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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.681.12942" ID-GBIF-Dataset="f05dcd99-6969-4fea-a189-681e959df550" ID-PMC="PMC5523879" ID-Pensoft-Pub="1313-2970-681-1" ID-PubMed="28769719" ID-ZBK="6A4B8B880C644465A443C6E8E454D41D" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-681-1" ModsDocOrigin="ZooKeys 681" ModsDocTitle="The subgenus Monotarsobius in the Iberian Peninsula with a description of a new pseudo-cryptic species from Northern Spain revealed by an integrative revision of Lithobiuscrassipes L. Koch, 1862 (Chilopoda, Lithobiomorpha, Lithobiidae)" checkinTime="1502744582383" checkinUser="pensoft" docAuthor="Voigtlaender, Karin, Iorio, Etienne, Decker, Peter &amp; Spelda, Joerg" docDate="2017" docId="40DE99BAB401F8CCB22E90FCCE1237EE" docLanguage="en" docName="ZooKeys 681: 1-38" docOrigin="ZooKeys 681" docSource="http://dx.doi.org/10.3897/zookeys.681.12942" docTitle="Lithobius crassipesoides Voigtlaender, Iorio, Decker &amp; Spelda, 2017, sp. n." docType="treatment" docUuid="EFB3B6F5-F82D-42AF-B1FC-6C656C0D18B7" docUuidSource="ZooBank" docVersion="5" lastPageNumber="19" masterDocId="B14FFFAA762BFFAC68207C286B1AFFAF" masterDocTitle="The subgenus Monotarsobius in the Iberian Peninsula with a description of a new pseudo-cryptic species from Northern Spain revealed by an integrative revision of Lithobiuscrassipes L. Koch, 1862 (Chilopoda, Lithobiomorpha, Lithobiidae)" masterLastPageNumber="38" masterPageNumber="1" pageNumber="14" updateTime="1668164555857" updateUser="ExternalLinkService">
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<mods:title>The subgenus Monotarsobius in the Iberian Peninsula with a description of a new pseudo-cryptic species from Northern Spain revealed by an integrative revision of Lithobiuscrassipes L. Koch, 1862 (Chilopoda, Lithobiomorpha, Lithobiidae)</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Voigtlaender, Karin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Iorio, Etienne</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Decker, Peter</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Spelda, Joerg</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>681</mods:number>
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<mods:start>1</mods:start>
<mods:end>38</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.681.12942</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.681.12942</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-681-1</mods:identifier>
<mods:identifier type="ZBK">6A4B8B880C644465A443C6E8E454D41D</mods:identifier>
<mods:identifier type="ZooBank">6A4B8B880C644465A443C6E8E454D41D</mods:identifier>
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<treatment ID-GBIF-Taxon="132649407" LSID="urn:lsid:zoobank.org:act:EFB3B6F5-F82D-42AF-B1FC-6C656C0D18B7" httpUri="http://treatment.plazi.org/id/40DE99BAB401F8CCB22E90FCCE1237EE" lastPageId="18" lastPageNumber="19" pageId="13" pageNumber="14">
<subSubSection pageId="13" pageNumber="14" type="nomenclature">
<paragraph pageId="13" pageNumber="14">
<taxonomicName LSID="http://zoobank.org/EFB3B6F5-F82D-42AF-B1FC-6C656C0D18B7" class="Chilopoda" family="Lithobiidae" genus="Lithobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithobius crassipesoides" order="Lithobiomorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="crassipesoides">Lithobius crassipesoides</taxonomicName>
<taxonomicNameLabel pageId="13" pageNumber="14">sp. n.</taxonomicNameLabel>
Figures 1, 2, 3, 4, 5, 6, 7, 8, 9, 10, 11, 12, 13, 14, 15, 16, 17, Tables 1, 2, 3, 4, 5, 6, 7, 8, 9, 10, 11, 12, 13, 14, 15, 16
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="14" type="reference_group">
<paragraph pageId="13" pageNumber="14">
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Lithobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithobius crassipes" order="Lithobiomorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="crassipes">Lithobius crassipes</taxonomicName>
-
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera 1980</bibRefCitation>
: 4 (listed), 5 (records, redescription), figs 3-5. -
<bibRefCitation author="Barace, J" journalOrPublisher="Graellsia" pageId="22" pageNumber="23" pagination="117 - 120" title="Contribucion al conocimiento de los Quilopodos de Navarra: relacion de especies y localidades (Myriapoda, Chilopoda)." volume="38" year="1982">Barace and Herrera 1982</bibRefCitation>
: 117 (records). -
<bibRefCitation author="Salinas, JA" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra, Serie Zoologica" pageId="24" pageNumber="25" pagination="1 - 70" title="Contribucion al conocimiento de los quilopodos de Navarra (Myriapoda: Chilopoda)." volume="20" year="1990">Salinas 1990</bibRefCitation>
: 2 (in checklist for Navarre), 59 (record), mapa 19 (map).
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="14" type="diagnosis">
<paragraph pageId="13" pageNumber="14">Diagnosis.</paragraph>
<paragraph pageId="13" pageNumber="14">
Small member (body length 6.4-11 mm) of the subgenus
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Lithobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithobius (Monotarsobius)" order="Lithobiomorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subGenus" subGenus="Monotarsobius">Monotarsobius</taxonomicName>
. Antennae with 20 articles, short, 2.6 times longer than head, 1/4 of body length. 5-11 ocelli, mostly 8 or 9, in two or three rows with one larger posterior ocellus. Legs with species-specific plectrotaxy; legpair 14 and 15 thickened in both sexes, much more so in males; legpair 15 without accessory apical claw, in males with a depression in the posterior half of tibia. Female gonopod claw tridentate.
