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<document id="1A7027903FDCD69254C3BAFC75E23BC7" ID-CLB-Dataset="286280" ID-DOI="10.11646/zootaxa.5403.4.2" ID-GBIF-Dataset="ef287197-ddaf-4d39-a05e-11a259d5bd44" ID-ISSN="1175-5326" ID-Zenodo-Dep="10562097" ID-ZooBank="1C98335E-4B25-426A-B058-A078EA5F3FE7" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1706105144170" checkinUser="plazi" docAuthor="Wang, Fa-Lei, Li, Yan-Long &amp; Gao, Chuan-Bu" docDate="2024" docId="3E6C87E6FFAFFFEDFF10FB86FF4AF9F2" docLanguage="en" docName="zootaxa.5403.4.2.pdf" docOrigin="Zootaxa 5403 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5403.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megistophylla keithi Wang &amp; Li &amp; Gao 2024, new species" docType="treatment" docUuid="9CACE86F-26D8-4BB7-8CA8-66AE07715031" docUuidSource="ZooBank" docVersion="4" lastPageNumber="440" masterDocId="C255FF9EFFAAFFE4FF87FFC5FFA8F85F" masterDocTitle="Description of two new Megistophylla Burmeister, 1855 species from China, with a redescription of M. andrewesi Moser, 1913 (Coleoptera: Scarabaeidae: Melolonthinae)" masterLastPageNumber="446" masterPageNumber="431" pageNumber="436" updateTime="1706202214864" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="A1F5FD8C0ECF19450FE526A1A38F909F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CA4117A64CE7E525503C3B8D2DFE7096">
<mods:title id="0FB04AAA8F6461AC1074C7BB858186DB">Description of two new Megistophylla Burmeister, 1855 species from China, with a redescription of M. andrewesi Moser, 1913 (Coleoptera: Scarabaeidae: Melolonthinae)</mods:title>
</mods:titleInfo>
<mods:name id="9A7882541F3EE012639B080C45C50F97" type="personal">
<mods:role id="BFBC13608F143912B2210CF0F0E2B802">
<mods:roleTerm id="4BBA24964A1CF122E9A7632F4D858E76">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="23870EDD49F1B866454E799452467143">Wang, Fa-Lei</mods:namePart>
<mods:affiliation id="4D4AAFE43B6488D79877CB910669AC6B">Dazhulin, Chongqing 621000, China.</mods:affiliation>
</mods:name>
<mods:name id="B5F2EDED7CA5BA4B92281012C67F9FA0" type="personal">
<mods:role id="1BC516B48976E16A43898B1FE9C4571B">
<mods:roleTerm id="5D5456989C4833BE50232808DB1AFB21">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0CA940708554785992E1312D5700DB5D">Li, Yan-Long</mods:namePart>
<mods:affiliation id="8EBE41B3FB2741D2AEF651A3A5FCA45A">177 Aoti Road W, Lixia District, Jinan City, Weifang 261000, China.</mods:affiliation>
</mods:name>
<mods:name id="3F96E9E4F2364B8AF8250C4C592EFEC6" type="personal">
<mods:role id="0B801F63DB60597C5A19FB2545297147">
<mods:roleTerm id="AE5DB7190380E7066AC0F8CF353ADF0F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="51FA6798B06D46769FD3CDA917AD6678">Gao, Chuan-Bu</mods:namePart>
<mods:affiliation id="94CCBBAD02189042B40F18FD31CCB44E">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang Road W., Guangzhou 510260, China.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="1AAA7B7C108EE635A4BBF60D40A31999">text</mods:typeOfResource>
<mods:relatedItem id="A944FE993A261BFF54854C0449BFD304" type="host">
<mods:titleInfo id="7A83FCCF1717CBEFCCCBCF20456DFA41">
<mods:title id="DF0BBE6912343BBF4AE17895BA478AF9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8383004338A43ABFC2EEA1F9D7FEBE27">
<mods:date id="AB697AF16CB44FCC465E003B9DB5BFF9">2024</mods:date>
<mods:detail id="7C8FBFC1CA3222D2C81871831E5C6D97" type="pubDate">
<mods:number id="0686096BB4E70AD40CEBF06203E45955">2024-01-23</mods:number>
</mods:detail>
<mods:detail id="0997B1A7BC170EB032E106460836CD4A" type="volume">
<mods:number id="A10E3811382A741B1E6AC6143943E544">5403</mods:number>
</mods:detail>
<mods:detail id="DCF829FB49E12D92489CAF5583A3228E" type="issue">
<mods:number id="0796C628ACAB1539DDE8D9662D8A0D13">4</mods:number>
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<mods:url id="21EFC23AF0CDD6ADDBED776BF456EDB2">http://dx.doi.org/10.11646/zootaxa.5403.4.2</mods:url>
</mods:location>
<mods:classification id="9081CF0EDAEBE407ED7127C478C41233">journal article</mods:classification>
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<mods:identifier id="60F92F678E530186EBD5035391B6793E" type="DOI">10.11646/zootaxa.5403.4.2</mods:identifier>
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</mods:mods>
<treatment id="3E6C87E6FFAFFFEDFF10FB86FF4AF9F2" ID-DOI="http://doi.org/10.5281/zenodo.10562121" ID-Zenodo-Dep="10562121" LSID="urn:lsid:zoobank.org:act:9CACE86F-26D8-4BB7-8CA8-66AE07715031" httpUri="http://treatment.plazi.org/id/3E6C87E6FFAFFFEDFF10FB86FF4AF9F2" lastPageId="9" lastPageNumber="440" pageId="5" pageNumber="436">
<subSubSection id="FEDF657BFFAFFFE1FF10FB86FD70FC01" box="[151,728,1091,1118]" pageId="5" pageNumber="436" type="nomenclature">
<paragraph id="B67A36F0FFAFFFE1FF10FB86FD70FC01" blockId="5.[151,728,1091,1154]" box="[151,728,1091,1118]" pageId="5" pageNumber="436">
<heading id="ED32819CFFAFFFE1FF10FB86FD70FC01" bold="true" box="[151,728,1091,1118]" fontSize="11" level="1" pageId="5" pageNumber="436" reason="1">
<emphasis id="84B1EAE2FFAFFFE1FF10FB86FD70FC01" bold="true" box="[151,728,1091,1118]" pageId="5" pageNumber="436">
