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<document id="09806C23BE6F6ABD69D8AA9B485C2EC0" ID-DOI="10.5281/zenodo.190657" ID-GBIF-Dataset="025e2ef4-42ca-47e0-b746-80c76aeb03fe" ID-ISSN="1175-5326" ID-Zenodo-Dep="190657" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461115598214" checkinUser="plazi" docAuthor="Lindquist, Evert E. &amp; Moraza, María L." docDate="2009" docId="03D10A69D445FFD2FF1AA64C01C2A4CE" docLanguage="en" docName="zt02243p039.pdf" docOrigin="Zootaxa 2243" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Anystipalpus labiduricola Lindquist &amp; Moraza, 2009, n. sp." docType="treatment" docVersion="8" lastPageNumber="27" masterDocId="FFE87211D453FFC8FF8DA21A034CA309" masterDocTitle="Anystipalpus, Antennoseius and Vitzthumia: a taxonomic and nomenclatural conundrum of genera (Acari: Mesostigmata: Dermanyssina), with description of four species of Anystipalpus" masterLastPageNumber="39" masterPageNumber="1" pageNumber="23" updateTime="1698595074968" updateUser="plazi">
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<mods:title id="F768A051E0DBA5E80C13D1AEC97D872A">Anystipalpus, Antennoseius and Vitzthumia: a taxonomic and nomenclatural conundrum of genera (Acari: Mesostigmata: Dermanyssina), with description of four species of Anystipalpus</mods:title>
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<mods:namePart id="108CB8AACA7B726123F01A340BDBDA7C">Lindquist, Evert E.</mods:namePart>
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<mods:namePart id="D3ADB7E6DC784BCE6A843B4B07D8CE09">Moraza, María L.</mods:namePart>
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<mods:date id="041679E88D058EF17E6AFE768ED8DA54">2009</mods:date>
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<mods:number id="2C310CE405885733EFEF4407E149D307">2243</mods:number>
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<treatment id="03D10A69D445FFD2FF1AA64C01C2A4CE" ID-DOI="http://doi.org/10.5281/zenodo.3502404" ID-GBIF-Taxon="119624781" ID-Zenodo-Dep="3502404" LSID="urn:lsid:plazi:treatment:03D10A69D445FFD2FF1AA64C01C2A4CE" httpUri="http://treatment.plazi.org/id/03D10A69D445FFD2FF1AA64C01C2A4CE" lastPageId="26" lastPageNumber="27" pageId="22" pageNumber="23">
<subSubSection id="C362E8F4D445FFDEFF1AA64C0261A79A" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8BC7BB7FD445FFDEFF1AA64C015BA779" blockId="22.[151,535,1110,1171]" box="[151,535,1110,1136]" pageId="22" pageNumber="23">
<heading id="D08F0C13D445FFDEFF1AA64C015BA779" bold="true" box="[151,535,1110,1136]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="B90C676DD445FFDEFF1AA64C015BA779" bold="true" box="[151,535,1110,1136]" pageId="22" pageNumber="23">
<taxonomicName id="4C78C0FCD445FFDEFF1AA64C0281A779" box="[151,461,1110,1136]" class="Arachnida" family="Laelapidae" genus="Anystipalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="labiduricola" status="sp. nov.">
<emphasis id="B90C676DD445FFDEFF1AA64C0281A779" bold="true" box="[151,461,1110,1136]" italics="true" pageId="22" pageNumber="23">Anystipalpus labiduricola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23FDA16D445FFDEFE59A64C015BA779" box="[468,535,1110,1136]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC7BB7FD445FFDEFF1AA6630261A79A" blockId="22.[151,535,1110,1171]" box="[151,301,1145,1171]" pageId="22" pageNumber="23">
(
<figureCitation id="1343A7FAD445FFDEFF12A6630269A79A" box="[159,293,1145,1171]" captionStart-0="FIGURES 29 34" captionStart-1="FIGURES 35 38" captionStart-2="FIGURES 39 42" captionStartId-0="24.[151,272,1750,1774]" captionStartId-1="25.[151,270,1908,1932]" captionStartId-2="26.[151,272,1440,1464]" captionTargetBox-0="[160,1436,187,1721]" captionTargetBox-1="[197,1409,462,1859]" captionTargetBox-2="[160,1431,194,1369]" captionTargetId-0="figure@24.[156,1441,187,1726]" captionTargetId-1="figure@25.[193,1415,455,1880]" captionTargetId-2="figure@26.[156,1441,194,1416]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="26" captionText-0="FIGURES 29 34. Anystipalpus labiduricola n. sp., adult female: 29, idiosoma, dorsal aspect; 30, enlarged separate views of setae denoted; 31, anterior margin and dorsal face of tectum; 32, cheliceral digits, dorsal view, showing dorsal antiaxial projection; 33, separate ventral view of movable digit, showing vestige of ventral process; 34, subcapitulum." captionText-1="FIGURES 35 38. Anystipalpus labiduricola n. sp., adult female: 35, idiosoma, ventral aspect; 36, palptarsus, enlarged view; 37, palpus, dorsal view; 38, leg I (excepting tarsus) posterolateral view." captionText-2="FIGURES 39 42. Figs 39 40, Anystipalpus labiduricola n. sp., adult female: 39, tarsus I, dorsal view; 40, tarsus I, detail of distal sensorial region. Figs 41 42, Anystipalpus kazemii n. sp., adult female: 41 a, palpus excepting tarsus, dorsal view; 41 b, palptarsus, dorsal view; 42 a, leg I excluding coxa and tarsus, dorsolateral view; 42 b, tarsus I, dorsal view." httpUri-0="https://zenodo.org/record/190664/files/figure.png" httpUri-1="https://zenodo.org/record/190665/files/figure.png" httpUri-2="https://zenodo.org/record/190666/files/figure.png" pageId="22" pageNumber="23">Figs 2940</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C362E8F4D445FFDEFF1AA6DC06D0A6AB" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8BC7BB7FD445FFDEFF1AA6DC06D0A6AB" blockId="22.[151,1437,1222,2022]" pageId="22" pageNumber="23">
<emphasis id="B90C676DD445FFDEFF1AA6DC02AFA7D6" bold="true" box="[151,483,1222,1248]" pageId="22" pageNumber="23">Diagnosis. ADULT FEMALE</emphasis>
. Dorsal shield setae collectively similar in slightly carinated, smooth or slightly barbed, pointed form and length (about 0.7 to 0.8 as long as longitudinal intervals between their bases). Epigynal shield drop-shaped, with lateral margins constricted anterior to genital setae and posterior margin bluntly pointed. Peritrematal-exopodal shield with four or five lines radiating from stigma to broadly scalloped posterior margin. Coxa I with one seta (
<emphasis id="B90C676DD445FFDEFD51A77B01B4A673" box="[732,760,1377,1402]" italics="true" pageId="22" pageNumber="23">av</emphasis>
) bluntly spinelike. Tarsus II lacking spinelike setae. Leg I with trochanter seta
<emphasis id="B90C676DD445FFDEFE08A79202D8A6A8" box="[389,404,1416,1441]" italics="true" pageId="22" pageNumber="23">d</emphasis>
and femoral seta
<emphasis id="B90C676DD445FFDEFDE9A79201DCA6A8" box="[612,656,1416,1441]" italics="true" pageId="22" pageNumber="23">pd2</emphasis>
barbed, slightly stout, pointed; other setae not markedly spinelike.
