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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.58.507" ID-GBIF-Dataset="1c4e9564-16cc-41fd-bdd2-d4a77484f100" ID-PMC="PMC3088339" ID-Pensoft-Pub="1313-2970-58-1" ID-PubMed="21594191" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-58-1" ModsDocOrigin="ZooKeys 58" ModsDocTitle="Australian gall-inducing scale insects on Eucalyptus: revision of Opisthoscelis Schrader (Coccoidea, Eriococcidae) and descriptions of a new genus and nine new species" checkinTime="1451250974324" checkinUser="pensoft" docAuthor="Hardy, Nate B. &amp; Gullan, Penny J." docDate="2010" docId="3E76686389857068A3B9ED213B82F270" docLanguage="en" docName="ZooKeys 58: 1-74" docOrigin="ZooKeys 58" docSource="http://dx.doi.org/10.3897/zookeys.58.507" docTitle="Tanyscelis pisiformis, comb. n." docType="treatment" docVersion="4" lastPageNumber="34" masterDocId="8968FFC5A95F7F195B2A10735E18FFA1" masterDocTitle="Australian gall-inducing scale insects on Eucalyptus: revision of Opisthoscelis Schrader (Coccoidea, Eriococcidae) and descriptions of a new genus and nine new species" masterLastPageNumber="74" masterPageNumber="1" pageNumber="32" updateTime="1668162972525" updateUser="ExternalLinkService">
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<mods:title>Australian gall-inducing scale insects on Eucalyptus: revision of Opisthoscelis Schrader (Coccoidea, Eriococcidae) and descriptions of a new genus and nine new species</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hardy, Nate B.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gullan, Penny J.</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2010</mods:date>
<mods:detail type="volume">
<mods:number>58</mods:number>
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<mods:start>1</mods:start>
<mods:end>74</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.58.507</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.58.507</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-58-1</mods:identifier>
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<treatment ID-GBIF-Taxon="159359975" LSID="urn:lsid:plazi:treatment:3E76686389857068A3B9ED213B82F270" httpUri="http://treatment.plazi.org/id/3E76686389857068A3B9ED213B82F270" lastPageId="34" lastPageNumber="34" pageId="31" pageNumber="32">
<subSubSection pageId="31" pageNumber="32" type="nomenclature">
<paragraph pageId="31" pageNumber="32">
<taxonomicName authority="Froggatt" class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis pisiformis" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pisiformis">Tanyscelis pisiformis (Froggatt)</taxonomicName>
<taxonomicNameLabel pageId="31" pageNumber="32">comb. n.</taxonomicNameLabel>
Figs 3b,c18, 19
</paragraph>
</subSubSection>
<subSubSection pageId="31" pageNumber="32" type="reference_group">
<paragraph pageId="31" pageNumber="32">
<taxonomicName class="Insecta" family="Eriococcidae" genus="Opisthoscelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthoscelis pisiformis" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pisiformis">Opisthoscelis pisiformis</taxonomicName>
<bibRefCitation author="Froggatt, WW" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="44" pageNumber="45" pagination="335 - 348" title="Notes on the family Brachyscelidae, with descriptions of new species. Part III." volume="8" year="1894 b">Froggatt 1894b: 343-344</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="31" pageNumber="32" type="general">
<paragraph pageId="31" pageNumber="32">General.</paragraph>
<paragraph pageId="31" pageNumber="32">
When
<bibRefCitation author="Froggatt, WW" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="44" pageNumber="45" pagination="335 - 348" title="Notes on the family Brachyscelidae, with descriptions of new species. Part III." volume="8" year="1894 b">Froggatt (1894b)</bibRefCitation>
described
<taxonomicName class="Insecta" family="Eriococcidae" genus="Opisthoscelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthoscelis pisiformis" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pisiformis">Opisthoscelis pisiformis</taxonomicName>
, he listed three host species of eucalypts -
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus melliodora" order="Myrtales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="species" species="melliodora">Eucalyptus melliodora</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus piperita" order="Myrtales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="species" species="piperita">Eucalyptus piperita</taxonomicName>
and
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus resinifera" order="Myrtales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="species" species="resinifera">Eucalyptus resinifera</taxonomicName>
- from three different localities in New South Wales -Bathurst, Thornleigh and Sutherland. Even though we designate a lectotype below, the type locality of this species continues to encompass these three localities because the collection locality of the lectotype is not known. Although
<bibRefCitation author="Miller, DR" journalOrPublisher="Intercept Ltd., Andover, UK" pageId="44" pageNumber="45" title="A Systematic Catalogue of the Eriococcidae (Felt Scales) (Hemiptera: Coccoidea) of the World." year="2000">Miller and Gimpel (2000)</bibRefCitation>
listed the BMNH as holding syntypes, the data associated with the four BMNH collections of dry galls labelled as
<taxonomicName class="Insecta" family="Eriococcidae" genus="Opisthoscelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthoscelis pisiformis" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pisiformis">Opisthoscelis pisiformis</taxonomicName>
either clearly show that the material was collected subsequent to description of this species or the data are inadequate to determine type status. Probably only two of the four BMNH collections belong to this species. Insects extracted from dry galls of one collection were slide-mounted by PJG in 1984 and are listed below in Material examined.