</paragraph>
<paragraph pageId="13" pageNumber="14">
<taxonomicName lsidName="L. crassipesoides" pageId="13" pageNumber="14" rank="species" species="crassipesoides">L. crassipesoides</taxonomicName>
sp. n. differs generally from other Iberian members of
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Lithobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithobius (Monotarsobius)" order="Lithobiomorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subGenus" subGenus="Monotarsobius">Monotarsobius</taxonomicName>
in the presence of a depression in the posterior half of the legpair 15 tibia. It differs from
<taxonomicName lsidName="L. osellai" pageId="13" pageNumber="14" rank="species" species="osellai">L. osellai</taxonomicName>
and
<taxonomicName lsidName="L. morenoi" pageId="13" pageNumber="14" rank="species" species="morenoi">L. morenoi</taxonomicName>
in having more than one row of ocelli; from
<taxonomicName lsidName="L. blascoi" pageId="13" pageNumber="14" rank="species" species="blascoi">L. blascoi</taxonomicName>
in having only 20 antennal articles; and from
<taxonomicName lsidName="L. crassipes" pageId="13" pageNumber="14" rank="species" species="crassipes">L. crassipes</taxonomicName>
in smaller body length, shorter antennae, lower number of ocelli, the DaP spine starting posteriorly from legpair (12) 13, and the different location and size of the male depression on legpair 15 tibia.
</paragraph>
<caption pageId="13" pageNumber="14">
<paragraph pageId="13" pageNumber="14">
Figure 12. Adult males, dorsal view A
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Lithobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithobius crassipesoides" order="Lithobiomorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="crassipesoides">Lithobius crassipesoides</taxonomicName>
sp. n. (SMNG VNR14828) B
<taxonomicName lsidName="L. crassipes" pageId="13" pageNumber="14" rank="species" species="crassipes">L. crassipes</taxonomicName>
(SMNG VNR15171). Scale bars: 2 mm (A, B).
</paragraph>
</caption>
<caption pageId="13" pageNumber="14">
<paragraph pageId="13" pageNumber="14">
Figure 13. Adult males, anterior body, dorsal view A
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Lithobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithobius crassipesoides" order="Lithobiomorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="crassipesoides">Lithobius crassipesoides</taxonomicName>
sp. n. (SMNG VNR14828) B
<taxonomicName lsidName="L. crassipes" pageId="13" pageNumber="14" rank="species" species="crassipes">L. crassipes</taxonomicName>
(SMNG VNR15171). Scale bars: 0.5 mm (A, B).
</paragraph>
</caption>
<caption pageId="13" pageNumber="14">
<paragraph pageId="13" pageNumber="14">
Figure 14. Coxosternum, ventral view A
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Lithobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithobius crassipesoides" order="Lithobiomorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="crassipesoides">Lithobius crassipesoides</taxonomicName>
sp. n. (MZNA 138944) B
<taxonomicName lsidName="L. crassipes" pageId="13" pageNumber="14" rank="species" species="crassipes">L. crassipes</taxonomicName>
(SMNG VNR015171). Scale bars: 0.1 mm (A, B).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="13" pageNumber="14" type="etymology">
<paragraph pageId="13" pageNumber="14">Etymology.</paragraph>
<paragraph pageId="13" pageNumber="14">
Derived from the morphological similarity to
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Lithobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithobius crassipes" order="Lithobiomorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="crassipes">Lithobius crassipes</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="materials_examined">
<paragraph pageId="13" pageNumber="14">Material examined.</paragraph>
<paragraph pageId="13" pageNumber="14">
Holotype. Spain: Navarre province: Leitza, Ariz Mendiak, between area
<normalizedToken originalValue="“Ustarleku”">&quot;Ustarleku&quot;</normalizedToken>
and
<normalizedToken originalValue="“Karobieta”">&quot;Karobieta&quot;</normalizedToken>
, above side stream to Gorriztaran; loamy and calcareous soil,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.0778">43.0778°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-1.8775">1.8775°W</geoCoordinate>
, 615 m a.s.l., 20 April 2009, leg. FT2009, grove of
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Castanea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Castanea" order="Fagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Castanea</taxonomicName>
, pollard trees on the slope with
<taxonomicName class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ranunculus ficaria" order="Ranunculales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="ficaria">Ranunculus ficaria</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Thymelaeaceae" genus="Daphne" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Daphne" order="Malvales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Daphne</taxonomicName>
,
<taxonomicName class="Liliopsida" family="Melanthiaceae" genus="Helleborus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Helleborus" order="Liliales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Helleborus</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Lamiaceae" genus="Salvia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salvia" order="Lamiales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Salvia</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Rubus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rubus" order="Rosales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Rubus</taxonomicName>
,
<taxonomicName genus="Lathrea" lsidName="Lathrea" pageId="13" pageNumber="14" rank="genus">Lathrea</taxonomicName>
; under leaves and bark of rotten trunk, 1 ♂ (SMNG VNR14744-4 d).
