treatments-xml/data/3E/75/36/3E753653C216FFC324BFFEEAFA19FC07.xml
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<document id="C599EEC2A5ABE87EBFD1A696EE23ACE3" ID-CLB-Dataset="6237" ID-DOI="10.11646/zootaxa.4974.1.3" ID-GBIF-Dataset="c7efa945-0e70-48aa-b2d4-e09beeffeb2b" ID-ISSN="1175-5326" ID-Zenodo-Dep="4772138" ID-ZooBank="0FD68AAC-F4C5-4926-A7C7-EA475A2FACA2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621413917890" checkinUser="plazi" docAuthor="Hoese, Douglass F. &amp; Hammer, Michael P." docDate="2021" docId="3E753653C216FFC324BFFEEAFA19FC07" docLanguage="en" docName="zootaxa.4974.1.3.pdf" docOrigin="Zootaxa 4974 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glossogobius" docType="key" docVersion="6" lastPageNumber="112" masterDocId="C24C4E2BC237FFE22428FFCDFF85FB3B" masterDocTitle="A review of the Glossogobius giuris complex in Australia, with wider discussion on nomenclature and possible synonymies" masterLastPageNumber="115" masterPageNumber="79" pageNumber="112" updateTime="1698940067824" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="ADFC16A3D8F14FC4DBC224336B15E721">A review of the Glossogobius giuris complex in Australia, with wider discussion on nomenclature and possible synonymies</mods:title>
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<mods:name id="DEF0E6B0EAE514CCD5D05016EF44E0CD" type="personal">
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<mods:namePart id="F6DC8114671450BDBF35773502B6A5B6">Hoese, Douglass F.</mods:namePart>
<mods:affiliation id="65EDB25C7F38075E56AE9EA4CDA96173">Australian Museum, 1 William St., Sydney, NSW 2010, Australia.</mods:affiliation>
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<mods:namePart id="1A1DF64DDFE3F91D0AE02502AC87961A">Hammer, Michael P.</mods:namePart>
<mods:nameIdentifier id="F1DD7CA7899B46E5FEFAA81F0F09083D" type="ORCID">0000-0002-0981-4647</mods:nameIdentifier>
<mods:affiliation id="B913D1B7FED0789099ABC7FC330DC538">Museum &amp; Art Gallery of the Northern Territory, PO Box 4646 Darwin, Northern Territory 0801, Australia. michael. hammer @ magnt. net. au; https: // orcid. org / 0000 - 0002 - 0981 - 4647</mods:affiliation>
<mods:nameIdentifier id="2439968668692F5D068C23A98B8A7C71" type="email">michael.hammer@magnt.net.au</mods:nameIdentifier>
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<mods:title id="F0449C1BD920A1704A61A91C0D5CAB8D">Zootaxa</mods:title>
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<mods:date id="C1314911C3349241F2563AEBB1F45932">2021</mods:date>
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<subSubSection id="FEC6D4CEC216FFC324BFFEEAFD41FA7B" box="[151,708,294,321]" pageId="33" pageNumber="112" type="nomenclature">
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<heading id="ED2B3029C216FFC324BFFEEAFD41FA7B" bold="true" box="[151,708,294,321]" fontSize="11" level="1" pageId="33" pageNumber="112" reason="1">
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Key to species of
<taxonomicName id="71DCFCC6C216FFC32546FEEBFD8CFA7A" ID-CoL="4P99" ID-ENA="150297" authority="Gill, 1859" box="[366,521,294,321]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="112" phylum="Chordata" rank="genus">
<emphasis id="84A85B57C216FFC32546FEEBFD8CFA7A" bold="true" box="[366,521,294,321]" italics="true" pageId="33" pageNumber="112">glossogobius</emphasis>
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from australia
</emphasis>
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<subSubSection id="FEC6D4CEC216FFC324BFFEA1FA19FC07" pageId="33" pageNumber="112" type="key">
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<paragraph id="B6638745C216FFC324BFFEA1FA19FAA5" blockId="33.[151,1437,364,1852]" pageId="33" pageNumber="112">
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1. A pair of barbels at tip of chin. Estuarine and lower reaches of rivers from Weipa to norther eastern Queensland, widespread in western Pacific northward to
<collectingCountry id="CECBC7D5C216FFC325D4FE44FDB6FAA5" box="[508,563,393,414]" name="Japan" pageId="33" pageNumber="112">Japan</collectingCountry>
and east to
<collectingCountry id="CECBC7D5C216FFC3268EFE44FD42FAA4" box="[678,711,393,415]" name="Fiji" pageId="33" pageNumber="112">Fiji</collectingCountry>
................................................
