509 lines
78 KiB
XML
509 lines
78 KiB
XML
<document id="E89A836D594A9518285FAB9C7FF2BFB5" ID-CLB-Dataset="7305" ID-DOI="10.1051/acarologia/20164158" ID-GBIF-Dataset="3d1b1ce8-cd07-4730-8e58-36433c6c3675" ID-ISSN="2107-7207" ID-Zenodo-Dep="4640248" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1616776605261" checkinUser="carolina" docAuthor="Witalinski, W." docDate="2017" docId="0391A308FFC6FFDCFE9CFC54FE4BFC42" docLanguage="en" docName="Acarologia.57.2.211-221.pdf" docOrigin="Acarologia 57 (2)" docSource="http://dx.doi.org/10.1051/acarologia/20164158" docStyle="DocumentStyle:8E0DCBB23C2CB3FD20935B7794B24F85.3:Acarologia.2010-2017.journal_article" docStyleId="8E0DCBB23C2CB3FD20935B7794B24F85" docStyleName="Acarologia.2010-2017.journal_article" docStyleVersion="3" docTitle="Holoparasitus fanes Witalinski 2017, n. sp." docType="treatment" docVersion="7" lastPageNumber="221" masterDocId="FFA8DB70FFC0FFD7FFACFFF9FFA0FF8C" masterDocTitle="Two new species of Holoparasitus Oudemans, 1936 from Europe (Parasitiformes: Parasitidae)" masterLastPageNumber="221" masterPageNumber="211" pageNumber="216" updateTime="1698935192083" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-ND-4.0">
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<mods:title id="4417CAA4A236F0A9E53A868BD4EADEB3">Two new species of Holoparasitus Oudemans, 1936 from Europe (Parasitiformes: Parasitidae)</mods:title>
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<mods:name id="F3B008E0D6B2376FA13C8B4E283932FC" type="personal">
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<mods:namePart id="BB4A01C1AAA3193CBF053DC7441AE015">Witalinski, W.</mods:namePart>
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<mods:title id="B048AD20A345494739074739A2C2C163">Acarologia</mods:title>
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<mods:part id="9125C170C6F772BD7BE3C9B5D7A6DDB1">
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<mods:date id="CF635B6C635D008356F6AACF17E94ACB">2017</mods:date>
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<mods:number id="F91C576E8E6B7741C2836F64D5CB12E5">2016-12-16</mods:number>
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<mods:number id="77A3E9587387152437F1BE862387A05E">57</mods:number>
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<treatment id="0391A308FFC6FFDCFE9CFC54FE4BFC42" ID-DOI="http://doi.org/10.5281/zenodo.4697140" ID-GBIF-Taxon="181020895" ID-Zenodo-Dep="4697140" LSID="urn:lsid:plazi:treatment:0391A308FFC6FFDCFE9CFC54FE4BFC42" httpUri="http://treatment.plazi.org/id/0391A308FFC6FFDCFE9CFC54FE4BFC42" lastPageId="11" lastPageNumber="221" pageId="6" pageNumber="216">
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<subSubSection id="C3224195FFC6FFD1FE9CFC54FDC8FC4A" box="[304,616,941,966]" pageId="6" pageNumber="216" type="nomenclature">
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<paragraph id="8B87121EFFC6FFD1FE9CFC54FDC8FC4A" blockId="6.[304,616,941,1001]" box="[304,616,941,966]" pageId="6" pageNumber="216">
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<heading id="D0CFA572FFC6FFD1FE9CFC54FDC8FC4A" bold="true" box="[304,616,941,966]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="216" reason="2">
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<emphasis id="B94CCE0CFFC6FFD1FE9CFC54FDC8FC4A" bold="true" box="[304,616,941,966]" pageId="6" pageNumber="216">
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<taxonomicName id="4C38699DFFC6FFD1FE9CFC54FD80FC49" authority="Witalinski, 2017" authorityName="Witalinski" authorityYear="2017" box="[304,544,941,965]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="216" phylum="Arthropoda" rank="species" species="fanes" status="sp. nov.">
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<emphasis id="B94CCE0CFFC6FFD1FE9CFC54FD80FC49" bold="true" box="[304,544,941,965]" italics="true" pageId="6" pageNumber="216">Holoparasitus fanes</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27F7377FFC6FFD1FD8AFC54FDC8FC4A" box="[550,616,941,966]" pageId="6" pageNumber="216" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3224195FFC6FFDCFE2DFC29FE4BFC42" lastPageId="11" lastPageNumber="221" pageId="6" pageNumber="216" type="description">
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<paragraph id="8B87121EFFC6FFD1FE2DFC29FDB7FC65" blockId="6.[304,616,941,1001]" box="[385,535,976,1001]" pageId="6" pageNumber="216">
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<emphasis id="B94CCE0CFFC6FFD1FE2DFC29FDB7FC65" bold="true" box="[385,535,976,1001]" pageId="6" pageNumber="216">
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(
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<figureCitation id="13030E9BFFC6FFD1FE24FC29FDAFFC65" box="[392,527,976,1001]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="7.[159,171,1799,1818]" captionStartId-1="8.[159,171,1264,1283]" captionStartId-2="10.[159,171,1149,1168]" captionTargetBox-0="[252,1342,318,1726]" captionTargetBox-1="[262,1342,323,1193]" captionTargetBox-2="[248,1329,313,1101]" captionTargetId-0="figure-6@7.[242,1345,313,1767]" captionTargetId-1="figure-3@8.[242,1345,302,1232]" captionTargetId-2="figure-3@10.[242,1345,302,1117]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="10" captionText-0="FIGURE 3: Holoparasitus fanes n. sp., female: A – Gnathotectum; B – Chelicera, antiaxial aspect and fixed digit in the lateroventral perspective; C – Praesternal plate with the lateral platelets; D – Sternal shield; E – Paragynial shields and apex of epigynium; F – Epigynium; G – Central epigynial prong with a subapical structure; H – Endogynium between paragynial adaxial borders; I – Endogynium. A,C,I: slide no. 1713; B: slide no. 1714, fixed digit slide no. 2002; D: slide no. 1641; E,F: slide no. 1620, holotype; G,H: slide no. 1714. Scale bars: 0.1 mm." captionText-1="FIGURE 4: Holoparasitus fanes n. sp., male: A – Gnathotectum, different aspects; B – Gnathosoma, ventrally; C – Chelicera, adaxial aspect; D – Chelicerae tips in ventral perspective, arrows indicate orifices at the base of minute pilus dentilis; E – Basal part of cheliceral digits, antiaxial aspect; F – Tip of fixed digit showing pilus dentilis; G – Pedipalpal trochanter anterolaterally, seta v1 also in ventral perspective; H – Trochanter I, arrow on anterolateral side. A: slides no. 1715, 1716, 2182; B,C: slide no. 2000; D-F: slide no. 1715; G: slide no. 1716, v1 slide no. 2000; H: slide no. 2182. Scale bars: A-E,G,H – 0.1 mm, F – 0.05 mm." captionText-2="FIGURE 5: Holoparasitus fanes n. sp., male: A – Sternal shield; B – Genital lamina; C – Leg II (Fe, Ge, Ti), anterolateral aspect; D – ibid., ventral aspect, arrow on anterolateral side; E – Leg II (Ge, Ti), ventral aspect, arrow on anterolateral side. A: slide no. 2001; B: slide no. 2179; C: slide no. 1640; D: slide no. 2002; E: slide no. 2000. Scale bars: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4640252" figureDoi-1="http://doi.org/10.5281/zenodo.4640254" figureDoi-2="http://doi.org/10.5281/zenodo.4640256" httpUri-0="https://zenodo.org/record/4640252/files/figure.png" httpUri-1="https://zenodo.org/record/4640254/files/figure.png" httpUri-2="https://zenodo.org/record/4640256/files/figure.png" pageId="6" pageNumber="216">Figures 3-5</figureCitation>
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)
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</emphasis>
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</paragraph>
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<paragraph id="8B87121EFFC6FFD1FF33FBEBFDFBFBAE" blockId="6.[159,762,1041,1948]" box="[159,603,1041,1059]" pageId="6" pageNumber="216">
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Zoobank:
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<uuid id="FF9E28CBFFC6FFD1FF5FFBE8FDFBFBAE" box="[243,603,1041,1058]" pageId="6" pageNumber="216">5326BC8D-B3D4-450C-B686-7F7B4F836AA0</uuid>
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</paragraph>
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<paragraph id="8B87121EFFC6FFD1FF69FBC2FA21FBDE" blockId="6.[159,762,1041,1948]" lastBlockId="6.[826,1429,311,1949]" pageId="6" pageNumber="216">
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Diagnosis — In both sexes gland pores
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<emphasis id="B94CCE0CFFC6FFD1FD7EFBC2FD59FBDF" box="[722,761,1083,1107]" italics="true" pageId="6" pageNumber="216">gv1</emphasis>
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are present, and pores
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<emphasis id="B94CCE0CFFC6FFD1FE15FBA7FE41FBFA" box="[441,481,1118,1142]" italics="true" pageId="6" pageNumber="216">gv2</emphasis>
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are located in unmodified flat cuticle. In the female, the gnathotectum is trispinate, presternal plate is ribbon-shaped, only slightly narrowed medially, with obliquely cut ends, no denticles or corrugation are present; lateral platelets free and wedge-shaped; sternal plate reticulation pronounced, axial granular or reticulated band absent, anterior sternal margin concave between
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<emphasis id="B94CCE0CFFC6FFD1FEABFA8EFE88FA03" box="[263,296,1399,1423]" italics="true" pageId="6" pageNumber="216">st1</emphasis>
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setae and frequently with small concavities laterally to sternal pores
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<emphasis id="B94CCE0CFFC6FFD1FDAAFA63FD8FFA3E" box="[518,559,1434,1458]" italics="true" pageId="6" pageNumber="216">iv1;</emphasis>
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posterior paragynial lobes in close proximity, metagynial sclerite heavily sclerotised and located close to adaxial edge of paragynium, its antiaxial margin straight whereas the adaxial margin forms a rounded protrusion matching the concavities in the anterior subapical thickening of the epigynium; epigynial central prong much wider than the subapical structure, which is narrow and anteriorly tripartite, with lateral hyaline protrusions rounded and extending slightly beyond the central prong margins; endogynium very small in ventral perspective, with heavily sclerotised roundish structure located centrally, and the lateral parts bearing minute denticles; anteriorly, endogynium forming a lamellar protru- sion. In the male, the gnathotectum the solid central prong is rounded apically, lateral prongs absent; corniculi conical; hypostome normal; hypognathal groove clearly visible; chelicera fixed digit straight and edentate with a characteristic lamellar protrusion laterally, movable digit solid and curved adaxially, bearing one dent; pilus dentilis minute, settled in orifice in lateral protrusion of fixed digit; palpcoxal setae scarcely pectinate and hypostomatics simple; palp trochanter with stout simple seta
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<emphasis id="B94CCE0CFFC6FFD1FC96FD6FFCF5FD22" box="[826,853,662,686]" italics="true" pageId="6" pageNumber="216">v1</emphasis>
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and longer seta
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<emphasis id="B94CCE0CFFC6FFD1FB8BFD6FFBE2FD22" box="[1063,1090,662,686]" italics="true" pageId="6" pageNumber="216">v2</emphasis>
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barbed terminally, both setae located in close proximity and separated by a prominence; palp femur with ventral tubercle near the anterolateral seta; sternal shield with a regular reticulation except the roundish area between coxa II and coxa III, where the reticulation is weak and the cuticle porous; genital lamina rounded, located in a concavity of the sternal margin, and flanked by the sclerotised prominences; apical segment of femur II main spur circular in the ventral perspective, axillar process wedge-like, spur on genu II moderate, the one on tibia II – low, roundish and broad; genu II extended anterolaterally in the distal part.