</paragraph>
</subSubSection>
<subSubSection id="C362E8F4D445FFDFFF4BA7B40184A45E" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
<paragraph id="8BC7BB7FD445FFDEFF4BA7B40395A47B" blockId="22.[151,1437,1222,2022]" pageId="22" pageNumber="23">
<emphasis id="B90C676DD445FFDEFF4BA7B4017BA6C1" bold="true" box="[198,567,1454,1480]" pageId="22" pageNumber="23">Description. ADULT FEMALE.</emphasis>
Dorsal shields together 358360 long, reticulated over entire surfaces, except posteromedial face between setae
<emphasis id="B90C676DD445FFDEFDF5A7CF01D8A6E7" box="[632,660,1493,1518]" italics="true" pageId="22" pageNumber="23">J5</emphasis>
and
<emphasis id="B90C676DD445FFDEFD46A7CF01A6A6E7" box="[715,746,1493,1518]" italics="true" pageId="22" pageNumber="23">Z5</emphasis>
primarily longitudinally lineate (
<figureCitation id="1343A7FAD445FFDEFBE0A7CF0788A6E6" box="[1133,1220,1493,1519]" captionStart="FIGURES 29 34" captionStartId="24.[151,272,1750,1774]" captionTargetBox="[160,1436,187,1721]" captionTargetId="figure@24.[156,1441,187,1726]" captionTargetPageId="24" captionText="FIGURES 29 34. Anystipalpus labiduricola n. sp., adult female: 29, idiosoma, dorsal aspect; 30, enlarged separate views of setae denoted; 31, anterior margin and dorsal face of tectum; 32, cheliceral digits, dorsal view, showing dorsal antiaxial projection; 33, separate ventral view of movable digit, showing vestige of ventral process; 34, subcapitulum." httpUri="https://zenodo.org/record/190664/files/figure.png" pageId="22" pageNumber="23">Fig. 29</figureCitation>
). Podonotal shield 190200 long, 182187 wide at level of setae
<emphasis id="B90C676DD445FFDEFD23A7E601A0A51C" box="[686,748,1532,1558]" italics="true" pageId="22" pageNumber="23">r2-r3</emphasis>
, with 21 pairs of smooth or slightly barbed setae (
<emphasis id="B90C676DD445FFDEFABEA7E60627A51C" box="[1331,1387,1532,1558]" italics="true" pageId="22" pageNumber="23">j1-j6</emphasis>
,
<emphasis id="B90C676DD445FFDEFAF5A7E603FDA535" italics="true" pageId="22" pageNumber="23">z1- z6</emphasis>
,
<emphasis id="B90C676DD445FFDEFF4CA43903B3A535" box="[193,255,1570,1596]" italics="true" pageId="22" pageNumber="23">s1-s6</emphasis>
,
<emphasis id="B90C676DD445FFDEFE9DA4390266A535" box="[272,298,1571,1596]" italics="true" pageId="22" pageNumber="23">r2</emphasis>
and two extra pairs in
<emphasis id="B90C676DD445FFDEFDCDA4390132A535" box="[576,638,1570,1596]" italics="true" pageId="22" pageNumber="23">s3-s6</emphasis>
area), none conspicuously spinelike. Opisthonotal shield 158166 long, 165168 wide at level of setae
<emphasis id="B90C676DD445FFDEFDCFA4530113A56B" box="[578,607,1609,1634]" italics="true" pageId="22" pageNumber="23">S2</emphasis>
, evenly rounded posteriorly, with 15 pairs of smooth or slightly barbed setae (
<emphasis id="B90C676DD445FFDEFF6FA46A026EA580" box="[226,290,1648,1674]" italics="true" pageId="22" pageNumber="23">J1-J5</emphasis>
,
<emphasis id="B90C676DD445FFDEFEBCA46A0235A580" box="[305,377,1648,1674]" italics="true" pageId="22" pageNumber="23">Z1-Z5</emphasis>
,
<emphasis id="B90C676DD445FFDEFE0AA46A0287A580" box="[391,459,1648,1674]" italics="true" pageId="22" pageNumber="23">S1-S5</emphasis>
). Setae
<emphasis id="B90C676DD445FFDEFDA4A46A010CA580" box="[553,576,1648,1673]" italics="true" pageId="22" pageNumber="23">j1</emphasis>
(12) and
<emphasis id="B90C676DD445FFDEFD22A46A0184A580" box="[687,712,1648,1673]" italics="true" pageId="22" pageNumber="23">z1</emphasis>
(8) short, slightly barbed (
<figureCitation id="1343A7FAD445FFDEFC72A46A071BA583" box="[1023,1111,1648,1674]" captionStart="FIGURES 29 34" captionStartId="24.[151,272,1750,1774]" captionTargetBox="[160,1436,187,1721]" captionTargetId="figure@24.[156,1441,187,1726]" captionTargetPageId="24" captionText="FIGURES 29 34. Anystipalpus labiduricola n. sp., adult female: 29, idiosoma, dorsal aspect; 30, enlarged separate views of setae denoted; 31, anterior margin and dorsal face of tectum; 32, cheliceral digits, dorsal view, showing dorsal antiaxial projection; 33, separate ventral view of movable digit, showing vestige of ventral process; 34, subcapitulum." httpUri="https://zenodo.org/record/190664/files/figure.png" pageId="22" pageNumber="23">Fig. 30</figureCitation>
),
<emphasis id="B90C676DD445FFDEFBE6A46A07CDA580" box="[1131,1153,1648,1673]" italics="true" pageId="22" pageNumber="23">j1</emphasis>
slightly spinelike; other setae on dorsal shields (1925) slightly thickened, narrowly keeled, acutely pointed (
<figureCitation id="1343A7FAD445FFDEFBF8A48C0780A5B9" box="[1141,1228,1686,1712]" captionStart="FIGURES 29 34" captionStartId="24.[151,272,1750,1774]" captionTargetBox="[160,1436,187,1721]" captionTargetId="figure@24.[156,1441,187,1726]" captionTargetPageId="24" captionText="FIGURES 29 34. Anystipalpus labiduricola n. sp., adult female: 29, idiosoma, dorsal aspect; 30, enlarged separate views of setae denoted; 31, anterior margin and dorsal face of tectum; 32, cheliceral digits, dorsal view, showing dorsal antiaxial projection; 33, separate ventral view of movable digit, showing vestige of ventral process; 34, subcapitulum." httpUri="https://zenodo.org/record/190664/files/figure.png" pageId="22" pageNumber="23">Fig. 30</figureCitation>
