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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.209.87681" ID-Pensoft-Pub="1314-2003-209-1" ID-Pensoft-UUID="2B9DFC7609F65C33A95B0065D5A5DBE2" ModsDocID="1314-2003-209-1" checkinTime="1663958886913" checkinUser="pensoft" docAuthor="Knapp, Sandra" docDate="2022" docId="BD958B41FA6F5F05A23B09F128B172C3" docLanguage="en" docName="PhytoKeys 209: 1-134" docOrigin="PhytoKeys 209" docPubDate="2022-09-23" docSource="http://dx.doi.org/10.3897/phytokeys.209.87681" docTitle="Lycianthes peranomala (Wernham ex. Ridl.) A. R. Bean, Austrobaileya 6 (3): 567. 2003." docType="treatment" docVersion="1" id="2B9DFC7609F65C33A95B0065D5A5DBE2" lastPageNumber="1" masterDocId="2B9DFC7609F65C33A95B0065D5A5DBE2" masterDocTitle="A revision of Lycianthes (Solanaceae) in Australia, New Guinea, and the Pacific" masterLastPageNumber="134" masterPageNumber="1" pageNumber="1" updateTime="1663958886913" updateUser="pensoft">
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<mods:title>A revision of Lycianthes (Solanaceae) in Australia, New Guinea, and the Pacific</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Knapp, Sandra</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7698-3945</mods:nameIdentifier>
<mods:affiliation>Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
<mods:nameIdentifier type="email">s.knapp@nhm.ac.uk</mods:nameIdentifier>
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<mods:date>2022</mods:date>
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<mods:number>2022-09-23</mods:number>
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<mods:number>209</mods:number>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.209.87681</mods:url>
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<treatment LSID="urn:lsid:plazi:treatment:BD958B41FA6F5F05A23B09F128B172C3" httpUri="http://treatment.plazi.org/id/BD958B41FA6F5F05A23B09F128B172C3" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
13.
<taxonomicName LSID="BD958B41-FA6F-5F05-A23B-09F128B172C3" authority="(Wernham ex. Ridl.) A. R. Bean, Austrobaileya 6 (3): 567. 2003." authorityName="(Wernham ex. Ridl.) A. R. Bean, Austrobaileya 6 (3): 567. 2003." class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes peranomala" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peranomala">Lycianthes peranomala (Wernham ex. Ridl.) A.R.Bean, Austrobaileya 6(3): 567. 2003.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Lycianthes peranomala (Wernhamam ex Ridl.) A. R. Bean. Reproduced from Ridley (1922: pl. 3062, as S. peranomalum Wernham ex Ridl.). Courtesy of permission of NHM Library and Archives, reproduced with permission." figureDoi="10.3897/phytokeys.209.87681.figure40" httpUri="https://binary.pensoft.net/fig/746689" pageId="0" pageNumber="1">Figs 40</figureCitation>
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Lycianthes peranomala herbarium specimen. Papua New Guinea. Chimbu: Takeuchi 11204 (A). Courtesy of the Herbarium of the Arnold Arboretum of Harvard University, reproduced with permission." figureDoi="10.3897/phytokeys.209.87681.figure41" httpUri="https://binary.pensoft.net/fig/746690" pageId="0" pageNumber="1">, 41</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum peranomalum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peranomalum">Solanum peranomalum</taxonomicName>
Wernham ex Ridl.,
<normalizedToken originalValue="Hookers">Hooker's</normalizedToken>
Icon. Pl. 31 (pt. 3): tab. 3062. 1922. Type. Indonesia. Papua: &quot;Mt. Carstenz [Canoe Camp on Utakwa River drainage]&quot; [Puncak Jaya or Mount Jaya], 45 m, 5 Dec 1912,
<emphasis italics="true" pageId="0" pageNumber="1">C.B. Kloss s.n.</emphasis>
(lectotype, designated by
<bibRefCitation author="Symon, DE" journalOrPublisher="Journal of the Adelaide Botanic Gardens" pageId="0" pageNumber="1" pagination="1 - 171" refId="B89" refString="Symon, DE, 1985. The Solanaceae of New Guinea. Journal of the Adelaide Botanic Gardens 8: 1 - 171" title="The Solanaceae of New Guinea." volume="8" year="1985">Symon 1985</bibRefCitation>
, pg. 58 [as holotype]: BM [BM001014584]).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Based on
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum peranomalum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peranomalum">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum peranomalum</emphasis>
</taxonomicName>
Wernham ex Ridl.
