNumber="115" stateProvince="Washington">Lawrence Co.</location>
</emphasis>
,
<specimenCount id="9D5CBED2FFB9552AFCDE6FF2FC1D55A8" box="[891,959,1469,1491]" pageId="2" pageNumber="115" type="male">1 male</specimenCount>
, JJ’s
<collectorName id="26AF108DFFB9552AFF0A6F98FF475596" box="[175,229,1495,1517]" pageId="2" pageNumber="115">Cave</collectorName>
,
<date id="FFE4539BFFB9552AFF546F98FED55596" box="[241,375,1495,1517]" pageId="2" pageNumber="115" value="2006-08-26">
<collectingDate id="EFA0AA73FFB9552AFF546F98FED55596" box="[241,375,1495,1517]" pageId="2" pageNumber="115" value="2006-08-26">VIII-26-2006</collectingDate>
</date>
,
<collectorName id="26AF108DFFB9552AFE276F98FDAF5596" box="[386,525,1495,1517]" pageId="2" pageNumber="115">Pierre Paquin</collectorName>
&amp;
<collectorName id="26AF108DFFB9552AFD8A6F98FD115596" box="[559,691,1495,1517]" pageId="2" pageNumber="115">Julian Lewis</collectorName>
;
<specimenCount id="9D5CBED2FFB9552AFD1B6F98FCB45596" box="[702,790,1495,1517]" pageId="2" pageNumber="115" type="female">1 female</specimenCount>
,
<location id="8E852380FFB9552AFC876F98FC4B5596" LSID="urn:lsid:plazi:treatment:03F3C44DFFB9552CFEFB6E75FC2B5229:8E852380FFB9552AFC876F98FC4B5596" box="[802,1001,1495,1517]" county="Crawford Co." municipality="Clark State Forest" name="Smith's Folly Cave" pageId="2" pageNumber="115" stateProvince="Washington">Smith’s Folly Cave</location>
,
<location id="8E852380FFB9552AFF0A6FBDFE755673" LSID="urn:lsid:plazi:treatment:03F3C44DFFB9552CFEFB6E75FC2B5229:8E852380FFB9552AFF0A6FBDFE755673" box="[175,471,1522,1544]" county="Crawford Co." municipality="Clark State Forest" name="Tincher Hollow Special Area" pageId="2" pageNumber="115" stateProvince="Washington">Tincher Hollow Special Area</location>
,
<location id="8E852380FFB9552AFE476FBDFD7F5673" LSID="urn:lsid:plazi:treatment:03F3C44DFFB9552CFEFB6E75FC2B5229:8E852380FFB9552AFE476FBDFD7F5673" box="[482,733,1522,1544]" county="Crawford Co." municipality="Clark State Forest" name="Hoosier National Forest" pageId="2" pageNumber="115" stateProvince="Washington">Hoosier National Forest</location>
,
<date id="FFE4539BFFB9552AFD426FBDFCC65673" box="[743,868,1522,1544]" pageId="2" pageNumber="115" value="2001-07-29">
<collectingDate id="EFA0AA73FFB9552AFD426FBDFCC65673" box="[743,868,1522,1544]" pageId="2" pageNumber="115" value="2001-07-29">VII-29-2001</collectingDate>
</date>
,
<collectorName id="26AF108DFFB9552AFCCB6FBDFC4E5673" box="[878,1004,1522,1544]" pageId="2" pageNumber="115">Julian Lewis</collectorName>
&amp;
<collectorName id="26AF108DFFB9552AFF626C43FEEB5659" box="[199,329,1548,1570]" pageId="2" pageNumber="115">Salisa Rafail</collectorName>
;
<specimenCount id="9D5CBED2FFB9552AFEF46C43FE3C5659" box="[337,414,1548,1570]" pageId="2" pageNumber="115" type="male">2 males</specimenCount>
, same locality,
<date id="FFE4539BFFB9552AFD906C43FD185659" box="[565,698,1548,1570]" pageId="2" pageNumber="115" value="2006-08-26">
<collectingDate id="EFA0AA73FFB9552AFD906C43FD185659" box="[565,698,1548,1570]" pageId="2" pageNumber="115" value="2006-08-26">VIII-26-2006</collectingDate>
</date>
,
<collectorName id="26AF108DFFB9552AFD666C43FCEE5659" box="[707,844,1548,1570]" pageId="2" pageNumber="115">Pierre Paquin</collectorName>
&amp;
<collectorName id="26AF108DFFB9552AFCCC6C43FC4B5659" box="[873,1001,1548,1570]" pageId="2" pageNumber="115">Julian Lewis</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE5755BFFB8552BFF0F6A8DFEDD5101" blockId="3.[146,976,194,1570]" pageId="3" pageNumber="116">
<emphasis id="B92EA949FFB8552BFF0F6A8DFE8150A3" bold="true" box="[170,291,194,216]" pageId="3" pageNumber="116">Description.</emphasis>
Length 2.3–2.5. Head with nude vertexal foveae; lateral occipital carinae extending anteriorly to point dorsal to anterior margin of eyes. Pronotum with antebasal sulcus biarcuate, antebasal foveae large, subequal, lateral antebasal foveae setose, median antebasal fovea nude; lateral longitudinal sulci distinct, median longitudinal sulcus weakly extending anteriorly to near middle of disc, disc convex; lateral antebasal tubercles low, bluntly pointed. Elytra with three antebasal foveae.