), most about 0.7 0.8 as long as longitudinal intervals between their bases; setae
<emphasis id="B90C676DD445FFDEFC1DA4A700E0A5DF" box="[912,940,1725,1750]" italics="true" pageId="22" pageNumber="23">J5</emphasis>
,
<emphasis id="B90C676DD445FFDEFC31A4A70090A5DF" box="[956,988,1725,1750]" italics="true" pageId="22" pageNumber="23">Z5</emphasis>
,
<emphasis id="B90C676DD445FFDEFC61A4A70746A5DF" box="[1004,1034,1725,1750]" italics="true" pageId="22" pageNumber="23">S5</emphasis>
and sometimes others of
<emphasis id="B90C676DD445FFDEFACCA4A7061DA5DF" box="[1345,1361,1725,1750]" italics="true" pageId="22" pageNumber="23">Z</emphasis>
and
<emphasis id="B90C676DD445FFDEFA00A4A706D0A5DF" box="[1421,1436,1725,1750]" italics="true" pageId="22" pageNumber="23">S</emphasis>
series slightly barbed; setae
<emphasis id="B90C676DD445FFDEFE74A4FE015AA5F4" box="[505,534,1764,1789]" italics="true" pageId="22" pageNumber="23">J2</emphasis>
and
<emphasis id="B90C676DD445FFDEFDD9A4FE013DA5F4" box="[596,625,1764,1789]" italics="true" pageId="22" pageNumber="23">J3</emphasis>
(2526) slightly longer than subequally long
<emphasis id="B90C676DD445FFDEFB20A4FE0786A5F4" box="[1197,1226,1764,1789]" italics="true" pageId="22" pageNumber="23">J4</emphasis>
,
<emphasis id="B90C676DD445FFDEFB56A4FE07B4A5F4" box="[1243,1272,1764,1789]" italics="true" pageId="22" pageNumber="23">J5</emphasis>
,
<emphasis id="B90C676DD445FFDEFA84A4FE0665A5F4" box="[1289,1321,1764,1789]" italics="true" pageId="22" pageNumber="23">Z5</emphasis>
(2022); transverse interval between setae
<emphasis id="B90C676DD445FFDEFDAAA511010FA42D" box="[551,579,1803,1828]" italics="true" pageId="22" pageNumber="23">J5</emphasis>
(4547) greater than those between
<emphasis id="B90C676DD445FFDEFC7AA5110758A42D" box="[1015,1044,1803,1828]" italics="true" pageId="22" pageNumber="23">J4</emphasis>
(2832) and
<emphasis id="B90C676DD445FFDEFB39A511079FA42D" box="[1204,1235,1803,1828]" italics="true" pageId="22" pageNumber="23">Z5</emphasis>
(3738). Lateral soft cuticle with 13 to15 pairs of smooth setae (1218), including
<emphasis id="B90C676DD445FFDEFC29A52B00AEA443" box="[932,994,1841,1867]" italics="true" pageId="22" pageNumber="23">r3-r6</emphasis>
,
<emphasis id="B90C676DD445FFDEFC7FA52B0771A443" box="[1010,1085,1841,1867]" italics="true" pageId="22" pageNumber="23">R1-R7</emphasis>
and three or four pairs of
<emphasis id="B90C676DD445FFDEFAF8A52B06D0A443" box="[1397,1436,1841,1866]" italics="true" pageId="22" pageNumber="23">UR</emphasis>
setae.
</paragraph>
<paragraph id="8BC7BB7FD445FFDFFF48A5640637A04A" blockId="22.[151,1437,1222,2022]" lastBlockId="23.[151,1438,152,2034]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
Tritosternum with laciniae (6870) free nearly to their bases, where fused for 7 µm and with denticulate fringe weakly formed above apical margin of tritosternal base. Presternal area with pair of narrow indistinct platelets adhered to anterior margin of sternal shield. Sternal shield 95105 long from anterior presternal margin to irregularly straight posterior margin, 72 at narrowest width between coxae II, shield continuous with endopodal extensions between coxae I and II, II and III; shield surface evenly sclerotized, with pair of faint parallel longitudinal lines near mid-surface, flanked by other faint lines laterally, and anterior margin notched medially to level of
<emphasis id="B90C676DD444FFDFFE66A3160141A22C" box="[491,525,268,293]" italics="true" pageId="23" pageNumber="24">st1</emphasis>
(
<figureCitation id="1343A7FAD444FFDFFD93A316013BA22F" box="[542,631,268,294]" captionStart="FIGURES 35 38" captionStartId="25.[151,270,1908,1932]" captionTargetBox="[197,1409,462,1859]" captionTargetId="figure@25.[193,1415,455,1880]" captionTargetPageId="25" captionText="FIGURES 35 38. Anystipalpus labiduricola n. sp., adult female: 35, idiosoma, ventral aspect; 36, palptarsus, enlarged view; 37, palpus, dorsal view; 38, leg I (excepting tarsus) posterolateral view." httpUri="https://zenodo.org/record/190665/files/figure.png" pageId="23" pageNumber="24">Fig. 35</figureCitation>
); sternal shield with one pair of poroids,
<date id="FFC69DBFD444FFDFFBD0A31607CDA22C" box="[1117,1153,268,293]" pageId="23" pageNumber="24">
<emphasis id="B90C676DD444FFDFFBD0A31607CDA22C" box="[1117,1153,268,293]" italics="true" pageId="23" pageNumber="24">iv2</emphasis>
</date>
, and usually three pairs of attenuate setae,
<emphasis id="B90C676DD444FFDFFEF2A32802EFA242" box="[383,419,306,331]" italics="true" pageId="23" pageNumber="24">st3</emphasis>
barely on posterolateral corners of shield or sometimes barely off it; sternal setae progressively slightly shorter from
<emphasis id="B90C676DD444FFDFFDC7A3430121A27B" box="[586,621,345,370]" italics="true" pageId="23" pageNumber="24">st1</emphasis>
(2325) to
<emphasis id="B90C676DD444FFDFFD72A343006FA27B" box="[767,803,345,370]" italics="true" pageId="23" pageNumber="24">st3</emphasis>
(1517). Setae
<emphasis id="B90C676DD444FFDFFC6AA3430746A27B" box="[999,1034,345,370]" italics="true" pageId="23" pageNumber="24">st4</emphasis>
(1517) inserted on soft cuticle. Endopodal strips weakly developed between coxae III and IV. Epigynal shield faintly lineated laterally, 92 100 long from anterior margin of broadly rounded hyaline rim to bluntly pointed posterior margin, 38 at widest level between setae