<taxonomicName id="71C54D73FFAFFFE1FF10FB86FD93FC01" authority="Wang &amp; Gao" authorityName="Wang &amp; Li &amp; Gao" authorityYear="2024" box="[151,571,1091,1118]" class="Insecta" family="Scarabaeidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="436" phylum="Arthropoda" rank="species" species="keithi" status="sp. nov.">
<emphasis id="84B1EAE2FFAFFFE1FF10FB86FE26FC01" bold="true" box="[151,398,1091,1118]" italics="true" pageId="5" pageNumber="436">Megistophylla keithi</emphasis>
Wang &amp; Gao
</taxonomicName>
,
<taxonomicNameLabel id="9F825799FFAFFFE1FDCFFB81FD70FC01" box="[584,728,1092,1118]" pageId="5" pageNumber="436" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FEDF657BFFAFFFE1FF10FBADFB97FC9B" pageId="5" pageNumber="436" type="description">
<paragraph id="B67A36F0FFAFFFE1FF10FBADFE86FCDD" blockId="5.[151,728,1091,1154]" box="[151,302,1128,1154]" pageId="5" pageNumber="436">
(
<figureCitation id="2EFE2A75FFAFFFE1FF18FBADFE8EFCDD" box="[159,294,1128,1154]" captionStart-0="FIGURES 20–22" captionStart-1="FIGUES 23–30" captionStart-2="FIGURES 31–36" captionStartId-0="6.[151,264,1868,1892]" captionStartId-1="7.[151,245,1707,1731]" captionStartId-2="8.[151,261,977,1001]" captionTargetBox-0="[161,1429,190,1838]" captionTargetBox-1="[159,1427,189,1676]" captionTargetBox-2="[154,1428,189,945]" captionTargetId-0="figure-29@6.[151,1436,181,1844]" captionTargetId-1="figure-18@7.[151,1436,181,1684]" captionTargetId-2="figure-29@8.[151,1436,181,953]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURES 20–22. Holotype of Megistophylla keithi Wang &amp; Gao, new species. 20, habitus in dorsal view; 21, habitus in ventral view; 22, habitus in lateral view. Scale bar: 5.0 mm." captionText-1="FIGUES 23–30. Holotype of M. keithi Wang &amp; Gao, new species. 23, head in dorsal view; 24, antennae; 25, scutellum; 26, pygidium; 27, parameres in dorsal view; 28, parameres in ventral view; 29, 30, male genitalia in lateral views. Scale bar: 1.0 mm." captionText-2="FIGURES 31–36. Details of Paratype of M. keithi Wang &amp; Gao, new species. 31, endophallus in ventral view; 32, labrum in dorsal view; 33, labrum in ventral view; 34, mandible of inner side; 35, maxilla; 36, mentum in ventral view. Scale bar: 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10562109" figureDoi-1="http://doi.org/10.5281/zenodo.10569246" figureDoi-2="http://doi.org/10.5281/zenodo.10562111" httpUri-0="https://zenodo.org/record/10562109/files/figure.png" httpUri-1="https://zenodo.org/record/10569246/files/figure.png" httpUri-2="https://zenodo.org/record/10562111/files/figure.png" pageId="5" pageNumber="436">Figs. 20–36</figureCitation>
)
</paragraph>
<paragraph id="B67A36F0FFAFFFE1FF10FB69FB97FC9B" blockId="5.[151,1087,1196,1220]" box="[151,1087,1196,1220]" pageId="5" pageNumber="436">
http://zoobank.org/
<uri id="C2543AF2FFAFFFE1FED9FB69FB97FC9B" box="[350,1087,1196,1220]" pageId="5" pageNumber="436">
urn:lsid:zoobank.org:act:
<uuid id="C2630C25FFAFFFE1FDD1FB69FB97FC9B" box="[598,1087,1196,1220]" pageId="5" pageNumber="436">9CACE86F-26D8-4BB7-8CA8-66AE07715031</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="FEDF657BFFAFFFE1FF10FB36FB7FFD6F" pageId="5" pageNumber="436" type="materials_examined">
<paragraph id="B67A36F0FFAFFFE1FF10FB36FB7FFD6F" blockId="5.[151,1436,1266,2012]" pageId="5" pageNumber="436">
<emphasis id="84B1EAE2FFAFFFE1FF10FB36FEE0FD53" bold="true" box="[151,328,1266,1293]" pageId="5" pageNumber="436">Type material.</emphasis>
<materialsCitation id="06AD3CADFFAFFFE1FED6FB36FE42FD6F" collectionCode="MYNU" collectorName="Wei-Zong Yang" country="China" county="Yingjiang County" location="Xima Village" municipality="Dehong" pageId="5" pageNumber="436" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="84B1EAE2FFAFFFE1FED6FB36FE16FD52" bold="true" box="[337,446,1267,1293]" pageId="5" pageNumber="436">
<typeStatus id="697E8852FFAFFFE1FED6FB36FE16FD52" box="[337,446,1267,1293]" pageId="5" pageNumber="436" type="holotype">Holotype</typeStatus>
</emphasis>
,
<specimenCount id="A0C3FD79FFAFFFE1FE48FB36FE4CFD52" box="[463,484,1267,1293]" count="1" pageId="5" pageNumber="436" type="male">♂</specimenCount>
(
<collectionCode id="D0D4AE35FFAFFFE1FE71FB36FDFBFD52" box="[502,595,1267,1293]" pageId="5" pageNumber="436">MYNU</collectionCode>
):
<collectingCountry id="CED27660FFAFFFE1FDE0FB36FD6CFD52" box="[615,708,1267,1293]" name="China" pageId="5" pageNumber="436">CHINA</collectingCountry>
:
<collectingRegion id="7401F812FFAFFFE1FD56FB36FC3CFD53" box="[721,916,1266,1293]" country="China" name="Yunnan" pageId="5" pageNumber="436">Yunnan Province</collectingRegion>
/
<collectingMunicipality id="561EAC8AFFAFFFE1FC28FB37FBA3FD53" box="[943,1035,1266,1292]" pageId="5" pageNumber="436">Dehong</collectingMunicipality>
,
<collectingCounty id="5F1B4E7CFFAFFFE1FB90FB37FB49FD52" box="[1047,1249,1266,1293]" pageId="5" pageNumber="436">Yingjiang County</collectingCounty>
/
<location id="B31A602BFFAFFFE1FB7AFB37FA30FD53" LSID="urn:lsid:plazi:treatment:3E6C87E6FFAFFFEDFF10FB86FF4AF9F2:B31A602BFFAFFFE1FB7AFB37FA30FD53" box="[1277,1432,1266,1292]" country="China" county="Yingjiang County" municipality="Dehong" name="Xima Village" pageId="5" pageNumber="436" stateProvince="Yunnan">Xima Village</location>
, 2019.VI /
<collectorName id="1B305326FFAFFFE1FE8EFAD3FE12FD6E" box="[265,442,1302,1329]" pageId="5" pageNumber="436">Wei-Zong Yang</collectorName>
leg.