</paragraph>
<paragraph pageId="13" pageNumber="14">
Paratype. Spain: Navarre province: Same data as holotype, 1 ♂ (SMNG VNR16739-1 b), 1 ♀ (SMNG VNR14744-4 b), 1 ♀ (SMNG VNR14752-1 c); Spain: Gipuzkoa province, Sierra de Aralar, Beasain, road from Lazkao to Etxarri-Aranaz, Pass Puerto de Lizarrusti, forest of
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus" order="Fagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
on the slope, under stones and in leaf litter,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="42.9614">42.9614°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-2.0983">2.0983°W</geoCoordinate>
, 690 m a.s.l, 21 April 2009, leg. FT2009, 1 ♂: (SMNG VNR14764-3).
</paragraph>
<paragraph lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
Other material examined.Spain: Navarre province: Leitza, Ariz Mendiak, between area
<normalizedToken originalValue="“Ustarleku”">&quot;Ustarleku&quot;</normalizedToken>
and
<normalizedToken originalValue="“Karobieta”">&quot;Karobieta&quot;</normalizedToken>
, above side stream to Gorriztaran; loamy and calcareous soil,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.0778">43.0778°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-1.8775">1.8775°W</geoCoordinate>
, 615 m a.s.l., 20 April 2009, leg. FT2009, grove of
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Castanea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Castanea" order="Fagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Castanea</taxonomicName>
, pollard trees on the slope with
<taxonomicName class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ranunculus ficaria" order="Ranunculales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="ficaria">Ranunculus ficaria</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Thymelaeaceae" genus="Daphne" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Daphne" order="Malvales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Daphne</taxonomicName>
,
<taxonomicName class="Liliopsida" family="Melanthiaceae" genus="Helleborus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Helleborus" order="Liliales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Helleborus</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Lamiaceae" genus="Salvia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salvia" order="Lamiales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Salvia</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Rubus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rubus" order="Rosales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Rubus</taxonomicName>
,
<taxonomicName genus="Lathrea" lsidName="Lathrea" pageId="13" pageNumber="14" rank="genus">Lathrea</taxonomicName>
; under leaves and bark of rotten trunk, 2 ♂♂, 3 ♀♀, 1 juv. ♂ (SMNG VNR14744-4), 1 ♂, 1 juv. ♂ (SMNG VNR16739-1), 1 ♀ (SMNG VNR14746-5), 1 ♀ (SMNG VNR17128-8); 100 meters down to a meadow along the way, 1 ♀ (SMNG VNR17126-4);
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus" order="Fagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Alnus</taxonomicName>
wood along stream, under bark of standing dead wood, 2 ♀♀, 1 juv. ♀ (SMNG VNR14752-1); with Carpinus betulus,
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Corylus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Corylus" order="Fagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Corylus</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus" order="Fagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
, Sambucus, 1 ♀ (SMNG VNR17129-3). - Leitza, Ariz Mendiak, between area
<normalizedToken originalValue="“Ustarleku”">&quot;Ustarleku&quot;</normalizedToken>
and
<normalizedToken originalValue="“Karobieta”">&quot;Karobieta&quot;</normalizedToken>
, next to road NA 4150 to Goizueta; on a slope with
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus" order="Fagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Corylus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Corylus" order="Fagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Corylus</taxonomicName>
, under a dead, mossy tree trunk,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="43.09092">43.09092°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-1.86613">1.86613°W</geoCoordinate>
, 593 m a.s.l., 20 April 2009, leg. FT2009, 2 ♀♀ (SMNG VNR17131-3). - Sierra de Aralar, south of Errazkin, north of area &quot;Axkarateko Malkorra&quot;, young
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus" order="Fagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
and old big
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus" order="Fagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ilex" order="Aquifoliales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Ilex</taxonomicName>
,
<taxonomicName class="Liliopsida" family="Asparagaceae" genus="Ruscus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ruscus" order="Asparagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Ruscus</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Araliaceae" genus="Hedera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hedera helix" order="Apiales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="helix">Hedera helix</taxonomicName>
, in leaf litter,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="42.9972">42.9972°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-1.9703">1.9703°W</geoCoordinate>
, 630 m a.s.l., 21 April 2009, leg. FT2009, 2 juv. ♂♂ (SMNG VNR14757-6), 1 ♂, 1 ♀ (SMNG VNR14758-3; SMNG VNR14813-1), 1 ♂, 1 ♀ (SMNG VNR17133-9). - Sierra de Aralar, south Baraibar, on road NA-7510 to Santuario de San Migel, at area
<normalizedToken originalValue="“Izáin”">&quot;Izain&quot;</normalizedToken>
, karst area with deep grykes and bare limestone rocks,
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus" order="Fagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
woodland and some