</paragraph>
<paragraph id="8BE5755BFFB8552BFF0F6B30FCC452A2" blockId="3.[146,976,194,1570]" pageId="3" pageNumber="116">
<emphasis id="B92EA949FFB8552BFF0F6B30FF4751EE" bold="true" box="[170,229,383,405]" pageId="3" pageNumber="116">Male.</emphasis>
Head with short broad sulci extending anteriorly from vertexal foveae; vertex prominently projecting on anterior margin between antennal bases, apex broadly and shallowly curved, sometimes appearing slightly sinuate laterally, apical portion slightly curving ventrally, densely punctate, with short anteriorlycurved setae; deep transverse sulcus between antennal insertions sharply defining underside of projection, sulcus with band of short dense setae between antennal insertions originating from vertexal overhang and clypeus; clypeus broadly projecting and truncate at border with sulcus. Eyes with about 45 facets. First antennomere unmodified, antennomeres III–VII subequal in length, nearly half again as long as wide, VIII as long as wide, IX transversely triangular, twice as wide as long, widest laterally, X enlarged and globular, as long as wide, nearly as wide as XI, flattened on ventral margin, with large setose fovea occupying about half of ventral surface; XI with ventral surface flattened in basal half.
</paragraph>
<paragraph id="8BE5755BFFB8552BFF0F6890FD2F533D" blockId="3.[146,976,194,1570]" pageId="3" pageNumber="116">Protibiae with apical half flattened, slightly widening to apex, with dense short setae on mesal margin; mesotibiae broadly angulate in apical half, densely setose from angulation to apex; second mesotarsomere sinuate; metatibiae simple, with dense short setae on mesal margin in apical half.</paragraph>
<paragraph id="8BE5755BFFB8552BFF0F6904FC72531A" blockId="3.[146,976,194,1570]" box="[170,976,843,865]" pageId="3" pageNumber="116">Fourth visible sternite broadly and slightly convex; fifth sternite flattened at middle.</paragraph>
<paragraph id="8BE5755BFFB8552BFF0F6928FDA253B5" blockId="3.[146,976,194,1570]" pageId="3" pageNumber="116">
<emphasis id="B92EA949FFB8552BFF0F6928FF5B5306" bold="true" box="[170,249,871,893]" pageId="3" pageNumber="116">Female.</emphasis>
Head with junction of vertex and frons distinctly angled between antennal bases, frons flat, more densely punctate than vertex, antennal tubercles equally densely punctate. Eyes with about 20 facets (
<bibRefCitation id="EFCB08AAFFB8552BFD6E69D2FCE553C8" author="Park" box="[715,839,925,947]" pageId="3" pageNumber="116" pagination="66 - 104" refId="ref3481" refString="Park, O. 1960. Cavernicolous pselaphid beetles of the United States. American Midland Naturalist 64 (1): 66 - 104." type="journal article" year="1960">Park (1960)</bibRefCitation>
observed 12 facets, but 20 is more appropriate).
</paragraph>
</subSubSection>
<subSubSection id="C34026D0FFB8552CFF0F699BFC2B5229" lastPageId="4" lastPageNumber="117" pageId="3" pageNumber="116" type="distribution">
<paragraph id="8BE5755BFFB8552BFF0F699BFD2E54F7" blockId="3.[146,976,194,1570]" pageId="3" pageNumber="116">
<emphasis id="B92EA949FFB8552BFF0F699BFE0C5391" bold="true" box="[170,430,980,1002]" pageId="3" pageNumber="116">Distribution and Ecology.</emphasis>
Specimens were found on the underside of rocks in caves from late July to October, with the specimen obtained from the salamander stomach found in May. In Hoton Canyon Cave the specimens were found under a stone far into the dark zone of the cave, while in the other caves the beetles were found in the twilight zone. Prolonged, repeated searching in the same part of Ashcraft Cave failed to secure a second specimen. The rocky slope outside the cave entrance was also searched without success.