</materialsCitation>
<materialsCitation id="06AD3CADFFAFFFE1FE76FAD2FB7BFD6F" box="[497,1235,1302,1329]" collectionCode="MYNU, CLYL" collectorName="Wei-Zong Yang" country="China" county="Yingjiang County" location="Xima Village" municipality="Dehong" pageId="5" pageNumber="436" specimenCount="2" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="84B1EAE2FFAFFFE1FE76FAD2FDC1FD6E" bold="true" box="[497,617,1303,1329]" pageId="5" pageNumber="436">
<typeStatus id="697E8852FFAFFFE1FE76FAD2FDC1FD6E" box="[497,617,1303,1329]" pageId="5" pageNumber="436" type="paratype">Paratypes</typeStatus>
</emphasis>
,
<specimenCount id="A0C3FD79FFAFFFE1FDF0FAD2FD1EFD6E" box="[631,694,1303,1329]" count="2" pageId="5" pageNumber="436" type="male">2 ♂♂</specimenCount>
(
<collectionCode id="D0D4AE35FFAFFFE1FD43FAD2FCB6FD6E" box="[708,798,1303,1329]" pageId="5" pageNumber="436">MYNU</collectionCode>
,
<collectionCode id="D0D4AE35FFAFFFE1FCAEFAD2FCDDFD6E" box="[809,885,1303,1329]" pageId="5" pageNumber="436">CLYL</collectionCode>
): same data as for the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FEDF657BFFAFFFECFF40FAFFFDA0FF39" lastPageId="8" lastPageNumber="439" pageId="5" pageNumber="436" type="description">
<paragraph id="B67A36F0FFAFFFE1FF40FAFFFEF0FD26" blockId="5.[151,1436,1266,2012]" pageId="5" pageNumber="436">
<emphasis id="84B1EAE2FFAFFFE1FF40FAFFFDB9FD0A" bold="true" box="[199,529,1338,1365]" pageId="5" pageNumber="436">
Description of the
<typeStatus id="697E8852FFAFFFE1FE2DFAFEFDB9FD0A" box="[426,529,1339,1365]" pageId="5" pageNumber="436" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="2EFE2A75FFAFFFE1FDA5FAFFFD04FD0B" box="[546,684,1338,1364]" captionStart-0="FIGURES 20–22" captionStart-1="FIGUES 23–30" captionStart-2="FIGURES 31–36" captionStartId-0="6.[151,264,1868,1892]" captionStartId-1="7.[151,245,1707,1731]" captionStartId-2="8.[151,261,977,1001]" captionTargetBox-0="[161,1429,190,1838]" captionTargetBox-1="[159,1427,189,1676]" captionTargetBox-2="[154,1428,189,945]" captionTargetId-0="figure-29@6.[151,1436,181,1844]" captionTargetId-1="figure-18@7.[151,1436,181,1684]" captionTargetId-2="figure-29@8.[151,1436,181,953]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURES 20–22. Holotype of Megistophylla keithi Wang &amp; Gao, new species. 20, habitus in dorsal view; 21, habitus in ventral view; 22, habitus in lateral view. Scale bar: 5.0 mm." captionText-1="FIGUES 23–30. Holotype of M. keithi Wang &amp; Gao, new species. 23, head in dorsal view; 24, antennae; 25, scutellum; 26, pygidium; 27, parameres in dorsal view; 28, parameres in ventral view; 29, 30, male genitalia in lateral views. Scale bar: 1.0 mm." captionText-2="FIGURES 31–36. Details of Paratype of M. keithi Wang &amp; Gao, new species. 31, endophallus in ventral view; 32, labrum in dorsal view; 33, labrum in ventral view; 34, mandible of inner side; 35, maxilla; 36, mentum in ventral view. Scale bar: 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10562109" figureDoi-1="http://doi.org/10.5281/zenodo.10569246" figureDoi-2="http://doi.org/10.5281/zenodo.10562111" httpUri-0="https://zenodo.org/record/10562109/files/figure.png" httpUri-1="https://zenodo.org/record/10569246/files/figure.png" httpUri-2="https://zenodo.org/record/10562111/files/figure.png" pageId="5" pageNumber="436">Figs. 20–36</figureCitation>
). Length
<quantity id="713D9B15FFAFFFE1FC9BFAFEFC2DFD0A" box="[796,901,1339,1365]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" pageId="5" pageNumber="436" unit="mm" value="20.2">20.2 mm</quantity>
, width
<quantity id="713D9B15FFAFFFE1FC5BFAFEFBEDFD0A" box="[988,1093,1339,1365]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="5" pageNumber="436" unit="mm" value="10.3">10.3 mm</quantity>
. Body oval-elongate, strongly convex (
<figureCitation id="2EFE2A75FFAFFFE1FF70FA9BFEE4FD26" box="[247,332,1374,1401]" captionStart="FIGURES 20–22" captionStartId="6.[151,264,1868,1892]" captionTargetBox="[161,1429,190,1838]" captionTargetId="figure-29@6.[151,1436,181,1844]" captionTargetPageId="6" captionText="FIGURES 20–22. Holotype of Megistophylla keithi Wang &amp; Gao, new species. 20, habitus in dorsal view; 21, habitus in ventral view; 22, habitus in lateral view. Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10562109" httpUri="https://zenodo.org/record/10562109/files/figure.png" pageId="5" pageNumber="436">Fig. 22</figureCitation>
).
</paragraph>
<paragraph id="B67A36F0FFAFFFE1FF40FA47FD10FDC3" blockId="5.[151,1436,1266,2012]" box="[199,696,1410,1437]" pageId="5" pageNumber="436">
<emphasis id="84B1EAE2FFAFFFE1FF40FA47FEA0FDC3" box="[199,264,1410,1436]" italics="true" pageId="5" pageNumber="436">Color</emphasis>
: The whole body dark reddish-brown.
</paragraph>
<paragraph id="B67A36F0FFAFFFE1FF40FA63FBC7FEC6" blockId="5.[151,1436,1266,2012]" pageId="5" pageNumber="436">
<emphasis id="84B1EAE2FFAFFFE1FF40FA63FEACFD9F" box="[199,260,1446,1472]" italics="true" pageId="5" pageNumber="436">Head</emphasis>
: Clypeus subtrapezoidal, as wide as frons; anterior margin reflexed, distinctly incurve at middle, surface with dense, deep punctures, fronto-clypeal suture complete and straight (
<figureCitation id="2EFE2A75FFAFFFE1FC3BFA0FFBA7FDBA" box="[956,1039,1482,1509]" captionStart="FIGUES 23–30" captionStartId="7.[151,245,1707,1731]" captionTargetBox="[159,1427,189,1676]" captionTargetId="figure-18@7.[151,1436,181,1684]" captionTargetPageId="7" captionText="FIGUES 23–30. Holotype of M. keithi Wang &amp; Gao, new species. 23, head in dorsal view; 24, antennae; 25, scutellum; 26, pygidium; 27, parameres in dorsal view; 28, parameres in ventral view; 29, 30, male genitalia in lateral views. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10569246" httpUri="https://zenodo.org/record/10569246/files/figure.png" pageId="5" pageNumber="436">Fig. 23</figureCitation>
). Frons densely punctate; punctures mostly connected to each other; compound eyes comparatively large, canthus with several setae, head approximately 0.56 times wider than interocular. Frontal carina distinctly raised. Vertex largely punctate anteriorly and setiferous laterally. Antennae with 10 antennomeres, antennal club composed of 8 antennomeres, strongly curved outwards, antennal club more than 4.0 times longer than antennomeres 1–2 combined; antennomere 1 twice as long as antennomere 2; antennomere 3 consist of the basal handle and extended blade (