<taxonomicName id="71DCFCC6C216FFC3213AFE44FA19FAA5" ID-CoL="6KJQX" baseAuthorityName="Weber" baseAuthorityYear="1894" box="[1298,1436,393,414]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="112" phylum="Chordata" rank="species" species="bicirrhosus">
<emphasis id="84A85B57C216FFC3213AFE44FA19FAA5" box="[1298,1436,393,414]" italics="true" pageId="33" pageNumber="112">G. bicirrhosus</emphasis>
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<keyLead id="0D2D2470C216FFC324BFFE6BFA19FA80" box="[151,1436,422,444]" pageId="33" pageNumber="112">1. No barbels at tip of chin (mental fraenum may be bilobed)..................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="0D289FE0C216FFC324BFFE09FA19F90A" pageId="33" pageNumber="112">
<paragraph id="B6638745C216FFC324BFFE09FA19FACC" blockId="33.[151,1437,364,1852]" pageId="33" pageNumber="112">
<keyLead id="0D2D2470C216FFC324BFFE09FA19FACC" pageId="33" pageNumber="112">
2. Cheek papillae arranged in 8 or more vertical rows. Gill rakers on outer face of first arch elongate, usually 1216. Mangroves from north-western
<collectingCountry id="CECBC7D5C216FFC32588FE2CFE7DFACC" box="[416,504,481,503]" name="Australia" pageId="33" pageNumber="112">Australia</collectingCountry>
to north-eastern eastern
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, widespread in western Pacific.........
<taxonomicName id="71DCFCC6C216FFC320DEFE2CFA19FACC" ID-CoL="3GDNJ" baseAuthorityName="Macleay" baseAuthorityYear="1883" box="[1270,1436,481,503]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="112" phylum="Chordata" rank="species" species="circumspectus">
<emphasis id="84A85B57C216FFC320DEFE2CFA19FACC" box="[1270,1436,481,503]" italics="true" pageId="33" pageNumber="112">G. circumspectus</emphasis>
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<paragraph id="B6638745C216FFC324BFFE33FA19F90A" blockId="33.[151,1437,364,1852]" pageId="33" pageNumber="112">
<keyLead id="0D2D2470C216FFC324BFFE33FA19F90A" pageId="33" pageNumber="112">2. Cheek papillae arranged in longitudinal rows, or rarely in few short vertical rows. Rakers on outer face of first arch short, 512................................................................................................ 3</keyLead>
</paragraph>
</keyStep>
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3. Predorsal naked or scaled forward to above operculum, scale rows 07. A conspicuous black spot at posterior end of first dorsal fin. Midline of body with thin elongate spots, sometimes forming a broken stripe. Lower attachment of gill membranes behind posterior preopercular margin. Maximum size
<quantity id="71242AA0C216FFC32689FDBEFD61F9B2" box="[673,740,627,649]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="33" pageNumber="112" unit="mm" value="50.0">50 mm</quantity>
SL...................................................... 4
</keyLead>
</paragraph>
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3. Predorsal scaled forward to above preoperculum, scale rows 1330. No spot or small black spot at posterior end of first dorsal fin. Midline of body with deep elongate spots or blotches. Lower attachment of gill membrane under or before posterior preopercular margin. Maximum size up to
<quantity id="71242AA0C216FFC32676FD01FD28F9DA" box="[606,685,716,737]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="112" unit="mm" value="300.0">300 mm</quantity>
SL.......................................................... 5
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<paragraph id="B6638745C216FFC324BFFD24FA19F86C" blockId="33.[151,1437,364,1852]" pageId="33" pageNumber="112">
<keyLead id="0D2D2470C216FFC324BFFD24FA19F86C" pageId="33" pageNumber="112">
4. Predorsal largely naked, a few scales at side of nape. Anal rays I, 67. No distinct lateral canal tube with pore at each end above operculum. Longitudinal scale count 2429. Transverse scale count 78.5. Body with elongate spots along midside forming a broken stripe. Western rivers of Cape York region, Queensland and Arnhem Land, Northern Territory and southern New
<collectingCountry id="CECBC7D5C216FFC324CAFC8CFEAFF86C" box="[226,298,833,855]" name="Guinea-Bissau" pageId="33" pageNumber="112">Guinea</collectingCountry>
.......................................................................................