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</paragraph>
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<paragraph id="8B87121EFFC6FFD1FCCCFB9FFC49FBF2" blockId="6.[826,1429,311,1949]" box="[864,1001,1126,1150]" pageId="6" pageNumber="216">Description</paragraph>
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<paragraph id="8B87121EFFC6FFD1FCCCFB6BFA86FABA" blockId="6.[826,1429,311,1949]" pageId="6" pageNumber="216">
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Female — Idiosoma brownish, 475-495 x
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<quantity id="4CC0BFFBFFC6FFD1FA90FB6BFCC0FB41" metricMagnitude="-1" metricUnit="m" metricValue="6.725" metricValueMax="6.75" metricValueMin="6.7" pageId="6" pageNumber="216" unit="mm" value="672.5" valueMax="675.0" valueMin="670.0">670-675 µm</quantity>
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(width range x length range) (n=5). Length of podonotal setae:
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<quantity id="4CC0BFFBFFC6FFD1FBAEFB21FBCCFB7C" box="[1026,1132,1240,1265]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="6" pageNumber="216" unit="mm" value="28.0" valueMax="30.0" valueMin="26.0">26-30 µm</quantity>
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(
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<emphasis id="B94CCE0CFFC6FFD1FBD7FB20FB30FB7D" box="[1147,1168,1241,1265]" italics="true" pageId="6" pageNumber="216">j1</emphasis>
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),
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<quantity id="4CC0BFFBFFC6FFD1FB09FB21FAB0FB7C" box="[1189,1296,1240,1265]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="6" pageNumber="216" unit="mm" value="25.5" valueMax="26.0" valueMin="25.0">25-26 µm</quantity>
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(
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<emphasis id="B94CCE0CFFC6FFD1FAB2FB20FA96FB7D" box="[1310,1334,1241,1265]" italics="true" pageId="6" pageNumber="216">r3</emphasis>
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), others
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<quantity id="4CC0BFFBFFC6FFD1FC96FB05FC09FA9F" box="[826,937,1275,1300]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="6" pageNumber="216" unit="mm" value="15.5" valueMax="17.0" valueMin="14.0">14-17 µm</quantity>
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; opisthonotal setae very short,
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<quantity id="4CC0BFFBFFC6FFD1FA80FB05FA2EFA9F" box="[1324,1422,1275,1300]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="6" pageNumber="216" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8-12 µm</quantity>
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; length of peritrematal groove
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<quantity id="4CC0BFFBFFC6FFD1FB35FAE7FABEFABB" box="[1177,1310,1310,1335]" metricMagnitude="-1" metricUnit="m" metricValue="2.12" metricValueMax="2.15" metricValueMin="2.09" pageId="6" pageNumber="216" unit="mm" value="212.0" valueMax="215.0" valueMin="209.0">209-215 µm</quantity>
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.
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</paragraph>
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<paragraph id="8B87121EFFC6FFD1FCCCFAB3FAACF968" blockId="6.[826,1429,311,1949]" pageId="6" pageNumber="216">
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Gnathosoma — Gnathotectum trispinate (
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<figureCitation id="13030E9BFFC6FFD1FAC4FAB3FCFEFA0A" captionStart="FIGURE" captionStartId="7.[159,171,1799,1818]" captionTargetBox="[252,1342,318,1726]" captionTargetId="figure-6@7.[242,1345,313,1767]" captionTargetPageId="7" captionText="FIGURE 3: Holoparasitus fanes n. sp., female: A – Gnathotectum; B – Chelicera, antiaxial aspect and fixed digit in the lateroventral perspective; C – Praesternal plate with the lateral platelets; D – Sternal shield; E – Paragynial shields and apex of epigynium; F – Epigynium; G – Central epigynial prong with a subapical structure; H – Endogynium between paragynial adaxial borders; I – Endogynium. A,C,I: slide no. 1713; B: slide no. 1714, fixed digit slide no. 2002; D: slide no. 1641; E,F: slide no. 1620, holotype; G,H: slide no. 1714. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4640252" httpUri="https://zenodo.org/record/4640252/files/figure.png" pageId="6" pageNumber="216">Fig. 3A</figureCitation>
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), corniculi conical. Hypognathal groove narrow, with
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<emphasis id="B94CCE0CFFC6FFD1FC1FFA68FC6AFA24" box="[947,970,1425,1448]" italics="true" pageId="6" pageNumber="216">ca</emphasis>
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. ten weakly pronounced rows of denticles. Palpcoxal and hypostomatic setae simple. Chelicera (
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<figureCitation id="13030E9BFFC6FFD1FC1AFA2EFBABFA63" box="[950,1035,1495,1519]" captionStart="FIGURE" captionStartId="7.[159,171,1799,1818]" captionTargetBox="[252,1342,318,1726]" captionTargetId="figure-6@7.[242,1345,313,1767]" captionTargetPageId="7" captionText="FIGURE 3: Holoparasitus fanes n. sp., female: A – Gnathotectum; B – Chelicera, antiaxial aspect and fixed digit in the lateroventral perspective; C – Praesternal plate with the lateral platelets; D – Sternal shield; E – Paragynial shields and apex of epigynium; F – Epigynium; G – Central epigynial prong with a subapical structure; H – Endogynium between paragynial adaxial borders; I – Endogynium. A,C,I: slide no. 1713; B: slide no. 1714, fixed digit slide no. 2002; D: slide no. 1641; E,F: slide no. 1620, holotype; G,H: slide no. 1714. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4640252" httpUri="https://zenodo.org/record/4640252/files/figure.png" pageId="6" pageNumber="216">Fig. 3B</figureCitation>
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). Fixed digit with two denticles in front of pilus dentilis, one larger and lamellar, covering pilus dentilis, and two small denticles behind pilus dentilis; the latter are located somewhat below the digit edge, which is lamellar and arcuate. The movable digit bears three teeth, pedipalp trochanter with seta
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<emphasis id="B94CCE0CFFC6FFD1FC03F950FC6AF94D" box="[943,970,1705,1729]" italics="true" pageId="6" pageNumber="216">v1</emphasis>
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simple,
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<emphasis id="B94CCE0CFFC6FFD1FB9CF950FBEBF94D" box="[1072,1099,1705,1729]" italics="true" pageId="6" pageNumber="216">v2</emphasis>
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longer and barbed distally; femur and genu setation unremarkable.
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</paragraph>
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<paragraph id="8B87121EFFC6FFDFFCCCF901FE3BF912" blockId="6.[826,1429,311,1949]" lastBlockId="8.[159,762,1459,1948]" lastPageId="8" lastPageNumber="218" pageId="6" pageNumber="216">
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Ventral idiosoma — Presternal plate (
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<figureCitation id="13030E9BFFC6FFD1FA86F901FA2CF89C" box="[1322,1420,1784,1808]" captionStart="FIGURE" captionStartId="7.[159,171,1799,1818]" captionTargetBox="[252,1342,318,1726]" captionTargetId="figure-6@7.[242,1345,313,1767]" captionTargetPageId="7" captionText="FIGURE 3: Holoparasitus fanes n. sp., female: A – Gnathotectum; B – Chelicera, antiaxial aspect and fixed digit in the lateroventral perspective; C – Praesternal plate with the lateral platelets; D – Sternal shield; E – Paragynial shields and apex of epigynium; F – Epigynium; G – Central epigynial prong with a subapical structure; H – Endogynium between paragynial adaxial borders; I – Endogynium. A,C,I: slide no. 1713; B: slide no. 1714, fixed digit slide no. 2002; D: slide no. 1641; E,F: slide no. 1620, holotype; G,H: slide no. 1714. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4640252" httpUri="https://zenodo.org/record/4640252/files/figure.png" pageId="6" pageNumber="216">Fig. 3C</figureCitation>
|
||
) ribbon-shaped with obliquely cut ends and only slightly narrowed medially, and with a sinuous anterior margin, no denticles or corrugation present. Lateral platelets are free and wedge-shaped. Ster- nal shield (
|
||
<figureCitation id="13030E9BFFC8FFDFFE8AFA4AFE26FA47" box="[294,390,1459,1483]" captionStart="FIGURE" captionStartId="7.[159,171,1799,1818]" captionTargetBox="[252,1342,318,1726]" captionTargetId="figure-6@7.[242,1345,313,1767]" captionTargetPageId="7" captionText="FIGURE 3: Holoparasitus fanes n. sp., female: A – Gnathotectum; B – Chelicera, antiaxial aspect and fixed digit in the lateroventral perspective; C – Praesternal plate with the lateral platelets; D – Sternal shield; E – Paragynial shields and apex of epigynium; F – Epigynium; G – Central epigynial prong with a subapical structure; H – Endogynium between paragynial adaxial borders; I – Endogynium. A,C,I: slide no. 1713; B: slide no. 1714, fixed digit slide no. 2002; D: slide no. 1641; E,F: slide no. 1620, holotype; G,H: slide no. 1714. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4640252" httpUri="https://zenodo.org/record/4640252/files/figure.png" pageId="8" pageNumber="218">Fig. 3D</figureCitation>
|
||
) anterior margin with shallow concavity between
|
||
<emphasis id="B94CCE0CFFC8FFDFFE2DFA2FFE02FA62" box="[385,418,1494,1518]" italics="true" pageId="8" pageNumber="218">st1</emphasis>
|
||
setae and usually with small concavities laterally to
|
||
<emphasis id="B94CCE0CFFC8FFDFFE05FA00FE6BF99D" box="[425,459,1529,1553]" italics="true" pageId="8" pageNumber="218">iv1</emphasis>
|
||
sternal pores, reticulation well developed. Pores
|
||
<emphasis id="B94CCE0CFFC8FFDFFE12F9E4FE46F9B9" box="[446,486,1565,1589]" italics="true" pageId="8" pageNumber="218">gv1</emphasis>
|
||
on sternal shield near posterior sternal margin. Setae of sternogenital region:
|
||
<quantity id="4CC0BFFBFFC8FFDFFF49F99AFEC5F9F7" box="[229,357,1634,1659]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.6" metricValueMin="3.9" pageId="8" pageNumber="218" unit="mm" value="42.5" valueMax="46.0" valueMin="39.0">39 – 46 µm</quantity>
|
||
(
|
||
<emphasis id="B94CCE0CFFC8FFDFFEDAF99AFE29F9F6" box="[374,393,1635,1658]" italics="true" pageId="8" pageNumber="218">st</emphasis>
|
||
1),
|
||
<quantity id="4CC0BFFBFFC8FFDFFE03F99BFD8FF9F7" box="[431,559,1634,1659]" metricMagnitude="-2" metricUnit="m" metricValue="5.45" metricValueMax="5.9" metricValueMin="5.0" pageId="8" pageNumber="218" unit="mm" value="54.5" valueMax="59.0" valueMin="50.0">50 – 59 µm</quantity>