<emphasis id="B90C676DD444FFDFFE5BA3D702B5A2EF" box="[470,505,461,486]" italics="true" pageId="23" pageNumber="24">st5</emphasis>
(15), and 20 at narrowest width between legs IV. Opisthosomatic venter with 1 pair of small postgenital platelets, and with metapodal platelets irregularly oval (
<date id="FFC69DBFD444FFDFFBDEA3EE07E3A107" box="[1107,1199,500,526]" pageId="23" pageNumber="24" value="1910-10-15">15 x 10</date>
), removed by about their greatest dimension from posterior margin of peritrematal-exopodal shields; anterior pair of small metapodal platelets amalgamated with expanded posterior margins of peritrematal-exopodal plates. Anal shield faintly reticulate-lineate, with paranal setae (1415) slightly shorter than postanal seta (1820); shield width (6570) subequal to its length including cribrum; paranal sigillae protruding from shields lateral margins. Soft cuticle with ten pairs of simple opisthogastric setae
<emphasis id="B90C676DD444FFDFFC10A0AF074EA1C7" box="[925,1026,693,719]" italics="true" pageId="23" pageNumber="24">JV1-JV5</emphasis>
,
<emphasis id="B90C676DD444FFDFFB9CA0AF0731A1C7" box="[1041,1149,693,719]" italics="true" pageId="23" pageNumber="24">ZV1-ZV5</emphasis>
flanked by three or four pairs of simple submarginal
<emphasis id="B90C676DD444FFDFFE7DA0C60154A1FC" box="[496,536,732,757]" italics="true" pageId="23" pageNumber="24">UR</emphasis>
and marginal
<emphasis id="B90C676DD444FFDFFD4AA0C60195A1FC" box="[711,729,732,757]" italics="true" pageId="23" pageNumber="24">R</emphasis>
setae. Peritrematal-exopodal shield with four or five lines radiating from stigma to broadly scalloped posteromedian margin (
<figureCitation id="1343A7FAD444FFDFFC37A1180758A015" box="[954,1044,770,796]" captionStart="FIGURES 35 38" captionStartId="25.[151,270,1908,1932]" captionTargetBox="[197,1409,462,1859]" captionTargetId="figure@25.[193,1415,455,1880]" captionTargetPageId="25" captionText="FIGURES 35 38. Anystipalpus labiduricola n. sp., adult female: 35, idiosoma, ventral aspect; 36, palptarsus, enlarged view; 37, palpus, dorsal view; 38, leg I (excepting tarsus) posterolateral view." httpUri="https://zenodo.org/record/190665/files/figure.png" pageId="23" pageNumber="24">Fig. 35</figureCitation>
); peritreme extends anteriorly to level of seta
<emphasis id="B90C676DD444FFDFFEA1A133020AA04B" box="[300,326,809,834]" italics="true" pageId="23" pageNumber="24">z1</emphasis>
, where vertex curved ventrally. Spermathecal structures indiscernible in specimens at hand.
</paragraph>
<paragraph id="8BC7BB7FD444FFDFFF48A14A014DA6DC" blockId="23.[151,1438,152,2034]" pageId="23" pageNumber="24">
Tectum with anterior margin finely denticulate, convex; mid surface of tectum with transverse line of denticles well defined, concave medially (
<figureCitation id="1343A7FAD444FFDFFD03A16C01ABA099" box="[654,743,886,912]" captionStart="FIGURES 29 34" captionStartId="24.[151,272,1750,1774]" captionTargetBox="[160,1436,187,1721]" captionTargetId="figure@24.[156,1441,187,1726]" captionTargetPageId="24" captionText="FIGURES 29 34. Anystipalpus labiduricola n. sp., adult female: 29, idiosoma, dorsal aspect; 30, enlarged separate views of setae denoted; 31, anterior margin and dorsal face of tectum; 32, cheliceral digits, dorsal view, showing dorsal antiaxial projection; 33, separate ventral view of movable digit, showing vestige of ventral process; 34, subcapitulum." httpUri="https://zenodo.org/record/190664/files/figure.png" pageId="23" pageNumber="24">Fig. 31</figureCitation>
). Cheliceral shaft, excluding basal section, 125 long, with slender digits; dorsal face of fixed digit on antiaxial side with pointed process extending to level of basal tooth of movable digit, seeming to cover movable digit in repose (
<figureCitation id="1343A7FAD444FFDFFCF9A1DE0083A0D7" box="[884,975,964,990]" captionStart="FIGURES 29 34" captionStartId="24.[151,272,1750,1774]" captionTargetBox="[160,1436,187,1721]" captionTargetId="figure@24.[156,1441,187,1726]" captionTargetPageId="24" captionText="FIGURES 29 34. Anystipalpus labiduricola n. sp., adult female: 29, idiosoma, dorsal aspect; 30, enlarged separate views of setae denoted; 31, anterior margin and dorsal face of tectum; 32, cheliceral digits, dorsal view, showing dorsal antiaxial projection; 33, separate ventral view of movable digit, showing vestige of ventral process; 34, subcapitulum." httpUri="https://zenodo.org/record/190664/files/figure.png" pageId="23" pageNumber="24">Fig. 32</figureCitation>
); fixed digit with row of five to seven weak teeth including barely offset subapical tooth along apical third and a tiny proximal tooth on masticatory margin (
<figureCitation id="1343A7FAD444FFDFFF77A60B021EA722" box="[250,338,1041,1067]" captionStart="FIGURES 29 34" captionStartId="24.[151,272,1750,1774]" captionTargetBox="[160,1436,187,1721]" captionTargetId="figure@24.[156,1441,187,1726]" captionTargetPageId="24" captionText="FIGURES 29 34. Anystipalpus labiduricola n. sp., adult female: 29, idiosoma, dorsal aspect; 30, enlarged separate views of setae denoted; 31, anterior margin and dorsal face of tectum; 32, cheliceral digits, dorsal view, showing dorsal antiaxial projection; 33, separate ventral view of movable digit, showing vestige of ventral process; 34, subcapitulum." httpUri="https://zenodo.org/record/190664/files/figure.png" pageId="23" pageNumber="24">Fig. 32</figureCitation>