</paragraph>
</subSubSection>
<subSubSection lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="gall">
<paragraph pageId="31" pageNumber="32">Gall</paragraph>
<paragraph lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
(Fig. 3b,c). Female (Fig. 3a). On leaf. Height 4.0-9.6 mm, width 4.0-6.4 mm, length of basal attachment 2.9-8.9 mm. Gall opening an almost closed slit in
<pageBreakToken pageId="32" pageNumber="33" start="start">young</pageBreakToken>
galls, in mature galls slit-like to oblong, 0.1-1.8 mm wide, 0.6-0.8 mm long; on abaxial (lower) leaf surface. Gall opening on small raised or low conical protrusion, opposite side of leaf sub-spherical, basal attachment may be constricted; surface green, sometimes with reddish tinge, leaf glands enlarged.
</paragraph>
<paragraph pageId="32" pageNumber="33">Male (Fig. 3b). On stem and leaf (probably either surface), height 1.1-4.0 mm, width 1.0-3.8 mm, length of basal attachment 1.4-3.9 mm. Gall conical, opening round to oblong, 0.2-1.8 mm wide.</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="33" type="adult female">
<paragraph pageId="32" pageNumber="33">Adult female</paragraph>
<paragraph pageId="32" pageNumber="33">
(Fig. 18) (n = 15). Body turbinate, margin incised at intersegmental boundaries, length 2.7-3.6 mm, greatest width 1.1-2.3 mm; abdomen tapered, about as long as head + thorax, extending far beyond femur. Eyespots each 15-25 mm wide, on dorsal margin. Antennal segmentation poorly developed; each antenna 80-228 mm long. Frontal lobes difficult to see, each 125-190
<normalizedToken originalValue="µm">µm</normalizedToken>
long, 180-240
<normalizedToken originalValue="µm">µm</normalizedToken>
wide. Tentorial box 300-495 mm long. Pump chamber 34-45
<normalizedToken originalValue="µm">µm</normalizedToken>
long, 33-53
<normalizedToken originalValue="µm">µm</normalizedToken>
wide. Labium 75-160 mm long, 65-220 mm wide. Spiracles 80-125 mm long, 40-75 mm wide across atrium. Fore and mid legs small sclerotic protuberances, 12-63
<normalizedToken originalValue="µm">µm</normalizedToken>
. Hind legs slender and elongate; coxa 270-440
<normalizedToken originalValue="µm">µm</normalizedToken>
long, trochanter + femur 405-630
<normalizedToken originalValue="µm">µm</normalizedToken>
long, tibia 860-1300
<normalizedToken originalValue="µm">µm</normalizedToken>
long, tarsus 250-380
<normalizedToken originalValue="µm">µm</normalizedToken>
long; translucent pores dense throughout dorsal and ventral surfaces of tibiotarsus, a few on distolateral part of coxa; trochanter with 2 campaniform sensilla on each side; femur-tibia articulation non-functional, tibia fixed in orientation parallel to long axis of femur; claw and digitules present but reduced. Anal opening 10-18
<normalizedToken originalValue="µm">µm</normalizedToken>
wide, with anal ring, 18-32
<normalizedToken originalValue="µm">µm</normalizedToken>
wide, sclerotization of ring uneven, weaker at posteroventral end, often appearing horseshoe-shaped, anal ring with 6 minute setae. Anal area with 4 stout spines, 25-35
<normalizedToken originalValue="µm">µm</normalizedToken>
long, each at the end of a fleshy protuberance; one spine on each side of body laterad of anal ring, another on each side of body posterolateral of anal ring, each anterior anal spine with small, auxiliary spine usually present near medial edge of base plus a few minute setae on surface of protuberance.