|
||
(
|
||
<emphasis id="B94CCE0CFFC8FFDFFDECF99AFDC1F9F7" box="[576,609,1635,1659]" italics="true" pageId="8" pageNumber="218">st2</emphasis>
|
||
),
|
||
<quantity id="4CC0BFFBFFC8FFDFFDD5F99AFD59F9F7" box="[633,761,1634,1659]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="5.1" metricValueMin="4.4" pageId="8" pageNumber="218" unit="mm" value="47.5" valueMax="51.0" valueMin="44.0">44 – 51 µm</quantity>
|
||
(
|
||
<emphasis id="B94CCE0CFFC8FFDFFF04F97FFF1BF911" box="[168,187,1670,1693]" italics="true" pageId="8" pageNumber="218">st</emphasis>
|
||
3),
|
||
<quantity id="4CC0BFFBFFC8FFDFFF72F97FFEFBF912" box="[222,347,1670,1694]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="3.9" metricValueMin="3.3" pageId="8" pageNumber="218" unit="mm" value="36.0" valueMax="39.0" valueMin="33.0">33 – 39 µm</quantity>
|
||
(
|
||
<emphasis id="B94CCE0CFFC8FFDFFEC7F97FFE2BF912" box="[363,395,1670,1694]" italics="true" pageId="8" pageNumber="218">st4</emphasis>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF474296FFC7FFD0FF33F8FEFDA1F805" ID-DOI="http://doi.org/10.5281/zenodo.4640252" ID-Zenodo-Dep="4640252" httpUri="https://zenodo.org/record/4640252/files/figure.png" pageId="7" pageNumber="217" startId="7.[159,171,1799,1818]" targetBox="[252,1342,318,1726]" targetPageId="7">
|
||
<paragraph id="8B87121EFFC7FFD0FF33F8FEFDA1F805" blockId="7.[159,1430,1798,1929]" pageId="7" pageNumber="217">
|
||
FIGURE 3:
|
||
<taxonomicName id="4C38699DFFC7FFD0FEA0F8FFFE19F895" authority="Witalinski, 2017" authorityName="Witalinski" authorityYear="2017" box="[268,441,1798,1817]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="217" phylum="Arthropoda" rank="species" species="fanes" status="sp. nov.">
|
||
<emphasis id="B94CCE0CFFC7FFD0FEA0F8FFFE19F895" box="[268,441,1798,1817]" italics="true" pageId="7" pageNumber="217">Holoparasitus fanes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94CCE0CFFC7FFD0FE6CF8FEFE5EF897" bold="true" box="[448,510,1799,1819]" pageId="7" pageNumber="217">
|
||
<taxonomicNameLabel id="A27F7377FFC7FFD0FE6CF8FEFE5EF897" box="[448,510,1799,1819]" pageId="7" pageNumber="217" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female: A – Gnathotectum; B – Chelicera, antiaxial aspect and fixed digit in the lateroventral perspective; C – Praesternal plate with the lateral platelets; D – Sternal shield; E – Paragynial shields and apex of epigynium; F – Epigynium; G – Central epigynial prong with a subapical structure; H – Endogynium between paragynial adaxial borders; I – Endogynium. A,C,I: slide no. 1713; B: slide no. 1714, fixed digit slide no. 2002; D: slide no. 1641; E,F: slide no. 1620, holotype; G,H: slide no. 1714. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF474296FFC8FFDFFF33FB09FB90FAFE" ID-DOI="http://doi.org/10.5281/zenodo.4640254" ID-Zenodo-Dep="4640254" httpUri="https://zenodo.org/record/4640254/files/figure.png" pageId="8" pageNumber="218" startId="8.[159,171,1264,1283]" targetBox="[262,1342,323,1193]" targetPageId="8">
|
||
<paragraph id="8B87121EFFC8FFDFFF33FB09FB90FAFE" blockId="8.[159,1430,1264,1394]" pageId="8" pageNumber="218">
|
||
FIGURE 4:
|
||
<taxonomicName id="4C38699DFFC8FFDFFEA8FB09FE0DFA8F" authority="Witalinski, 2017" authorityName="Witalinski" authorityYear="2017" box="[260,429,1264,1283]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="218" phylum="Arthropoda" rank="species" species="fanes" status="sp. nov.">
|
||
<emphasis id="B94CCE0CFFC8FFDFFEA8FB09FE0DFA8F" box="[260,429,1264,1283]" italics="true" pageId="8" pageNumber="218">Holoparasitus fanes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94CCE0CFFC8FFDFFE1DFB09FE46FA88" bold="true" box="[433,486,1264,1284]" pageId="8" pageNumber="218">
|
||
<taxonomicNameLabel id="A27F7377FFC8FFDFFE1DFB09FE46FA88" box="[433,486,1264,1284]" pageId="8" pageNumber="218" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male: A – Gnathotectum, different aspects; B – Gnathosoma, ventrally; C – Chelicera, adaxial aspect; D – Chelicerae tips in ventral perspective, arrows indicate orifices at the base of minute pilus dentilis; E – Basal part of cheliceral digits, antiaxial aspect; F – Tip of fixed digit showing pilus dentilis; G – Pedipalpal trochanter anterolaterally, seta
|
||
<emphasis id="B94CCE0CFFC8FFDFFB58FADEFAA9FAB6" box="[1268,1289,1319,1338]" italics="true" pageId="8" pageNumber="218">v1</emphasis>
|
||
also in ventral perspective; H – Trochanter I, arrow on anterolateral side. A: slides no. 1715, 1716, 2182; B,C: slide no. 2000; D-F: slide no. 1715; G: slide no. 1716,
|
||
<emphasis id="B94CCE0CFFC8FFDFFEE0FAA6FEC1FAFE" box="[332,353,1375,1394]" italics="true" pageId="8" pageNumber="218">v1</emphasis>
|
||
slide no. 2000; H: slide no. 2182. Scale bars: A-E,G,H – 0.1 mm, F – 0.05 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B87121EFFC8FFDEFF69F94BFDB7FE19" blockId="8.[159,762,1459,1948]" lastBlockId="9.[159,762,311,1948]" lastPageId="9" lastPageNumber="219" pageId="8" pageNumber="218">
|
||
Genital region — Paragynial shields (
|
||
<figureCitation id="13030E9BFFC8FFDFFD3DF94BFD51F946" box="[657,753,1714,1738]" captionStart="FIGURE" captionStartId="7.[159,171,1799,1818]" captionTargetBox="[252,1342,318,1726]" captionTargetId="figure-6@7.[242,1345,313,1767]" captionTargetPageId="7" captionText="FIGURE 3: Holoparasitus fanes n. sp., female: A – Gnathotectum; B – Chelicera, antiaxial aspect and fixed digit in the lateroventral perspective; C – Praesternal plate with the lateral platelets; D – Sternal shield; E – Paragynial shields and apex of epigynium; F – Epigynium; G – Central epigynial prong with a subapical structure; H – Endogynium between paragynial adaxial borders; I – Endogynium. A,C,I: slide no. 1713; B: slide no. 1714, fixed digit slide no. 2002; D: slide no. 1641; E,F: slide no. 1620, holotype; G,H: slide no. 1714. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4640252" httpUri="https://zenodo.org/record/4640252/files/figure.png" pageId="8" pageNumber="218">Fig. 3E</figureCitation>
|
||
) reticulated, posterolateral protrusions well developed, roundish or sometimes asymmetrical. Posterior paragynial lobes in close proximity. Adaxial paragynial edge thickened and metagynial sclerite heavily sclerotised, located close to the adaxial paragynial edge; antiaxial margin of metagynial sclerite straight or slightly curved, adaxial margin forming a rounded protrusion matching concavity in the anterior subapical thickening of the epigynium (
|
||
<figureCitation id="13030E9BFFC8FFDFFC24F9E5FC7EF9B9" box="[904,990,1564,1589]" captionStart="FIGURE" captionStartId="7.[159,171,1799,1818]" captionTargetBox="[252,1342,318,1726]" captionTargetId="figure-6@7.[242,1345,313,1767]" captionTargetPageId="7" captionText="FIGURE 3: Holoparasitus fanes n. sp., female: A – Gnathotectum; B – Chelicera, antiaxial aspect and fixed digit in the lateroventral perspective; C – Praesternal plate with the lateral platelets; D – Sternal shield; E – Paragynial shields and apex of epigynium; F – Epigynium; G – Central epigynial prong with a subapical structure; H – Endogynium between paragynial adaxial borders; I – Endogynium. A,C,I: slide no. 1713; B: slide no. 1714, fixed digit slide no. 2002; D: slide no. 1641; E,F: slide no. 1620, holotype; G,H: slide no. 1714. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4640252" httpUri="https://zenodo.org/record/4640252/files/figure.png" pageId="8" pageNumber="218">Fig. 3E</figureCitation>
|
||
). Thickening facing coxa III is absent. Central prong of epigynium (
|
||
<figureCitation id="13030E9BFFC8FFDFFB22F9C6FB58F9D4" box="[1166,1272,1599,1624]" captionStart="FIGURE" captionStartId="7.[159,171,1799,1818]" captionTargetBox="[252,1342,318,1726]" captionTargetId="figure-6@7.[242,1345,313,1767]" captionTargetPageId="7" captionText="FIGURE 3: Holoparasitus fanes n. sp., female: A – Gnathotectum; B – Chelicera, antiaxial aspect and fixed digit in the lateroventral perspective; C – Praesternal plate with the lateral platelets; D – Sternal shield; E – Paragynial shields and apex of epigynium; F – Epigynium; G – Central epigynial prong with a subapical structure; H – Endogynium between paragynial adaxial borders; I – Endogynium. A,C,I: slide no. 1713; B: slide no. 1714, fixed digit slide no. 2002; D: slide no. 1641; E,F: slide no. 1620, holotype; G,H: slide no. 1714. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4640252" httpUri="https://zenodo.org/record/4640252/files/figure.png" pageId="8" pageNumber="218">Fig. 3F,G</figureCitation>
|
||
) with convex margins, wider than the subapical structure, which is narrow and anteriorly tripartite. Lateral hyaline protrusions are rounded and extend only slightly beyond the margins of the central prong. Endogynium (
|
||
<figureCitation id="13030E9BFFC8FFDFFC30F916FBB7F88B" box="[924,1047,1775,1799]" captionStart="FIGURE" captionStartId="7.[159,171,1799,1818]" captionTargetBox="[252,1342,318,1726]" captionTargetId="figure-6@7.[242,1345,313,1767]" captionTargetPageId="7" captionText="FIGURE 3: Holoparasitus fanes n. sp., female: A – Gnathotectum; B – Chelicera, antiaxial aspect and fixed digit in the lateroventral perspective; C – Praesternal plate with the lateral platelets; D – Sternal shield; E – Paragynial shields and apex of epigynium; F – Epigynium; G – Central epigynial prong with a subapical structure; H – Endogynium between paragynial adaxial borders; I – Endogynium. A,C,I: slide no. 1713; B: slide no. 1714, fixed digit slide no. 2002; D: slide no. 1641; E,F: slide no. 1620, holotype; G,H: slide no. 1714. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4640252" httpUri="https://zenodo.org/record/4640252/files/figure.png" pageId="8" pageNumber="218">Fig. 3H,I</figureCitation>
|
||
) is small, with heavily sclerotised roundish structure located centrally and with the lateral, less sclerotised parts bearing minute denticles; anteriorly, endogynium forms a lamellar tongue-like protrusion directed anteriorly and frequently indented apically. Gland pores
|
||
<emphasis id="B94CCE0CFFC9FFDEFD2CFECEFD08FEC3" box="[640,680,311,335]" italics="true" pageId="9" pageNumber="219">
|
||
gv
|
||
<quantity id="4CC0BFFBFFC9FFDEFD37FECEFD08FEC3" box="[667,680,311,335]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="9" pageNumber="219" unit="in" value="2.0">2</quantity>
|
||
</emphasis>
|
||
in unmodified flat cuticle. Opisthogaster with eight pairs of ventral setae
|
||
<quantity id="4CC0BFFBFFC9FFDEFEF4FE84FE75FE19" box="[344,469,381,405]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="4.2" metricValueMin="1.5" pageId="9" pageNumber="219" unit="mm" value="28.5" valueMax="42.0" valueMin="15.0">15 – 42 µm</quantity>
|
||
long.