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Crataegus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Crataegus" order="Rosales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Crataegus</taxonomicName>
bushes, in leaf litter, partly sieved out,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="42.9714">42.9714°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-1.9384">1.9384°W</geoCoordinate>
, 790 m a.s.l., 22 April 2009, leg. FT2009, 1 ♀ (SMNG VNR14770-12), 1 ♀ (SMNG VNR14771-6). - Sierra de San Miguel, mountain point Artxueta at radio mast, karst area with bare limestone rocks, grove of low growing
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus" order="Fagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
, in litter and under bark,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="42.9525">42.9525°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-1.9668">1.9668°W</geoCoordinate>
, 1300 m a.s.l., 22 April 2009, leg. FT2009, 3 ♂♂, 1 ♀ (SMNG VNR14773-1). - Sierra de Urbasa, road NA-718 from Olazi /
<normalizedToken originalValue="Olazagutía">Olazagutia</normalizedToken>
to Estella, at the end of the hairpin curves, under the bark of a very large old beech with a lot of moos; limestone,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="42.86031">42.86031°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-2.18055">2.18055°W</geoCoordinate>
, 888 m a.s.l., 23 April 2009, leg. FT2009, 1 ♂, 1 ♀ (SMNG VNR17135-4). - Sierra de Urbasa, 1 km east of road junction to road NA-7182 at site
<normalizedToken originalValue="“Bentakaita”">&quot;Bentakaita&quot;</normalizedToken>
,
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus" order="Fagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
forest with
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Prunus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Prunus" order="Rosales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Prunus</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Papaveraceae" genus="Corydalis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Corydalis" order="Ranunculales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Corydalis</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Mercurialis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Mercurialis" order="Malpighiales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Mercurialis</taxonomicName>
,
<taxonomicName class="Liliopsida" family="Liliaceae" genus="Erythronium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Erythronium" order="Liliales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Erythronium</taxonomicName>
; in leaf litter,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="42.854">42.8540°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-2.1595">2.1595°W</geoCoordinate>
, 890 m a.s.l., 23 April 2009, leg. FT2009, 1 ♂ (SMNG VNR16734-1), 1 ♂ (SMNG VNR17136-4). - Urroz, hayedo [beech forest], 25 September 1995, leg. Javier
<normalizedToken originalValue="Sáenz">Saenz</normalizedToken>
de
<normalizedToken originalValue="Cabezón">Cabezon</normalizedToken>
, 1 ♀ (MZNA MZ-19951125 a). - Imbuluzqueta,
<normalizedToken originalValue="tocón">tocon</normalizedToken>
[stub], 21 February 1993, 2 ♂♂, 4 ♀♀ (MZNA MZ-19931121a). - Eguaras, Vedado de,
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus coccifera" order="Fagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="coccifera">Quercus coccifera</taxonomicName>
, 10 January 1980, leg. R. Jordana, 1 juv. ♀ (MZNA VE1041AS). - Irati, suelo de hayedo [floor of beech forest], 17 September 1982, leg. J. Barace, 1 ♀ (MZNA MZ-19820917,
<bibRefCitation author="Salinas, JA" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra, Serie Zoologica" pageId="24" pageNumber="25" pagination="1 - 70" title="Contribucion al conocimiento de los quilopodos de Navarra (Myriapoda: Chilopoda)." volume="20" year="1990">Salinas 1990</bibRefCitation>
). - Velate, hayedo [beech forest], 25 September 1995, leg. Javier
<normalizedToken originalValue="Sáenz">Saenz</normalizedToken>
de
<normalizedToken originalValue="Cabezón">Cabezon</normalizedToken>
, 1 ♂ (MZNA MZ-19951125). - Lagos de Urroz, hayedo [beech forest], 25 September 1995, leg. Javier
<normalizedToken originalValue="Sáenz">Saenz</normalizedToken>
de
<normalizedToken originalValue="Cabezón">Cabezon</normalizedToken>
, 1 ♀ (MZNA MZ-19951125 a). - Tirapegui,
<normalizedToken originalValue="tocón">tocon</normalizedToken>
[stub], 21 February 1993, 3 ♂♂, 1 ♀ (MZNA MZ-19931121 b). - Aquerreta, corteza [bark], 15 January 1994, 1 ♂, 6 ♀♀ (MZNA
<pageBreakToken pageId="14" pageNumber="15" start="start">MZ-</pageBreakToken>
19940119a). - Quinto Real, 29 June 1977, 1 juv. ♀ (MZNA MZ-19770629d,
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera 1980</bibRefCitation>
). - Quinto Real, carretera [road], 11 May 1977, leg. R. Jordana, 1 ♀ (MZNA MZ-19770511d,
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera 1980</bibRefCitation>
). - Quinto Real,
<normalizedToken originalValue="tocón">tocon</normalizedToken>
[stub], 4 August 1977, leg. Vierna, 4 ♂♂ (MZNA MZ-19770804e,
<bibRefCitation author="Barace, J" journalOrPublisher="Graellsia" pageId="22" pageNumber="23" pagination="117 - 120" title="Contribucion al conocimiento de los Quilopodos de Navarra: relacion de especies y localidades (Myriapoda, Chilopoda)." volume="38" year="1982">Barace and Herrera 1982</bibRefCitation>
). - Quinto Real, carretera [road], 11 May 1977, 1 ♂ (MZNA MZ-19770511d,
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera 1980</bibRefCitation>
). - Quinto Real, pinar [pine forest], 14 September 1977, 1 ♂ (MZNA MZ-19770914,
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera 1980</bibRefCitation>
). - Quinto Real, 24 November 1976, leg. Labiano, 1 ♂ (MZNA MZ-19761124a,
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera 1980</bibRefCitation>
). - Quinto Real, 29 June 1977, leg. Monreal, 1 ♂ (MZNA MZ-19770629d,
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera 1980</bibRefCitation>