</paragraph>
<paragraph pageId="32" pageNumber="33">
Dorsum. Derm membranous. Dorsal setae ranging from minute and conical to long and flagellate, 8-95 mm long; arranged in a transverse row or narrow band across each body segment, longer setae present on abdominal segments and along margin. Spinose seta, 5-10
<normalizedToken originalValue="µm">µm</normalizedToken>
long, found on margin of each posterior abdominal segment, absent from margin of anterior abdominal segments, head and thorax. Macrotubular ducts 15 mm long, dermal orifice with a 5 mm wide rim; in transverse row across each abdominal segment, scattered over thorax, absent from head. Microtubular ducts absent. Quinquelocular pores 5-7
<normalizedToken originalValue="µm">µm</normalizedToken>
in diameter, scattered over dorsum.
</paragraph>
<paragraph pageId="32" pageNumber="33">Venter. Setae as on dorsum, each 12-175 mm long, in a transverse row or narrow band across each abdominal segment as well as meta- and mesothorax, along margin and submargin of head and prothorax. Macrotubular ducts similar to those on dorsum, restricted to submarginal areas of abdominal segments. Quinquelocular pores similar to those on dorsum, similar in distribution to ventral setae; pores often occurring in pairs.</paragraph>
</subSubSection>
<subSubSection lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="first-instar nymph">
<paragraph pageId="32" pageNumber="33">First-instar nymph</paragraph>
<paragraph lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
(Fig. 19) (n = 4). Body outline elliptical, anterior margin incised at midline, length 263-283
<normalizedToken originalValue="µm">µm</normalizedToken>
, greatest width 158-188
<normalizedToken originalValue="µm">µm</normalizedToken>
. Eyespots
<pageBreakToken pageId="33" pageNumber="34" start="start">on</pageBreakToken>
margin, each 9-15 mm wide. Antennae 3-segmented, ca 60 mm long, with 4 fleshy setae. Tentorial box 50 mm long. Labium 20-28 mm long, 18-33 mm wide. Spiracles 11-18 mm long, 8 mm wide across atrium. Legs subequal in size: coxa 18-25
<normalizedToken originalValue="µm">µm</normalizedToken>
, with 5 setae, trochanter + femur 53-58
<normalizedToken originalValue="µm">µm</normalizedToken>
long, trochanter with 4 setae, femur with 2 setae, tibia 20-33
<normalizedToken originalValue="µm">µm</normalizedToken>
long, with 4 setae, tarsus 25-35
<normalizedToken originalValue="µm">µm</normalizedToken>
long, with 4 setae, claw 10-13
<normalizedToken originalValue="µm">µm</normalizedToken>
long; tarsal digitules capitate, unequal length, short digitule 18-20
<normalizedToken originalValue="µm">µm</normalizedToken>
long, long digitule 25-28
<normalizedToken originalValue="µm">µm</normalizedToken>
long, claw digitules capitate, each ca 15
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Anal ring 9-15
<normalizedToken originalValue="µm">µm</normalizedToken>
wide, with 6 fine setae, each ca 8
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Apical seta 55-65
<normalizedToken originalValue="µm">µm</normalizedToken>
long.