|
||
</paragraph>
|
||
<paragraph id="8B87121EFFC9FFDEFF69FE51FF76FE6E" blockId="9.[159,762,311,1948]" pageId="9" pageNumber="219">Legs — Leg structure and setation unremarkable.</paragraph>
|
||
<paragraph id="8B87121EFFC9FFDEFF69FE0DFE35FD14" blockId="9.[159,762,311,1948]" pageId="9" pageNumber="219">
|
||
Male — Idiosoma brownish, 387 – 425 × 580 – 640 (n=5). Length of podonotal setae:
|
||
<quantity id="4CC0BFFBFFC9FFDEFDD8FDEEFD59FDA3" box="[628,761,535,560]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="9" pageNumber="219" unit="mm" value="27.5" valueMax="29.0" valueMin="26.0">26 – 29 µm</quantity>
|
||
(
|
||
<emphasis id="B94CCE0CFFC9FFDEFF04FDC2FF1CFDDF" box="[168,188,571,595]" italics="true" pageId="9" pageNumber="219">j1</emphasis>
|
||
),
|
||
<quantity id="4CC0BFFBFFC9FFDEFF7AFDC3FEFBFDDF" box="[214,347,570,595]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="9" pageNumber="219" unit="mm" value="17.0" valueMax="18.0" valueMin="16.0">16 – 18 µm</quantity>
|
||
(
|
||
<emphasis id="B94CCE0CFFC9FFDEFEC1FDC2FE25FDDF" box="[365,389,571,595]" italics="true" pageId="9" pageNumber="219">r3</emphasis>
|
||
), others
|
||
<quantity id="4CC0BFFBFFC9FFDEFE5DFDC2FDC9FDDF" box="[497,617,571,595]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.4" metricValueMin="0.9" pageId="9" pageNumber="219" unit="mm" value="11.5" valueMax="14.0" valueMin="9.0">9 – 14 µm</quantity>
|
||
; opisthonotal setae short,
|
||
<quantity id="4CC0BFFBFFC9FFDEFEF5FDA7FE40FDFA" box="[345,480,606,630]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="9" pageNumber="219" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10 – 12 µm</quantity>
|
||
; length of peritrematal groove
|
||
<quantity id="4CC0BFFBFFC9FFDEFF5BFD78FE2EFD15" box="[247,398,641,665]" metricMagnitude="-1" metricUnit="m" metricValue="2.03" metricValueMax="2.07" metricValueMin="1.9899999999999998" pageId="9" pageNumber="219" unit="mm" value="203.0" valueMax="207.0" valueMin="199.0">199 – 207 µm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B87121EFFC9FFDEFF69FD52FE4FFAD1" blockId="9.[159,762,311,1948]" pageId="9" pageNumber="219">
|
||
Gnathosoma — Gnathotectum with a solid central prong rounded apically, lateral prongs absent, gnathotectum margins convex (
|
||
<figureCitation id="13030E9BFFC9FFDEFDBBFD08FDD7FC85" box="[535,631,753,777]" captionStart="FIGURE" captionStartId="8.[159,171,1264,1283]" captionTargetBox="[262,1342,323,1193]" captionTargetId="figure-3@8.[242,1345,302,1232]" captionTargetPageId="8" captionText="FIGURE 4: Holoparasitus fanes n. sp., male: A – Gnathotectum, different aspects; B – Gnathosoma, ventrally; C – Chelicera, adaxial aspect; D – Chelicerae tips in ventral perspective, arrows indicate orifices at the base of minute pilus dentilis; E – Basal part of cheliceral digits, antiaxial aspect; F – Tip of fixed digit showing pilus dentilis; G – Pedipalpal trochanter anterolaterally, seta v1 also in ventral perspective; H – Trochanter I, arrow on anterolateral side. A: slides no. 1715, 1716, 2182; B,C: slide no. 2000; D-F: slide no. 1715; G: slide no. 1716, v1 slide no. 2000; H: slide no. 2182. Scale bars: A-E,G,H – 0.1 mm, F – 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4640254" httpUri="https://zenodo.org/record/4640254/files/figure.png" pageId="9" pageNumber="219">Fig. 4A</figureCitation>
|
||
). Corniculi (
|
||
<figureCitation id="13030E9BFFC9FFDEFF0BFCEDFF5DFCA0" box="[167,253,788,812]" captionStart="FIGURE" captionStartId="8.[159,171,1264,1283]" captionTargetBox="[262,1342,323,1193]" captionTargetId="figure-3@8.[242,1345,302,1232]" captionTargetPageId="8" captionText="FIGURE 4: Holoparasitus fanes n. sp., male: A – Gnathotectum, different aspects; B – Gnathosoma, ventrally; C – Chelicera, adaxial aspect; D – Chelicerae tips in ventral perspective, arrows indicate orifices at the base of minute pilus dentilis; E – Basal part of cheliceral digits, antiaxial aspect; F – Tip of fixed digit showing pilus dentilis; G – Pedipalpal trochanter anterolaterally, seta v1 also in ventral perspective; H – Trochanter I, arrow on anterolateral side. A: slides no. 1715, 1716, 2182; B,C: slide no. 2000; D-F: slide no. 1715; G: slide no. 1716, v1 slide no. 2000; H: slide no. 2182. Scale bars: A-E,G,H – 0.1 mm, F – 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4640254" httpUri="https://zenodo.org/record/4640254/files/figure.png" pageId="9" pageNumber="219">Fig. 4B</figureCitation>
|
||
) conical. Hypostome normal, hypognathal groove rather wide, and ca. 10 rows of denticles moderately visible. Palpcoxal setae scarcely pectinate, hypostomal setae simple. Chelicera (
|
||
<figureCitation id="13030E9BFFC9FFDEFD3BFC84FD7FFC19" box="[663,735,893,917]" captionStart="FIGURE" captionStartId="8.[159,171,1264,1283]" captionTargetBox="[262,1342,323,1193]" captionTargetId="figure-3@8.[242,1345,302,1232]" captionTargetPageId="8" captionText="FIGURE 4: Holoparasitus fanes n. sp., male: A – Gnathotectum, different aspects; B – Gnathosoma, ventrally; C – Chelicera, adaxial aspect; D – Chelicerae tips in ventral perspective, arrows indicate orifices at the base of minute pilus dentilis; E – Basal part of cheliceral digits, antiaxial aspect; F – Tip of fixed digit showing pilus dentilis; G – Pedipalpal trochanter anterolaterally, seta v1 also in ventral perspective; H – Trochanter I, arrow on anterolateral side. A: slides no. 1715, 1716, 2182; B,C: slide no. 2000; D-F: slide no. 1715; G: slide no. 1716, v1 slide no. 2000; H: slide no. 2182. Scale bars: A-E,G,H – 0.1 mm, F – 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4640254" httpUri="https://zenodo.org/record/4640254/files/figure.png" pageId="9" pageNumber="219">Fig. 4</figureCitation>
|
||
C- F). Fixed digit straight and edentate with a characteristic lateral lamellar protrusion showing a small orifice at the base of minute pilus dentilis (
|
||
<figureCitation id="13030E9BFFC9FFDEFD61FC1FFF79FBAE" captionStart="FIGURE" captionStartId="8.[159,171,1264,1283]" captionTargetBox="[262,1342,323,1193]" captionTargetId="figure-3@8.[242,1345,302,1232]" captionTargetPageId="8" captionText="FIGURE 4: Holoparasitus fanes n. sp., male: A – Gnathotectum, different aspects; B – Gnathosoma, ventrally; C – Chelicera, adaxial aspect; D – Chelicerae tips in ventral perspective, arrows indicate orifices at the base of minute pilus dentilis; E – Basal part of cheliceral digits, antiaxial aspect; F – Tip of fixed digit showing pilus dentilis; G – Pedipalpal trochanter anterolaterally, seta v1 also in ventral perspective; H – Trochanter I, arrow on anterolateral side. A: slides no. 1715, 1716, 2182; B,C: slide no. 2000; D-F: slide no. 1715; G: slide no. 1716, v1 slide no. 2000; H: slide no. 2182. Scale bars: A-E,G,H – 0.1 mm, F – 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4640254" httpUri="https://zenodo.org/record/4640254/files/figure.png" pageId="9" pageNumber="219">Fig. 4D,F</figureCitation>
|
||
). The protrusion is clearly visible in the ventral perspective (
|
||
<figureCitation id="13030E9BFFC9FFDEFECBFBD4FE67FBC9" box="[359,455,1069,1093]" captionStart="FIGURE" captionStartId="8.[159,171,1264,1283]" captionTargetBox="[262,1342,323,1193]" captionTargetId="figure-3@8.[242,1345,302,1232]" captionTargetPageId="8" captionText="FIGURE 4: Holoparasitus fanes n. sp., male: A – Gnathotectum, different aspects; B – Gnathosoma, ventrally; C – Chelicera, adaxial aspect; D – Chelicerae tips in ventral perspective, arrows indicate orifices at the base of minute pilus dentilis; E – Basal part of cheliceral digits, antiaxial aspect; F – Tip of fixed digit showing pilus dentilis; G – Pedipalpal trochanter anterolaterally, seta v1 also in ventral perspective; H – Trochanter I, arrow on anterolateral side. A: slides no. 1715, 1716, 2182; B,C: slide no. 2000; D-F: slide no. 1715; G: slide no. 1716, v1 slide no. 2000; H: slide no. 2182. Scale bars: A-E,G,H – 0.1 mm, F – 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4640254" httpUri="https://zenodo.org/record/4640254/files/figure.png" pageId="9" pageNumber="219">Fig. 4D</figureCitation>
|
||
). Movable digit solid and curved adaxially; bearing one tooth, in some specimens followed distally by 1-2 minute denticles. Pedipalp trochanter (
|
||
<figureCitation id="13030E9BFFC9FFDEFE30FB6FFDB3FB22" box="[412,531,1174,1198]" captionStart="FIGURE" captionStartId="8.[159,171,1264,1283]" captionTargetBox="[262,1342,323,1193]" captionTargetId="figure-3@8.[242,1345,302,1232]" captionTargetPageId="8" captionText="FIGURE 4: Holoparasitus fanes n. sp., male: A – Gnathotectum, different aspects; B – Gnathosoma, ventrally; C – Chelicera, adaxial aspect; D – Chelicerae tips in ventral perspective, arrows indicate orifices at the base of minute pilus dentilis; E – Basal part of cheliceral digits, antiaxial aspect; F – Tip of fixed digit showing pilus dentilis; G – Pedipalpal trochanter anterolaterally, seta v1 also in ventral perspective; H – Trochanter I, arrow on anterolateral side. A: slides no. 1715, 1716, 2182; B,C: slide no. 2000; D-F: slide no. 1715; G: slide no. 1716, v1 slide no. 2000; H: slide no. 2182. Scale bars: A-E,G,H – 0.1 mm, F – 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4640254" httpUri="https://zenodo.org/record/4640254/files/figure.png" pageId="9" pageNumber="219">Fig. 4B,G</figureCitation>
|
||
) with stout seta
|
||
<emphasis id="B94CCE0CFFC9FFDEFD73FB6FFD5AFB22" box="[735,762,1174,1198]" italics="true" pageId="9" pageNumber="219">v1</emphasis>
|
||
and longer seta
|
||
<emphasis id="B94CCE0CFFC9FFDEFEF7FB40FED6FB5D" box="[347,374,1209,1233]" italics="true" pageId="9" pageNumber="219">v2</emphasis>
|
||
barbed terminally, both setae located in close proximity and separated by a small prominence. Pedipalp femur (
|
||
<figureCitation id="13030E9BFFC9FFDEFDA8FB06FDFAFA9B" box="[516,602,1279,1303]" captionStart="FIGURE" captionStartId="8.[159,171,1264,1283]" captionTargetBox="[262,1342,323,1193]" captionTargetId="figure-3@8.[242,1345,302,1232]" captionTargetPageId="8" captionText="FIGURE 4: Holoparasitus fanes n. sp., male: A – Gnathotectum, different aspects; B – Gnathosoma, ventrally; C – Chelicera, adaxial aspect; D – Chelicerae tips in ventral perspective, arrows indicate orifices at the base of minute pilus dentilis; E – Basal part of cheliceral digits, antiaxial aspect; F – Tip of fixed digit showing pilus dentilis; G – Pedipalpal trochanter anterolaterally, seta v1 also in ventral perspective; H – Trochanter I, arrow on anterolateral side. A: slides no. 1715, 1716, 2182; B,C: slide no. 2000; D-F: slide no. 1715; G: slide no. 1716, v1 slide no. 2000; H: slide no. 2182. Scale bars: A-E,G,H – 0.1 mm, F – 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4640254" httpUri="https://zenodo.org/record/4640254/files/figure.png" pageId="9" pageNumber="219">Fig. 4B</figureCitation>
|
||
) with ventral tubercle near the anterolateral seta; both femur and genu setation unremarkable.