); movable digit (38) bidentate, with vestige of process on midventral hardly perceptible (
<figureCitation id="1343A7FAD444FFDFFAE1A60B03F5A75B" captionStart="FIGURES 29 34" captionStartId="24.[151,272,1750,1774]" captionTargetBox="[160,1436,187,1721]" captionTargetId="figure@24.[156,1441,187,1726]" captionTargetPageId="24" captionText="FIGURES 29 34. Anystipalpus labiduricola n. sp., adult female: 29, idiosoma, dorsal aspect; 30, enlarged separate views of setae denoted; 31, anterior margin and dorsal face of tectum; 32, cheliceral digits, dorsal view, showing dorsal antiaxial projection; 33, separate ventral view of movable digit, showing vestige of ventral process; 34, subcapitulum." httpUri="https://zenodo.org/record/190664/files/figure.png" pageId="23" pageNumber="24">Fig. 33</figureCitation>
). Subcapitulum (
<figureCitation id="1343A7FAD444FFDFFE0BA6220292A75B" box="[390,478,1080,1106]" captionStart="FIGURES 29 34" captionStartId="24.[151,272,1750,1774]" captionTargetBox="[160,1436,187,1721]" captionTargetId="figure@24.[156,1441,187,1726]" captionTargetPageId="24" captionText="FIGURES 29 34. Anystipalpus labiduricola n. sp., adult female: 29, idiosoma, dorsal aspect; 30, enlarged separate views of setae denoted; 31, anterior margin and dorsal face of tectum; 32, cheliceral digits, dorsal view, showing dorsal antiaxial projection; 33, separate ventral view of movable digit, showing vestige of ventral process; 34, subcapitulum." httpUri="https://zenodo.org/record/190664/files/figure.png" pageId="23" pageNumber="24">Fig. 34</figureCitation>
) slender, elongate, its greatest width (72) at base about 0.7 its length (98) from capitular base to apex of corniculi; hypostome on a neck-like projection, such that longitudinal distance between
<emphasis id="B90C676DD444FFDFFE8CA69F0261A797" box="[257,301,1157,1182]" italics="true" pageId="23" pageNumber="24">hp3</emphasis>
and
<emphasis id="B90C676DD444FFDFFEE5A69F02C8A797" box="[360,388,1157,1182]" italics="true" pageId="23" pageNumber="24">pc</emphasis>
(35) over four times that between
<emphasis id="B90C676DD444FFDFFCACA69F0001A797" box="[801,845,1157,1182]" italics="true" pageId="23" pageNumber="24">hp1</emphasis>
and
<emphasis id="B90C676DD444FFDFFC05A69F00F8A797" box="[904,948,1157,1182]" italics="true" pageId="23" pageNumber="24">hp3</emphasis>
(8). Deutosternum with seven transverse rows of denticles, all but seventh row connected laterally, similar in width, each with a single median denticle but sixth and sometimes seventh row with one or two denticles on either side (
<figureCitation id="1343A7FAD444FFDFFBCCA6C907D0A7E4" box="[1089,1180,1235,1261]" captionStart="FIGURES 29 34" captionStartId="24.[151,272,1750,1774]" captionTargetBox="[160,1436,187,1721]" captionTargetId="figure@24.[156,1441,187,1726]" captionTargetPageId="24" captionText="FIGURES 29 34. Anystipalpus labiduricola n. sp., adult female: 29, idiosoma, dorsal aspect; 30, enlarged separate views of setae denoted; 31, anterior margin and dorsal face of tectum; 32, cheliceral digits, dorsal view, showing dorsal antiaxial projection; 33, separate ventral view of movable digit, showing vestige of ventral process; 34, subcapitulum." httpUri="https://zenodo.org/record/190664/files/figure.png" pageId="23" pageNumber="24">Fig. 34</figureCitation>
). Corniculi normal in form, slightly longer than internal malae; subcapitular setae simple,
<emphasis id="B90C676DD444FFDFFC35A6E300A8A61B" box="[952,996,1273,1298]" italics="true" pageId="23" pageNumber="24">hp1</emphasis>
(2022) longer than capitular seta
<emphasis id="B90C676DD444FFDFFA0DA6E306D7A61B" box="[1408,1435,1273,1298]" italics="true" pageId="23" pageNumber="24">pc</emphasis>
(1516), and
<emphasis id="B90C676DD444FFDFFEB9A73A022CA630" box="[308,352,1312,1337]" italics="true" pageId="23" pageNumber="24">hp3</emphasis>
short (10), scarcely longer than
<emphasis id="B90C676DD444FFDFFD51A73A004BA630" box="[732,775,1312,1337]" italics="true" pageId="23" pageNumber="24">hp2</emphasis>
(68). Palpus elongate (170175), about 0.53 as long as leg I, each of palpal trochanter (4243), femur (38), genu (3638), tibia (4041) similarly elongated (
<figureCitation id="1343A7FAD444FFDFFAB1A75C03F4A68E" captionStart="FIGURES 35 38" captionStartId="25.[151,270,1908,1932]" captionTargetBox="[197,1409,462,1859]" captionTargetId="figure@25.[193,1415,455,1880]" captionTargetPageId="25" captionText="FIGURES 35 38. Anystipalpus labiduricola n. sp., adult female: 35, idiosoma, ventral aspect; 36, palptarsus, enlarged view; 37, palpus, dorsal view; 38, leg I (excepting tarsus) posterolateral view." httpUri="https://zenodo.org/record/190665/files/figure.png" pageId="23" pageNumber="24">Figs 36, 37</figureCitation>
), about 2.52.8 times as long as palptarsus (1517); palptibia 3.8 times as long as wide; palptrochanter with more basal seta (15) half as long as more distal seta (28); apical
<emphasis id="B90C676DD444FFDFFC44A78E00AAA6A4" box="[969,998,1428,1453]" italics="true" pageId="23" pageNumber="24">pd</emphasis>
seta of palptibia recurved, blunt (13) in distinction to adjacent setae.