</paragraph>
<paragraph id="8BE5755BFFB8552BFF0F6EDEFC1C551F" blockId="3.[146,976,194,1570]" pageId="3" pageNumber="116">
The range spans a linear distance of approximately
<quantity id="4CA2D8BEFFB8552BFD6A6EDEFCBC54DC" box="[719,798,1169,1191]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="116" unit="km" value="100.0">100 km</quantity>
, from Clark Co. west to Crawford Co., north to Lawrence and Greene Counties. Cave Spring Cave (Clark Co.) was formed in limestone of the southeastern
<collectingRegion id="499EBBB9FFB8552BFCB06E88FCC654A6" box="[789,868,1223,1245]" country="United States of America" name="Indiana" pageId="3" pageNumber="116">Indiana</collectingRegion>
karst that was covered by the Illinoian glaciation. The other three caves lie in the unglaciated south-central
<collectingRegion id="499EBBB9FFB8552BFE386EB2FE4E5568" box="[413,492,1277,1299]" country="United States of America" name="Indiana" pageId="3" pageNumber="116">Indiana</collectingRegion>
karst region. The only known epigean locality of
<taxonomicName id="4C5A0ED8FFB8552BFF106F57FE935556" authorityName="Park" authorityYear="1947" box="[181,305,1304,1325]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="hairstoni">
<emphasis id="B92EA949FFB8552BFF106F57FE935556" box="[181,305,1304,1325]" italics="true" pageId="3" pageNumber="116">B. hairstoni</emphasis>
</taxonomicName>
is from a non-karst area of rugged hill topography in the intervening Scottsburg Lowland, a region separating the two karst belts. The zoogeography of
<collectingRegion id="499EBBB9FFB8552BFEE26F01FE34551F" box="[327,406,1358,1380]" country="United States of America" name="Indiana" pageId="3" pageNumber="116">Indiana</collectingRegion>
cavernicoles was discussed in detail by
<bibRefCitation id="EFCB08AAFFB8552BFC916F01FC18551F" author="Lewis" box="[820,954,1358,1380]" pageId="3" pageNumber="116" pagination="34 - 40" refId="ref3375" refString="Lewis, J. J. 1983. The obligatory subterranean invertebrates of glaciated southeastern Indiana. National Speleological Society Bulletin 45: 34 - 40." type="journal article" year="1983">Lewis (1983)</bibRefCitation>
.
</paragraph>
<paragraph id="8BE5755BFFB8552BFF0F6F25FCE95597" blockId="3.[146,976,194,1570]" pageId="3" pageNumber="116">The eyes of the male are somewhat reduced (circa 45 facets), and those of the female distinctly so (circa 20 facets), indicating an adaptation to caves or possibly deep soil habitats. One specimen taken in August is teneral, and another is not fully sclerotized. Flight wings of the male are fully developed, while those of the female are slightly shortened, extending only to the abdomen apex.</paragraph>
<paragraph id="8BE5755BFFB8552CFF0F6FBEFC2C51EE" blockId="3.[146,976,194,1570]" lastBlockId="4.[175,1004,194,594]" lastPageId="4" lastPageNumber="117" pageId="3" pageNumber="116">
<emphasis id="B92EA949FFB8552BFF0F6FBEFE94567C" bold="true" box="[170,310,1521,1543]" pageId="3" pageNumber="116">Relationships.</emphasis>
Superficially similar to the locally common
<taxonomicName id="4C5A0ED8FFB8552BFCA96FBDFC3F567C" baseAuthorityName="Aube" baseAuthorityYear="1833" box="[780,925,1522,1543]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="lineaticollis">
<emphasis id="B92EA949FFB8552BFCA96FBDFC3F567C" box="[780,925,1522,1543]" italics="true" pageId="3" pageNumber="116">B. lineaticollis</emphasis>
</taxonomicName>
, but separable by the weak median longitudinal sulcus that barely reaches the middle of the pronotal disc in
<taxonomicName id="4C5A0ED8FFBF552CFE056A8DFDB450AC" authorityName="Park" authorityYear="1947" box="[416,534,194,215]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="species" species="hairstoni">
<emphasis id="B92EA949FFBF552CFE056A8DFDB450AC" box="[416,534,194,215]" italics="true" pageId="4" pageNumber="117">B. hairstoni</emphasis>
</taxonomicName>
, while in
<taxonomicName id="4C5A0ED8FFBF552CFDDA6A8DFCAC50AC" baseAuthorityName="Aube" baseAuthorityYear="1833" box="[639,782,194,215]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="species" species="lineaticollis">