|
||
</paragraph>
|
||
<paragraph id="8B87121EFFC9FFDEFF69FA96FD63F8A0" blockId="9.[159,762,311,1948]" pageId="9" pageNumber="219">
|
||
Ventral idiosoma — Presternal platelets small and roughly triangular (
|
||
<figureCitation id="13030E9BFFC9FFDEFE65FA6BFD90FA26" box="[457,560,1426,1450]" captionStart="FIGURE" captionStartId="10.[159,171,1149,1168]" captionTargetBox="[248,1329,313,1101]" captionTargetId="figure-3@10.[242,1345,302,1117]" captionTargetPageId="10" captionText="FIGURE 5: Holoparasitus fanes n. sp., male: A – Sternal shield; B – Genital lamina; C – Leg II (Fe, Ge, Ti), anterolateral aspect; D – ibid., ventral aspect, arrow on anterolateral side; E – Leg II (Ge, Ti), ventral aspect, arrow on anterolateral side. A: slide no. 2001; B: slide no. 2179; C: slide no. 1640; D: slide no. 2002; E: slide no. 2000. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4640256" httpUri="https://zenodo.org/record/4640256/files/figure.png" pageId="9" pageNumber="219">Fig. 5A</figureCitation>
|
||
). Genital lamina (
|
||
<figureCitation id="13030E9BFFC9FFDEFF0BFA4CFEAFFA41" box="[167,271,1461,1485]" captionStart="FIGURE" captionStartId="10.[159,171,1149,1168]" captionTargetBox="[248,1329,313,1101]" captionTargetId="figure-3@10.[242,1345,302,1117]" captionTargetPageId="10" captionText="FIGURE 5: Holoparasitus fanes n. sp., male: A – Sternal shield; B – Genital lamina; C – Leg II (Fe, Ge, Ti), anterolateral aspect; D – ibid., ventral aspect, arrow on anterolateral side; E – Leg II (Ge, Ti), ventral aspect, arrow on anterolateral side. A: slide no. 2001; B: slide no. 2179; C: slide no. 1640; D: slide no. 2002; E: slide no. 2000. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4640256" httpUri="https://zenodo.org/record/4640256/files/figure.png" pageId="9" pageNumber="219">Fig. 5B</figureCitation>
|
||
) roundish with sclerotised margin and heavily sclerotised subgenital microsclerite. The lamina is located in the concavity of the anterior sternal margin, flanked by the sclerotised tubercles on both sides of the concavity. Reticulation of the sternogenital shield (
|
||
<figureCitation id="13030E9BFFC9FFDEFE37F99CFE5CF9F1" box="[411,508,1637,1661]" captionStart="FIGURE" captionStartId="10.[159,171,1149,1168]" captionTargetBox="[248,1329,313,1101]" captionTargetId="figure-3@10.[242,1345,302,1117]" captionTargetPageId="10" captionText="FIGURE 5: Holoparasitus fanes n. sp., male: A – Sternal shield; B – Genital lamina; C – Leg II (Fe, Ge, Ti), anterolateral aspect; D – ibid., ventral aspect, arrow on anterolateral side; E – Leg II (Ge, Ti), ventral aspect, arrow on anterolateral side. A: slide no. 2001; B: slide no. 2179; C: slide no. 1640; D: slide no. 2002; E: slide no. 2000. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4640256" httpUri="https://zenodo.org/record/4640256/files/figure.png" pageId="9" pageNumber="219">Fig. 5A</figureCitation>
|
||
) clearly visible laterally, but in central part between coxa II and coxa III weakly pronounced, and the sternal cuticle is porous. Excipulum absent. Sternal setae length:
|
||
<quantity id="4CC0BFFBFFC9FFDEFD73F937FF59F885" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="3.9" metricValueMin="3.4" pageId="9" pageNumber="219" unit="mm" value="36.5" valueMax="39.0" valueMin="34.0">34 – 39 µm</quantity>
|
||
(
|
||
<emphasis id="B94CCE0CFFC9FFDEFEA4F908FE89F885" box="[264,297,1777,1801]" italics="true" pageId="9" pageNumber="219">st1</emphasis>
|
||
),
|
||
<quantity id="4CC0BFFBFFC9FFDEFE92F908FE19F885" box="[318,441,1777,1802]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="9" pageNumber="219" unit="mm" value="31.5" valueMax="33.0" valueMin="30.0">30 – 33 µm</quantity>
|
||
(
|
||
<emphasis id="B94CCE0CFFC9FFDEFE64F908FE49F885" box="[456,489,1777,1801]" italics="true" pageId="9" pageNumber="219">st2</emphasis>
|
||
),
|
||
<quantity id="4CC0BFFBFFC9FFDEFE53F908FDDAF885" box="[511,634,1777,1802]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.5" metricValueMin="3.1" pageId="9" pageNumber="219" unit="mm" value="33.0" valueMax="35.0" valueMin="31.0">31 – 35 µm</quantity>
|
||
(
|
||
<emphasis id="B94CCE0CFFC9FFDEFD25F908FD0AF885" box="[649,682,1777,1801]" italics="true" pageId="9" pageNumber="219">st3</emphasis>
|
||
), setae
|
||
<emphasis id="B94CCE0CFFC9FFDEFF33F8EDFF1FF8A0" box="[159,191,1812,1836]" italics="true" pageId="9" pageNumber="219">st4</emphasis>
|
||
<quantity id="4CC0BFFBFFC9FFDEFF6AF8EDFEE3F8A0" box="[198,323,1812,1837]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="9" pageNumber="219" unit="mm" value="28.5" valueMax="30.0" valueMin="27.0">27 – 30 µm</quantity>
|
||
, opisthogastral setae
|
||
<quantity id="4CC0BFFBFFC9FFDEFDECF8EDFD1CF8A0" box="[576,700,1812,1837]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.7" metricValueMin="2.1" pageId="9" pageNumber="219" unit="mm" value="29.0" valueMax="37.0" valueMin="21.0">21 – 37 µm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B87121EFFC9FFDEFF69F8C6FA99FE72" blockId="9.[159,762,311,1948]" lastBlockId="9.[826,1429,311,1949]" pageId="9" pageNumber="219">
|
||
Legs — Leg I, III, IV largely unremarkable, with simple setae, but anterolateral setae on trochanter I and trochanter II, which are thicker and pectinate distally (
|
||
<figureCitation id="13030E9BFFC9FFDEFC0DFECEFC5FFEC3" box="[929,1023,311,335]" captionStart="FIGURE" captionStartId="8.[159,171,1264,1283]" captionTargetBox="[262,1342,323,1193]" captionTargetId="figure-3@8.[242,1345,302,1232]" captionTargetPageId="8" captionText="FIGURE 4: Holoparasitus fanes n. sp., male: A – Gnathotectum, different aspects; B – Gnathosoma, ventrally; C – Chelicera, adaxial aspect; D – Chelicerae tips in ventral perspective, arrows indicate orifices at the base of minute pilus dentilis; E – Basal part of cheliceral digits, antiaxial aspect; F – Tip of fixed digit showing pilus dentilis; G – Pedipalpal trochanter anterolaterally, seta v1 also in ventral perspective; H – Trochanter I, arrow on anterolateral side. A: slides no. 1715, 1716, 2182; B,C: slide no. 2000; D-F: slide no. 1715; G: slide no. 1716, v1 slide no. 2000; H: slide no. 2182. Scale bars: A-E,G,H – 0.1 mm, F – 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4640254" httpUri="https://zenodo.org/record/4640254/files/figure.png" pageId="9" pageNumber="219">Fig. 4H</figureCitation>
|
||
). Leg II (
|
||
<figureCitation id="13030E9BFFC9FFDEFBC4FECEFB0BFEC3" box="[1128,1195,311,335]" captionStart="FIGURE" captionStartId="10.[159,171,1149,1168]" captionTargetBox="[248,1329,313,1101]" captionTargetId="figure-3@10.[242,1345,302,1117]" captionTargetPageId="10" captionText="FIGURE 5: Holoparasitus fanes n. sp., male: A – Sternal shield; B – Genital lamina; C – Leg II (Fe, Ge, Ti), anterolateral aspect; D – ibid., ventral aspect, arrow on anterolateral side; E – Leg II (Ge, Ti), ventral aspect, arrow on anterolateral side. A: slide no. 2001; B: slide no. 2179; C: slide no. 1640; D: slide no. 2002; E: slide no. 2000. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4640256" httpUri="https://zenodo.org/record/4640256/files/figure.png" pageId="9" pageNumber="219">Fig. 5</figureCitation>
|
||
C-E) spurred as follows: apical segment of femur II main spur circular in the ventral perspective, whereas the axillar process wedge-like; spur on genu II moderate, that on tibia II low, roundish and broad. genu II is extended anterolaterally in the distal part (
|
||
<figureCitation id="13030E9BFFC9FFDEFB10FE1FFA8CFE72" box="[1212,1324,486,510]" captionStart="FIGURE" captionStartId="10.[159,171,1149,1168]" captionTargetBox="[248,1329,313,1101]" captionTargetId="figure-3@10.[242,1345,302,1117]" captionTargetPageId="10" captionText="FIGURE 5: Holoparasitus fanes n. sp., male: A – Sternal shield; B – Genital lamina; C – Leg II (Fe, Ge, Ti), anterolateral aspect; D – ibid., ventral aspect, arrow on anterolateral side; E – Leg II (Ge, Ti), ventral aspect, arrow on anterolateral side. A: slide no. 2001; B: slide no. 2179; C: slide no. 1640; D: slide no. 2002; E: slide no. 2000. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4640256" httpUri="https://zenodo.org/record/4640256/files/figure.png" pageId="9" pageNumber="219">Fig. 5D,E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B87121EFFC9FFDEFCCCFDE9FA2FFDC7" blockId="9.[826,1429,311,1949]" pageId="9" pageNumber="219">