</paragraph>
<paragraph id="8BC7BB7FD444FFDFFF48A7FB01D9A439" blockId="23.[151,1438,152,2034]" pageId="23" pageNumber="24">
Legs I (320325) about 0.9 as long as collective length of the two dorsal shields; other leg lengths (excluding pretarsi): II (250256), III (225230), IV (288300). Leg I length ratios, genu: tibia: tarsus, about 1.0: 0.9: 1.41.5; tarsus (7577) slightly longer than femur (7274). Coxa I with two or three small, raised bumps on ventral face (
<figureCitation id="1343A7FAD444FFDFFE24A44F014CA566" box="[425,512,1621,1647]" captionStart="FIGURES 35 38" captionStartId="25.[151,270,1908,1932]" captionTargetBox="[197,1409,462,1859]" captionTargetId="figure@25.[193,1415,455,1880]" captionTargetPageId="25" captionText="FIGURES 35 38. Anystipalpus labiduricola n. sp., adult female: 35, idiosoma, ventral aspect; 36, palptarsus, enlarged view; 37, palpus, dorsal view; 38, leg I (excepting tarsus) posterolateral view." httpUri="https://zenodo.org/record/190665/files/figure.png" pageId="23" pageNumber="24">Fig. 38</figureCitation>
); coxae IIV lineate on posterior ventral surfaces (as in
<figureCitation id="1343A7FAD444FFDFFB00A44F079FA566" box="[1165,1235,1621,1647]" captionStart="FIGURES 7 10" captionStartId="16.[151,274,1737,1761]" captionTargetBox="[153,1436,317,1682]" captionTargetId="figure@16.[153,1438,317,1707]" captionTargetPageId="16" captionText="FIGURES 7 10. Anystipalpus percicola Berlese, adult female: 7, idiosoma, ventral aspect; 8, tarsus II excluding pretarsus, dorsal view; 9, leg I excluding coxa and tarsus, dorsal view; 10 a, chelicera, dorsal view; 10 b, separate ventral view of distal part of fixed digit; 10 c, separate antiaxial view of movable digit." httpUri="https://zenodo.org/record/190659/files/figure.png" pageId="23" pageNumber="24">Fig. 7</figureCitation>
); basal seta
<emphasis id="B90C676DD444FFDFFAECA44F0631A567" box="[1377,1405,1621,1646]" italics="true" pageId="23" pageNumber="24">av</emphasis>
of coxa I and
<emphasis id="B90C676DD444FFDFFE95A466027FA59C" box="[280,307,1660,1685]" italics="true" pageId="23" pageNumber="24">pv</emphasis>
of coxa II modified as thick, conical, blunt spines (78);
<emphasis id="B90C676DD444FFDFFC59A46600BCA59C" box="[980,1008,1660,1685]" italics="true" pageId="23" pageNumber="24">pv</emphasis>
of coxa I,
<emphasis id="B90C676DD444FFDFFBE0A46607C5A59C" box="[1133,1161,1660,1685]" italics="true" pageId="23" pageNumber="24">av</emphasis>
of coxa II, and setae of coxae IIIIV normal, slender,
<emphasis id="B90C676DD444FFDFFD9DA4B90161A5B5" box="[528,557,1699,1724]" italics="true" pageId="23" pageNumber="24">pv</emphasis>
of coxa III and
<emphasis id="B90C676DD444FFDFFD75A4B90049A5B5" box="[760,773,1699,1724]" italics="true" pageId="23" pageNumber="24">v</emphasis>
of coxa IV short (12). Legs I to IV with chaetotactic formulae of femora, genua, tibiae as described for genus. Leg I with setae
<emphasis id="B90C676DD444FFDFFC71A4D30747A5EB" box="[1020,1035,1737,1762]" italics="true" pageId="23" pageNumber="24">d</emphasis>
(17) of trochanter and
<emphasis id="B90C676DD444FFDFFA95A4D30609A5EB" box="[1304,1349,1737,1762]" italics="true" pageId="23" pageNumber="24">pd2</emphasis>
(22) of femur barbed, slightly stout, pointed,
<emphasis id="B90C676DD444FFDFFDD5A4EA0139A400" box="[600,629,1776,1801]" italics="true" pageId="23" pageNumber="24">pd</emphasis>
1 of femur slightly thickened, pointed (
<figureCitation id="1343A7FAD444FFDFFBC4A4EA07EEA403" box="[1097,1186,1776,1802]" captionStart="FIGURES 35 38" captionStartId="25.[151,270,1908,1932]" captionTargetBox="[197,1409,462,1859]" captionTargetId="figure@25.[193,1415,455,1880]" captionTargetPageId="25" captionText="FIGURES 35 38. Anystipalpus labiduricola n. sp., adult female: 35, idiosoma, ventral aspect; 36, palptarsus, enlarged view; 37, palpus, dorsal view; 38, leg I (excepting tarsus) posterolateral view." httpUri="https://zenodo.org/record/190665/files/figure.png" pageId="23" pageNumber="24">Fig. 38</figureCitation>
); other setae normal, slender. Legs IIIV without modified setae.
</paragraph>
<paragraph id="8BC7BB7FD444FFDFFF4BA5270184A45E" blockId="23.[151,1438,152,2034]" box="[198,712,1853,1879]" pageId="23" pageNumber="24">
<emphasis id="B90C676DD444FFDFFF4BA5270109A45E" bold="true" box="[198,581,1853,1879]" pageId="23" pageNumber="24">ADULT MALE and IMMATURES.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C362E8F4D444FFDFFF4BA5790093A4FB" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="8BC7BB7FD444FFDFFF4BA5790093A4FB" blockId="23.[151,1438,152,2034]" pageId="23" pageNumber="24">
<emphasis id="B90C676DD444FFDFFF4BA5790230A474" bold="true" box="[198,380,1891,1917]" pageId="23" pageNumber="24">
<typeStatus id="54C305DDD444FFDFFF4BA579024FA474" box="[198,259,1891,1917]" pageId="23" pageNumber="24">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54C305DDD444FFDFFE0EA57E014FA477" box="[387,515,1892,1918]" pageId="23" pageNumber="24" type="holotype">HOLOTYPE</typeStatus>
, adult female:
<collectingCountry id="F36FFBEFD444FFDFFD3CA57E01B0A477" box="[689,764,1892,1918]" name="Iran" pageId="23" pageNumber="24">IRAN</collectingCountry>
, North Khorasan Province, Esfarayen (
<geoCoordinate id="EE4CDDB8D444FFDFFB48A57E062BA477" box="[1221,1383,1892,1918]" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="15" value="36.924725">36° 55' 29&quot; N</geoCoordinate>
,
<geoCoordinate id="EE4CDDB8D444FFDFFAFEA57E0242A4AD" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="15" value="57.735558">57° 44' 08&quot; E</geoCoordinate>
) elevation
<quantity id="4C80169AD444FFDFFE10A590014BA4AD" box="[413,519,1930,1956]" metricMagnitude="0" metricUnit="m" metricValue="1.471" pageId="23" pageNumber="24" unit="m" value="1.471">1,471 m</quantity>
,
<date id="FFC69DBFD444FFDFFD98A590018DA4AD" box="[533,705,1930,1956]" pageId="23" pageNumber="24" value="2007-06-26">26 June 2007</date>
, coll. S. Kazemi, ex. “earwig” (
<taxonomicName id="4C78C0FCD444FFDFFBD8A59007BCA4AD" box="[1109,1264,1930,1956]" class="Insecta" kingdom="Animalia" order="Dermaptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="order">Dermaptera</taxonomicName>
:
<taxonomicName id="4C78C0FCD444FFDFFB71A59006D5A4AD" box="[1276,1433,1930,1956]" class="Insecta" family="Labiduridae" kingdom="Animalia" order="Dermaptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="family">Labiduridae</taxonomicName>
:
<taxonomicName id="4C78C0FCD444FFDFFF1AA5AB024AA4C3" box="[151,262,1969,1994]" class="Insecta" family="Labiduridae" genus="Labidura" kingdom="Animalia" order="Dermaptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B90C676DD444FFDFFF1AA5AB024AA4C3" box="[151,262,1969,1994]" italics="true" pageId="23" pageNumber="24">Labidura</emphasis>
</taxonomicName>
sp.).
<typeStatus id="54C305DDD444FFDFFEC6A5AB029BA4C2" box="[331,471,1969,1995]" pageId="23" pageNumber="24" type="paratype">PARATYPES</typeStatus>
: nine adult females, with same data as
<typeStatus id="54C305DDD444FFDFFC39A5AB076EA4C2" box="[948,1058,1969,1995]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
: two deposited in the
<collectionCode id="ED6923BAD444FFDFFAA6A5AB06D4A4C2" box="[1323,1432,1969,1995]" pageId="23" pageNumber="24">AETMU</collectionCode>
, Tehran, five in the MZUNAV, Pamplona, and two in the
<collectionCode id="ED6923BAD444FFDFFCA3A5C2003BA4FB" box="[814,887,2008,2034]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="23" pageNumber="24">CNCI</collectionCode>
, Ottawa.