). - Quinto Real, pinar [pine forest], 14 September 1977, 1 ♀ (MZNA MZ-19770914,
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera 1980</bibRefCitation>
). - Quinto Real, pinar [pine forest], 14 September 1977, 1 ♀ (MZNA MZ-19770914,
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera 1980</bibRefCitation>
). - Quinto Real, corteza de haya [bark of beech], 3 September 1977, leg. J. Barace, 1 ♂ (MZNA MZ-19770903 d,
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera 1980</bibRefCitation>
). - Quinto Real, pinar [pine forest], 14 September 1977, 1 ♂ (MZNA MZ-19770914,
<bibRefCitation author="Barace, J" journalOrPublisher="Graellsia" pageId="22" pageNumber="23" pagination="117 - 120" title="Contribucion al conocimiento de los Quilopodos de Navarra: relacion de especies y localidades (Myriapoda, Chilopoda)." volume="38" year="1982">Barace and Herrera 1982</bibRefCitation>
). - Quinto Real, corteza [bark], 29 June 1977, 1 ♂ (MZNA MZ-19770629 e,
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera 1980</bibRefCitation>
). - Quinto Real, corteza [bark], 29 June 1977, 1 (juv.?) ♀ (MZNA MZ-19770629 e,
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera 1980</bibRefCitation>
). - Quinto Real,
<normalizedToken originalValue="tocón">tocon</normalizedToken>
[stub], 4 August 1977, 1 ♀ (MZNA MZ-19770804 f,
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera 1980</bibRefCitation>
). - Quinto Real,
<normalizedToken originalValue="tocón">tocon</normalizedToken>
[stub], 16 March 1977, leg. Labiano, 1 ♀ (MZNA MZ-19770316 b,
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera 1980</bibRefCitation>
). Spain: Gipuzkoa province: Sierra de Aralar, Beasain, road from Lazkao to Etxarri-Aranaz, west of the Pass Puerto de Lizarrusti, forest of
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus" order="Fagales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
, in leaf litter,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="42.9572">42.9572°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-2.1122">2.1122°W</geoCoordinate>
, 550 m a.s.l., 21 April 2009, leg. FT2009, 1 ♂ (SMNG VNR14763-9), 1 juv. ♂ (SMNG VNR14763-11). - Natural Park Aizkorri-Aratz, near Onati; close to the monastery Arantzazu, beechwood in limestone with
<taxonomicName class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ilex" order="Aquifoliales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="genus">Ilex</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus" order="Fagales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Rubus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rubus" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="genus">Rubus</taxonomicName>
fructicosus,
<taxonomicName class="Anthozoa" family="Alcyoniidae" genus="Hedera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hedera" order="Alcyonacea" pageId="14" pageNumber="15" phylum="Cnidaria" rank="genus">Hedera</taxonomicName>
, Larix,
<taxonomicName class="Liliopsida" family="Melanthiaceae" genus="Helleborus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Helleborus" order="Liliales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="genus">Helleborus</taxonomicName>
, under bark of a dead tree trunk,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="42.97766">42.97766°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-2.38989">2.38989°W</geoCoordinate>
, 850 m a.s.l., 24 April 2009, leg. FT2009, 1 ♀ (SMNG VNR17139-5). - Natural Park Aizkorri-Aratz, Sierra de Urquilla, Montes de Altzania, south of mountain chain Aikorriko Mendikatea, south west of Mountain Aitzgorri, northwest Zumarraundi, high plain Alizania, north of Portua Zarra, karst plain with small growing trees of
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus" order="Fagales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
and open grasslands,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="42.9354">42.9354°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-2.3298">2.3298°W</geoCoordinate>
, 1160 m a.s.l., 24 April 2009, leg. FT2009, 1 ♂, 1 ♀, 1 juv. ♂, 1 juv. ♀ (SMNG VNR14786-3), 1 ♂, 1 ♀ (SMNG VNR14828-3), 1 ♂ (SMNG VNR14791-5). - Natural Park Aizkorri-Aratz, Sierra de Urquilla, near Zumaraundi, deep doline with creek discharge near cave entry; much leaf litter; sieved from
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus" order="Fagales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
leaves
<geoCoordinate direction="north" orientation="latitude" precision="5" value="42.9241">42.9241°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-2.3224">2.3224°W</geoCoordinate>
, 980 m a.s.l., 24 April 2009, leg. FT2009, 1 juv. ♂ (SMNG VNR14793-5).
</paragraph>
</subSubSection>
<subSubSection lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15" type="description">
<paragraph pageId="14" pageNumber="15">Description.</paragraph>
<paragraph pageId="14" pageNumber="15">Habitus. Slightly fusiform, widest around tergite 10 (Fig. 12A).</paragraph>
<paragraph pageId="14" pageNumber="15">
Colour. General body colouration varies from pale and light buff (most individuals), through yellow ochre to tawny olive (Fig. 12A). Mostly last third of the body and usually also the head a little darker. Head light yellow ochre, sometimes darker in front and around the ocellar area. Last antennal articles yellowish. Some or all tergites more or less darker on posterior margin. Most specimens are paler than
<taxonomicName lsidName="L. crassipes" pageId="14" pageNumber="15" rank="species" species="crassipes">L. crassipes</taxonomicName>
.
</paragraph>
<paragraph pageId="14" pageNumber="15">Length. 6.4-11.0 mm (Fig. 2, Table 2).</paragraph>
<paragraph pageId="15" pageNumber="16">
<pageBreakToken pageId="15" pageNumber="16" start="start">Head</pageBreakToken>
. Head roundish, mostly as broad as long or little broader than long and head broader or as broad as T5. Head length 0.6-1.08 mm (Fig. 3, Table 4).