|
||
Etymology — The specific name refers to its
|
||
<typeStatus id="5483ACBCFFC9FFDEFACDFDE8FA35FDA5" box="[1377,1429,529,553]" pageId="9" pageNumber="219">type</typeStatus>
|
||
locality, Fanes in Val Travenanzes, Dolomites,
|
||
<collectingCountry id="F32F528EFFC9FFDEFAF9FDCAFA2BFDC7" box="[1365,1419,563,587]" name="Italy" pageId="9" pageNumber="219">Italy</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B87121EFFC9FFDEFCCCFDA7FC55FB96" blockId="9.[826,1429,311,1949]" pageId="9" pageNumber="219">
|
||
<materialsCitation id="3B501843FFC9FFDEFCCCFDA7FC55FB96" ID-GBIF-Occurrence="3077740304" collectingDate="2001-08-18" collectingDateMax="2007-09-07" collectingDateMin="2001-08-18" collectorName="di Fanes" country="Italy" county="Dolomites" elevation="1311" latitude="46.58545" location="Val Travenanzes" longLatPrecision="1" longitude="12.07675" municipality="Val Travenanzes" pageId="9" pageNumber="219" specimenCount="28" specimenCount-female="12" specimenCount-male="16" typeStatus="holotype">
|
||
Material examined —
|
||
<typeStatus id="5483ACBCFFC9FFDEFBCDFDA7FB70FDFA" box="[1121,1232,606,630]" pageId="9" pageNumber="219" type="holotype">Holotype</typeStatus>
|
||
female (slide no. 1620),
|
||
<date id="FF8634DEFFC9FFDEFC27FD78FBA5FD15" box="[907,1029,641,665]" pageId="9" pageNumber="219" value="2001-08-18">
|
||
<collectingDate id="EFC2CD36FFC9FFDEFC27FD78FBA5FD15" box="[907,1029,641,665]" pageId="9" pageNumber="219" value="2001-08-18">18.08.2001</collectingDate>
|
||
</date>
|
||
,
|
||
<taxonomicName id="4C38699DFFC9FFDEFBB9FD78FA31FD15" authority="d'Ampezzo, Dolomites" authorityName="d'Ampezzo, Dolomites" box="[1045,1425,641,665]" class="Polycystina" family="Stephaniidae" genus="Cortina" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="9" pageNumber="219" phylum="Radiozoa" rank="genus">Cortina d’Ampezzo, Dolomites</taxonomicName>
|
||
,
|
||
<collectingCountry id="F32F528EFFC9FFDEFC96FD5DFCCFFD30" box="[826,879,676,700]" name="Italy" pageId="9" pageNumber="219">Italy</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE0C74D9FFC9FFDEFCD0FD5DFBAAFD30" box="[892,1034,676,700]" degrees="46" direction="north" minutes="34.329" orientation="latitude" pageId="9" pageNumber="219" precision="1" value="46.57215">46°34.329’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0C74D9FFC9FFDEFBBBFD5DFB32FD30" box="[1047,1170,676,700]" degrees="12" direction="east" minutes="6.789" orientation="longitude" pageId="9" pageNumber="219" precision="1" value="12.11315">12°6.789’E</geoCoordinate>
|
||
, alt.
|
||
<emphasis id="B94CCE0CFFC9FFDEFB7DFD5DFB48FD37" box="[1233,1256,676,699]" italics="true" pageId="9" pageNumber="219">ca</emphasis>
|
||
.
|
||
<elevation id="0015F52DFFC9FFDEFB50FD5DFA2FFD30" box="[1276,1423,676,700]" metricMagnitude="3" metricUnit="m" metricValue="1.311" pageId="9" pageNumber="219" unit="m" value="1311.0">
|
||
<quantity id="4CC0BFFBFFC9FFDEFB50FD5DFAF1FD37" box="[1276,1361,676,700]" metricMagnitude="3" metricUnit="m" metricValue="1.311" pageId="9" pageNumber="219" unit="m" value="1311.0">1311 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, moss on tree stump in a spruce forest near the road to Olympia Camping;
|
||
<specimenCount id="9D3ED997FFC9FFDEFB22FD13FB55FC8E" box="[1166,1269,746,770]" pageId="9" pageNumber="219" type="female">8 female</specimenCount>
|
||
and
|
||
<specimenCount id="9D3ED997FFC9FFDEFA9BFD13FA34FC8E" box="[1335,1428,746,770]" pageId="9" pageNumber="219" type="male">12 male</specimenCount>
|
||
<typeStatus id="5483ACBCFFC9FFDEFC96FCF4FC0EFCA9" box="[826,942,781,805]" pageId="9" pageNumber="219" type="paratype">paratypes</typeStatus>
|
||
(slides no. 2000, 2001, 2002),
|
||
<date id="FF8634DEFFC9FFDEFABAFCF4FA31FCA9" box="[1302,1425,781,805]" pageId="9" pageNumber="219" value="2005-12-09">
|
||
<collectingDate id="EFC2CD36FFC9FFDEFABAFCF4FA31FCA9" box="[1302,1425,781,805]" pageId="9" pageNumber="219" value="2005-12-09">12.09.2005</collectingDate>
|
||
</date>
|
||
,
|
||
<location id="8EE744C5FFC9FFDEFC96FCC9FC5EFCCB" LSID="urn:lsid:plazi:treatment:0391A308FFC6FFDCFE9CFC54FE4BFC42:8EE744C5FFC9FFDEFC96FCC9FC5EFCCB" box="[826,1022,816,840]" country="Italy" county="Dolomites" latitude="46.58545" longLatPrecision="1" longitude="12.07675" municipality="Val Travenanzes" name="Val Travenanzes" pageId="9" pageNumber="219">Val Travenanzes</location>
|
||
, Dolomites, close to Fanes waterfalls (
|
||
<location id="8EE744C5FFC9FFDEFCD6FCAAFBE4FCE7" LSID="urn:lsid:plazi:treatment:0391A308FFC6FFDCFE9CFC54FE4BFC42:8EE744C5FFC9FFDEFCD6FCAAFBE4FCE7" box="[890,1092,851,875]" country="Italy" county="Dolomites" latitude="46.58545" longLatPrecision="1" longitude="12.07675" municipality="Val Travenanzes" name="Cascate di Fanes" pageId="9" pageNumber="219">
|
||
Cascate
|
||
<collectorName id="26CD77C8FFC9FFDEFC71FCAAFBE4FCE7" box="[989,1092,851,875]" pageId="9" pageNumber="219">di Fanes</collectorName>
|
||
</location>
|
||
), moss and litter in a spruce forest,
|
||
<geoCoordinate id="EE0C74D9FFC9FFDEFC26FC8FFBB8FC02" box="[906,1048,886,910]" degrees="46" direction="north" minutes="35.723" orientation="latitude" pageId="9" pageNumber="219" precision="1" value="46.595383">46°35.723’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0C74D9FFC9FFDEFB8FFC8FFB3FFC02" box="[1059,1183,886,910]" degrees="12" direction="east" minutes="5.185" orientation="longitude" pageId="9" pageNumber="219" precision="1" value="12.086416">12°5.185’E</geoCoordinate>
|
||
, alt.
|
||
<emphasis id="B94CCE0CFFC9FFDEFB74FC8EFB4FFC02" box="[1240,1263,887,910]" italics="true" pageId="9" pageNumber="219">ca</emphasis>
|
||
.
|
||
<elevation id="0015F52DFFC9FFDEFB53FC8FFA2FFC02" box="[1279,1423,886,910]" metricMagnitude="3" metricUnit="m" metricValue="1.455" pageId="9" pageNumber="219" unit="m" value="1455.0">
|
||
<quantity id="4CC0BFFBFFC9FFDEFB53FC8FFAF2FC02" box="[1279,1362,886,910]" metricMagnitude="3" metricUnit="m" metricValue="1.455" pageId="9" pageNumber="219" unit="m" value="1455.0">1455 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<specimenCount id="9D3ED997FFC9FFDEFC96FC60FC02FC3D" box="[826,930,921,945]" pageId="9" pageNumber="219" type="female">4 female</specimenCount>
|
||
and
|
||
<specimenCount id="9D3ED997FFC9FFDEFC4AFC60FB99FC3D" box="[998,1081,921,945]" pageId="9" pageNumber="219" type="male">4 male</specimenCount>
|
||
<typeStatus id="5483ACBCFFC9FFDEFBE9FC63FB19FC3E" box="[1093,1209,922,946]" pageId="9" pageNumber="219" type="paratype">paratypes</typeStatus>
|
||
(slides no. 2162, 2163),
|
||
<date id="FF8634DEFFC9FFDEFC2AFC45FC5DFC58" box="[902,1021,956,980]" pageId="9" pageNumber="219" value="2007-09-07">
|
||
<collectingDate id="EFC2CD36FFC9FFDEFC2AFC45FC5DFC58" box="[902,1021,956,980]" pageId="9" pageNumber="219" value="2007-09-07">7.09. 2007</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMunicipality id="6BE38864FFC9FFDEFBA4FC45FB6AFC58" box="[1032,1226,956,980]" pageId="9" pageNumber="219">Val Travenanzes</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62E66A92FFC9FFDEFB7AFC45FAF2FC58" box="[1238,1362,956,980]" pageId="9" pageNumber="219">Dolomites</collectingCounty>
|
||
, litter and moss in a spruce forest,
|
||
<geoCoordinate id="EE0C74D9FFC9FFDEFBD2FC26FAACFC7B" box="[1150,1292,991,1015]" degrees="46" direction="north" minutes="35.127" orientation="latitude" pageId="9" pageNumber="219" precision="1" value="46.58545">46°35.127’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0C74D9FFC9FFDEFAB9FC26FA31FC7B" box="[1301,1425,991,1015]" degrees="12" direction="east" minutes="4.605" orientation="longitude" pageId="9" pageNumber="219" precision="1" value="12.07675">12°4.605’E</geoCoordinate>
|
||
, alt.