</paragraph>
</subSubSection>
<caption id="DF07EBF7D44BFFD0FF1AA4CC06CCA43A" httpUri="https://zenodo.org/record/190664/files/figure.png" pageId="24" pageNumber="25" targetBox="[160,1436,187,1721]" targetPageId="24">
<paragraph id="8BC7BB7FD44BFFD0FF1AA4CC06CCA43A" blockId="24.[151,1436,1750,1843]" pageId="24" pageNumber="25">
<emphasis id="B90C676DD44BFFD0FF1AA4CC022FA5E7" bold="true" box="[151,355,1750,1774]" pageId="24" pageNumber="25">FIGURES 2934.</emphasis>
<taxonomicName id="4C78C0FCD44BFFD0FEE6A4CD01C4A5E7" box="[363,648,1751,1774]" class="Arachnida" family="Laelapidae" genus="Anystipalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="labiduricola" status="sp. nov.">
<emphasis id="B90C676DD44BFFD0FEE6A4CD01C4A5E7" box="[363,648,1751,1774]" italics="true" pageId="24" pageNumber="25">Anystipalpus labiduricola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23FDA16D44BFFD0FD1DA4CC0182A5E7" box="[656,718,1750,1774]" pageId="24" pageNumber="25" rank="species">
<emphasis id="B90C676DD44BFFD0FD1DA4CC0182A5E7" bold="true" box="[656,718,1750,1774]" pageId="24" pageNumber="25">n. sp.</emphasis>
</taxonomicNameLabel>
, adult female: 29, idiosoma, dorsal aspect; 30, enlarged separate views of setae denoted; 31, anterior margin and dorsal face of tectum; 32, cheliceral digits, dorsal view, showing dorsal antiaxial projection; 33, separate ventral view of movable digit, showing vestige of ventral process; 34, subcapitulum.
</paragraph>
</caption>
<subSubSection id="C362E8F4D44BFFD0FF4BA57B0120A4AB" pageId="24" pageNumber="25" type="etymology">
<paragraph id="8BC7BB7FD44BFFD0FF4BA57B0120A4AB" blockId="24.[151,1436,1889,2032]" pageId="24" pageNumber="25">
<emphasis id="B90C676DD44BFFD0FF4BA57B021FA472" bold="true" box="[198,339,1889,1915]" pageId="24" pageNumber="25">Etymology.</emphasis>
The specific name,
<taxonomicName id="4C78C0FCD44BFFD0FDB0A5780182A472" box="[573,718,1890,1915]" class="Arachnida" family="Laelapidae" genus="Anystipalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="labiduricola">
<emphasis id="B90C676DD44BFFD0FDB0A5780182A472" box="[573,718,1890,1915]" italics="true" pageId="24" pageNumber="25">labiduricola</emphasis>
</taxonomicName>
, is based on the name of the genus of labidurid earwigs under whose tegmina these mites are phoretic.
</paragraph>
</subSubSection>
<subSubSection id="C362E8F4D44BFFD2FF4BA5B501C2A4CE" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" type="discussion">
<paragraph id="8BC7BB7FD44BFFD1FF4BA5B501C0A293" blockId="24.[151,1436,1889,2032]" lastBlockId="25.[151,1436,152,410]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="B90C676DD44BFFD0FF4BA5B5020CA4C0" bold="true" box="[198,320,1967,1993]" pageId="24" pageNumber="25">Remarks.</emphasis>
Adult females of
<taxonomicName id="4C78C0FCD44BFFD0FD9FA5B5000DA4C1" box="[530,833,1967,1992]" class="Arachnida" family="Laelapidae" genus="Anystipalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="labiduricola">
<emphasis id="B90C676DD44BFFD0FD9FA5B5000DA4C1" box="[530,833,1967,1992]" italics="true" pageId="24" pageNumber="25">Anystipalpus labiduricola</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="4C78C0FCD44BFFD0FBC3A5B50218A4F9" authority="Eidelberg, 2000" authorityName="Eidelberg" authorityYear="2000" class="Arachnida" family="Ascidae" genus="Antennoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="longisetus" subGenus="Antennoseius">
<emphasis id="B90C676DD44BFFD0FBC3A5B507A6A4C1" box="[1102,1258,1967,1992]" italics="true" pageId="24" pageNumber="25">Antennoseius</emphasis>
(
<emphasis id="B90C676DD44BFFD0FB76A5B50641A4C1" box="[1275,1293,1967,1992]" italics="true" pageId="24" pageNumber="25">A</emphasis>
.)
<emphasis id="B90C676DD44BFFD0FAA8A5B506D0A4C1" box="[1317,1436,1967,1992]" italics="true" pageId="24" pageNumber="25">longisetus</emphasis>
<bibRefCitation id="EFE9C68ED44BFFD0FF1AA5CC0218A4F9" author="Eidelberg" box="[151,340,2006,2032]" pageId="24" pageNumber="37" refString="Eidelberg, M. M. (2000) Three new mite species of the family Antennoseiidae (Parasitiformes, Gamasina). Zoologicheskii Zhurnal, 79, 1396 - 1401. [in Russian]" type="journal article" year="2000">Eidelberg, 2000</bibRefCitation>
</taxonomicName>
. Both forms are similar in the number and relatively long length of setae on the dorsal shields, in the forms of the sternal and epigynal shields, and in the scarcity of modified spinelike setae on leg I. Those of
<taxonomicName id="4C78C0FCD44AFFD1FF35A2A5029AA3D1" box="[184,470,191,216]" class="Arachnida" family="Ascidae" genus="Antennoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="longisetus">
<emphasis id="B90C676DD44AFFD1FF35A2A5029AA3D1" box="[184,470,191,216]" italics="true" pageId="25" pageNumber="26">Antennoseius longisetus</emphasis>
</taxonomicName>
differ in the posterior dorsal shield having setae
<emphasis id="B90C676DD44AFFD1FBAEA2A50729A3D1" box="[1059,1125,190,216]" italics="true" pageId="25" pageNumber="26">J4-J5</emphasis>
,
<emphasis id="B90C676DD44AFFD1FBF9A2A507F1A3D1" box="[1140,1213,190,216]" italics="true" pageId="25" pageNumber="26">Z4-Z5</emphasis>
,
<emphasis id="B90C676DD44AFFD1FB41A2A507A5A3D1" box="[1228,1257,191,216]" italics="true" pageId="25" pageNumber="26">S5</emphasis>
conspicuously barbed, the narrowed neck of the epigynal shield more constricted between legs IV, none of the setae on coxae I and II modified, spinelike, and the chelicerae typical of
<taxonomicName id="4C78C0FCD44AFFD1FCCEA31600ACA22C" box="[835,992,268,293]" class="Arachnida" family="Ascidae" genus="Antennoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B90C676DD44AFFD1FCCEA31600ACA22C" box="[835,992,268,293]" italics="true" pageId="25" pageNumber="26">Antennoseius</emphasis>
</taxonomicName>
, with the fixed digit multidenticulate along nearly the entire masticatory margin and the movable digit having a gutterlike process along its midventral face. No information was provided on the length of the palpi and their segments and on the degree of subcapitular elongation in
<taxonomicName id="4C78C0FCD44AFFD1FE63A39A01CAA290" box="[494,646,384,410]" class="Arachnida" family="Ascidae" genus="Antennoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="longisetus">
<emphasis id="B90C676DD44AFFD1FE63A39A014CA290" box="[494,512,384,409]" italics="true" pageId="25" pageNumber="26">A</emphasis>
.