<figureCitation id="2EFE2A75FFAFFFE1FB89F9BBFBCBFEC6" box="[1038,1123,1662,1689]" captionStart="FIGUES 23–30" captionStartId="7.[151,245,1707,1731]" captionTargetBox="[159,1427,189,1676]" captionTargetId="figure-18@7.[151,1436,181,1684]" captionTargetPageId="7" captionText="FIGUES 23–30. Holotype of M. keithi Wang &amp; Gao, new species. 23, head in dorsal view; 24, antennae; 25, scutellum; 26, pygidium; 27, parameres in dorsal view; 28, parameres in ventral view; 29, 30, male genitalia in lateral views. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10569246" httpUri="https://zenodo.org/record/10569246/files/figure.png" pageId="5" pageNumber="436">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="B67A36F0FFAFFFE1FF40F967FEA8FF13" blockId="5.[151,1436,1266,2012]" pageId="5" pageNumber="436">
<emphasis id="84B1EAE2FFAFFFE1FF40F967FEE2FEE3" box="[199,330,1698,1724]" italics="true" pageId="5" pageNumber="436">Mouthparts</emphasis>
: Labrum strongly depressed at middle; ventral side setiferous; dorsal side with dense short setae along depression; both sides triangular-shaped (
<figureCitation id="2EFE2A75FFAFFFE1FD20F903FC85FEBE" box="[679,813,1734,1761]" captionStart="FIGURES 31–36" captionStartId="8.[151,261,977,1001]" captionTargetBox="[154,1428,189,945]" captionTargetId="figure-29@8.[151,1436,181,953]" captionTargetPageId="8" captionText="FIGURES 31–36. Details of Paratype of M. keithi Wang &amp; Gao, new species. 31, endophallus in ventral view; 32, labrum in dorsal view; 33, labrum in ventral view; 34, mandible of inner side; 35, maxilla; 36, mentum in ventral view. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10562111" httpUri="https://zenodo.org/record/10562111/files/figure.png" pageId="5" pageNumber="436">Figs. 32, 33</figureCitation>
). Mandible with wrinkled molar lobe and rounded notch in incisor lobe (
<figureCitation id="2EFE2A75FFAFFFE1FECDF92FFE08FF5A" box="[330,416,1770,1797]" captionStart="FIGURES 31–36" captionStartId="8.[151,261,977,1001]" captionTargetBox="[154,1428,189,945]" captionTargetId="figure-29@8.[151,1436,181,953]" captionTargetPageId="8" captionText="FIGURES 31–36. Details of Paratype of M. keithi Wang &amp; Gao, new species. 31, endophallus in ventral view; 32, labrum in dorsal view; 33, labrum in ventral view; 34, mandible of inner side; 35, maxilla; 36, mentum in ventral view. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10562111" httpUri="https://zenodo.org/record/10562111/files/figure.png" pageId="5" pageNumber="436">Fig. 34</figureCitation>
). Distal galea of maxilla with 6 sharp teeth (
<figureCitation id="2EFE2A75FFAFFFE1FC1EF92FFC46FF5A" box="[921,1006,1770,1797]" captionStart="FIGURES 31–36" captionStartId="8.[151,261,977,1001]" captionTargetBox="[154,1428,189,945]" captionTargetId="figure-29@8.[151,1436,181,953]" captionTargetPageId="8" captionText="FIGURES 31–36. Details of Paratype of M. keithi Wang &amp; Gao, new species. 31, endophallus in ventral view; 32, labrum in dorsal view; 33, labrum in ventral view; 34, mandible of inner side; 35, maxilla; 36, mentum in ventral view. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10562111" httpUri="https://zenodo.org/record/10562111/files/figure.png" pageId="5" pageNumber="436">Fig. 35</figureCitation>
).
<collectingRegion id="7401F812FFAFFFE1FB84F92EFBCAFF5A" box="[1027,1122,1771,1797]" country="Myanmar" name="Chin" pageId="5" pageNumber="436">Mentum</collectingRegion>
distinctly bilobate at apical half and formed a small notch at middle of anterior margin; each paraglossa with 6–7 setae near base of labial palpus (
<figureCitation id="2EFE2A75FFAFFFE1FF18F8F7FF5CFF13" box="[159,244,1842,1868]" captionStart="FIGURES 31–36" captionStartId="8.[151,261,977,1001]" captionTargetBox="[154,1428,189,945]" captionTargetId="figure-29@8.[151,1436,181,953]" captionTargetPageId="8" captionText="FIGURES 31–36. Details of Paratype of M. keithi Wang &amp; Gao, new species. 31, endophallus in ventral view; 32, labrum in dorsal view; 33, labrum in ventral view; 34, mandible of inner side; 35, maxilla; 36, mentum in ventral view. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10562111" httpUri="https://zenodo.org/record/10562111/files/figure.png" pageId="5" pageNumber="436">Fig. 36</figureCitation>
).
</paragraph>
<paragraph id="B67A36F0FFAFFFE1FF40F892FE82FF83" blockId="5.[151,1436,1266,2012]" pageId="5" pageNumber="436">
<emphasis id="84B1EAE2FFAFFFE1FF40F892FE9EFF2F" box="[199,310,1879,1904]" italics="true" pageId="5" pageNumber="436">Pronotum</emphasis>
glabrous and punctate, sparser around posterior corners; with a smooth middle longitudinal line on posterior half. Anterior margin complete. Both sides flat in dorsal view, lateral margins crenulate and setiferous; posterior margin bear some short setae laterally on lateral punctures. Anterior angles obtuse, posterior angles obtuse and rounded.
</paragraph>
<caption id="E2BA6678FFACFFE2FF10F889FD0FFFD7" ID-DOI="http://doi.org/10.5281/zenodo.10562109" ID-Zenodo-Dep="10562109" httpUri="https://zenodo.org/record/10562109/files/figure.png" pageId="6" pageNumber="437" startId="6.[151,264,1868,1892]" targetBox="[161,1429,190,1838]" targetPageId="6" targetType="figure">
<paragraph id="B67A36F0FFACFFE2FF10F889FD0FFFD7" blockId="6.[151,1436,1868,1928]" pageId="6" pageNumber="437">
<emphasis id="84B1EAE2FFACFFE2FF10F889FEE4FF3B" bold="true" box="[151,332,1868,1892]" pageId="6" pageNumber="437">FIGURES 20–22</emphasis>
. Holotype of
<taxonomicName id="71C54D73FFACFFE2FE51F889FC87FF3B" authority="Wang &amp; Gao" authorityName="Wang &amp; Li &amp; Gao" authorityYear="2024" box="[470,815,1868,1893]" class="Insecta" family="Scarabaeidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="437" phylum="Arthropoda" rank="species" species="keithi" status="sp. nov.">
<emphasis id="84B1EAE2FFACFFE2FE51F889FD0CFF3B" box="[470,676,1868,1892]" italics="true" pageId="6" pageNumber="437">Megistophylla keithi</emphasis>
Wang &amp; Gao
</taxonomicName>
,
<taxonomicNameLabel id="9F825799FFACFFE2FCBFF888FC1BFF3B" box="[824,947,1868,1892]" pageId="6" pageNumber="437" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="84B1EAE2FFACFFE2FC3BF889FC74FF3B" bold="true" box="[956,988,1868,1892]" pageId="6" pageNumber="437">20,</emphasis>
habitus in dorsal view;
<emphasis id="84B1EAE2FFACFFE2FB4EF889FB41FF3B" bold="true" box="[1225,1257,1868,1892]" pageId="6" pageNumber="437">21,</emphasis>
habitus in ventral view;
<emphasis id="84B1EAE2FFACFFE2FF51F8B5FF5DFFD7" bold="true" box="[214,245,1904,1928]" pageId="6" pageNumber="437">22,</emphasis>
habitus in lateral view. Scale bar: 5.0 mm.