</paragraph>
<paragraph pageId="15" pageNumber="16">Antennae. 20 antennal articles, short and stout, 1.6-3.2 mm long, 2.6 times longer than head (Figs 4, 13A, Table 6).</paragraph>
<paragraph pageId="15" pageNumber="16">Ocelli. 5 to 11 ocelli on each side of the head, mostly 8 or 9, rarely 5 or 11, arranged in two or three rows (Fig. 5, Table 8), posteriorly with one greater, longitudinally oval ocellus, clearly separated from the others. The most common arrangements (n = 47) are 1 + 4, 3 (21 %), 1 + 4, 3, 1 (14 %) and 1 + 4, 3, 2 (14 %). There is no significant difference between males and females in number and arrangement. In 54 % the number of ocelli differs between right and left side in one or two ocelli.</paragraph>
<paragraph pageId="15" pageNumber="16">Coxosternum. Anterior border with 2+2 teeth, upper part slender, acuminate, lateral borders without shoulders. Middle notch narrow to moderate width. Sometimes coxosternum slenderer and the middle notch narrower (Fig. 14A).</paragraph>
<paragraph lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
Tergites. Surface slightly rough, glossy. Posterior border of T1 feebly concave or straight, T3 to T5 feebly concave, T8 to T15 distinctly concave, T16 feebly to dis
<pageBreakToken pageId="16" pageNumber="17" start="start">tinctly</pageBreakToken>
concave. Posterior angles of T9, T11 and T13 mostly obtuse or rounded with no trace of lobes or triangular projections.
</paragraph>
<paragraph pageId="16" pageNumber="17">Legs. Tarsus and metatarsus fused on legpair 1 to 11. On legpair 12 and 13 the tarsal-metatarsal articulation is indistinct. Penultimate and ultimate legpairs (14, 15) are densely covered with pores. Last two legpairs are thickened in both sexes, much more so in males. Without accessory apical claw on legpair 15.</paragraph>
<paragraph pageId="16" pageNumber="17">Legpair 15 tibia of males with a more setiferous depression (fossa), which is distinct and well-developed in specimens in later developmental stages (Fig. 15A), in younger developmental stages only indicated. The depression starts more or less in the half of the tibia (55-75 % of tibia length) and reaches nearly up to the end of the tibia (88-93 % of tibia length) and has a relative length of about 30 to 40 % of tibia (n = 28).</paragraph>
<paragraph pageId="16" pageNumber="17">Coxal pores. Round, 2-4 (sometimes 5) pores on each coxa (Fig. 6, Table 9). Mostly 2, 3, 3, 2 or 3, 3, 3, 2 (coxae 15-12) with the highest observed number of 4, 5, 4, 3 in a female (Table 15).</paragraph>
<paragraph lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
Plectrotaxy. The plectrotaxy of legs of
<taxonomicName lsidName="L. crassipesoides" pageId="16" pageNumber="17" rank="species" species="crassipesoides">L. crassipesoides</taxonomicName>
sp. n. is given in Table 16. It differs from
<taxonomicName lsidName="L. crassipes" pageId="16" pageNumber="17" rank="species" species="crassipes">L. crassipes</taxonomicName>
in the absence of a DaP spine up to legpair 11, DaP very rarely
<pageBreakToken pageId="17" pageNumber="18" start="start">present</pageBreakToken>
on legpair 12, rarely present on legpair 13 to 14, while almost always present in
<taxonomicName lsidName="L. crassipes" pageId="17" pageNumber="18" rank="species" species="crassipes">L. crassipes</taxonomicName>
on legpair 10 to 15, frequently on legpair 9, rarely also on legpair 7 and 8 or only on legpair 11 to 15 (Table 17).
</paragraph>
<caption pageId="17" pageNumber="18">
<paragraph pageId="17" pageNumber="18">
Table 16. Plectrotaxy of
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Lithobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithobius crassipesoides" order="Lithobiomorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crassipesoides">Lithobius crassipesoides</taxonomicName>
sp. n. (n = 11 ♂♂ and 11 ♀♀). In brackets spines absent in more than 50 % of individuals.
</paragraph>
</caption>
<paragraph pageId="17" pageNumber="18">
<table pageId="17" pageNumber="18">
<tr pageId="17" pageNumber="18">
<th colspan="1" pageId="17" pageNumber="18" rowspan="2">Legpair</th>
<th colspan="5" pageId="17" pageNumber="18" rowspan="1">Ventral</th>
<th colspan="5" pageId="17" pageNumber="18" rowspan="1">Dorsal</th>
</tr>
<tr pageId="17" pageNumber="18">
<th colspan="1" pageId="17" pageNumber="18" rowspan="1">
<abbrev id="ABBRID0EYACK" pageId="17" pageNumber="18">C</abbrev>
</th>
<th colspan="1" pageId="17" pageNumber="18" rowspan="1">
<abbrev id="ABBRID0EABCK" pageId="17" pageNumber="18">t</abbrev>
</th>
<th colspan="1" pageId="17" pageNumber="18" rowspan="1">
<abbrev id="ABBRID0EIBCK" pageId="17" pageNumber="18">P</abbrev>
</th>
<th colspan="1" pageId="17" pageNumber="18" rowspan="1">
<abbrev id="ABBRID0EQBCK" pageId="17" pageNumber="18">F</abbrev>
</th>
<th colspan="1" pageId="17" pageNumber="18" rowspan="1">
<abbrev id="ABBRID0EYBCK" pageId="17" pageNumber="18">T</abbrev>
</th>
<th colspan="1" pageId="17" pageNumber="18" rowspan="1">
<abbrev id="ABBRID0EACCK" pageId="17" pageNumber="18">C</abbrev>
</th>
<th colspan="1" pageId="17" pageNumber="18" rowspan="1">
<abbrev id="ABBRID0EICCK" pageId="17" pageNumber="18">t</abbrev>
</th>
<th colspan="1" pageId="17" pageNumber="18" rowspan="1">
<abbrev id="ABBRID0EQCCK" pageId="17" pageNumber="18">P</abbrev>
</th>
<th colspan="1" pageId="17" pageNumber="18" rowspan="1">
<abbrev id="ABBRID0EYCCK" pageId="17" pageNumber="18">F</abbrev>
</th>
<th colspan="1" pageId="17" pageNumber="18" rowspan="1">
<abbrev id="ABBRID0EADCK" pageId="17" pageNumber="18">T</abbrev>
</th>
</tr>
<tr pageId="17" pageNumber="18">
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">1</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">p</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">m</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">mp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">a</td>
</tr>
<tr pageId="17" pageNumber="18">
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">2</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">p</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">am</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">mp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
</tr>
<tr pageId="17" pageNumber="18">
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">3</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">p</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">am</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">mp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
</tr>
<tr pageId="17" pageNumber="18">
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">4</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">p</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">am</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">mp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
</tr>
<tr pageId="17" pageNumber="18">
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">5</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">p</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">am</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">mp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
</tr>
<tr pageId="17" pageNumber="18">
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">6</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">p</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">am</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">mp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
</tr>
<tr pageId="17" pageNumber="18">
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">7</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">p</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">am</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">mp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
</tr>
<tr pageId="17" pageNumber="18">
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">8</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">p</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">am</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">mp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
</tr>
<tr pageId="17" pageNumber="18">
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">9</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">p</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">am</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">mp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
</tr>
<tr pageId="17" pageNumber="18">
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">10</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">mp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">am</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">mp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
</tr>
<tr pageId="17" pageNumber="18">
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">11</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">mp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">am</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">mp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
</tr>
<tr pageId="17" pageNumber="18">
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">12</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">m</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">am</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">a</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">p</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">ap</td>
</tr>
<tr pageId="17" pageNumber="18">
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">13</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">m</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">am</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">a</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">p</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">p</td>
</tr>
<tr pageId="17" pageNumber="18">
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">14</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">m</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">am</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">a</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">p</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">p</td>
</tr>
<tr pageId="17" pageNumber="18">
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">15</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">m</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">a</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">amp</td>
<td colspan="1" pageId="17" pageNumber="18" rowspan="1">p</td>
</tr>
</table>
</paragraph>
<paragraph pageId="17" pageNumber="18">Male gonopods. Uni-articulated.</paragraph>
<paragraph pageId="17" pageNumber="18">Female gonopods. Basal article with two conical spurs on each side, their apical edge serrated (Fig. 16A). With 5-9 ventrolateral setae with nearly the same length. No dorsomedial setae. One specimen (SMNG VNR14770) with two minute setae. Article II with 4-6 dorsolateral setae, stout, straight and fairly long, evenly distributed over the whole length of the article. Ventrolaterally with 4-6 setae without characteristic arrangement. Article III with 1 dorsolateral seta, 1 ventrolateral, 1 ventral and 2 ventromedial setae. Claw tridentated, dorsal denticle longer than the ventral denticle. At high magnification (SEM) with distinct pores of glandulae.</paragraph>
<caption pageId="17" pageNumber="18">
<paragraph pageId="17" pageNumber="18">
Figure 15. Tibia of male legpair 15, dorsal view A
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Lithobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithobius crassipesoides" order="Lithobiomorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crassipesoides">Lithobius crassipesoides</taxonomicName>
sp. n. (SMNG VNR14773), right leg B
<taxonomicName lsidName="L. crassipes" pageId="17" pageNumber="18" rank="species" species="crassipes">L. crassipes</taxonomicName>
(SMNG VNR10335), left leg. Abbreviations: F = femur; Ts = tarsus. Scale bars: 0.2 mm (A, B).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="distribution">
<paragraph pageId="18" pageNumber="19">
<pageBreakToken pageId="18" pageNumber="19" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph pageId="18" pageNumber="19">
So far only known from Navarre and Gipuzkoa provinces, northern Spain (Fig. 17). Some records are only a few kilometers from the French border.
<taxonomicName lsidName="L. crassipesoides" pageId="18" pageNumber="19" rank="species" species="crassipesoides">L. crassipesoides</taxonomicName>
sp. n. is therefore expected also in the Western Pyrenees in France.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="habitat">
<paragraph pageId="18" pageNumber="19">Habitat.</paragraph>
<paragraph pageId="18" pageNumber="19">The species was mostly found in the leaf litter and under the bark of dead wood in mountain deciduous forests from 550 to 1300 m a.s.l.</paragraph>
<caption pageId="18" pageNumber="19">
<paragraph pageId="18" pageNumber="19">
Figure 16. Left female gonopod, ventral view A
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Lithobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithobius crassipesoides" order="Lithobiomorpha" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipesoides">Lithobius crassipesoides</taxonomicName>
sp. n. (SMNG VNR14828) B
<taxonomicName lsidName="L. crassipes" pageId="18" pageNumber="19" rank="species" species="crassipes">L. crassipes</taxonomicName>
(SMNG VNR15171). Scale bars: 0.1 mm (A, B).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="remarks">
<paragraph pageId="18" pageNumber="19">Remarks.</paragraph>
<paragraph pageId="18" pageNumber="19">
<taxonomicName lsidName="L. crassipes" pageId="18" pageNumber="19" rank="species" species="crassipes">L. crassipes</taxonomicName>
was recorded from the Navarre region from Irati (
<bibRefCitation author="Salinas, JA" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra, Serie Zoologica" pageId="24" pageNumber="25" pagination="1 - 70" title="Contribucion al conocimiento de los quilopodos de Navarra (Myriapoda: Chilopoda)." volume="20" year="1990">Salinas 1990</bibRefCitation>
) and several locations near Quinto Real (
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera 1980</bibRefCitation>
,
<bibRefCitation author="Barace, J" journalOrPublisher="Graellsia" pageId="22" pageNumber="23" pagination="117 - 120" title="Contribucion al conocimiento de los Quilopodos de Navarra: relacion de especies y localidades (Myriapoda, Chilopoda)." volume="38" year="1982">1982</bibRefCitation>
).