|
||
<elevation id="0015F52DFFC9FFDEFCCAFBFBFC55FB96" box="[870,1013,1026,1050]" metricMagnitude="3" metricUnit="m" metricValue="1.635" pageId="9" pageNumber="219" unit="m" value="1635.0">
|
||
<quantity id="4CC0BFFBFFC9FFDEFCCAFBFBFC19FB96" box="[870,953,1026,1050]" metricMagnitude="3" metricUnit="m" metricValue="1.635" pageId="9" pageNumber="219" unit="m" value="1635.0">1635 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B87121EFFC9FFDEFCCCFBD4FADDF9BC" blockId="9.[826,1429,311,1949]" pageId="9" pageNumber="219">
|
||
Other material:
|
||
<specimenCount id="9D3ED997FFC9FFDEFB91FBD4FB19FBC9" box="[1085,1209,1069,1093]" pageId="9" pageNumber="219" type="female">3 females</specimenCount>
|
||
,
|
||
<specimenCount id="9D3ED997FFC9FFDEFB7FFBD4FA96FBC9" box="[1235,1334,1069,1093]" pageId="9" pageNumber="219" type="male">4 males</specimenCount>
|
||
(slides no. 1640, 1641, 1713-1717),
|
||
<date id="FF8634DEFFC9FFDEFB01FBA9FA86FBE4" box="[1197,1318,1104,1128]" pageId="9" pageNumber="219" value="2002-10-09">10.09.2002</date>
|
||
,
|
||
<taxonomicName id="4C38699DFFC9FFDEFA90FBA9FBF6FB07" authority="d'Ampezzo, Dolomites" authorityName="d'Ampezzo, Dolomites" class="Polycystina" family="Stephaniidae" genus="Cortina" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="9" pageNumber="219" phylum="Radiozoa" rank="genus">Cortina d’Ampezzo, Dolomites</taxonomicName>
|
||
, near Olympia Camping, litter in a spruce forest,
|
||
<geoCoordinate id="EE0C74D9FFC9FFDEFBD9FB6FFAA3FB22" box="[1141,1283,1174,1198]" degrees="46" direction="north" minutes="34.329" orientation="latitude" pageId="9" pageNumber="219" precision="1" value="46.57215">46°34.329’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0C74D9FFC9FFDEFAB9FB6FFA31FB22" box="[1301,1425,1174,1198]" degrees="12" direction="east" minutes="6.789" orientation="longitude" pageId="9" pageNumber="219" precision="1" value="12.11315">12°6.789’E</geoCoordinate>
|
||
, alt.
|
||
<emphasis id="B94CCE0CFFC9FFDEFC3CFB43FC07FB5D" box="[912,935,1210,1233]" italics="true" pageId="9" pageNumber="219">ca</emphasis>
|
||
.
|
||
<quantity id="4CC0BFFBFFC9FFDEFC73FB40FBE0FB5C" box="[991,1088,1209,1233]" metricMagnitude="3" metricUnit="m" metricValue="1.311" pageId="9" pageNumber="219" unit="m" value="1311.0">1311 m</quantity>
|
||
a.s.l.;
|
||
<specimenCount id="9D3ED997FFC9FFDEFB07FB40FAA6FB5D" box="[1195,1286,1209,1233]" pageId="9" pageNumber="219" type="male">1 male</specimenCount>
|
||
(slide no. 2011),
|
||
<date id="FF8634DEFFC9FFDEFC24FB25FBA3FB78" box="[904,1027,1244,1268]" pageId="9" pageNumber="219" value="2005-09-13">13.09.2005</date>
|
||
, near Olympia Camping,
|
||
<taxonomicName id="4C38699DFFC9FFDEFA90FB25FBF0FA9B" authority="d'Ampezzo, Dolomites" authorityName="d'Ampezzo, Dolomites" class="Polycystina" family="Stephaniidae" genus="Cortina" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="9" pageNumber="219" phylum="Radiozoa" rank="genus">Cortina d’Ampezzo, Dolomites</taxonomicName>
|
||
, thick layer of moss covering soil and logs in a spruce forest,
|
||
<geoCoordinate id="EE0C74D9FFC9FFDEFAAEFADBFA31FAB6" box="[1282,1425,1314,1338]" degrees="46" direction="north" minutes="34.335" orientation="latitude" pageId="9" pageNumber="219" precision="1" value="46.57225">46°34.335’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0C74D9FFC9FFDEFC96FABCFC15FAD1" box="[826,949,1349,1373]" degrees="12" direction="east" minutes="6.746" orientation="longitude" pageId="9" pageNumber="219" precision="1" value="12.112433">12°6.746’E</geoCoordinate>
|
||
, alt.
|
||
<emphasis id="B94CCE0CFFC9FFDEFC47FABFFBA2FAD1" box="[1003,1026,1350,1373]" italics="true" pageId="9" pageNumber="219">ca</emphasis>
|
||
.
|
||
<quantity id="4CC0BFFBFFC9FFDEFBBCFABCFBC2FAD1" box="[1040,1122,1349,1373]" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" pageId="9" pageNumber="219" unit="m" value="1320.0">1320 m</quantity>
|
||
a.s.l.;
|
||
<specimenCount id="9D3ED997FFC9FFDEFB05FABCFA85FAD1" box="[1193,1317,1349,1373]" pageId="9" pageNumber="219" type="female">13 females</specimenCount>
|
||
,
|
||
<specimenCount id="9D3ED997FFC9FFDEFA9CFABCFA34FAD1" box="[1328,1428,1349,1373]" pageId="9" pageNumber="219" type="male">24 males</specimenCount>
|
||
(slides no. 2179, 2180, 2182),
|
||
<date id="FF8634DEFFC9FFDEFB3BFA91FAB1FA0C" box="[1175,1297,1384,1408]" pageId="9" pageNumber="219" value="2008-09-30">30.09.2008</date>
|
||
, near Dobbiaco, Dolomites, moss under trees,
|
||
<geoCoordinate id="EE0C74D9FFC9FFDEFAAEFA72FA31FA2F" box="[1282,1425,1419,1443]" degrees="46" direction="north" minutes="41.520" orientation="latitude" pageId="9" pageNumber="219" precision="1" value="46.692">46°41.520’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0C74D9FFC9FFDEFC96FA57FC62FA4A" box="[826,962,1454,1478]" degrees="12" direction="east" minutes="12.450" orientation="longitude" pageId="9" pageNumber="219" precision="1" value="12.2075">12°12.450’E</geoCoordinate>
|
||
, alt.
|
||
<emphasis id="B94CCE0CFFC9FFDEFBA1FA56FB84FA4A" box="[1037,1060,1455,1478]" italics="true" pageId="9" pageNumber="219">ca</emphasis>
|
||
.
|
||
<quantity id="4CC0BFFBFFC9FFDEFBEDFA57FB38FA4A" box="[1089,1176,1454,1478]" metricMagnitude="3" metricUnit="m" metricValue="1.626" pageId="9" pageNumber="219" unit="m" value="1626.0">1626 m</quantity>
|
||
a.s.l.
|
||
<materialsCitation id="3B501843FFC9FFDEFB43FA56FAD9F9BC" ID-GBIF-Occurrence="3077740303" country="Poland" location="Poland" pageId="9" pageNumber="219" specimenCount="2">
|
||
<typeStatus id="5483ACBCFFC9FFDEFB43FA56FA94FA4B" box="[1263,1332,1455,1479]" pageId="9" pageNumber="219">Types</typeStatus>
|
||
are deposited in the Zoological Museum of the Jagiellonian University, Cracow,
|
||
<collectingCountry id="F32F528EFFC9FFDEFB38FA0DFB4AF980" box="[1172,1258,1524,1548]" name="Poland" pageId="9" pageNumber="219">Poland</collectingCountry>
|
||
, whereas the other material remains in the Author’s collection
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B87121EFFC9FFDEFCCCF9BBFBCEF9F1" blockId="9.[826,1429,311,1949]" pageId="9" pageNumber="219">
|
||
<emphasis id="B94CCE0CFFC9FFDEFCCCF9BBFB2BF9D5" box="[864,1163,1602,1626]" italics="true" pageId="9" pageNumber="219">
|
||
<taxonomicName id="4C38699DFFC9FFDEFCCCF9BBFC57F9D6" baseAuthorityName="Oudemans" baseAuthorityYear="1936" box="[864,1015,1602,1626]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="219" phylum="Arthropoda" rank="genus">Holoparasitus</taxonomicName>
|
||
crassisetosus
|
||
</emphasis>
|
||
species-group, newly defined. Species included:
|
||
</paragraph>
|
||
<paragraph id="8B87121EFFC9FFDEFC96F971FC15F94F" blockId="9.[826,1429,311,1949]" pageId="9" pageNumber="219">
|
||
<emphasis id="B94CCE0CFFC9FFDEFC96F971FBCAF92C" box="[826,1130,1672,1696]" italics="true" pageId="9" pageNumber="219">
|
||
<taxonomicName id="4C38699DFFC9FFDEFC96F971FC71F92C" baseAuthorityName="Oudemans" baseAuthorityYear="1936" box="[826,977,1672,1696]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="219" phylum="Arthropoda" rank="genus">Holoparasitus</taxonomicName>
|
||
crassisetosus
|
||
</emphasis>
|
||
Juvara-Bals and Witali´nski, 2000
|
||
</paragraph>
|
||
<paragraph id="8B87121EFFC9FFDEFC96F937FCCFF885" blockId="9.[826,1429,311,1949]" pageId="9" pageNumber="219">
|
||
<emphasis id="B94CCE0CFFC9FFDEFC96F937FBF3F96A" box="[826,1107,1742,1766]" italics="true" pageId="9" pageNumber="219">
|
||
<taxonomicName id="4C38699DFFC9FFDEFC96F937FC71F96A" baseAuthorityName="Oudemans" baseAuthorityYear="1936" box="[826,977,1742,1766]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="219" phylum="Arthropoda" rank="genus">Holoparasitus</taxonomicName>
|
||
digitiformis
|
||
</emphasis>
|
||
Juvara-Bals and Witali´nski, 2000
|
||
</paragraph>
|
||
<paragraph id="8B87121EFFC9FFDDFCCCF8E2FAF1F90C" blockId="9.[826,1429,311,1949]" lastBlockId="10.[826,1429,1288,1664]" lastPageId="10" lastPageNumber="220" pageId="9" pageNumber="219">
|
||
Diagnosis of the species group — In both sexes (1) gnathotectum trispinate; (2) gland pores
|
||
<emphasis id="B94CCE0CFFC9FFDEFAC1F8C6FA34F8DB" box="[1389,1428,1855,1879]" italics="true" pageId="9" pageNumber="219">gv1</emphasis>
|
||
present; (3) gland pores
|
||
<emphasis id="B94CCE0CFFC9FFDEFBF1F89BFB25F8F6" box="[1117,1157,1890,1914]" italics="true" pageId="9" pageNumber="219">gv2</emphasis>
|
||
located in unmodified flat cuticle. Female — (1) gnathotectum regularly trispinate; (2) presternal plate with the anterior margin smooth, lateral platelets free, or one-sidedly accreted to presternal plate; (3) sternum entire, with regular reticulation, axial granular or reticulated band absent; (4) sternal anterior margin developed normally; (5) posterior paragynial lobes in close proximity to each other; (6) the thickening of the anterior paragynial edge facing coxa III present; (7) central epigynial prong moderate and triangular, the thickening of subapical epigynial structure well visible, compact, anteriorly rounded; (8) well sclerotised endogynium very small and roundish, with a protrusion directed dorsally. Male — (1) gnathotectum trispinate-type, but the central prong well pronounced and acute, whereas the lateral prongs minute; (2) hypostome regularly triangular and moderately sclerotised; (3) hypostomatic setae on entire cuticle, but not on the piece of cuticle separated by lateral and posterior soft-cuticular inci- sions; (4) corniculi with a circular lamellar protrusion located adaxially; (5) excipulum on the sternum absent; (6) genital lamina with a well sclerotised subgenital microsclerite and long, acute hyaline lateral protrusions; (7) pedipalp trochanter with seta
|
||
<emphasis id="B94CCE0CFFCAFFDDFCC2FA41FC29FA5C" box="[878,905,1464,1488]" italics="true" pageId="10" pageNumber="220">v1</emphasis>
|
||
simple and thickened basally, seta
|
||
<emphasis id="B94CCE0CFFCAFFDDFA8EFA41FA9DFA5C" box="[1314,1341,1464,1488]" italics="true" pageId="10" pageNumber="220">v2</emphasis>
|
||
barbed terminally; (8) pedipalp femur bears a ventral tubercle; (9) femur II spur conical, with a small terminal segment, axillary process settled on the slope of femoral spur; (10) spur on genu II small and semicircular, located in the middle of the segment.