</paragraph>
</caption>
<caption id="E2BA6678FFADFFE3FF10F96EFF6EFF54" ID-DOI="http://doi.org/10.5281/zenodo.10569246" ID-Zenodo-Dep="10569246" httpUri="https://zenodo.org/record/10569246/files/figure.png" pageId="7" pageNumber="438" startId="7.[151,245,1707,1731]" targetBox="[159,1427,189,1676]" targetPageId="7">
<paragraph id="B67A36F0FFADFFE3FF10F96EFF6EFF54" blockId="7.[151,1437,1707,1803]" pageId="7" pageNumber="438">
<emphasis id="84B1EAE2FFADFFE3FF10F96EFE95FE9B" bold="true" box="[151,317,1707,1732]" pageId="7" pageNumber="438">FIGUES 23–30</emphasis>
.
<typeStatus id="697E8852FFADFFE3FECCF969FE01FE9B" box="[331,425,1708,1732]" pageId="7" pageNumber="438" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="71C54D73FFADFFE3FE49F969FD15FE9B" authority="Wang &amp; Gao" authorityName="Wang &amp; Li &amp; Gao" authorityYear="2024" box="[462,701,1708,1732]" class="Insecta" family="Scarabaeidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="keithi" status="new species">
<emphasis id="84B1EAE2FFADFFE3FE49F969FD82FE9B" box="[462,554,1708,1732]" italics="true" pageId="7" pageNumber="438">M. keithi</emphasis>
Wang &amp; Gao
</taxonomicName>
,
<taxonomicNameLabel id="9F825799FFADFFE3FD4FF969FCEEFE9B" box="[712,838,1708,1732]" pageId="7" pageNumber="438" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="84B1EAE2FFADFFE3FCD5F969FCD9FE9B" bold="true" box="[850,881,1708,1732]" pageId="7" pageNumber="438">23,</emphasis>
head in dorsal view;
<emphasis id="84B1EAE2FFADFFE3FBD4F969FBDAFE9B" bold="true" box="[1107,1138,1708,1732]" pageId="7" pageNumber="438">24,</emphasis>
antennae;
<emphasis id="84B1EAE2FFADFFE3FB64F969FAABFE9B" bold="true" box="[1251,1283,1708,1732]" pageId="7" pageNumber="438">25,</emphasis>
scutellum;
<emphasis id="84B1EAE2FFADFFE3FAFAF969FA34FE9B" bold="true" box="[1405,1436,1708,1732]" pageId="7" pageNumber="438">26,</emphasis>
pygidium;
<emphasis id="84B1EAE2FFADFFE3FE80F915FE8EFEB7" bold="true" box="[263,294,1744,1768]" pageId="7" pageNumber="438">27,</emphasis>
parameres in dorsal view;
<emphasis id="84B1EAE2FFADFFE3FDB9F915FDF6FEB7" bold="true" box="[574,606,1744,1768]" pageId="7" pageNumber="438">28,</emphasis>
parameres in ventral view;
<emphasis id="84B1EAE2FFADFFE3FCF9F915FC6DFEB7" bold="true" box="[894,965,1744,1768]" pageId="7" pageNumber="438">29, 30,</emphasis>
male genitalia in lateral views. Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph id="B67A36F0FFADFFE3FF40F89BFB33FF26" blockId="7.[151,1436,1886,2020]" box="[199,1179,1886,1913]" pageId="7" pageNumber="438">
<emphasis id="84B1EAE2FFADFFE3FF40F89BFE9DFF27" box="[199,309,1886,1912]" italics="true" pageId="7" pageNumber="438">Scutellum</emphasis>
: Scutellum glabrous and smooth, with several or without punctures (
<figureCitation id="2EFE2A75FFADFFE3FBBDF89BFB27FF26" box="[1082,1167,1886,1913]" captionStart="FIGUES 23–30" captionStartId="7.[151,245,1707,1731]" captionTargetBox="[159,1427,189,1676]" captionTargetId="figure-18@7.[151,1436,181,1684]" captionTargetPageId="7" captionText="FIGUES 23–30. Holotype of M. keithi Wang &amp; Gao, new species. 23, head in dorsal view; 24, antennae; 25, scutellum; 26, pygidium; 27, parameres in dorsal view; 28, parameres in ventral view; 29, 30, male genitalia in lateral views. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10569246" httpUri="https://zenodo.org/record/10569246/files/figure.png" pageId="7" pageNumber="438">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="B67A36F0FFADFFE3FF40F847FA23FF9F" blockId="7.[151,1436,1886,2020]" pageId="7" pageNumber="438">
<emphasis id="84B1EAE2FFADFFE3FF40F847FEA5FFC3" box="[199,269,1922,1948]" italics="true" pageId="7" pageNumber="438">Elytra</emphasis>
: Surface of elytra deep and dense punctate, slightly wrinkled beside sutural costa. Sutural costa developed, indistinct on anterior 1/3. Epipleuron disappears before hind angles; bearing a row of long soft setae before apex.
</paragraph>
<paragraph id="B67A36F0FFADFFE3FF40F80FFAAFFFBB" blockId="7.[151,1436,1886,2020]" box="[199,1287,1994,2020]" pageId="7" pageNumber="438">
<emphasis id="84B1EAE2FFADFFE3FF40F80FFE9AFFBB" box="[199,306,1994,2020]" italics="true" pageId="7" pageNumber="438">Pygidium</emphasis>
: Apex broadly rounded, surface with deep, sparse, inhomogeneous punctures (
<figureCitation id="2EFE2A75FFADFFE3FB21F80FFB52FFBB" box="[1190,1274,1994,2020]" captionStart="FIGUES 23–30" captionStartId="7.[151,245,1707,1731]" captionTargetBox="[159,1427,189,1676]" captionTargetId="figure-18@7.[151,1436,181,1684]" captionTargetPageId="7" captionText="FIGUES 23–30. Holotype of M. keithi Wang &amp; Gao, new species. 23, head in dorsal view; 24, antennae; 25, scutellum; 26, pygidium; 27, parameres in dorsal view; 28, parameres in ventral view; 29, 30, male genitalia in lateral views. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10569246" httpUri="https://zenodo.org/record/10569246/files/figure.png" pageId="7" pageNumber="438">Fig. 26</figureCitation>
).
</paragraph>
<caption id="E2BA6678FFA2FFECFF10FC14FAE7FC52" ID-DOI="http://doi.org/10.5281/zenodo.10562111" ID-Zenodo-Dep="10562111" httpUri="https://zenodo.org/record/10562111/files/figure.png" pageId="8" pageNumber="439" startId="8.[151,261,977,1001]" targetBox="[154,1428,189,945]" targetPageId="8" targetType="figure">
<paragraph id="B67A36F0FFA2FFECFF10FC14FAE7FC52" blockId="8.[151,1437,977,1038]" pageId="8" pageNumber="439">
<emphasis id="84B1EAE2FFA2FFECFF10FC14FEE1FBB6" bold="true" box="[151,329,977,1001]" pageId="8" pageNumber="439">FIGURES 31–36</emphasis>
. Details of Paratype of
<taxonomicName id="71C54D73FFA2FFECFDA8FC17FCA6FBB5" authority="Wang &amp; Gao" authorityName="Wang &amp; Li &amp; Gao" authorityYear="2024" box="[559,782,977,1002]" class="Insecta" family="Scarabaeidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="keithi" status="new species">
<emphasis id="84B1EAE2FFA2FFECFDA8FC17FD2EFBB6" box="[559,646,977,1001]" italics="true" pageId="8" pageNumber="439">M. keithi</emphasis>
Wang &amp; Gao
</taxonomicName>
,
<taxonomicNameLabel id="9F825799FFA2FFECFC91FC17FC38FBB6" box="[790,912,977,1001]" pageId="8" pageNumber="439" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="84B1EAE2FFA2FFECFC1FFC14FC1FFBB6" bold="true" box="[920,951,977,1001]" pageId="8" pageNumber="439">31,</emphasis>
endophallus in ventral view;
<emphasis id="84B1EAE2FFA2FFECFB5FFC14FB5FFBB6" bold="true" box="[1240,1271,977,1001]" pageId="8" pageNumber="439">32,</emphasis>
labrum in dorsal view;
<emphasis id="84B1EAE2FFA2FFECFF52FC30FF5CFC52" bold="true" box="[213,244,1013,1037]" pageId="8" pageNumber="439">33,</emphasis>
labrum in ventral view;
<emphasis id="84B1EAE2FFA2FFECFE6FFC30FDAFFC52" bold="true" box="[488,519,1013,1037]" pageId="8" pageNumber="439">34,</emphasis>
mandible of inner side;
<emphasis id="84B1EAE2FFA2FFECFD7EFC30FCB0FC52" bold="true" box="[761,792,1013,1037]" pageId="8" pageNumber="439">35,</emphasis>
maxilla;
<emphasis id="84B1EAE2FFA2FFECFCF1FC30FC3DFC52" bold="true" box="[886,917,1013,1037]" pageId="8" pageNumber="439">36,</emphasis>
mentum in ventral view. Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph id="B67A36F0FFA2FFECFF40FBA5FC8BFD56" blockId="8.[151,1437,1120,2040]" pageId="8" pageNumber="439">
<emphasis id="84B1EAE2FFA2FFECFF40FBA5FEA7FC26" box="[199,271,1120,1145]" italics="true" pageId="8" pageNumber="439">Venter</emphasis>
: Ventral thorax setiferous. Entire surface of metepisternum and metepimeron covered with dense and long yellow hair; metepisternum narrow. Abdominal sternite 1and posterior sternite 2 with dense punctures and dense shot setae; middle of abdominal sternites 3 and 4 sparsely punctate, punctures moderately dense on lateral sides; posterior half of abdominal sternite 5 with long soft setae; sternite 6 with dense punctures and short setae (
<figureCitation id="2EFE2A75FFA2FFECFAE8FB0EFF1EFD56" captionStart="FIGURES 20–22" captionStartId="6.[151,264,1868,1892]" captionTargetBox="[161,1429,190,1838]" captionTargetId="figure-29@6.[151,1436,181,1844]" captionTargetPageId="6" captionText="FIGURES 20–22. Holotype of Megistophylla keithi Wang &amp; Gao, new species. 20, habitus in dorsal view; 21, habitus in ventral view; 22, habitus in lateral view. Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10562109" httpUri="https://zenodo.org/record/10562109/files/figure.png" pageId="8" pageNumber="439">Fig. 21</figureCitation>
). Surface of propygidium with dense rugose punctures.
</paragraph>
<paragraph id="B67A36F0FFA2FFECFF40FAD6FED8FE5C" blockId="8.[151,1437,1120,2040]" pageId="8" pageNumber="439">
<emphasis id="84B1EAE2FFA2FFECFF40FAD6FF54FD73" box="[199,252,1299,1324]" italics="true" pageId="8" pageNumber="439">Legs</emphasis>
: Meso- and metafemora setiferous ventrally. Protibia tridentate, terminal tooth long and crooked, middle tooth acute, basal tooth nearly right angle, anterior inner margin with a row of long setae, inner spur long and sharp; ventral side with a longitudinal sharp ridge. Apex of mesotibia and metatibia widened, inner surface with a row of strong setae; apex of mesotibia with two equal length spurs; apex of metatibia with two unequal length spurs. Outer surface of meso- and metatibia with a transverse complete transverse carina; setae on carina longer than those on apex. Inner margin of dorsal surface of metatibia with 3 teeth. Each pro-, meso- and metatarsal claw with a tooth vertically medially.
</paragraph>
<paragraph id="B67A36F0FFA2FFECFF40F9C9FCA4FEA2" blockId="8.[151,1437,1120,2040]" pageId="8" pageNumber="439">
<emphasis id="84B1EAE2FFA2FFECFF40F9C9FE9CFE79" box="[199,308,1548,1574]" italics="true" pageId="8" pageNumber="439">Aedeagus</emphasis>
: Phallobase shorter than parameres and connected to each other by wrinkle membranous area on dorsum; anterior 1/4 of phallobase reduced. Parameres slender with two long, symmetric branches; branches curved about 45° downward at middle; posterior half of left and right branch connected to each other by membrane ventrally, with dense, irregular longitudinal sulcus dorsally and laterally; lateral sides of posterior half of two branches with a long depression at middle laterally; anterior half of two branches separated; apex crooked (in lateral view) and bloated, rounded (
<figureCitation id="2EFE2A75FFA2FFECFEE4F97AFE46FE86" box="[355,494,1727,1753]" captionStart="FIGUES 23–30" captionStartId="7.[151,245,1707,1731]" captionTargetBox="[159,1427,189,1676]" captionTargetId="figure-18@7.[151,1436,181,1684]" captionTargetPageId="7" captionText="FIGUES 23–30. Holotype of M. keithi Wang &amp; Gao, new species. 23, head in dorsal view; 24, antennae; 25, scutellum; 26, pygidium; 27, parameres in dorsal view; 28, parameres in ventral view; 29, 30, male genitalia in lateral views. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10569246" httpUri="https://zenodo.org/record/10569246/files/figure.png" pageId="8" pageNumber="439">Figs. 27–30</figureCitation>
). Endophallus with two long temones apophysis nearly parallel at dorsum, and with epithelium bears sparsely punctate a sensillae at middle.
</paragraph>
<paragraph id="B67A36F0FFA2FFECFF40F8C3FB1EFF7E" blockId="8.[151,1437,1120,2040]" box="[199,1206,1798,1825]" pageId="8" pageNumber="439">
<emphasis id="84B1EAE2FFA2FFECFF40F8C3FEE5FF7F" bold="true" box="[199,333,1798,1824]" pageId="8" pageNumber="439">Variability.</emphasis>
<typeStatus id="697E8852FFA2FFECFED3F8C2FE69FF7E" box="[340,449,1799,1825]" pageId="8" pageNumber="439" type="paratype">Paratypes</typeStatus>
differ in size (body length
<quantity id="713D9B15FFA2FFECFD76F8C3FC3CFF7E" box="[753,916,1798,1825]" metricMagnitude="-2" metricUnit="m" metricValue="1.83" metricValueMax="1.98" metricValueMin="1.68" pageId="8" pageNumber="439" unit="mm" value="18.3" valueMax="19.8" valueMin="16.8">16.8–19.8 mm</quantity>
and width 9.1–10.0 mm).
</paragraph>
<paragraph id="B67A36F0FFA2FFECFF40F8EFFE0AFF1B" blockId="8.[151,1437,1120,2040]" box="[199,418,1834,1860]" pageId="8" pageNumber="439">
<emphasis id="84B1EAE2FFA2FFECFF40F8EFFE8DFF1B" bold="true" box="[199,293,1834,1860]" pageId="8" pageNumber="439">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="B67A36F0FFA2FFECFF40F88BFDA0FF39" blockId="8.[151,1437,1120,2040]" box="[199,520,1870,1896]" pageId="8" pageNumber="439">
<emphasis id="84B1EAE2FFA2FFECFF40F88BFEDDFF37" bold="true" box="[199,373,1870,1896]" pageId="8" pageNumber="439">
<taxonomicName id="71C54D73FFA2FFECFF40F88BFE8EFF37" box="[199,294,1870,1896]" class="Aves" family="Anatidae" genus="Chinese" kingdom="Animalia" order="Anseriformes" pageId="8" pageNumber="446" phylum="Chordata" rank="genus">Chinese</taxonomicName>
name.
</emphasis>
DZƶDzẅm
</paragraph>
</subSubSection>
<subSubSection id="FEDF657BFFA2FFEDFF40F8B7FCF5F8A6" lastPageId="9" lastPageNumber="440" pageId="8" pageNumber="439" type="diagnosis">
<paragraph id="B67A36F0FFA2FFEDFF40F8B7FCF5F8A6" blockId="8.[151,1437,1120,2040]" lastBlockId="9.[151,1437,151,429]" lastPageId="9" lastPageNumber="440" pageId="8" pageNumber="439">
<emphasis id="84B1EAE2FFA2FFECFF40F8B7FE67FFD3" bold="true" box="[199,463,1906,1932]" pageId="8" pageNumber="439">Differential diagnosis</emphasis>
.Based on the antennal club composed of eight antennomeres and body length,
<taxonomicName id="71C54D73FFA2FFECFF10F850FED4FFF0" authority="Wang &amp; Li &amp; Gao, 2024" authorityName="Wang &amp; Li &amp; Gao" authorityYear="2024" box="[151,380,1941,1967]" class="Insecta" family="Scarabaeidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="keithi" status="sp. nov.">
<emphasis id="84B1EAE2FFA2FFECFF10F850FED4FFF0" box="[151,380,1941,1967]" italics="true" pageId="8" pageNumber="439">Megistophylla keithi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F825799FFA2FFECFE05F853FDA0FFF0" box="[386,520,1941,1968]" pageId="8" pageNumber="439" rank="species">new species</taxonomicNameLabel>
shares similarities with two congeners:
<taxonomicName id="71C54D73FFA2FFECFC41F853FBF5FFF0" authorityName="Moser" authorityYear="1913" box="[966,1117,1941,1967]" class="Insecta" family="Scarabaeidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="84B1EAE2FFA2FFECFC41F853FBF5FFF0" box="[966,1117,1941,1967]" italics="true" pageId="8" pageNumber="439">M. andrewesi</emphasis>
</taxonomicName>
and
<taxonomicName id="71C54D73FFA2FFECFB15F853FF67FF8C" authority="Keith, 2020" authorityName="Keith" authorityYear="2020" class="Insecta" family="Scarabaeidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="lamdongensis">
<emphasis id="84B1EAE2FFA2FFECFB15F853FAF9FFF0" box="[1170,1361,1941,1967]" italics="true" pageId="8" pageNumber="439">M. lamdongensis</emphasis>
<bibRefCitation id="D2544B01FFA2FFECFAD0F850FF67FF8C" author="Keith, D." pageId="8" pageNumber="439" pagination="269 - 276" refId="ref7065" refString="Keith, D. (2020) Description de nouvelles especes du genre Megistophylla Burmeister, 1855. L'Entomologiste, 76 (5), 269 - 276." type="journal article" year="2020">Keith, 2020</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="D2544B01FFA2FFECFF67F87CFECDFF8C" author="Keith, D." box="[224,357,1977,2003]" pageId="8" pageNumber="439" pagination="269 - 276" refId="ref7065" refString="Keith, D. (2020) Description de nouvelles especes du genre Megistophylla Burmeister, 1855. L'Entomologiste, 76 (5), 269 - 276." type="journal article" year="2020">Keith 2020</bibRefCitation>
;
<bibRefCitation id="D2544B01FFA2FFECFEF5F87CFDBCFF8C" author="Li, C. - L. &amp; Yang, P. - S. &amp; Wang, C. - C." box="[370,532,1977,2004]" pageId="8" pageNumber="439" pagination="240 - 250" refId="ref6994" refString="Li, C. - L., Yang, P. - S. &amp; Wang, C. - C. (2016) Two new species of Megistophylla Burmeister (Coleoptera: Scarabaeidae: Melolonthinae) from Taiwan, with notes on the genus. Zootaxa, 4126 (2), 240 - 250. https: // doi. org / 10.11646 / zootaxa. 4126.2.4" type="journal article" year="2016">
Li
<emphasis id="84B1EAE2FFA2FFECFE13F87FFE60FF8C" box="[404,456,1977,2003]" italics="true" pageId="8" pageNumber="439">et al</emphasis>
. 2016
</bibRefCitation>
).