<taxonomicName id="71DCFCC6C216FFC3216EFC8CFA19F86C" authorityName="Hoese, Allen &amp; Hadiaty" authorityYear="2017" box="[1350,1436,833,855]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="112" phylum="Chordata" rank="species" species="nanus">
<emphasis id="84A85B57C216FFC3216EFC8CFA19F86C" box="[1350,1436,833,855]" italics="true" pageId="33" pageNumber="112">G. nanus</emphasis>
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<keyLead id="0D2D2470C216FFC324BFFC93FA19F8F0" pageId="33" pageNumber="112">
4. Midline of predorsal with 27 scale rows, reaching forward to above operculum. A lateral canal tube with pore at each end above operculum. Anal rays usually I, 8. Longitudinal scale count 2931. Transverse scale count usually 8.510.5. Body with slightly elongate thin spots, with bands extending from spots. Endemic to Wet Tropics (Mossman to Russell-Mulgrave System), north-eastern
<collectingRegion id="741849A7C216FFC3254EFC7BFE5FF8F7" box="[358,474,950,972]" country="Australia" name="Queensland" pageId="33" pageNumber="112">Queensland</collectingRegion>
.`...............................................................
<taxonomicName id="71DCFCC6C216FFC320DAFC7AFA19F8F0" ID-CoL="3GDN9" authorityName="Hoese &amp; Allen" authorityYear="2009" box="[1266,1436,950,972]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="112" phylum="Chordata" rank="species" species="bellendenensis">
<emphasis id="84A85B57C216FFC320DAFC7AFA19F8F0" box="[1266,1436,950,972]" italics="true" pageId="33" pageNumber="112">G. bellendenensis</emphasis>
</taxonomicName>
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<paragraph id="B6638745C216FFC324BFFC1EFA19FF18" blockId="33.[151,1437,364,1852]" pageId="33" pageNumber="112">
<keyLead id="0D2D2470C216FFC324BFFC1EFA19FF18" pageId="33" pageNumber="112">
5. Head compressed. Second dorsal rays I,1012. No gill rakers developed on upper part of outer face of first arch, lower rakers 68. Precaudal vertebrae 13-15. Western rivers of Cape York, Queensland, Cato and Goyder river systems of the Northern Territory and southern rivers of New
<collectingCountry id="CECBC7D5C216FFC32600FBC3FDF5FF1F" box="[552,624,1038,1060]" name="Guinea-Bissau" pageId="33" pageNumber="112">Guinea</collectingCountry>
...................................................……
<taxonomicName id="71DCFCC6C216FFC3212CFBC2FA19FF18" ID-CoL="3GDNM" baseAuthorityName="Ramsay &amp; Ogilby" baseAuthorityYear="1886" box="[1284,1436,1038,1060]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="112" phylum="Chordata" rank="species" species="concavifrons">
<emphasis id="84A85B57C216FFC3212CFBC2FA19FF18" box="[1284,1436,1038,1060]" italics="true" pageId="33" pageNumber="112">G. concavifrons</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="B6638745C216FFC324BFFBE6FA19FF64" blockId="33.[151,1437,364,1852]" pageId="33" pageNumber="112">
<keyLead id="0D2D2470C216FFC324BFFBE6FA19FF64" pageId="33" pageNumber="112">5. Head depressed. Second dorsal I,9. Usually 12 rakers on upper part of outer face of first arch, rakers usually 812. Precaudal vertebrae 10-11....................................................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="0D289FE0C216FFC324BFFBABFA19FFEF" pageId="33" pageNumber="112">
<paragraph id="B6638745C216FFC324BFFBABFA19FFA2" blockId="33.[151,1437,364,1852]" pageId="33" pageNumber="112">
<keyLead id="0D2D2470C216FFC324BFFBABFA19FFA2" pageId="33" pageNumber="112">
6. Predorsal scales large, usually in 1216 rows. An infraorbital pore behind eye and 4 lateral canal pores above preoperculum and operculum. Coastal streams of north-eastern Queensland and widespread in northern New
<collectingCountry id="CECBC7D5C216FFC3206FFB49FB0AFFA1" box="[1095,1167,1156,1178]" name="Guinea-Bissau" pageId="33" pageNumber="112">Guinea</collectingCountry>
, ranging to
<collectingCountry id="CECBC7D5C216FFC32128FB49FABFFFA2" box="[1280,1338,1156,1177]" name="Japan" pageId="33" pageNumber="112">Japan</collectingCountry>
.