<emphasis id="B92EA949FFBF552CFDDA6A8DFCAC50AC" box="[639,782,194,215]" italics="true" pageId="4" pageNumber="117">B. lineaticollis</emphasis>
</taxonomicName>
this sulcus is distinct and reaches the pronotal apex. Males of
<taxonomicName id="4C5A0ED8FFBF552CFDF96A92FD735089" authorityName="Park" authorityYear="1947" box="[604,721,221,242]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="species" species="hairstoni">
<emphasis id="B92EA949FFBF552CFDF96A92FD735089" box="[604,721,221,242]" italics="true" pageId="4" pageNumber="117">B. hairstoni</emphasis>
</taxonomicName>
are easily separable by the broadly rounded vertexal projection, antennomeres X and XI being equally wide, the large fovea on the underside of antennomere X, and the sinuate second mesotarsomere, the definitive character for the subgenus
<taxonomicName id="4C5A0ED8FFBF552CFD546B61FCDA5138" baseAuthorityName="Park" baseAuthorityYear="1956" box="[753,888,302,323]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="subGenus" subGenus="Babnormodes">
<emphasis id="B92EA949FFBF552CFD546B61FCDA5138" box="[753,888,302,323]" italics="true" pageId="4" pageNumber="117">Babnormodes</emphasis>
</taxonomicName>
.
<taxonomicName id="4C5A0ED8FFBF552CFC216B61FE825125" baseAuthorityName="Aube" baseAuthorityYear="1833" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="species" species="lineaticollis">
<emphasis id="B92EA949FFBF552CFC216B61FE825125" italics="true" pageId="4" pageNumber="117">Batrisodes lineaticollis</emphasis>
</taxonomicName>
males are characterized by a bluntly angular vertexal projection, antennomere X is clearly wider than XI, the fovea on the underside of antennomere X is punctiform, and the second mesotarsomere is linear.
</paragraph>
<paragraph id="8BE5755BFFBF552CFF626BD5FC2B5229" blockId="4.[175,1004,194,594]" pageId="4" pageNumber="117">
This species appears to be closest to the widespread eastern
<taxonomicName id="4C5A0ED8FFBF552CFC216BD4FDF751B7" authority="(Casey, 1887)" baseAuthorityName="Casey" baseAuthorityYear="1887" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="species" species="foveicornis" subGenus="Babnormodes">
<emphasis id="B92EA949FFBF552CFC216BD4FC4E51CB" box="[900,1004,411,432]" italics="true" pageId="4" pageNumber="117">Batrisodes</emphasis>
(
<emphasis id="B92EA949FFBF552CFF126BF9FE9C51B0" box="[183,318,438,459]" italics="true" pageId="4" pageNumber="117">Babnormodes</emphasis>
)
<emphasis id="B92EA949FFBF552CFEF46BF9FE1951B0" box="[337,443,438,459]" italics="true" pageId="4" pageNumber="117">foveicornis</emphasis>
(Casey, 1887)
</taxonomicName>
by the similar antennal and pronotal characters of the males. In
<taxonomicName id="4C5A0ED8FFBF552CFE686B9EFDF4519D" baseAuthorityName="Casey" baseAuthorityYear="1887" box="[461,598,465,486]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="species" species="foveicornis">
<emphasis id="B92EA949FFBF552CFE686B9EFDF4519D" box="[461,598,465,486]" italics="true" pageId="4" pageNumber="117">B. foveicornis</emphasis>
</taxonomicName>
the vertexal apex smoothly declines to the frons with a slight transverse sulcus indicated at the junction of the frons and vertex, in contrast to the prominent and well-defined vertexal apex with a deep transverse indentation at the junction with the frons in
<taxonomicName id="4C5A0ED8FFBF552CFCB5686DFC2B524C" authorityName="Park" authorityYear="1947" box="[784,905,546,567]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="species" species="hairstoni">
<emphasis id="B92EA949FFBF552CFCB5686DFC2B524C" box="[784,905,546,567]" italics="true" pageId="4" pageNumber="117">B. hairstoni</emphasis>
</taxonomicName>
, and the mesotibiae lack the mesal angulation that is prominent in
<taxonomicName id="4C5A0ED8FFBF552CFCA96872FC215229" authorityName="Park" authorityYear="1960" box="[780,899,573,594]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="species" species="krekeleri">
<emphasis id="B92EA949FFBF552CFCA96872FC215229" box="[780,899,573,594]" italics="true" pageId="4" pageNumber="117">B. krekeleri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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