</paragraph>
<caption doi="10.3897/phytokeys.209.87681.figure40" httpUri="https://binary.pensoft.net/fig/746689" pageId="0" pageNumber="1" start="Figure 40" startId="F40">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 40.</emphasis>
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes peranomala" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peranomala">
<emphasis italics="true" pageId="0" pageNumber="1">Lycianthes peranomala</emphasis>
</taxonomicName>
(Wernhamam ex Ridl.) A.R.Bean. Reproduced from
<bibRefCitation author="Ridley, HN" journalOrPublisher="Botanische Jahrbuecher fuer Systematik, Pflanzengeschichte und Pflanzengeographie" pageId="0" pageNumber="1" refId="B70" refString="Ridley, HN, 1922. Solanum peranomalum. Hooker's Icones Plantarum 31(pt. 3): tab. 3062." title="Solanum peranomalum. Hooker's Icones Plantarum 31 (pt. 3): tab. 3062." year="1922">Ridley (1922</bibRefCitation>
: pl. 3062, as
<taxonomicName family="Solanaceae" kingdom="Plantae" lsidName="S. peranomalum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="peranomalum">
<emphasis italics="true" pageId="0" pageNumber="1">S. peranomalum</emphasis>
</taxonomicName>
Wernham ex Ridl.). Courtesy of permission of NHM Library and Archives, reproduced with permission.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Shrub to 3 m, or a woody climber with length not recorded; stems terete, sparsely pubescent with appressed stiff antrorse simple uniseriate 1-6-celled trichomes to 1 mm long, the basal cell of each enlarged and sometimes remaining as a pustule or bump; new growth densely stiff-pubescent, the trichomes simple, uniseriate and strongly antrorse; bark of older stems brown, glabrescent, somewhat rugose and corky. Sympodial units unifoliate or difoliate, if difoliate the leaves geminate, the leaves of a pair different in size and sometimes shape. Leaves simple; blades of major leaves 9-16 cm long, 4-7 cm wide, elliptic, widest at the middle, the two sides occasionally uneven in size with the basiscopic half narrower, concolorous or somewhat discolorous, chartaceous or coriaceous; adaxial surfaces bullate (fide
<emphasis italics="true" pageId="0" pageNumber="1">Takeuchi 11204</emphasis>
), glabrous, the midrib keeled; abaxial surfaces glabrous, the veins prominent; principal veins 8-9 pairs, yellowish abaxially; base acute, oblique; margins entire; apex acuminate with an elongate drip-tip; petiole 0.7-1.1 cm long, sparsely pubescent with a few scattered stiff antrorse simple uniseriate trichomes on the adaxial surfaced and near the base; blades of minor leaves 1-2.5(4) cm long, 1-2.2 cm wide, elliptic to orbicular or heart-shaped, often apparently clasping the stem (&quot;retrorsely directed&quot; fide
<emphasis italics="true" pageId="0" pageNumber="1">Takeuchi 11204</emphasis>
), similar in texture and pubescent to the major leaves; base cordate or rounded; margins entire, usually revolute, sometimes markedly so; apex abruptly acute; petioles 0.1-0.2 cm long, glabrous or sometimes with a few simple trichomes like those of the stems. Inflorescences axillary fascicles of 8-10 flowers, several open at once, sparsely pubescent with stiff antrorse trichomes to 0.5 mm long like those of the stems; pedicels at anthesis 0.6-0.7 cm long, ca. 0.5 mm in diameter at the base, ca. 1.5 mm in diameter at the apex, spreading (?), white or pale purple, sparsely pubescent with stiff antrorse simple uniseriate trichomes like those of the stems, ca. 0.5 mm long; pedicel scars clustered in the leaf axils; buds ellipsoid, the corolla ca. halfway exserted from the calyx tube before anthesis. Flowers 5-merous, heterostylous and unisexual, specimens with either short-styled flowers or long-styled flowers and fruit, the plants possibly dioecious. Calyx tube 2.5-3 mm long, 2-2.5 mm wide, urn-shaped, the rim somewhat constricted, thick and woody (dry) or fleshy (live plants?), with no appendages, sparsely pubescent with stiff trichomes like those of the pedicels, these deciduous. Corolla 0.6-0.8 cm in diameter, purple, stellate, lobed nearly to the base, interpetalar tissue absent, the lobes 3-4 mm wide, 1.2-2 mm wide, reflexed, thick and fleshy, adaxially glabrous with a prominent ridged midvein, abaxially glabrous or sparsely papillate, densely papillate on tips and margins. Stamens equal; filament tube minute; free portion of the filaments ca. 1 mm long, glabrous; anthers 1.5-2 mm long, 1-1.5 mm wide, plumply ellipsoid, yellow, poricidal at the tips, the pores distally directed, circular, not elongating to slits with age. Ovary conical, glabrous; style in short-styled flowers ca. 0.5 mm long, in long-styled flowers 4-4.5 mm long, glabrous; stigma capitate or minutely bilobed, the surfaces minutely papillate. Fruit a globose berry, 0.5-0.9 cm in diameter, colour at maturity not known, the pericarp brittle in dry material, glabrous, matte, opaque, fruiting pedicels 0.5-0.8 cm long, ca. 1 mm in diameter at the base, ca. 1.5 mm in diameter at the apex, stiff and somewhat woody and tuberculate, spreading; fruiting calyx a cup at the base of the fruit, covering less than 1/4 of the berry, somewhat tuberculate, with a few stiff trichomes, but these usually deciduous. Seeds 10-20 per berry, ca. 2 mm long, ca. 1.5 mm wide, flattened reniform or slightly tear-shaped, reddish brown, the surfaces deeply pitted especially on the thickened margins, the testal cells pentagonal in outline. Stone cells absent. Chromosome number not known.
</paragraph>
<caption doi="10.3897/phytokeys.209.87681.figure41" httpUri="https://binary.pensoft.net/fig/746690" pageId="0" pageNumber="1" start="Figure 41" startId="F41">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 41.</emphasis>
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes peranomala" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peranomala">
<emphasis italics="true" pageId="0" pageNumber="1">Lycianthes peranomala</emphasis>
</taxonomicName>
herbarium specimen. Papua New Guinea. Chimbu:
<emphasis italics="true" pageId="0" pageNumber="1">Takeuchi 11204</emphasis>
(A). Courtesy of the Herbarium of the Arnold Arboretum of Harvard University, reproduced with permission.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Distribution of Lycianthes peranomala." figureDoi="10.3897/phytokeys.209.87681.figure42" httpUri="https://binary.pensoft.net/fig/746691" pageId="0" pageNumber="1">42</figureCitation>
).
</emphasis>
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes peranomala" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peranomala">
<emphasis italics="true" pageId="0" pageNumber="1">Lycianthes peranomala</emphasis>
</taxonomicName>
is endemic to the island of New Guinea; known from Papua New Guinea (Chimbu, Madang, Oro) and Indonesia (Papua).
</paragraph>
<caption doi="10.3897/phytokeys.209.87681.figure42" httpUri="https://binary.pensoft.net/fig/746691" pageId="0" pageNumber="1" start="Figure 42" startId="F42">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 42.</emphasis>
Distribution of
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes peranomala" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peranomala">
<emphasis italics="true" pageId="0" pageNumber="1">Lycianthes peranomala</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology and habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes peranomala" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peranomala">
<emphasis italics="true" pageId="0" pageNumber="1">Lycianthes peranomala</emphasis>
</taxonomicName>
occurs in lowland rainforest and riverine forests, between 50 and 640 m elevation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="common names">
<paragraph pageId="0" pageNumber="1">Common names.</paragraph>
<paragraph pageId="0" pageNumber="1">None recorded.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="preliminary conservation assessment">
<paragraph pageId="0" pageNumber="1">Preliminary conservation assessment</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(
<bibRefCitation author="Hunziker, AT" journalOrPublisher="The Linnean Society of London, Academic Press, London" pageId="0" pageNumber="1" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B38" refString="2020. . http://www.iucnredlist.org/documents/RedListGuidelines.pdf" url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2020">IUCN 2020</bibRefCitation>
).
</emphasis>
EOO (189,160 km2 - LC); AOO (16km2 - EN).