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera (1980)</bibRefCitation>
also gave a brief description of 12 males and eight females. Most of the material studied by
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera (1980</bibRefCitation>
,
<bibRefCitation author="Barace, J" journalOrPublisher="Graellsia" pageId="22" pageNumber="23" pagination="117 - 120" title="Contribucion al conocimiento de los Quilopodos de Navarra: relacion de especies y localidades (Myriapoda, Chilopoda)." volume="38" year="1982">1982</bibRefCitation>
) and
<bibRefCitation author="Salinas, JA" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra, Serie Zoologica" pageId="24" pageNumber="25" pagination="1 - 70" title="Contribucion al conocimiento de los quilopodos de Navarra (Myriapoda: Chilopoda)." volume="20" year="1990">Salinas (1990)</bibRefCitation>
was available for re-examination at MZNA (38 specimens) and was confirmed to be
<taxonomicName lsidName="L. crassipesoides" pageId="18" pageNumber="19" rank="species" species="crassipesoides">L. crassipesoides</taxonomicName>
sp. n. Therefore, all records of
<bibRefCitation author="Barace, J" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra" pageId="22" pageNumber="23" pagination="3 - 30" title="Estudio faunistico del macizo de Quinto Real III: Miriapodos Quilopodos (Myriapoda, Chilopoda)." volume="4" year="1980">Barace and Herrera (1980</bibRefCitation>
,
<bibRefCitation author="Barace, J" journalOrPublisher="Graellsia" pageId="22" pageNumber="23" pagination="117 - 120" title="Contribucion al conocimiento de los Quilopodos de Navarra: relacion de especies y localidades (Myriapoda, Chilopoda)." volume="38" year="1982">1982</bibRefCitation>
) and
<bibRefCitation author="Salinas, JA" journalOrPublisher="Publicaciones de Biologia de la Universidad de Navarra, Serie Zoologica" pageId="24" pageNumber="25" pagination="1 - 70" title="Contribucion al conocimiento de los quilopodos de Navarra (Myriapoda: Chilopoda)." volume="20" year="1990">Salinas (1990)</bibRefCitation>
of
<taxonomicName lsidName="L. crassipes" pageId="18" pageNumber="19" rank="species" species="crassipes">L. crassipes</taxonomicName>
are hereby assigned to
<taxonomicName lsidName="L. crassipesoides" pageId="18" pageNumber="19" rank="species" species="crassipesoides">L. crassipesoides</taxonomicName>
sp. n. Finally, as mentioned above, the description of
<bibRefCitation author="Serra, A" journalOrPublisher="Universidad de Barcelona, Barcelona, Spain" pageId="24" pageNumber="25" title="Contribucion al conocimiento de los Lithobiomorpha (Chilopoda) de la Peninsula Iberica" volumeTitle="PhS thesis" year="1980">Serra (1980)</bibRefCitation>
for
<taxonomicName lsidName="L. crassipes" pageId="18" pageNumber="19" rank="species" species="crassipes">L. crassipes</taxonomicName>
is possibly composite and perhaps partially concerns
<taxonomicName lsidName="L. crassipesoides" pageId="18" pageNumber="19" rank="species" species="crassipesoides">L. crassipesoides</taxonomicName>
sp. n.
</paragraph>
<caption pageId="18" pageNumber="19">
<paragraph pageId="18" pageNumber="19">
Figure 17. Distribution map of
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Lithobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithobius (Monotarsobius)" order="Lithobiomorpha" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subGenus" subGenus="Monotarsobius">Lithobius (Monotarsobius)</taxonomicName>
on the Iberian Peninsula:
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Lithobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithobius blascoi" order="Lithobiomorpha" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="blascoi">Lithobius blascoi</taxonomicName>
= diamond;
<taxonomicName lsidName="L. crassipes" pageId="18" pageNumber="19" rank="species" species="crassipes">L. crassipes</taxonomicName>
= square;
<taxonomicName lsidName="L. crassipesoides" pageId="18" pageNumber="19" rank="species" species="crassipesoides">L. crassipesoides</taxonomicName>
sp. n. = circle;
<taxonomicName lsidName="L. morenoi" pageId="18" pageNumber="19" rank="species" species="morenoi">L. morenoi</taxonomicName>
stat. n. = triangle;
<taxonomicName lsidName="L. osellai" pageId="18" pageNumber="19" rank="species" species="osellai">L. osellai</taxonomicName>
= star.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>