</paragraph>
<paragraph pageId="33" pageNumber="34">
Dorsum. Derm membranous. Dorsal setae fine and minute £1 mm long; arranged in submedial longitudinal row on each side of body, 1 seta on each side of head, prothorax, and each of abdominal segments
<normalizedToken originalValue="IVII">I-VII</normalizedToken>
. Microtubular ducts 4
<normalizedToken originalValue="µm">µm</normalizedToken>
long, each side of body with 1 duct on submargin of each thoracic segment plus each of abdominal segments I and V; also with 1 duct on submedial area of head, each thoracic segment and abdominal segment VIII. Marginal setae sagittate, 2-5
<normalizedToken originalValue="µm">µm</normalizedToken>
long, each side of body with ca 6 setae between midline and eyes, 4 on prothorax, 3 on mesothorax, 2 on metathorax, 1 on each of abdominal segments
<normalizedToken originalValue="IVII">I-VII</normalizedToken>
, and on abdominal segment VIII 1 lateral and 2 medial of apical seta, these most likely homologous to anal lobe setae.
</paragraph>
<paragraph pageId="33" pageNumber="34">
Venter. Setae hair-like, each 1-20 mm long, each side of body with 3 setae medial of scape, 1 seta medial of each coxa, 3 longitudinal rows on abdomen, each row with 1 seta on each of abdominal segments
<normalizedToken originalValue="IIVII">II-VII</normalizedToken>
; suranal and ventral lobe setae hair-like, each ca 15
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Trilocular pores 3
<normalizedToken originalValue="µm">µm</normalizedToken>
in diameter, 1 pore near each spiracle.
</paragraph>
<caption pageId="33" pageNumber="34">
<paragraph pageId="33" pageNumber="34">
Figure 18. Adult female of
<taxonomicName class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis pisiformis" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="pisiformis">Tanyscelis pisiformis</taxonomicName>
(Froggatt).
</paragraph>
</caption>
<caption pageId="33" pageNumber="34">
<paragraph pageId="33" pageNumber="34">
Figure 19. First-instar nymph of
<taxonomicName class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis pisiformis" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="pisiformis">Tanyscelis pisiformis</taxonomicName>
(Froggatt).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="33" pageNumber="34" type="material examined">
<paragraph pageId="33" pageNumber="34">Material examined.</paragraph>
<paragraph pageId="33" pageNumber="34">
Lectotype of
<taxonomicName class="Insecta" family="Eriococcidae" genus="Opisthoscelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthoscelis pisiformis" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="pisiformis">Opisthoscelis pisiformis</taxonomicName>
(here designated): AUSTRALIA: New South Wales: 1 adult female (3.5 mm long, 2.3 mm wide): ex dry gall on leaf, with printed label: &quot;No. 1786 E / GALL MAKING COCCIDS. /
<taxonomicName class="Insecta" family="Eriococcidae" genus="Opisthoscelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthoscelis pisiformis" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="pisiformis">Opisthoscelis pisiformis</taxonomicName>
, Frogtt. / Male and female galls on Eucalyptus sp. / N.S.W.&quot; ASCT00004858 (ASCU).
</paragraph>
<paragraph pageId="33" pageNumber="34">Paralectotypes: AUSTRALIA: New South Wales: 2 adult female, ca 400 first-instar nymphs &amp; embryos [10 slides], ex dry galls on leaves and stems: same data as lectotype (ASCU).</paragraph>
<subSection pageId="33" pageNumber="34" type="additional material">
<paragraph pageId="33" pageNumber="34">Additional material:</paragraph>
<paragraph pageId="33" pageNumber="34">
AUSTRALIA: Australian Capital Territory: 6 adult females: ex galls,
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus botryoides" order="Myrtales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="botryoides">Eucalyptus botryoides</taxonomicName>
, Jervis Bay, Jervis Bay Rd, 5 Sep., 1984, PJG (ANIC). New South Wales: 5 adult females: ex stem galls,
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus" order="Myrtales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="genus">Eucalyptus</taxonomicName>
sp., Wonboyn, 21 Feb, 1993, PJG (ANIC); 3 adult female, 25 first-instar nymphs: ex galls on leaves,
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus" order="Myrtales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="genus">Eucalyptus</taxonomicName>
sp., nr Wonboyn Lake Resort, N shore, -37.24°; 149.93°, 15 Jan., 1994, PJG (ANIC); 6 adult females: ex leaf galls,
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus saligna" order="Myrtales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="saligna">Eucalyptus saligna</taxonomicName>
, S. Brooman,
<normalizedToken originalValue="“Strathclyde”">&quot;Strathclyde&quot;</normalizedToken>
property, bank of Clyde River, -35.52°; 150.22°, 10 Jan., 1996, PJG (ANIC); 4 adult females (on 1 slide): ex
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus robusta" order="Myrtales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="robusta">Eucalyptus robusta</taxonomicName>
, Sydney, 25 Dec. 1929, WWF, #1867 (ANIC); 2 adult females (1 slide): ex
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus robusta" order="Myrtales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="robusta">Eucalyptus robusta</taxonomicName>
, Sydney Harbour, WWF, #1868 (ANIC). Locality unknown: 5 adult females, ex dry galls with labels: &quot;Australia / W.W. Froggatt / 95-74&quot; and
<normalizedToken originalValue="“15”">&quot;15&quot;</normalizedToken>
(BMNH).