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes peranomala" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peranomala">
<emphasis italics="true" pageId="0" pageNumber="1">Lycianthes peranomala</emphasis>
</taxonomicName>
is known from four localities, some of which are within protected areas. Based on EOO alone, it would merit a status of Least Concern, but given the few collections, its forest habitat and general lack of knowledge about the species, I propose a preliminary threat status of Vulnerable (VU [B2a, b(iii,iv)]) for
<taxonomicName class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" lsidName="L. peranomala" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peranomala">
<emphasis italics="true" pageId="0" pageNumber="1">L. peranomala</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="discussion">
<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes peranomala" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peranomala">
<emphasis italics="true" pageId="0" pageNumber="1">Lycianthes peranomala</emphasis>
</taxonomicName>
is vegetatively similar to
<taxonomicName class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" lsidName="L. impar" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="impar">
<emphasis italics="true" pageId="0" pageNumber="1">L. impar</emphasis>
</taxonomicName>
in its orbicular to heart-shaped minor leaves and relatively large, elliptic major leaves with prominent venation. They differ in trichome morphology,
<taxonomicName class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" lsidName="L. peranomala" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peranomala">
<emphasis italics="true" pageId="0" pageNumber="1">L. peranomala</emphasis>
</taxonomicName>
has sparse stiff, strongly antrorse trichomes on stems, veins near the leaf base and calyces, while those of
<taxonomicName class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" lsidName="L. impar" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="impar">
<emphasis italics="true" pageId="0" pageNumber="1">L. impar</emphasis>
</taxonomicName>
are soft and curling and found on stems only. Inflorescences of
<taxonomicName class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" lsidName="L. impar" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="impar">
<emphasis italics="true" pageId="0" pageNumber="1">L. impar</emphasis>
</taxonomicName>
are elongate with paired pedicel scars, while those of
<taxonomicName class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" lsidName="L. peranomala" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peranomala">
<emphasis italics="true" pageId="0" pageNumber="1">L. peranomala</emphasis>
</taxonomicName>
are strictly axillary.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Like
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes kaernbachii" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="kaernbachii">
<emphasis italics="true" pageId="0" pageNumber="1">Lycianthes kaernbachii</emphasis>
</taxonomicName>
and
<taxonomicName class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" lsidName="L. oliveriana" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="oliveriana">
<emphasis italics="true" pageId="0" pageNumber="1">L. oliveriana</emphasis>
</taxonomicName>
the flowers of
<taxonomicName class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" lsidName="L. peranomala" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peranomala">
<emphasis italics="true" pageId="0" pageNumber="1">L. peranomala</emphasis>
</taxonomicName>
are usually less than 1 cm in diameter with fleshy corolla lobes lacking any interpetalar tissue and the plants are possibly dioecious with short-styled flowers and long-styled flowers (and fruit) on different plants.
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes peranomala" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peranomala">
<emphasis italics="true" pageId="0" pageNumber="1">Lycianthes peranomala</emphasis>
</taxonomicName>
can be distinguished from those taxa in its minor leaves that are very dissimilar in shape to the major leaves; both
<taxonomicName class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" lsidName="L. kaernbachii" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="kaernbachii">
<emphasis italics="true" pageId="0" pageNumber="1">L. kaernbachii</emphasis>
</taxonomicName>
had
<taxonomicName class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" lsidName="L. oliveriana" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="oliveriana">
<emphasis italics="true" pageId="0" pageNumber="1">L. oliveriana</emphasis>
</taxonomicName>
have minor leaves that are different in size but not so strongly dissimilar in shape.
</paragraph>
<paragraph pageId="0" pageNumber="1">
The name
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum peranomalum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="peranomalum">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum peranomalum</emphasis>
</taxonomicName>
was proposed by Herbert Fuller Wernham, an assistant in the Botany Department of the British Museum, for plants collected by Cecil Boden Kloss on the 1912-1913 expedition led by the ornithologist Arthur Wollaston in a second attempt to reach the high peaks of the &quot;Snow Mountains&quot; (Mount Jaya) in the central range (
<bibRefCitation DOI="https://doi.org/10.1111/j.1095-8339.1916.tb00009.x" author="Ridley, HN" journalOrPublisher="Botanische Jahrbuecher fuer Systematik, Pflanzengeschichte und Pflanzengeographie" pageId="0" pageNumber="1" refId="B69" refString="Ridley, HN, 1916. https://doi.org/10.1111/j.1095-8339.1916.tb00009.x" url="https://doi.org/10.1111/j.1095-8339.1916.tb00009.x" year="1916">Ridley 1916</bibRefCitation>
). His publication of
<taxonomicName family="Solanaceae" kingdom="Plantae" lsidName="S. peranomalum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="peranomalum">
<emphasis italics="true" pageId="0" pageNumber="1">S. peranomalum</emphasis>
</taxonomicName>
was superseded by Nicholas
<normalizedToken originalValue="Ridleys">Ridley's</normalizedToken>
publication of the same name, with an illustration (Fig.