<taxonomicName id="71C54D73FFA2FFECFDAEF87CFCB8FF8C" authority="Wang &amp; Li &amp; Gao, 2024" authorityName="Wang &amp; Li &amp; Gao" authorityYear="2024" box="[553,784,1977,2003]" class="Insecta" family="Scarabaeidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="keithi" status="sp. nov.">
<emphasis id="84B1EAE2FFA2FFECFDAEF87CFCB8FF8C" box="[553,784,1977,2003]" italics="true" pageId="8" pageNumber="439">Megistophylla keithi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F825799FFA2FFECFC9DF87FFC0BFF8C" box="[794,931,1977,2004]" pageId="8" pageNumber="439" rank="species">new species</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="71C54D73FFA2FFECFB5AF87FFADFFF8C" authorityName="Moser" authorityYear="1913" box="[1245,1399,1977,2003]" class="Insecta" family="Scarabaeidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="84B1EAE2FFA2FFECFB5AF87FFADFFF8C" box="[1245,1399,1977,2003]" italics="true" pageId="8" pageNumber="439">M. andrewesi</emphasis>
</taxonomicName>
by its darker body color (
<figureCitation id="2EFE2A75FFA2FFECFE17F818FE45FFA8" box="[400,493,2013,2039]" captionStart="FIGUES 23–30" captionStartId="7.[151,245,1707,1731]" captionTargetBox="[159,1427,189,1676]" captionTargetId="figure-18@7.[151,1436,181,1684]" captionTargetPageId="7" captionText="FIGUES 23–30. Holotype of M. keithi Wang &amp; Gao, new species. 23, head in dorsal view; 24, antennae; 25, scutellum; 26, pygidium; 27, parameres in dorsal view; 28, parameres in ventral view; 29, 30, male genitalia in lateral views. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10569246" httpUri="https://zenodo.org/record/10569246/files/figure.png" pageId="8" pageNumber="439">Figs. 24</figureCitation>
, 41), a longer clypeus, antennomere 2 lacking a spike, antennomere 3 not thickened (
<figureCitation id="2EFE2A75FFA3FFEDFF18FF52FE83F8EE" box="[159,299,151,177]" captionStart="FIGURES 20–22" captionStartId="6.[151,264,1868,1892]" captionTargetBox="[161,1429,190,1838]" captionTargetId="figure-29@6.[151,1436,181,1844]" captionTargetPageId="6" captionText="FIGURES 20–22. Holotype of Megistophylla keithi Wang &amp; Gao, new species. 20, habitus in dorsal view; 21, habitus in ventral view; 22, habitus in lateral view. Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10562109" httpUri="https://zenodo.org/record/10562109/files/figure.png" pageId="9" pageNumber="440">Figs. 20–22</figureCitation>
,
<figureCitation id="2EFE2A75FFA3FFEDFEBDFF52FE2CF8EE" box="[314,388,151,177]" captionStart="FIGURES 37–40" captionStartId="10.[151,264,1860,1884]" captionTargetBox="[163,1426,189,1828]" captionTargetId="figure-29@10.[151,1436,181,1836]" captionTargetPageId="10" captionText="FIGURES 37–40. Syntype of Megistophylla andrewesi Moser, 1913. 37, habitus in dorsal view; 38, habitus in ventral view; 39, habitus in lateral view; 40, labels. Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10562113" httpUri="https://zenodo.org/record/10562113/files/figure.png" pageId="9" pageNumber="440">37–39</figureCitation>
), and displaying slender male genitalia (
<figureCitation id="2EFE2A75FFA3FFEDFCDAFF52FC44F8EE" box="[861,1004,151,177]" captionStart="FIGUES 23–30" captionStartId="7.[151,245,1707,1731]" captionTargetBox="[159,1427,189,1676]" captionTargetId="figure-18@7.[151,1436,181,1684]" captionTargetPageId="7" captionText="FIGUES 23–30. Holotype of M. keithi Wang &amp; Gao, new species. 23, head in dorsal view; 24, antennae; 25, scutellum; 26, pygidium; 27, parameres in dorsal view; 28, parameres in ventral view; 29, 30, male genitalia in lateral views. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10569246" httpUri="https://zenodo.org/record/10569246/files/figure.png" pageId="9" pageNumber="440">Figs. 29, 30</figureCitation>
, 47, 48). Furthermore, it deviates
<taxonomicName id="71C54D73FFA3FFEDFAF9FF52FE99F88A" authorityName="Keith" authorityYear="2020" class="Insecta" family="Scarabaeidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="440" phylum="Arthropoda" rank="species" species="lamdongensis">
<emphasis id="84B1EAE2FFA3FFEDFAF9FF52FE99F88A" italics="true" pageId="9" pageNumber="440">M. lamdongensis</emphasis>
</taxonomicName>
due to its shorter antennal club, pronouncedly curved sides of pronotum (
<figureCitation id="2EFE2A75FFA3FFEDFBEFFF7EFB13F88A" box="[1128,1211,187,213]" captionStart="FIGURES 20–22" captionStartId="6.[151,264,1868,1892]" captionTargetBox="[161,1429,190,1838]" captionTargetId="figure-29@6.[151,1436,181,1844]" captionTargetPageId="6" captionText="FIGURES 20–22. Holotype of Megistophylla keithi Wang &amp; Gao, new species. 20, habitus in dorsal view; 21, habitus in ventral view; 22, habitus in lateral view. Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10562109" httpUri="https://zenodo.org/record/10562109/files/figure.png" pageId="9" pageNumber="440">Fig. 20</figureCitation>
;
<bibRefCitation id="D2544B01FFA3FFEDFB40FF7EFAEEF88A" author="Keith, D." box="[1223,1350,187,213]" pageId="9" pageNumber="440" pagination="269 - 276" refId="ref7065" refString="Keith, D. (2020) Description de nouvelles especes du genre Megistophylla Burmeister, 1855. L'Entomologiste, 76 (5), 269 - 276." type="journal article" year="2020">Keith 2020</bibRefCitation>
, fig. 5), and slender male genitalia (
<figureCitation id="2EFE2A75FFA3FFEDFE4BFF1AFDFCF8A6" box="[460,596,223,249]" captionStart="FIGUES 23–30" captionStartId="7.[151,245,1707,1731]" captionTargetBox="[159,1427,189,1676]" captionTargetId="figure-18@7.[151,1436,181,1684]" captionTargetPageId="7" captionText="FIGUES 23–30. Holotype of M. keithi Wang &amp; Gao, new species. 23, head in dorsal view; 24, antennae; 25, scutellum; 26, pygidium; 27, parameres in dorsal view; 28, parameres in ventral view; 29, 30, male genitalia in lateral views. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10569246" httpUri="https://zenodo.org/record/10569246/files/figure.png" pageId="9" pageNumber="440">Figs. 29, 30</figureCitation>
;
<bibRefCitation id="D2544B01FFA3FFEDFDE7FF1AFD48F8A6" author="Keith, D." box="[608,736,223,249]" pageId="9" pageNumber="440" pagination="269 - 276" refId="ref7065" refString="Keith, D. (2020) Description de nouvelles especes du genre Megistophylla Burmeister, 1855. L'Entomologiste, 76 (5), 269 - 276." type="journal article" year="2020">Keith 2020</bibRefCitation>
, figs. 6, 7).
</paragraph>
</subSubSection>
<subSubSection id="FEDF657BFFA3FFEDFF40FEC6FF72F93A" pageId="9" pageNumber="440" type="etymology">
<paragraph id="B67A36F0FFA3FFEDFF40FEC6FF72F93A" blockId="9.[151,1437,151,429]" pageId="9" pageNumber="440">
<emphasis id="84B1EAE2FFA3FFEDFF40FEC6FEE6F942" bold="true" box="[199,334,259,285]" pageId="9" pageNumber="440">Etymology.</emphasis>
The new species is named after Denis Keith, a renowned scarab researcher from
<collectingCountry id="CED27660FFA3FFEDFA89FEC6FAF5F942" box="[1294,1373,259,285]" name="France" pageId="9" pageNumber="440">France</collectingCountry>
, who graciously provided us with his taxonomic articles. These articles have proved to be of significant assistance to our study.
</paragraph>
</subSubSection>
<subSubSection id="FEDF657BFFA3FFEDFF40FEABFF4AF9F2" pageId="9" pageNumber="440" type="distribution">
<paragraph id="B67A36F0FFA3FFEDFF40FEABFF4AF9F2" blockId="9.[151,1437,151,429]" pageId="9" pageNumber="440">
<emphasis id="84B1EAE2FFA3FFEDFF40FEABFEC9F9D7" bold="true" box="[199,353,366,392]" pageId="9" pageNumber="440">Distribution.</emphasis>
Known only from the
<typeStatus id="697E8852FFA3FFEDFDDEFEAAFD21F9D6" box="[601,649,367,393]" pageId="9" pageNumber="440">type</typeStatus>
locality in Yingjiang County, southwestern region of
<collectingRegion id="7401F812FFA3FFEDFB50FEAAFA30F9D6" box="[1239,1432,367,393]" country="China" name="Yunnan" pageId="9" pageNumber="440">Yunnan Province</collectingRegion>
,
<collectingCountry id="CED27660FFA3FFEDFF10FE56FF76F9F2" box="[151,222,403,429]" name="China" pageId="9" pageNumber="440">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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