</paragraph>
</subSection>
</subSubSection>
<subSubSection lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="comments">
<paragraph pageId="33" pageNumber="34">Comments.</paragraph>
<paragraph lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
Adult females of
<taxonomicName class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis pisiformis" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="pisiformis">Tanyscelis pisiformis</taxonomicName>
are very similar to those of
<taxonomicName class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis verrucula" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="verrucula">Tanyscelis verrucula.</taxonomicName>
Both species have 4 anal spines and a small but distinct anal ring that often appears horseshoe-shaped and has ca 6 setae. Both also have marginal abdominal spines that are smaller than the anal spines, and they frequently have paired quinquelocular pores.
<pageBreakToken pageId="34" pageNumber="35" start="start">Adult</pageBreakToken>
females of
<taxonomicName class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis pisiformis" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="pisiformis">Tanyscelis pisiformis</taxonomicName>
can be distinguished from those of
<taxonomicName class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis verrucula" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="verrucula">Tanyscelis verrucula</taxonomicName>
by having (1) marginal spines restricted to the posterior abdominal segments (marginal spines also occurring along margin on head and thorax on
<taxonomicName class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis verrucula" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="verrucula">Tanyscelis verrucula</taxonomicName>
); (2) much smaller eyes (15-25
<normalizedToken originalValue="µm">µm</normalizedToken>
wide, compared to
<taxonomicName class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis verrucula" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="verrucula">Tanyscelis verrucula</taxonomicName>
with eyes 30-65
<normalizedToken originalValue="µm">µm</normalizedToken>
wide); (3) elongate flagellate setae up to 95
<normalizedToken originalValue="µm">µm</normalizedToken>
long on dorsal surface of posterior abdominal segments (shorter than 25
<normalizedToken originalValue="µm">µm</normalizedToken>
in
<taxonomicName class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis verrucula" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="verrucula">Tanyscelis verrucula</taxonomicName>
); and (4) no weakly-sclerotised pads or protuberances near the spiracles (present in
<taxonomicName class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis verrucula" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="verrucula">Tanyscelis verrucula</taxonomicName>
). Both
<taxonomicName class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis pisiformis" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="pisiformis">Tanyscelis pisiformis</taxonomicName>
and
<taxonomicName class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis verrucula" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="verrucula">Tanyscelis verrucula</taxonomicName>
are known from southeastern Australia, although
<taxonomicName class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis verrucula" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="verrucula">Tanyscelis verrucula</taxonomicName>
is much more widely distributed and
<taxonomicName class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis pisiformis" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="pisiformis">Tanyscelis pisiformis</taxonomicName>
appears to have a strictly coastal distribution and has not been collected in Victoria. The two have never been collected from the same site, although they both occur in New South Wales.
<taxonomicName class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis verrucula" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="verrucula">Tanyscelis verrucula</taxonomicName>
is known only from eucalypt species in the section
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Maidenaria" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Maidenaria" order="Myrtales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">Maidenaria</taxonomicName>
. Most of the
<taxonomicName class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tanyscelis pisiformis" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="pisiformis">Tanyscelis pisiformis</taxonomicName>
material was collected from unidentified hosts, but one sample was from
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus botryoides" order="Myrtales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="botryoides">Eucalyptus botryoides</taxonomicName>
and another from
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus saligna" order="Myrtales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="saligna">Eucalyptus saligna</taxonomicName>
(both in section
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Latoangulatae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Latoangulatae" order="Myrtales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">Latoangulatae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>