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<document ID-DOI="10.11646/palaeoentomology.5.4.12" ID-ISSN="2624-2834" ID-Zenodo-Dep="7333367" ID-ZooBank="E4F9017E-513F-4607-833D-92782FB23DA1" approvalRequired="5" approvalRequired_for_treatments="5" checkinTime="1668764398756" checkinUser="plazi" docAuthor="LI, YAN-DA, HUANG, DI‑YING &amp; CAI, CHEN‑YANG" docDate="2022" docId="CB2EEE4BFFAFFFE79C31E0DCFA02F782" docLanguage="en" docName="palaeoentomology.5.4.12.pdf" docOrigin="Palaeoentomology 5 (4)" docSource="http://dx.doi.org/10.11646/palaeoentomology.5.4.12" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.2:Palaeontomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeontomology.2018-.journal_article" docStyleVersion="2" docTitle="Nothattagenus LI &amp; HUANG &amp; CAI 2022, gen. nov." docType="treatment" docVersion="2" lastPageNumber="390" masterDocId="37179633FFAFFFE79F1BE53AFF95FF96" masterDocTitle="“ Attagenus ” burmiticus from mid-Cretaceous amber reinterpreted as a member of Orphilinae (Coleoptera: Dermestidae)" masterLastPageNumber="394" masterPageNumber="390" pageNumber="390" updateTime="1668764769353" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>“ Attagenus ” burmiticus from mid-Cretaceous amber reinterpreted as a member of Orphilinae (Coleoptera: Dermestidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>LI, YAN-DA</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>HUANG, DI‑YING</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>CAI, CHEN‑YANG</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Palaeoentomology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>390</mods:start>
<mods:end>394</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/palaeoentomology.5.4.12</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/palaeoentomology.5.4.12</mods:identifier>
<mods:identifier type="ISSN">2624-2834</mods:identifier>
<mods:identifier type="Zenodo-Dep">7333367</mods:identifier>
<mods:identifier type="ZooBank">E4F9017E-513F-4607-833D-92782FB23DA1</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7333371" ID-Zenodo-Dep="7333371" LSID="urn:lsid:plazi:treatment:CB2EEE4BFFAFFFE79C31E0DCFA02F782" httpUri="http://treatment.plazi.org/id/CB2EEE4BFFAFFFE79C31E0DCFA02F782" lastPageNumber="390" pageId="0" pageNumber="390">
<subSubSection box="[810,1287,1510,1537]" pageId="0" pageNumber="390" type="nomenclature">
<paragraph blockId="0.[810,1287,1510,1537]" box="[810,1287,1510,1537]" pageId="0" pageNumber="390">
<heading box="[810,1287,1510,1537]" fontSize="10" level="2" pageId="0" pageNumber="390" reason="2">
<emphasis bold="true" box="[810,1287,1510,1537]" pageId="0" pageNumber="390">
Genus
<taxonomicName authority="Li &amp; Cai" authorityName="LI &amp; HUANG &amp; CAI" authorityYear="2022" box="[893,1175,1510,1537]" class="Insecta" family="Dermestidae" genus="Nothattagenus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="390" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis bold="true" box="[893,1063,1510,1537]" italics="true" pageId="0" pageNumber="390">Nothattagenus</emphasis>
Li &amp; Cai
</taxonomicName>
<taxonomicNameLabel box="[1182,1287,1511,1537]" pageId="0" pageNumber="390" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="390" type="type_taxon">
<paragraph blockId="0.[810,1452,1588,2068]" pageId="0" pageNumber="390">
<emphasis bold="true" box="[810,963,1588,1612]" pageId="0" pageNumber="390">
<typeStatus box="[810,866,1588,1612]" pageId="0" pageNumber="390">Type</typeStatus>
species.
</emphasis>
<taxonomicName authority="Cai, Hava &amp; Huang, 2016" authorityName="Cai, Hava &amp; Huang" authorityYear="2016" class="Insecta" family="Dermestidae" genus="Attagenus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="390" phylum="Arthropoda" rank="species" species="burmiticus">
<emphasis box="[974,1206,1588,1612]" italics="true" pageId="0" pageNumber="390">Attagenus burmiticus</emphasis>
Cai, Háva &amp; Huang, 2016
</taxonomicName>
(
<emphasis box="[879,900,1627,1650]" italics="true" pageId="0" pageNumber="390">in</emphasis>
<bibRefCitation author="Cai, C. Y. &amp; Hava, J. &amp; Huang, D. Y." box="[906,1071,1626,1650]" pageId="0" pageNumber="390" pagination="95 - 99" refId="ref1669" refString="Cai, C. Y., Hava, J. &amp; Huang, D. Y. (2017) The earliest Attagenus species (Coleoptera: Dermestidae: Attageninae) from Upper Cretaceous Burmese amber. Cretaceous Research, 72, 95 - 99. https: // doi. org / 10.1016 / j. cretres. 2016.12.018" type="journal article" year="2017">
Cai
<emphasis box="[950,997,1626,1650]" italics="true" pageId="0" pageNumber="390">et al</emphasis>
., 2017
</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="390" type="etymology">
<paragraph blockId="0.[810,1452,1588,2068]" pageId="0" pageNumber="390">
<emphasis bold="true" box="[863,992,1664,1688]" pageId="0" pageNumber="390">Etymology.</emphasis>
The generic name is derived from the Greek “
<emphasis box="[822,893,1702,1726]" italics="true" pageId="0" pageNumber="390">nothos</emphasis>
”, false, and the generic name
<taxonomicName authority="Latreille." authorityName="Latreille." box="[1237,1452,1702,1726]" class="Insecta" family="Dermestidae" genus="Attagenus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="390" phylum="Arthropoda" rank="genus">
<emphasis box="[1237,1344,1702,1725]" italics="true" pageId="0" pageNumber="390">Attagenus</emphasis>
Latreille.
</taxonomicName>
The name is masculine in gender.
</paragraph>
<paragraph blockId="0.[810,1452,1588,2068]" pageId="0" pageNumber="390">
<emphasis bold="true" box="[863,1015,1778,1802]" pageId="0" pageNumber="390">Composition.</emphasis>
Only
<taxonomicName authorityName="LI &amp; HUANG &amp; CAI" authorityYear="2022" baseAuthorityName="Cai, Hava &amp; Huang" baseAuthorityYear="2022" box="[1099,1383,1778,1802]" class="Insecta" family="Dermestidae" genus="Nothattagenus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="390" phylum="Arthropoda" rank="species" species="burmiticus">
<emphasis box="[1099,1383,1778,1802]" italics="true" pageId="0" pageNumber="390">Nothattagenus burmiticus</emphasis>
</taxonomicName>
(Cai, Háva &amp; Huang,
<quantity box="[984,1065,1816,1840]" metricMagnitude="1" metricUnit="m" metricValue="5.12064" pageId="0" pageNumber="390" unit="in" value="2016.0">
2016
<emphasis box="[1044,1065,1817,1840]" italics="true" pageId="0" pageNumber="390">in</emphasis>
</quantity>
<bibRefCitation author="Cai, C. Y. &amp; Hava, J. &amp; Huang, D. Y." box="[1072,1231,1816,1840]" pageId="0" pageNumber="390" pagination="95 - 99" refId="ref1669" refString="Cai, C. Y., Hava, J. &amp; Huang, D. Y. (2017) The earliest Attagenus species (Coleoptera: Dermestidae: Attageninae) from Upper Cretaceous Burmese amber. Cretaceous Research, 72, 95 - 99. https: // doi. org / 10.1016 / j. cretres. 2016.12.018" type="journal article" year="2017">
Cai
<emphasis box="[1116,1162,1816,1840]" italics="true" pageId="0" pageNumber="390">et al</emphasis>
. 2017
</bibRefCitation>
)
<emphasis bold="true" box="[1244,1367,1816,1840]" pageId="0" pageNumber="390">comb. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="390" type="diagnosis">
<paragraph blockId="0.[810,1452,1588,2068]" pageId="0" pageNumber="390">
<emphasis bold="true" box="[863,980,1854,1878]" pageId="0" pageNumber="390">Diagnosis.</emphasis>
Dorsal surface pubescent (
<figureCitation box="[1294,1388,1854,1878]" captionStart="FIGURE 1" captionStartId="1.[136,235,1191,1215]" captionTargetBox="[151,1436,193,1168]" captionTargetId="figure-364@1.[151,1436,193,1168]" captionTargetPageId="1" captionText="FIGURE 1. General habitus of Nothattagenus burmiticus (Cai et al.) comb. nov., holotype, NIGP164890, under incident light. A, Dorsal view. B, Ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7333369" httpUri="https://zenodo.org/record/7333369/files/figure.png" pageId="0" pageNumber="390">Figs 1A</figureCitation>
,
<figureCitation box="[1406,1440,1854,1878]" captionStart="FIGURE 2" captionStartId="2.[136,235,1359,1383]" captionTargetBox="[151,1436,204,1336]" captionTargetId="figure-189@2.[151,1436,204,1336]" captionTargetPageId="2" captionText="FIGURE 2. General habitus of Nothattagenus burmiticus (Cai et al.) comb. nov., holotype, NIGP164890, under confocal microscopy. A, Head and thorax, ventral view. B, Head and thorax, ventrolateral view. C, Abdominal base, ventrolateral view. D, Abdominal apex, ventral view. E, Scutellum, dorsal view. Abbreviations: an, antenna; el, elytron; ey, compound eye; lbp, labial palp; md, mandible; msf, mesofemur; mstb, mesotibia; mtc, metacoxa; mtes, metanepisternum; mtf, metafemur; mxp, maxillary palp; oc, ocellus; pf, profemur; pn, pronotum; ptb, protibia; v1–5, ventrites 1–5. Scale bars: 300 μm." figureDoi="http://doi.org/10.5281/zenodo.7333377" httpUri="https://zenodo.org/record/7333377/files/figure.png" pageId="0" pageNumber="390">2E</figureCitation>
). Antennomere 11 abruptly narrowed in apical half (
<figureCitation captionStart="FIGURE 2" captionStartId="2.[136,235,1359,1383]" captionTargetBox="[151,1436,204,1336]" captionTargetId="figure-189@2.[151,1436,204,1336]" captionTargetPageId="2" captionText="FIGURE 2. General habitus of Nothattagenus burmiticus (Cai et al.) comb. nov., holotype, NIGP164890, under confocal microscopy. A, Head and thorax, ventral view. B, Head and thorax, ventrolateral view. C, Abdominal base, ventrolateral view. D, Abdominal apex, ventral view. E, Scutellum, dorsal view. Abbreviations: an, antenna; el, elytron; ey, compound eye; lbp, labial palp; md, mandible; msf, mesofemur; mstb, mesotibia; mtc, metacoxa; mtes, metanepisternum; mtf, metafemur; mxp, maxillary palp; oc, ocellus; pf, profemur; pn, pronotum; ptb, protibia; v1–5, ventrites 1–5. Scale bars: 300 μm." figureDoi="http://doi.org/10.5281/zenodo.7333377" httpUri="https://zenodo.org/record/7333377/files/figure.png" pageId="0" pageNumber="390">Fig. 2A, B</figureCitation>
). Mandibular apex normally situated (rather than displaced proximally) (
<figureCitation box="[963,1062,1968,1992]" captionStart="FIGURE 2" captionStartId="2.[136,235,1359,1383]" captionTargetBox="[151,1436,204,1336]" captionTargetId="figure-189@2.[151,1436,204,1336]" captionTargetPageId="2" captionText="FIGURE 2. General habitus of Nothattagenus burmiticus (Cai et al.) comb. nov., holotype, NIGP164890, under confocal microscopy. A, Head and thorax, ventral view. B, Head and thorax, ventrolateral view. C, Abdominal base, ventrolateral view. D, Abdominal apex, ventral view. E, Scutellum, dorsal view. Abbreviations: an, antenna; el, elytron; ey, compound eye; lbp, labial palp; md, mandible; msf, mesofemur; mstb, mesotibia; mtc, metacoxa; mtes, metanepisternum; mtf, metafemur; mxp, maxillary palp; oc, ocellus; pf, profemur; pn, pronotum; ptb, protibia; v1–5, ventrites 1–5. Scale bars: 300 μm." figureDoi="http://doi.org/10.5281/zenodo.7333377" httpUri="https://zenodo.org/record/7333377/files/figure.png" pageId="0" pageNumber="390">Fig. 2A</figureCitation>
). Metacoxae strongly transverse, reaching elytral margins laterally (
<figureCitation box="[1212,1305,2006,2030]" captionStart="FIGURE 2" captionStartId="2.[136,235,1359,1383]" captionTargetBox="[151,1436,204,1336]" captionTargetId="figure-189@2.[151,1436,204,1336]" captionTargetPageId="2" captionText="FIGURE 2. General habitus of Nothattagenus burmiticus (Cai et al.) comb. nov., holotype, NIGP164890, under confocal microscopy. A, Head and thorax, ventral view. B, Head and thorax, ventrolateral view. C, Abdominal base, ventrolateral view. D, Abdominal apex, ventral view. E, Scutellum, dorsal view. Abbreviations: an, antenna; el, elytron; ey, compound eye; lbp, labial palp; md, mandible; msf, mesofemur; mstb, mesotibia; mtc, metacoxa; mtes, metanepisternum; mtf, metafemur; mxp, maxillary palp; oc, ocellus; pf, profemur; pn, pronotum; ptb, protibia; v1–5, ventrites 1–5. Scale bars: 300 μm." figureDoi="http://doi.org/10.5281/zenodo.7333377" httpUri="https://zenodo.org/record/7333377/files/figure.png" pageId="0" pageNumber="390">Fig. 2C</figureCitation>
). Abdominal ventrites completely lined by sharp outer carina (
<figureCitation box="[1332,1419,2044,2068]" captionStart="FIGURE 2" captionStartId="2.[136,235,1359,1383]" captionTargetBox="[151,1436,204,1336]" captionTargetId="figure-189@2.[151,1436,204,1336]" captionTargetPageId="2" captionText="FIGURE 2. General habitus of Nothattagenus burmiticus (Cai et al.) comb. nov., holotype, NIGP164890, under confocal microscopy. A, Head and thorax, ventral view. B, Head and thorax, ventrolateral view. C, Abdominal base, ventrolateral view. D, Abdominal apex, ventral view. E, Scutellum, dorsal view. Abbreviations: an, antenna; el, elytron; ey, compound eye; lbp, labial palp; md, mandible; msf, mesofemur; mstb, mesotibia; mtc, metacoxa; mtes, metanepisternum; mtf, metafemur; mxp, maxillary palp; oc, ocellus; pf, profemur; pn, pronotum; ptb, protibia; v1–5, ventrites 1–5. Scale bars: 300 μm." figureDoi="http://doi.org/10.5281/zenodo.7333377" httpUri="https://zenodo.org/record/7333377/files/figure.png" pageId="0" pageNumber="390">Fig. 2D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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