<taxonomicName id="71DCFCC6C216FFC3216BFB49FA19FFA2" ID-CoL="3GDNY" authorityName="Hoese &amp; Allen" authorityYear="2012" box="[1347,1436,1156,1177]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="112" phylum="Chordata" rank="species" species="illimis">
<emphasis id="84A85B57C216FFC3216BFB49FA19FFA2" box="[1347,1436,1156,1177]" italics="true" pageId="33" pageNumber="112">G. illimis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B6638745C216FFC324BFFB6CFA19FFEF" blockId="33.[151,1437,364,1852]" pageId="33" pageNumber="112">
<keyLead id="0D2D2470C216FFC324BFFB6CFA19FFEF" pageId="33" pageNumber="112">6. Predorsal scales small, usually in 1730 rows. An infraorbital pore and 2 lateral canal pores above preoperculum and operculum................................................................................................ 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="0D289FE0C216FFC324BFFB11FA19FE72" pageId="33" pageNumber="112">
<paragraph id="B6638745C216FFC324BFFB11FA19FE35" blockId="33.[151,1437,364,1852]" pageId="33" pageNumber="112">
<keyLead id="0D2D2470C216FFC324BFFB11FA19FE35" pageId="33" pageNumber="112">7. A line of papillae from middle of upper jaw to infraorbital pore behind eye with distinct posterior branch under eye (line 6). Longitudinal lines 7, 9 and 10 on cheek composed of multiple rows of papillae, except in small juveniles................ 8</keyLead>
</paragraph>
<paragraph id="B6638745C216FFC324BFFADAFA19FE72" blockId="33.[151,1437,364,1852]" pageId="33" pageNumber="112">
<keyLead id="0D2D2470C216FFC324BFFADAFA19FE72" pageId="33" pageNumber="112">7. Line of papillae from middle of upper jaw to infraorbital pore without branch. Longitudinal lines 7, 9 and 10 on cheek composed of single row of papillae........................................................................... 9</keyLead>
</paragraph>
</keyStep>
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<paragraph id="B6638745C216FFC324BFFA9CFA19FE84" blockId="33.[151,1437,364,1852]" pageId="33" pageNumber="112">
<keyLead id="0D2D2470C216FFC324BFFA9CFA19FE84" pageId="33" pageNumber="112">
8. Operculum with few or no scales anteriorly on operculum. Predorsal scales reach to above preoperculum with a gap behind the posterior interorbital pore. Chin uniformly pigmented, often with a V-shaped dark bar behind chin; often with black marking on distal margins of first dorsal fin. North-western
<collectingRegion id="741849A7C216FFC326E6FA41FC05FE99" box="[718,896,1420,1442]" country="Australia" name="Northern Territory" pageId="33" pageNumber="112">Northern Territory</collectingRegion>
and
<collectingRegion id="741849A7C216FFC32787FA41FBD9FE99" box="[943,1116,1420,1442]" country="Australia" name="Western Australia" pageId="33" pageNumber="112">Western Australia</collectingRegion>
, with populations extending well inland........................................................................................
<taxonomicName id="71DCFCC6C216FFC3216FFA64FA19FE84" ID-CoL="3GDNT" box="[1351,1436,1449,1471]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="112" phylum="Chordata" rank="species" species="giuris">
<emphasis id="84A85B57C216FFC3216FFA64FA19FE84" box="[1351,1436,1449,1471]" italics="true" pageId="33" pageNumber="112">G. giuris</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B6638745C216FFC324BFFA0BFA18FD6A" blockId="33.[151,1437,364,1852]" pageId="33" pageNumber="112">
<keyLead id="0D2D2470C216FFC324BFFA0BFA18FD6A" pageId="33" pageNumber="112">
8. Operculum with scales extending about ⅔ to ¾ of upper operculum in two to four rows. Predorsal scaled to or almost up to posterior interorbital pore, often obscuring postorbital pore. Chin with no melanophores centrally or scattered melanophores and bordered laterally by dark longitudinal stripes; pigment behind chin light or dark without a distinct transverse band, no prominent black marks on distal margin of first dorsal fin. Coastal rivers of eastern
<collectingRegion id="741849A7C216FFC32040F9D3FB5CFD0F" box="[1128,1241,1566,1588]" country="Australia" name="Queensland" pageId="33" pageNumber="112">Queensland</collectingRegion>
and
<collectingRegion id="741849A7C216FFC32121F9D3FE8EFD6A" country="Australia" name="Northern Territory" pageId="33" pageNumber="112">Northern Territory</collectingRegion>
………..…….