<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Lycianthes peranomala (Wernhamam ex Ridl.) A. R. Bean. Reproduced from Ridley (1922: pl. 3062, as S. peranomalum Wernham ex Ridl.). Courtesy of permission of NHM Library and Archives, reproduced with permission." figureDoi="10.3897/phytokeys.209.87681.figure40" httpUri="https://binary.pensoft.net/fig/746689" pageId="0" pageNumber="1">40</figureCitation>
) but a slightly different and less complete description a few months earlier. Ridley perhaps was inpatient for the Transactions to appear or had little regard for Wernham at the Museum, who had a tragic career cut short by mental health issues and alcoholism (
<bibRefCitation DOI="https://doi.org/10.2307/1219381" author="Stearn, WT" journalOrPublisher="Taxon" pageId="0" pageNumber="1" pagination="1 - 6" refId="B82" refString="Stearn, WT, 1981. Herbert Fuller Wernham (1879-1941), a centenary commemoration. Taxon 30 (1): 1 - 6, DOI: https://doi.org/10.2307/1219381" title="Herbert Fuller Wernham (1879 - 1941), a centenary commemoration." url="https://doi.org/10.2307/1219381" volume="30" year="1981">Stearn 1981</bibRefCitation>
). It is strange that neither of the two other species described by
<bibRefCitation author="Wernham, HF" editor="Ridley, HN" journalOrPublisher="Transactions of the Linnean Society of London" pageId="0" pageNumber="1" pagination="119 - 120" refId="B101" refString="Wernham, HF, 1916. Solanaceae. In: Ridley, HN, Ed., Report on the botany of the Wollaston expedition to Dutch New Guinea, 1912-1913. Transactions of the Linnean Society of London 9: 119 - 120" title="Solanaceae." volume="9" volumeTitle="Report on the botany of the Wollaston expedition to Dutch New Guinea, 1912 - 1913." year="1916">Wernham (1916)</bibRefCitation>
were published by Ridley (
<taxonomicName family="Solanaceae" kingdom="Plantae" lsidName="S. ridleyanum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="ridleyanum">
<emphasis italics="true" pageId="0" pageNumber="1">S. ridleyanum</emphasis>
</taxonomicName>
and
<taxonomicName family="Solanaceae" kingdom="Plantae" lsidName="S. wollastonii" order="Solanales" pageId="0" pageNumber="1" rank="species" species="wollastonii">
<emphasis italics="true" pageId="0" pageNumber="1">S. wollastonii</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Indonesia" location="Indonesia" specimenCount="1" typeStatus="type">
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
. [
<typeStatus>type</typeStatus>
only]
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="1936-03-22" collectingDateMax="2008-08-07" collectingDateMin="1936-03-22" country="US" county="Ohu Village" elevation="640" location="Crater Mountain Wildlife Management Area" municipality="Wara" specimenCount="1" stateProvince="Madang">
Papua New Guinea.
<emphasis bold="true" pageId="0" pageNumber="1">Chimbu</emphasis>
:
<location LSID="urn:lsid:plazi:treatment:BD958B41FA6F5F05A23B09F128B172C3:37C7367E08529C7D5A04F087D62156C7" country="US" county="Ohu Village" municipality="Wara" name="Crater Mountain Wildlife Management Area" stateProvince="Madang">Crater Mountain Wildlife Management Area</location>
,
<location LSID="urn:lsid:plazi:treatment:BD958B41FA6F5F05A23B09F128B172C3:15C132BCE7DD04BE14E4B2F48D6E07A7" country="US" county="Ohu Village" municipality="Wara" name="Vicinity of Haia" stateProvince="Madang">Vicinity of Haia</location>
, along the
<collectingMunicipality>Wara</collectingMunicipality>
oo streamcourse (first river
<locationDeviation location="Mt. Widau">E of Mt. Widau</locationDeviation>
),
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.4" unit="m" value="640.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.4" unit="m" value="640.0">640 m</elevation>
</quantity>
,
<collectingDate value="1996-09-16">16 Sep 1996</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="1">Takeuchi 11204</emphasis>
(A, BM, K, L, LAE, US).
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Madang">Madang</collectingRegion>
</emphasis>
:
<collectingCounty>Ohu Village</collectingCounty>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
</quantity>
,
<collectingDate value="2008-08-07">7 Aug 2008</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="1">Ctvrtecka 1644</emphasis>
(
<collectingCountry name="United States of America">US</collectingCountry>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Oro">Oro</collectingRegion>
</emphasis>
: Kokoda,
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.65" unit="m" value="365.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.65" unit="m" value="365.0">365 m</elevation>
</quantity>
,
<collectingDate value="1936-03-22">22 Mar 1936</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="1">Carr 16195</emphasis>
(BM, K, NY)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>