|
||
</paragraph>
|
||
<caption id="DF474296FFCAFFDDFF33FB84FC1CFB4B" ID-DOI="http://doi.org/10.5281/zenodo.4640256" ID-Zenodo-Dep="4640256" httpUri="https://zenodo.org/record/4640256/files/figure.png" pageId="10" pageNumber="220" startId="10.[159,171,1149,1168]" targetBox="[248,1329,313,1101]" targetPageId="10">
|
||
<paragraph id="8B87121EFFCAFFDDFF33FB84FC1CFB4B" blockId="10.[159,1429,1148,1223]" pageId="10" pageNumber="220">
|
||
FIGURE 5:
|
||
<taxonomicName id="4C38699DFFCAFFDDFEABFB85FE12FB1C" authority="Witalinski, 2017" authorityName="Witalinski" authorityYear="2017" box="[263,434,1148,1168]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="fanes" status="sp. nov.">
|
||
<emphasis id="B94CCE0CFFCAFFDDFEABFB85FE12FB1C" box="[263,434,1148,1168]" italics="true" pageId="10" pageNumber="220">Holoparasitus fanes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94CCE0CFFCAFFDDFE14FB84FE50FB1D" bold="true" box="[440,496,1149,1169]" pageId="10" pageNumber="220">
|
||
<taxonomicNameLabel id="A27F7377FFCAFFDDFE14FB84FE50FB1D" box="[440,496,1149,1169]" pageId="10" pageNumber="220" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male: A – Sternal shield; B – Genital lamina; C – Leg II (Fe, Ge, Ti), anterolateral aspect; D –
|
||
<emphasis id="B94CCE0CFFCAFFDDFAC6FB85FA2AFB03" box="[1386,1418,1148,1167]" italics="true" pageId="10" pageNumber="220">ibid</emphasis>
|
||
., ventral aspect, arrow on anterolateral side; E – Leg II (Ge, Ti), ventral aspect, arrow on anterolateral side. A: slide no. 2001; B: slide no. 2179; C: slide no. 1640; D: slide no. 2002; E: slide no. 2000. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B87121EFFCAFFDDFCCCF967FB91F893" blockId="10.[826,1429,1694,1823]" pageId="10" pageNumber="220">
|
||
Distribution — Both species of this group are found in the central part of
|
||
<collectingCountry id="F32F528EFFCAFFDDFB29F938FB1AF955" box="[1157,1210,1729,1753]" name="Italy" pageId="10" pageNumber="220">Italy</collectingCountry>
|
||
,
|
||
<collectingRegion id="49FCDCFCFFCAFFDDFB6AF93BFA86F956" box="[1222,1318,1730,1754]" country="Italy" name="Toscana" pageId="10" pageNumber="220">Tuscany</collectingRegion>
|
||
, whereas
|
||
<emphasis id="B94CCE0CFFCAFFDDFC96F91CFC79F970" box="[826,985,1764,1788]" italics="true" pageId="10" pageNumber="220">H. digitiformis</emphasis>
|
||
is also found in
|
||
<collectingRegion id="49FCDCFCFFCAFFDDFB31F91DFB5FF970" box="[1181,1279,1764,1788]" country="Italy" name="Sardegna" pageId="10" pageNumber="220">Sardinia</collectingRegion>
|
||
(Juvara-Bals and Witali´nski 2000).
|
||
</paragraph>
|
||
<paragraph id="8B87121EFFCAFFDCFCCCF8C7FE53FDAD" blockId="10.[826,1429,1854,1949]" lastBlockId="11.[159,762,311,974]" lastPageId="11" lastPageNumber="221" pageId="10" pageNumber="220">
|
||
Remarks — The new species
|
||
<taxonomicName id="4C38699DFFCAFFDDFB02F8C6FAA5F8DA" authorityName="Witalinski" authorityYear="2017" box="[1198,1285,1854,1878]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="fanes">
|
||
<emphasis id="B94CCE0CFFCAFFDDFB02F8C6FAA5F8DA" box="[1198,1285,1854,1878]" italics="true" pageId="10" pageNumber="220">H. fanes</emphasis>
|
||
</taxonomicName>
|
||
is most similar to the
|
||
<emphasis id="B94CCE0CFFCAFFDDFC02F89BFBF9F8F5" box="[942,1113,1890,1913]" italics="true" pageId="10" pageNumber="220">H. crassisetosus</emphasis>
|
||
species-group. It shows all the character states shared by both sexes (1-3). The females of
|
||
<taxonomicName id="4C38699DFFCBFFDCFE8FFEC1FEDFFEC3" authorityName="Witalinski" authorityYear="2017" box="[291,383,311,335]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="221" phylum="Arthropoda" rank="species" species="fanes">
|
||
<emphasis id="B94CCE0CFFCBFFDCFE8FFEC1FEDFFEC3" box="[291,383,311,335]" italics="true" pageId="11" pageNumber="221">H. fanes</emphasis>
|
||
</taxonomicName>
|
||
possess some characteristics of the group (1-5), but details of the genital orifice area are different. The males of
|
||
<taxonomicName id="4C38699DFFCBFFDCFE5EFE87FDEEFE19" authorityName="Witalinski" authorityYear="2017" box="[498,590,381,405]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="221" phylum="Arthropoda" rank="species" species="fanes">
|
||
<emphasis id="B94CCE0CFFCBFFDCFE5EFE87FDEEFE19" box="[498,590,381,405]" italics="true" pageId="11" pageNumber="221">H. fanes</emphasis>
|
||
</taxonomicName>
|
||
are similar to the
|
||
<emphasis id="B94CCE0CFFCBFFDCFF60FE58FEDAFE34" box="[204,378,417,440]" italics="true" pageId="11" pageNumber="221">H. crassisetosus</emphasis>
|
||
group in gnathotectum and hypostome structure (1-3), sternum (5) and pedipalp (7, 8), but corniculi (4), genital lamina (6) and leg II structure (9, 10) are different.
|
||
</paragraph>
|
||
<paragraph id="8B87121EFFCBFFDCFF69FDCDFE4BFC42" blockId="11.[159,762,311,974]" pageId="11" pageNumber="221">
|
||
The female of
|
||
<taxonomicName id="4C38699DFFCBFFDCFEC6FDCCFE63FDC1" authorityName="Witalinski" authorityYear="2017" box="[362,451,565,589]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="221" phylum="Arthropoda" rank="species" species="fanes">
|
||
<emphasis id="B94CCE0CFFCBFFDCFEC6FDCCFE63FDC1" box="[362,451,565,589]" italics="true" pageId="11" pageNumber="221">H. fanes</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from the
|
||
<emphasis id="B94CCE0CFFCBFFDCFF61FDA1FEDBFDE3" box="[205,379,600,623]" italics="true" pageId="11" pageNumber="221">H. crassisetosus</emphasis>
|
||
species-group by the anteriorly tripartite, rather than rounded, thickening of the subapical epigynial structure. In the male, a porous area in the sternum center is present only in
|
||
<taxonomicName id="4C38699DFFCBFFDCFD30FD38FD53FD55" authorityName="Witalinski" authorityYear="2017" box="[668,755,705,729]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="221" phylum="Arthropoda" rank="species" species="fanes">
|
||
<emphasis id="B94CCE0CFFCBFFDCFD30FD38FD53FD55" box="[668,755,705,729]" italics="true" pageId="11" pageNumber="221">H. fanes</emphasis>
|
||
</taxonomicName>
|
||
. Moreover, the corniculi are conical and tibia II regularly spurred. These characteristics differentiate
|
||
<taxonomicName id="4C38699DFFCBFFDCFF33FCD2FF58FCCE" authorityName="Witalinski" authorityYear="2017" box="[159,248,810,834]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="221" phylum="Arthropoda" rank="species" species="fanes">
|
||
<emphasis id="B94CCE0CFFCBFFDCFF33FCD2FF58FCCE" box="[159,248,810,834]" italics="true" pageId="11" pageNumber="221">H. fanes</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<emphasis id="B94CCE0CFFCBFFDCFEECFCD2FE6CFCCE" box="[320,460,811,834]" italics="true" pageId="11" pageNumber="221">H. inornatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA96FEFFFCBFFDCFE77FCD3FDD4FCCE" author="Berlese A." box="[475,628,810,834]" pageId="11" pageNumber="221" pagination="66 - 304" refId="ref5904" refString="Berlese A. 1906 - Monografia del genere Gamasus Latr. - Redia, 3: 66 - 304." type="journal article" year="1906">Berlese, 1906</bibRefCitation>
|
||
) and
|
||
<emphasis id="B94CCE0CFFCBFFDCFD14FCD2FEA9FCE9" italics="true" pageId="11" pageNumber="221">H. intermedius</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA96FEFFFCBFFDCFEB0FCB4FE45FCE9" author="Holzmann C." box="[284,485,845,869]" pageId="11" pageNumber="221" pagination="3 - 55" refId="ref5924" refString="Holzmann C. 1969 - Die Familie Parasitidae Oudemans, 1901 - Acarologie, 13: 3 - 55 + 23 Plates." type="book chapter" year="1969">Holzmann, 1969</bibRefCitation>
|
||
), both species with indented corniculi.
|
||
<taxonomicName id="4C38699DFFCBFFDCFEC3FC88FD59FC04" authority="Schmolzer, 1995" authorityName="Schmolzer" authorityYear="1995" box="[367,761,880,904]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="221" phylum="Arthropoda" rank="species" species="megacalcaratus">
|
||
<emphasis id="B94CCE0CFFCBFFDCFEC3FC88FD96FC04" box="[367,566,880,904]" italics="true" pageId="11" pageNumber="221">H. megacalcaratus</emphasis>
|
||
<bibRefCitation id="EFA96FEFFFCBFFDCFD91FC89FD59FC04" author="Schmolzer K." box="[573,761,880,904]" pageId="11" pageNumber="221" pagination="499 - 512" refId="ref6266" refString="Schmolzer K. 1995 - Landmilben aus den ostlichen Karawanken (Acarina, Parasitiformes) - Carinthia II, 185: 499 - 512." type="journal article" year="1995">Schmölzer, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
possesses conical corniculi, but with only a single, large main spur on femur II.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |