185 lines
18 KiB
XML
185 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1139.94586" ID-Pensoft-Pub="1313-2970-1139-137" ID-Pensoft-UUID="2D3F572308DA5BAEA2F11417AE507FD7" ID-ZooBank="57CDA88415C542A39CF789DBC60150C4" ModsDocID="1313-2970-1139-137" checkinTime="1673667859025" checkinUser="pensoft" docAuthor="Gattolliat, Jean-Luc, Samraoui, Boudjema, Benhadji, Nadhira, Kechemir, Lina, Zrelli, Sonia, El Yaagoubi, Sara, El Moutaouakil, Majida El Alami & Sartori, Michel" docDate="2023" docId="40FA990841FF56B289059090862C031A" docLanguage="en" docName="ZooKeys 1139: 137-163" docOrigin="ZooKeys 1139" docPubDate="2023-01-13" docSource="http://dx.doi.org/10.3897/zookeys.1139.94586" docTitle="Alainites Waltz & McCafferty 1994" docType="treatment" docVersion="1" id="2D3F572308DA5BAEA2F11417AE507FD7" lastPageNumber="137" masterDocId="2D3F572308DA5BAEA2F11417AE507FD7" masterDocTitle="Baetidae (Baetidae, Ephemeroptera) in the Maghreb: state of the art, key, and perspectives" masterLastPageNumber="163" masterPageNumber="137" pageNumber="137" updateTime="1673667859025" updateUser="pensoft">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>Baetidae (Baetidae, Ephemeroptera) in the Maghreb: state of the art, key, and perspectives</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Gattolliat, Jean-Luc</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5873-5083</mods:nameIdentifier>
|
|
<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1014 Lausanne, Switzerland & Department of Ecology and Evolution, Biophore, University of Lausanne, CH- 1015 Lausanne, Switzerland</mods:affiliation>
|
|
<mods:nameIdentifier type="email">jean-luc.gattolliat@vd.ch</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Samraoui, Boudjema</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0608-9021</mods:nameIdentifier>
|
|
<mods:affiliation>Laboratoire de Conservation des Zones Humides, University 8 mai 1945, Guelma, Algeria & Department of Biology, University Badji Mokhtar Annaba, Annaba, Algeria</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Benhadji, Nadhira</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9252-8432</mods:nameIdentifier>
|
|
<mods:affiliation>Laboratoire de Recherche Valorisation des Actions de L'homme Pour la Protection de L'environnement et Application en Sante Publique, University of Tlemcen, BP 119, 13000 Tlemcen, Algeria & Department of Hydrobiology, Institute of Biology, University of Szczecin, Felczaka street 3 c, 71 - 412 Szczecin, Poland</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Kechemir, Lina</mods:namePart>
|
|
<mods:affiliation>Faculte des Sciences agronomiques et des sciences biologiques, University of Mouloud Mammeri, Tizi-Ouzou, Algeria</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Zrelli, Sonia</mods:namePart>
|
|
<mods:affiliation>Unit of Hydrobiology, Laboratory of Environment Biomonitoring (LBE), Faculty of Sciences of Bizerta, University of Bizerta, 7021 Jarzouna, Tunisia & Honoris United Universities, Ecole Polytechnique Centrale, Avenue Mohamed V, 1002 Tunis, Tunisia</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>El Yaagoubi, Sara</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1860-6433</mods:nameIdentifier>
|
|
<mods:affiliation>Laboratory of Ecology, Systematics and Conservation of the Biodiversity, department of Biology, Faculty of Science, University of Abdelmalek Essaadi, Avenue Sebta, 93002 Tetouan, Morocco</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>El Moutaouakil, Majida El Alami</mods:namePart>
|
|
<mods:affiliation>Laboratory of Ecology, Systematics and Conservation of the Biodiversity, department of Biology, Faculty of Science, University of Abdelmalek Essaadi, Avenue Sebta, 93002 Tetouan, Morocco</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Sartori, Michel</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3397-3397</mods:nameIdentifier>
|
|
<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1014 Lausanne, Switzerland & Department of Ecology and Evolution, Biophore, University of Lausanne, CH- 1015 Lausanne, Switzerland</mods:affiliation>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2023</mods:date>
|
|
<mods:detail type="pubDate">
|
|
<mods:number>2023-01-13</mods:number>
|
|
</mods:detail>
|
|
<mods:detail type="volume">
|
|
<mods:number>1139</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>137</mods:start>
|
|
<mods:end>163</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/zookeys.1139.94586</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1139.94586</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1313-2970-1139-137</mods:identifier>
|
|
<mods:identifier type="ZooBank">57CDA88415C542A39CF789DBC60150C4</mods:identifier>
|
|
<mods:identifier type="Pensoft-UUID">2D3F572308DA5BAEA2F11417AE507FD7</mods:identifier>
|
|
</mods:mods>
|
|
<treatment LSID="urn:lsid:plazi:treatment:40FA990841FF56B289059090862C031A" httpUri="http://treatment.plazi.org/id/40FA990841FF56B289059090862C031A" lastPageNumber="137" pageId="0" pageNumber="137">
|
|
<subSubSection pageId="0" pageNumber="137" type="nomenclature">
|
|
<paragraph pageId="0" pageNumber="137">
|
|
2.
|
|
<taxonomicName ID-CoL="RCZ" LSID="40FA9908-41FF-56B2-8905-9090862C031A" authority="Waltz & McCafferty, 1994" authorityName="Waltz & McCafferty" authorityYear="1994" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites" order="Ephemeroptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">Alainites Waltz & McCafferty, 1994</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="137" type="diagnosis">
|
|
<paragraph pageId="0" pageNumber="137">Diagnosis.</paragraph>
|
|
<paragraph pageId="0" pageNumber="137">1) Antennae located close together with a well-developed carina in between; 2) paraproct with unique elongate prolongation on distal margin; 3) mouthparts in a hypognathous position giving to the body a characteristic curved posture in lateral view; 4) right mandible with a bifid prostheca.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="137" type="remarks">
|
|
<paragraph pageId="0" pageNumber="137">Remarks.</paragraph>
|
|
<paragraph pageId="0" pageNumber="137">
|
|
The position of the antenna and the general posture in lateral view (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Baetidae: larvae in toto in dorsal view (except 3 C lateral view) A Acentrella cf. sinaica B Alainites oukaimeden C Alainites sadati D Baetis (Patites) sp. E Baetis (Baetis) pavidus." figureDoi="10.3897/zookeys.1139.94586.figure3" httpUri="https://binary.pensoft.net/fig/795194" pageId="0" pageNumber="137">3C</figureCitation>
|
|
) easily separate
|
|
<taxonomicName authorityName="Waltz & McCafferty" authorityYear="1994" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites" order="Ephemeroptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="137">Alainites</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName authorityName="Novikova & Kluge" authorityYear="1987" class="Insecta" family="Baetidae" genus="Nigrobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nigrobaetis" order="Ephemeroptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="137">Nigrobaetis</emphasis>
|
|
</taxonomicName>
|
|
from other genera in the Maghreb. The prolongation of the distal margin of paraproct (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Baetidae: larval characters A right mandible: Alainites sadati B right mandible: Nigrobaetis rhithralis C right mandible: Baetis (Rhodobaetis) sp. D right mandible: Baetis (Patites) sp. E head frontal view: Alainites sp. F head frontal view: Baetis sp. G costal margin of gill IV: Baetis (Baetis) sp. H gill IV: Baetis (Rhodobaetis) sp. I costal margin of gill IV: Baetis (Rhodobaetis) sp. J paraproct: Nigrobaetis rhithralis K paraproct: Alainites sadati L distal margin tergite IV: Baetis (Rhodobaetis) sp. M distal margin tergite IV: Baetis (Patites) sp. N distal margin tergite IV: Baetis (Baetis) Baetis pavidus O foretibia: Acentrella cf. sinaica P foretibia: Baetis (Rhodobaetis) sp." figureDoi="10.3897/zookeys.1139.94586.figure2" httpUri="https://binary.pensoft.net/fig/795201" pageId="0" pageNumber="137">2K</figureCitation>
|
|