<emphasis id="B90C676DD44AFFD1FD82A39A01CAA290" box="[527,646,384,409]" italics="true" pageId="25" pageNumber="26">longisetus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF07EBF7D44AFFD1FF1AA56E009EA4A7" httpUri="https://zenodo.org/record/190665/files/figure.png" pageId="25" pageNumber="26" targetBox="[197,1409,462,1859]" targetPageId="25">
<paragraph id="8BC7BB7FD44AFFD1FF1AA56E009EA4A7" blockId="25.[151,1435,1908,1966]" pageId="25" pageNumber="26">
<emphasis id="B90C676DD44AFFD1FF1AA56E0212A485" bold="true" box="[151,350,1908,1932]" pageId="25" pageNumber="26">FIGURES 3538.</emphasis>
<taxonomicName id="4C78C0FCD44AFFD1FEE8A56F0137A485" box="[357,635,1909,1932]" class="Arachnida" family="Laelapidae" genus="Anystipalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="labiduricola" status="sp. nov.">
<emphasis id="B90C676DD44AFFD1FEE8A56F0137A485" box="[357,635,1909,1932]" italics="true" pageId="25" pageNumber="26">Anystipalpus labiduricola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23FDA16D44AFFD1FD0FA56E01F3A485" box="[642,703,1908,1932]" pageId="25" pageNumber="26" rank="species">
<emphasis id="B90C676DD44AFFD1FD0FA56E01F3A485" bold="true" box="[642,703,1908,1932]" pageId="25" pageNumber="26">n. sp.</emphasis>
</taxonomicNameLabel>
, adult female: 35, idiosoma, ventral aspect; 36, palptarsus, enlarged view; 37, palpus, dorsal view; 38, leg I (excepting tarsus) posterolateral view.
</paragraph>
</caption>
<caption id="DF07EBF7D449FFD2FF1AA7BA039DA516" httpUri="https://zenodo.org/record/190666/files/figure.png" pageId="26" pageNumber="27" targetBox="[160,1431,194,1369]" targetPageId="26">
<paragraph id="8BC7BB7FD449FFD2FF1AA7BA039DA516" blockId="26.[151,1436,1440,1567]" pageId="26" pageNumber="27">
<emphasis id="B90C676DD449FFD2FF1AA7BA022FA6B1" bold="true" box="[151,355,1440,1464]" pageId="26" pageNumber="27">FIGURES 3942.</emphasis>
Figs 3940,
<taxonomicName id="4C78C0FCD449FFD2FE79A7BA005EA6BE" box="[500,786,1440,1463]" class="Arachnida" family="Laelapidae" genus="Anystipalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="labiduricola" status="sp. nov.">
<emphasis id="B90C676DD449FFD2FE79A7BA005EA6BE" box="[500,786,1440,1463]" italics="true" pageId="26" pageNumber="27">Anystipalpus labiduricola</emphasis>
</taxonomicName>
<emphasis id="B90C676DD449FFD2FC97A7BA0014A6B1" bold="true" box="[794,856,1440,1464]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="A23FDA16D449FFD2FC97A7BA0014A6B1" box="[794,856,1440,1464]" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult female: 39, tarsus I, dorsal view; 40, tarsus I, detail of distal sensorial region. Figs 4142,
<taxonomicName id="4C78C0FCD449FFD2FD0BA7D9003CA6D3" box="[646,880,1475,1498]" class="Arachnida" family="Laelapidae" genus="Anystipalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="kazemii" status="sp. nov.">
<emphasis id="B90C676DD449FFD2FD0BA7D9003CA6D3" box="[646,880,1475,1498]" italics="true" pageId="26" pageNumber="27">Anystipalpus kazemii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23FDA16D449FFD2FCFAA7D800F9A6D3" box="[887,949,1474,1498]" pageId="26" pageNumber="27" rank="species">
<emphasis id="B90C676DD449FFD2FCFAA7D800F9A6D3" bold="true" box="[887,949,1474,1498]" pageId="26" pageNumber="27">n. sp.</emphasis>
</taxonomicNameLabel>
, adult female: 41a, palpus excepting tarsus, dorsal view; 41b, palptarsus, dorsal view; 42a, leg I excluding coxa and tarsus, dorsolateral view; 42b, tarsus I, dorsal view.
</paragraph>
</caption>
<paragraph id="8BC7BB7FD449FFD2FF48A44801C2A4CE" blockId="26.[151,1436,1618,1991]" pageId="26" pageNumber="27">
Based on how the earwigs were collected and the location of a considerable number of the mites tightly positioned on several host specimens, the extraordinary finding of adult females of
<taxonomicName id="4C78C0FCD449FFD2FBEDA463065DA59B" box="[1120,1297,1656,1682]" class="Arachnida" family="Laelapidae" genus="Anystipalpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="labiduricola">
<emphasis id="B90C676DD449FFD2FBEDA463073EA59B" box="[1120,1138,1657,1682]" italics="true" pageId="26" pageNumber="27">A</emphasis>
.
<emphasis id="B90C676DD449FFD2FB0DA463065DA59B" box="[1152,1297,1657,1682]" italics="true" pageId="26" pageNumber="27">labiduricola</emphasis>
</taxonomicName>
phoretic on labidurid earwigs does not seem to be accidental. The earwigs were collected individually by hand under stones, and not in the presence of carabid beetles (S. Kazemi, personal communication). The mites were found by one of us (MLM) to occupy three different positions on the earwigs: (a) on the dorsal surface of the folded membranous hind wing, in the small depressions of the axillar region; (b) on the small depressions of the dorsal surface of the meso- and meta-notum, beneath the folded hind wing; and (c) between the plications in the distal region of the folded membranous hind wing (where these mites were at the same frontal level as the others, perhaps enabling them to move from one position to another). Further thoughts on this phoretic association are presented in the discussion.
</paragraph>
</subSubSection>
</treatment>
</document>