<emphasis id="84A85B57C216FFC325B8F9F1FA18FD6A" box="[400,1437,1596,1617]" italics="true" pageId="33" pageNumber="112">
…….........................................................................
<taxonomicName id="71DCFCC6C216FFC3211CF9F1FA18FD6A" authorityName="De Vis" authorityYear="1884" box="[1332,1437,1596,1617]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="112" phylum="Chordata" rank="species" species="laticeps">G. laticeps</taxonomicName>
</emphasis>
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</paragraph>
</keyStep>
<keyStep id="0D289FE0C216FFC324BFF994FA19FC07" pageId="33" pageNumber="112">
<paragraph id="B6638745C216FFC324BFF994FA19FDFC" blockId="33.[151,1437,364,1852]" pageId="33" pageNumber="112">
<keyLead id="0D2D2470C216FFC324BFF994FA19FDFC" pageId="33" pageNumber="112">
9. Chin without a distinct bilobed mental frenum. Snout long 1011% SL. Head large, 3436% SL. Mouth large reaching to below anterior margin of pupil, 1113% SL. Distance from posterior end of eye to posterior preopercular margin long, 78% SL. No bump on snout under anterior nostril. Predorsal scales usually more than 24. Rivers of northern
<collectingCountry id="CECBC7D5C216FFC320A9F959FB5CFD91" box="[1153,1241,1684,1706]" name="Australia" pageId="33" pageNumber="112">Australia</collectingCountry>
east of Darwin area across to north-eastern
<collectingRegion id="741849A7C216FFC325E9F97CFDB7FDFC" box="[449,562,1713,1735]" country="Australia" name="Queensland" pageId="33" pageNumber="112">Queensland</collectingRegion>
................................................................
<taxonomicName id="71DCFCC6C216FFC32116F97CFA19FDFC" ID-CoL="3GDN8" authorityName="Akihito &amp; Meguro" authorityYear="1975" box="[1342,1436,1713,1735]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="112" phylum="Chordata" rank="species" species="aureus">
<emphasis id="84A85B57C216FFC32116F97CFA19FDFC" box="[1342,1436,1713,1735]" italics="true" pageId="33" pageNumber="112">G. aureus</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="B6638745C216FFC324BFF903FA19FC07" blockId="33.[151,1437,364,1852]" pageId="33" pageNumber="112">
<keyLead id="0D2D2470C216FFC324BFF903FA19FC07" pageId="33" pageNumber="112">
9. Chin with a distinct bilobed mental frenum. Snout short, 910% SL. Head short, 3032% SL. Mouth reaching to below anterior margin of eye, 911% SL. Distance from posterior end of eye to posterior preopercular margin short, 56% SL. A small bump on snout below anterior nostril. Predorsal scales usually less than 24. Confined to rivers of northern
<collectingCountry id="CECBC7D5C216FFC320BAF8C4FB6FFC24" box="[1170,1258,1801,1823]" name="Australia" pageId="33" pageNumber="112">Australia</collectingCountry>
and southern New
<collectingCountry id="CECBC7D5C216FFC324CAF8EBFEAFFC07" box="[226,298,1830,1852]" name="Guinea-Bissau" pageId="33" pageNumber="112">Guinea</collectingCountry>
, but not known from eastern Queensland.....................................................
<taxonomicName id="71DCFCC6C216FFC32112F8EAFA19FC07" ID-CoL="6KK39" authorityName="Hoese &amp; Allen" authorityYear="2012" box="[1338,1436,1831,1852]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="112" phylum="Chordata" rank="species" species="munroi">
<emphasis id="84A85B57C216FFC32112F8EAFA19FC07" box="[1338,1436,1831,1852]" italics="true" pageId="33" pageNumber="112">G. munroi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</subSection>
</document>