) and the bifid prostheca (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Baetidae: larval characters A right mandible: Alainites sadati B right mandible: Nigrobaetis rhithralis C right mandible: Baetis (Rhodobaetis) sp. D right mandible: Baetis (Patites) sp. E head frontal view: Alainites sp. F head frontal view: Baetis sp. G costal margin of gill IV: Baetis (Baetis) sp. H gill IV: Baetis (Rhodobaetis) sp. I costal margin of gill IV: Baetis (Rhodobaetis) sp. J paraproct: Nigrobaetis rhithralis K paraproct: Alainites sadati L distal margin tergite IV: Baetis (Rhodobaetis) sp. M distal margin tergite IV: Baetis (Patites) sp. N distal margin tergite IV: Baetis (Baetis) Baetis pavidus O foretibia: Acentrella cf. sinaica P foretibia: Baetis (Rhodobaetis) sp." figureDoi="10.3897/zookeys.1139.94586.figure2" httpUri="https://binary.pensoft.net/fig/795201" pageId="0" pageNumber="137">2A</figureCitation>
|
|
) unambiguously separate
|
|
<taxonomicName authorityName="Waltz & McCafferty" authorityYear="1994" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites" order="Ephemeroptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="137">Alainites</emphasis>
|
|
</taxonomicName>
|
|
from all the other Maghreb genera including
|
|
<taxonomicName authorityName="Novikova & Kluge" authorityYear="1987" class="Insecta" family="Baetidae" genus="Nigrobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nigrobaetis" order="Ephemeroptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="137">Nigrobaetis</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="137">
|
|
Three species of
|
|
<taxonomicName authorityName="Waltz & McCafferty" authorityYear="1994" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites" order="Ephemeroptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="137">Alainites</emphasis>
|
|
</taxonomicName>
|
|
are reported in the Maghreb:
|
|
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites muticus" order="Ephemeroptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="muticus">Alainites cf. muticus</taxonomicName>
|
|
(Linnaeus, 1758),
|
|
<taxonomicName baseAuthorityName="Thomas & Sartori" baseAuthorityYear="1991" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites oukaimeden" order="Ephemeroptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="oukaimeden">
|
|
<emphasis italics="true" pageId="0" pageNumber="137">Alainites oukaimeden</emphasis>
|
|
</taxonomicName>
|
|
(Thomas & Sartori, 1992) (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Baetidae: larvae in toto in dorsal view (except 3 C lateral view) A Acentrella cf. sinaica B Alainites oukaimeden C Alainites sadati D Baetis (Patites) sp. E Baetis (Baetis) pavidus." figureDoi="10.3897/zookeys.1139.94586.figure3" httpUri="https://binary.pensoft.net/fig/795194" pageId="0" pageNumber="137">3B</figureCitation>
|
|
) and
|
|
<taxonomicName authorityName="Thomas" authorityYear="1994" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites sadati" order="Ephemeroptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="sadati">
|
|
<emphasis italics="true" pageId="0" pageNumber="137">Alainites sadati</emphasis>
|
|
</taxonomicName>
|
|
Thomas, 1994 (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Baetidae: larvae in toto in dorsal view (except 3 C lateral view) A Acentrella cf. sinaica B Alainites oukaimeden C Alainites sadati D Baetis (Patites) sp. E Baetis (Baetis) pavidus." figureDoi="10.3897/zookeys.1139.94586.figure3" httpUri="https://binary.pensoft.net/fig/795194" pageId="0" pageNumber="137">3C</figureCitation>
|
|
).
|
|
<taxonomicName baseAuthorityName="Thomas & Sartori" baseAuthorityYear="1991" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites oukaimeden" order="Ephemeroptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="oukaimeden">
|
|
<emphasis italics="true" pageId="0" pageNumber="137">Alainites oukaimeden</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName lsidName="A. sadati" pageId="0" pageNumber="137" rank="species" species="sadati">
|
|
<emphasis italics="true" pageId="0" pageNumber="137">A. sadati</emphasis>
|
|
</taxonomicName>
|
|
are endemic to the Maghreb and present allopatric distribution:
|
|
<taxonomicName lsidName="A. sadati" pageId="0" pageNumber="137" rank="species" species="sadati">
|
|
<emphasis italics="true" pageId="0" pageNumber="137">A. sadati</emphasis>
|
|
</taxonomicName>
|
|
is widely distributed in north Algeria and north Tunisia (
|
|
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3497.1.6" author="Zrelli, S" journalOrPublisher="Zootaxa" pageId="0" pageNumber="137" pagination="60 - 68" refId="B86" refString="Zrelli, S, Gattolliat, J-L, Boumaiza, M, Thomas, A, 2012. First record of Alainites sadati Thomas, 1994 (Ephemeroptera: Baetidae) in Tunisia, description of the larval stage and ecology. Zootaxa 3497 (1): 60 - 68, DOI: https://doi.org/10.11646/zootaxa.3497.1.6" title="First record of Alainites sadati Thomas, 1994 (Ephemeroptera: Baetidae) in Tunisia, description of the larval stage and ecology." url="https://doi.org/10.11646/zootaxa.3497.1.6" volume="3497" year="2012">Zrelli et al. 2012</bibRefCitation>
|
|
), while
|
|
<taxonomicName lsidName="A. oukaimeden" pageId="0" pageNumber="137" rank="species" species="oukaimeden">
|
|
<emphasis italics="true" pageId="0" pageNumber="137">A. oukaimeden</emphasis>
|
|
</taxonomicName>
|
|
is only reported from the High Atlas, Morocco (
|
|
<bibRefCitation author="Thomas, AGB" journalOrPublisher="Mitteilungen der Schweizerische Entomologische Gesellschaft" pageId="0" pageNumber="137" pagination="369 - 377" refId="B77" refString="Thomas, AGB, Bouzidi, A, Sartori, M, Assef, S, Ajakane, A, 1992. Complement et corrections a la faune des Ephemeropteres d'Afrique du Nord. 5. Baetis oukaimeden n. sp. du Haut Atlas marocain: Description et ecologie (Ephemeroptera, Baetidae). Mitteilungen der Schweizerische Entomologische Gesellschaft 65: 369 - 377" title="Complement et corrections a la faune des Ephemeropteres d'Afrique du Nord. 5. Baetis oukaimeden n. sp. du Haut Atlas marocain: Description et ecologie (Ephemeroptera, Baetidae)." volume="65" year="1992">Thomas et al. 1992</bibRefCitation>
|
|
;
|
|
<bibRefCitation DOI="https://doi.org/10.3390/d14060498" author="El Alami, M" journalOrPublisher="Diversity" pageId="0" pageNumber="137" pagination=": 498" refId="B22" refString="El Alami, M, El Yaagoubi, S, Gattolliat, JL, Sartori, M, Dakki, M, 2022a. Diversity and Distribution of Mayflies from Morocco (Ephemeroptera, Insecta). Diversity 14 (6): 498, DOI: https://doi.org/10.3390/d14060498" title="Diversity and Distribution of Mayflies from Morocco (Ephemeroptera, Insecta)." url="https://doi.org/10.3390/d14060498" volume="14" year="2022 a">El Alami et al. 2022a</bibRefCitation>
|
|
). The two endemic species have six pairs of gills; they can be only separated by intricate characters such as the reticulation of the surface of tergites and mandibles, the shape of the spines of distal margin of tergite IV and the number of strong setae on the dorsal margin of fore femora. A still undescribed new species, closely related to the West Palaearctic species
|
|
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites muticus" order="Ephemeroptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="muticus">
|
|
<emphasis italics="true" pageId="0" pageNumber="137">Alainites muticus</emphasis>
|
|
</taxonomicName>
|
|
, with seven pairs of gills, is present in Maghreb but with a restricted distribution limited to northern Morocco (
|
|
<bibRefCitation DOI="https://doi.org/10.3390/d14060498" author="El Alami, M" journalOrPublisher="Diversity" pageId="0" pageNumber="137" pagination=": 498" refId="B22" refString="El Alami, M, El Yaagoubi, S, Gattolliat, JL, Sartori, M, Dakki, M, 2022a. Diversity and Distribution of Mayflies from Morocco (Ephemeroptera, Insecta). Diversity 14 (6): 498, DOI: https://doi.org/10.3390/d14060498" title="Diversity and Distribution of Mayflies from Morocco (Ephemeroptera, Insecta)." url="https://doi.org/10.3390/d14060498" volume="14" year="2022 a">El Alami et al. 2022a</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |