220 lines
24 KiB
XML
220 lines
24 KiB
XML
<document id="FDCD383706E059E57E8AB31DDF2CEF73" ID="10.11646/zootaxa.4184.1.3" ID-DOI="10.11646/zootaxa.4184.1.3" ID-GBIF-Dataset="20cd1682-9b9d-4f4f-8b83-16cdde47142a" ID-ISSN="1175-5326" ID-Zenodo-Dep="164475" ID-ZooBank="B5E0AB75-5EA2-4C60-9017-9DB4A82A49BB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1478157609891" checkinUser="plazi" docAuthor="Last, Peter R., Weigmann, Simon & Dumale, Don" docDate="2016" docId="405987E440285258FF2CF63F19E9FE54" docLanguage="en" docName="zootaxa.4184.1.3.pdf" docOrigin="Zootaxa 4184 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Orbiraja Last, Weigmann & Dumale, 2016, gen. nov." docType="treatment" docVersion="8" masterDocId="BC60FF9C4029525AFFBBF5081915FF84" masterDocTitle="A new skate genus Orbiraja (Rajiformes: Rajidae) from the Indo-West Pacific" updateTime="1698391765355" updateUser="plazi">
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<mods:mods id="8EA5C5A3C92CCCC76326F9489E6F56AE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="ABB665BD57A4DFDDDB090344E4079883">
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<mods:title id="E47EEBDE0E6997F2EA9A96F709D07052">A new skate genus Orbiraja (Rajiformes: Rajidae) from the Indo-West Pacific</mods:title>
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</mods:titleInfo>
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<mods:name id="38222881D13ED1BE70D10B62254A2E77" type="personal">
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<mods:role id="EC4853ABBDA3E3EDF6D9E7C00FA495BA">
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<mods:roleTerm id="65DF8C899D557F23C1498830145BFF2E">Author</mods:roleTerm>
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<mods:namePart id="C76B45190A4B6964640BDC14FE07B3CF">Last, Peter R.</mods:namePart>
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<mods:name id="4D7B9D288C77584163470269C4503598" type="personal">
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<mods:roleTerm id="4EF534D60082010F4FCC1137AEAC4108">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B10C24CD2E15960E96190DD2F2918261">Weigmann, Simon</mods:namePart>
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</mods:name>
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<mods:name id="0A79C89F863DB58913134C5F6696FBF1" type="personal">
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<mods:role id="0E67687EA06A9B91640EDA9EC19B2E03">
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<mods:roleTerm id="4D391804533190219EF26732BF8032FD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C1BC8AB8FF76184DC9304DDD098ACF18">Dumale, Don</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="1F58EA711CAB834CBB932D492C3F6D95">text</mods:typeOfResource>
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<mods:title id="5EF52DAA36785C180A30AA0FFC3FB80E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="9BF1B4B35BD63CA017F88398575C756C">
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<mods:date id="A20CE5DD143A939B5F0FB2C82914D6F9">2016</mods:date>
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<mods:detail id="0DADBCF00F0EC5CF3AB59379AAAEEB26" type="volume">
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<mods:number id="370F2DBF88CF497A6FC47FC311177705">4184</mods:number>
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</mods:detail>
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<mods:detail id="F0AD5278285256BA39DE83B8551EA01C" type="issue">
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<mods:number id="037CB9CCAFEB56AC84E056CDAF9C01B1">1</mods:number>
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</mods:detail>
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<mods:classification id="982F0CAD2548DD45FB715BC38EF1779B">journal volume</mods:classification>
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<mods:identifier id="A9F8C78AE21BB81A14C2F0244E7C4B0A" type="DOI">10.11646/zootaxa.4184.1.3</mods:identifier>
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<mods:identifier id="8C22BE2D6781FEF6445010048A003394" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="7477222F943147195C42CE2169E0D86F" type="Zenodo-Dep">164475</mods:identifier>
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</mods:mods>
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<treatment id="405987E440285258FF2CF63F19E9FE54" ID-DOI="http://doi.org/10.5281/zenodo.6085171" ID-GBIF-Taxon="124562741" ID-Zenodo-Dep="6085171" LSID="urn:lsid:plazi:treatment:405987E440285258FF2CF63F19E9FE54" httpUri="http://treatment.plazi.org/id/405987E440285258FF2CF63F19E9FE54" lastPageId="2" pageId="1">
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<subSubSection id="80EA65794028525BFF2CF63F1861FCD6" box="[151,372,823,850]" pageId="1" type="nomenclature">
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<paragraph id="C84F36F24028525BFF2CF63F1861FCD6" blockId="1.[151,372,823,850]" box="[151,372,823,850]" pageId="1">
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<heading id="9307819E4028525BFF2CF63F1861FCD6" bold="true" box="[151,372,823,850]" fontSize="11" level="1" pageId="1" reason="1">
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<emphasis id="FA84EAE04028525BFF2CF63F1861FCD6" bold="true" box="[151,372,823,850]" pageId="1">
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<taxonomicName id="0FF04D714028525BFF2CF63F19EAFCD5" ID-CoL="69DK" ID-ENA="2497397" box="[151,255,823,849]" class="Elasmobranchii" family="Rajidae" genus="Orbiraja" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="FA84EAE04028525BFF2CF63F19EAFCD5" bold="true" box="[151,255,823,849]" italics="true" pageId="1">Orbiraja</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E1B7579B4028525BFEBDF6301861FCD6" box="[262,372,824,850]" pageId="1" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="80EA65794028525BFF2CF67719F9FA2A" pageId="1" type="materials_examined">
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<paragraph id="C84F36F24028525BFF2CF6771BBDFC10" blockId="1.[151,680,894,917]" box="[151,680,894,917]" pageId="1">
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Type species.
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<taxonomicName id="0FF04D714028525BFE9EF6761BB1FC10" authority="Last & Lim, 2010" authorityName="Last & Lim" authorityYear="2010" box="[293,676,894,917]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="species" species="jensenae">
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<emphasis id="FA84EAE04028525BFE9EF67618F1FC11" box="[293,484,894,917]" italics="true" pageId="1">Okamejei jensenae</emphasis>
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<bibRefCitation id="AC614B034028525BFE50F6771BB1FC10" author="Last" box="[491,676,894,916]" pageId="1" refString="Last, P. R. & Lim, A. P. K. (2010) A new species of skate Okamejei jensenae sp. nov. (Rajoidei: Rajidae) from the seas off Borneo, with a redescription of the Kwangtung Skate, Dipturus kwangtungensis (Chu), pp. 101 - 114. In: P. R. Last, W. T. White, J. J. Pogonoski (Eds.), Descriptions of New Sharks and Rays from Borneo. CSIRO Marine and Atmospheric Research Paper, 0 32, 165 pp." type="book" year="2010">Last & Lim, 2010</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="C84F36F24028525BFF2CF6C919F9FA2A" blockId="1.[151,1437,961,2030]" pageId="1">
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<emphasis id="FA84EAE04028525BFF2CF6C91803FC5E" bold="true" box="[151,278,961,986]" pageId="1">Definition.</emphasis>
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Small to medium-sized rostrorajin skates (to about
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<quantity id="0F089B174028525BFCC5F6C91AD2FC5D" box="[894,967,961,986]" metricMagnitude="-1" metricUnit="m" metricValue="5.3" pageId="1" unit="cm" value="53.0">53 cm</quantity>
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TL) with the following combination of characters: disc rhombic, width 1–1.2 times length; snout moderately elongate, 3.9–5 times orbit length; spiracle large and broadly oval, well removed from orbit, its anteriormost margin not or barely extending forward below eye; tooth rows in upper jaw 61–76; median thorns on tail in three rows, those of outer rows usually larger than central row; procaudal length long to very long (0.9–1.4 times snout length). Clasper moderately slender and depressed, glans expanded; no dermal denticles or pseudosiphon; dorsal lobe components consisting of clefts, terminal bridge; ventral lobe components consisting of eperon, rhipidion, sentinel, shield and spike; dorsal terminal 2 cartilage long and attached to distal tip of axial cartilage (without dorsal terminal 3 cartilage); accessory terminal 1 cartilage massive rather than y-shaped with a prominent groove near its lateral margin and a slender, s-shaped distal extension through most of glans; accessory terminal 2 cartilage with a very long, curved lateral distal extension forming a component spike, and a strongly articulated, medio-distal extension (accessory terminal 3 cartilage) partly overlapping axial cartilage medially. Pectoral ocelli moderately large and almost circular; no darkedged ventral pores. Pectoral-fin radials 77–82; predorsal vertebrae 79–86; northern Indian Ocean and North-West Pacific.
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</paragraph>
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</subSubSection>
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<subSubSection id="80EA65794028525BFF7CF0B11ABFF94A" pageId="1" type="etymology">
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<paragraph id="C84F36F24028525BFF7CF0B11D15FA72" blockId="1.[151,1437,961,2030]" pageId="1">
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<emphasis id="FA84EAE04028525BFF7CF0B1185BFA56" bold="true" box="[199,334,1465,1490]" pageId="1">Etymology.</emphasis>
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Epithet based on the Latin word
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<emphasis id="FA84EAE04028525BFD67F0B11A03FA55" box="[732,790,1465,1489]" italics="true" pageId="1">orbis</emphasis>
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(meaning circle) with reference to the paired, ring-like pectoral-fin ocelli in all three nominal species. Vernacular name: Ring Skates.
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</paragraph>
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<paragraph id="C84F36F24028525BFF7CF3091ABFF94A" blockId="1.[151,1437,961,2030]" pageId="1">
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<materialsCitation id="78983CAF4028525BFF7CF3091AB2F94A" ID-GBIF-Occurrence="1324562737" country="Myanmar" location="Myanmar" pageId="1" specimenCount="1" typeStatus="holotype">
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<emphasis id="FA84EAE04028525BFF7CF3091830F99E" bold="true" box="[199,293,1537,1562]" pageId="1">Species.</emphasis>
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<taxonomicName id="0FF04D714028525BFE97F3091BCCF99E" authority="Last & Lim, 2010" authorityName="Last & Lim" authorityYear="2010" box="[300,729,1537,1562]" class="Elasmobranchii" family="Rajidae" genus="Orbiraja" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="species" species="jensenae">
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<emphasis id="FA84EAE04028525BFE97F30918EDF99D" box="[300,504,1537,1561]" italics="true" pageId="1">Orbiraja jensenae</emphasis>
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(
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<bibRefCitation id="AC614B034028525BFDBDF30A1BC4F99E" author="Last" box="[518,721,1537,1562]" pageId="1" refString="Last, P. R. & Lim, A. P. K. (2010) A new species of skate Okamejei jensenae sp. nov. (Rajoidei: Rajidae) from the seas off Borneo, with a redescription of the Kwangtung Skate, Dipturus kwangtungensis (Chu), pp. 101 - 114. In: P. R. Last, W. T. White, J. J. Pogonoski (Eds.), Descriptions of New Sharks and Rays from Borneo. CSIRO Marine and Atmospheric Research Paper, 0 32, 165 pp." type="book" year="2010">Last & Lim, 2010</bibRefCitation>
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)
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</taxonomicName>
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and
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<taxonomicName id="0FF04D714028525BFCB5F30A1D23F99E" authority="Alcock, 1898" authorityName="Alcock" authorityYear="1898" box="[782,1078,1537,1562]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="species" species="powelli">
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<emphasis id="FA84EAE04028525BFCB5F30A1A96F99D" box="[782,899,1537,1561]" italics="true" pageId="1">O. powelli</emphasis>
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(
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<bibRefCitation id="AC614B034028525BFC2AF3091D38F99E" author="Alcock" box="[913,1069,1537,1562]" pageId="1" refString="Alcock, A. W. (1898) Natural history notes from H. M. Indian marine survey ship ' Investigator', Commander T. H. Heming, R. N., commanding. Series II., No. 25. A note on the deep-sea fishes, with descriptions of some new genera and species, including another probably viviparous ophidioid. Annals and Magazine of Natural History (Series 7), 2 (8), 136 - 156." type="journal article" year="1898">Alcock, 1898</bibRefCitation>
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)
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</taxonomicName>
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. The status of
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<taxonomicName id="0FF04D714028525BFB6CF30A19CDF9BA" authority="Lloyd, 1906" authorityName="Lloyd" authorityYear="1906" class="Elasmobranchii" family="Rajidae" genus="Orbiraja" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="species" species="philipi">
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<emphasis id="FA84EAE04028525BFB6CF30A1C57F99D" box="[1239,1346,1537,1561]" italics="true" pageId="1">O. philipi</emphasis>
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(
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<bibRefCitation id="AC614B034028525BFAEAF30919C5F9BA" author="Lloyd" pageId="1" refString="Lloyd, R. E. (1906) Natural history notes from the R. I. M. S. ship ' lnvestigator'. Notes on the skull of the genus Aulostomatomorpha, with descriptions of some new deep-sea fish. Annals and Magazine of Natural History, 18 (7), 306 - 311. http: // dx. doi. org / 10.1080 / 00222930608562615" type="journal article" year="1906">Lloyd, 1906</bibRefCitation>
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)
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</taxonomicName>
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needs to be reassessed as it has been considered a possible junior synonym of
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<taxonomicName id="0FF04D714028525BFBEEF32E1DDEF9B9" box="[1109,1227,1573,1597]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="species" species="powelli">
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<emphasis id="FA84EAE04028525BFBEEF32E1DDEF9B9" box="[1109,1227,1573,1597]" italics="true" pageId="1">O. powelli</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="AC614B034028525BFB61F32D19C6F9E6" author="Weigmann" pageId="1" refString="Weigmann, S., Stehmann, M. F. W. & Thiel, R. (2015) Okamejei ornata n. sp., a new deep-water skate (Elasmobranchii, Rajidae) from the northwestern Indian Ocean off Socotra Islands. Deep-Sea Research II, 115, 18 - 29. http: // dx. doi. org / 10.1016 / j. dsr 2.2013.09.005" type="journal article" year="2015">
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Weigmann
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<emphasis id="FA84EAE04028525BFAE7F32E1C9BF9B9" box="[1372,1422,1573,1597]" italics="true" pageId="1">et al</emphasis>
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., 2015
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</bibRefCitation>
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;
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<bibRefCitation id="AC614B034028525BFF5EF34118A4F9E6" author="Weigmann" box="[229,433,1609,1634]" pageId="1" refString="Weigmann, S. (2016) Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. Journal of Fish Biology, 88 (3), 837 - 1037." type="journal article" year="2016">Weigmann, 2016</bibRefCitation>
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). New material of
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<taxonomicName id="0FF04D714028525BFD1BF3421A08F9E5" box="[672,797,1609,1633]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="species" species="powelli">
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<emphasis id="FA84EAE04028525BFD1BF3421A08F9E5" box="[672,797,1609,1633]" italics="true" pageId="1">O. powelli</emphasis>
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</taxonomicName>
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collected off
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<collectingCountry id="B0E776624028525BFC74F3421D29F9E6" box="[975,1084,1610,1634]" name="Myanmar" pageId="1">Myanmar</collectingCountry>
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suggests that this species is morphologically variable, including the shape of the pectoral marking. Lloyd’s original figure of the
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<typeStatus id="174B88504028525BFAA8F3661C56F902" box="[1299,1347,1646,1670]" pageId="1">type</typeStatus>
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has the pectoral marking placed more posteriorly on the disc than in typical
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<emphasis id="FA84EAE04028525BFC28F39A1D1AF92D" box="[915,1039,1681,1705]" italics="true" pageId="1">
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<taxonomicName id="0FF04D714028525BFC28F39A1D1EF92D" box="[915,1035,1681,1705]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="species" species="powelli">O. powelli</taxonomicName>
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.
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</emphasis>
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Preliminary investigations indicate that an additional, un-named species of
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<taxonomicName id="0FF04D714028525BFDEFF3BD1BADF949" box="[596,696,1717,1741]" class="Elasmobranchii" family="Rajidae" genus="Orbiraja" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="FA84EAE04028525BFDEFF3BD1BADF949" box="[596,696,1717,1741]" italics="true" pageId="1">Orbiraja</emphasis>
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</taxonomicName>
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occurs off
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<collectingCountry id="B0E776624028525BFC8DF3BD1AB2F94A" box="[822,935,1717,1742]" name="Indonesia" pageId="1">Indonesia</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="80EA657940285258FF7CF3D119E9FE54" lastPageId="2" pageId="1" type="discussion">
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<paragraph id="C84F36F240285258FF7CF3D119E9FE54" blockId="1.[151,1437,961,2030]" lastBlockId="2.[151,1437,151,464]" lastPageId="2" pageId="1">
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<emphasis id="FA84EAE04028525BFF7CF3D1182EF976" bold="true" box="[199,315,1753,1778]" pageId="1">Remarks.</emphasis>
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A substantial body of literature has been assembled on the clasper morphology of skates, but no comprehensive coverage of all extant genera has been published and details of the adult clasper are unknown for some genera (e.g.
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<taxonomicName id="0FF04D714028525BFEDAF22918F7F8BD" box="[353,482,1825,1849]" class="Elasmobranchii" family="Arhynchobatidae" genus="Pseudoraja" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="genus">
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<emphasis id="FA84EAE04028525BFEDAF22918F7F8BD" box="[353,482,1825,1849]" italics="true" pageId="1">Pseudoraja</emphasis>
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</taxonomicName>
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).
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<taxonomicName id="0FF04D714028525BFE42F2291B48F8BD" box="[505,605,1825,1849]" class="Elasmobranchii" family="Rajidae" genus="Orbiraja" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="FA84EAE04028525BFE42F2291B48F8BD" box="[505,605,1825,1849]" italics="true" pageId="1">Orbiraja</emphasis>
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</taxonomicName>
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resembles
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<taxonomicName id="0FF04D714028525BFD61F2291A50F8BD" box="[730,837,1825,1849]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="genus">
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<emphasis id="FA84EAE04028525BFD61F2291A50F8BD" box="[730,837,1825,1849]" italics="true" pageId="1">Okamejei</emphasis>
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</taxonomicName>
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in external morphology, but members of these genera differ from each other in clasper morphology, squamation and in their molecular sequences. In
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<taxonomicName id="0FF04D714028525BFB07F24D1C35F8D9" box="[1212,1312,1861,1885]" class="Elasmobranchii" family="Rajidae" genus="Orbiraja" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="FA84EAE04028525BFB07F24D1C35F8D9" box="[1212,1312,1861,1885]" italics="true" pageId="1">Orbiraja</emphasis>
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</taxonomicName>
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there are 3 well-developed rows of thorns along the middle of the tail and these are positioned unusually close together and well away from the lateral thorn rows. Some
|
||
<taxonomicName id="0FF04D714028525BFD2AF2851BE9F821" box="[657,764,1933,1957]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="genus">
|
||
<emphasis id="FA84EAE04028525BFD2AF2851BE9F821" box="[657,764,1933,1957]" italics="true" pageId="1">Okamejei</emphasis>
|
||
</taxonomicName>
|
||
(e.g.
|
||
<taxonomicName id="0FF04D714028525BFC87F2861ABAF821" box="[828,943,1933,1957]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="species" species="kenojei">
|
||
<emphasis id="FA84EAE04028525BFC87F2861ABAF821" box="[828,943,1933,1957]" italics="true" pageId="1">O. kenojei</emphasis>
|
||
</taxonomicName>
|
||
) have three prominent thorn rows along the median tail but these are well separated (lateral-most rows usually dorsolateral in position).
|
||
<taxonomicName id="0FF04D714028525BFB1FF2B91C1AF84D" box="[1188,1295,1969,1993]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="genus">
|
||
<emphasis id="FA84EAE04028525BFB1FF2B91C1AF84D" box="[1188,1295,1969,1993]" italics="true" pageId="1">Okamejei</emphasis>
|
||
</taxonomicName>
|
||
species also have dark-edged pores on the ventral head compared to not being dark-edged in
|
||
<taxonomicName id="0FF04D714028525BFBF3F2DD1DB9F869" box="[1096,1196,2005,2029]" class="Elasmobranchii" family="Rajidae" genus="Orbiraja" kingdom="Animalia" order="Rajiformes" pageId="1" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="FA84EAE04028525BFBF3F2DD1DB9F869" box="[1096,1196,2005,2029]" italics="true" pageId="1">Orbiraja</emphasis>
|
||
</taxonomicName>
|
||
. The claspers of the
|
||
<taxonomicName id="0FF04D71402B5258FF2CF5901817FF34" box="[151,258,152,176]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="2" phylum="Chordata" rank="genus">
|
||
<emphasis id="FA84EAE0402B5258FF2CF5901817FF34" box="[151,258,152,176]" italics="true" pageId="2">Okamejei</emphasis>
|
||
</taxonomicName>
|
||
species have a component funnel (absent in
|
||
<taxonomicName id="0FF04D71402B5258FCAFF5901A6DFF34" box="[788,888,152,176]" class="Elasmobranchii" family="Rajidae" genus="Orbiraja" kingdom="Animalia" order="Rajiformes" pageId="2" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="FA84EAE0402B5258FCAFF5901A6DFF34" box="[788,888,152,176]" italics="true" pageId="2">Orbiraja</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="0FF04D71402B5258FC0DF59019C1FF51" authority="Weigmann, Stehmann & Thiel" authorityName="Weigmann, Stehmann & Thiel" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="2" phylum="Chordata" rank="species" species="ornata">
|
||
<emphasis id="FA84EAE0402B5258FC0DF5901D62FF34" box="[950,1143,152,176]" italics="true" pageId="2">Okamejei ornata</emphasis>
|
||
Weigmann, Stehmann & Thiel
|
||
</taxonomicName>
|
||
) and the ventral terminal cartilage is claw shaped (not claw-shaped in
|
||
<taxonomicName id="0FF04D71402B5258FC45F5B41D77FF50" box="[1022,1122,188,212]" class="Elasmobranchii" family="Rajidae" genus="Orbiraja" kingdom="Animalia" order="Rajiformes" pageId="2" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="FA84EAE0402B5258FC45F5B41D77FF50" box="[1022,1122,188,212]" italics="true" pageId="2">Orbiraja</emphasis>
|
||
</taxonomicName>
|
||
). Neither
|
||
<taxonomicName id="0FF04D71402B5258FB63F5B41C56FF50" box="[1240,1347,188,212]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="2" phylum="Chordata" rank="genus">
|
||
<emphasis id="FA84EAE0402B5258FB63F5B41C56FF50" box="[1240,1347,188,212]" italics="true" pageId="2">Okamejei</emphasis>
|
||
</taxonomicName>
|
||
nor the other rostrorajin genera (
|
||
<taxonomicName id="0FF04D71402B5258FE16F5E81B24FF7C" box="[429,561,224,248]" class="Elasmobranchii" family="Rajidae" genus="Malacoraja" kingdom="Animalia" order="Rajiformes" pageId="2" phylum="Chordata" rank="genus">
|
||
<emphasis id="FA84EAE0402B5258FE16F5E81B24FF7C" box="[429,561,224,248]" italics="true" pageId="2">Malacoraja</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0FF04D71402B5258FD84F5E81B8EFF73" box="[575,667,224,247]" class="Elasmobranchii" family="Rajidae" genus="Neoraja" kingdom="Animalia" order="Rajiformes" pageId="2" phylum="Chordata" rank="genus">
|
||
<emphasis id="FA84EAE0402B5258FD84F5E81B8EFF73" box="[575,667,224,247]" italics="true" pageId="2">Neoraja</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="0FF04D71402B5258FD6AF5E81A5CFF73" box="[721,841,224,247]" class="Elasmobranchii" family="Rajidae" genus="Rostroraja" kingdom="Animalia" order="Rajiformes" pageId="2" phylum="Chordata" rank="genus">
|
||
<emphasis id="FA84EAE0402B5258FD6AF5E81A5CFF73" box="[721,841,224,247]" italics="true" pageId="2">Rostroraja</emphasis>
|
||
</taxonomicName>
|
||
) are known to have an accessory terminal 3 cartilage (arising from the disto-medial margin of the accessory terminal 2 cartilage); an accessory terminal 3 cartilage is present in the ‘amphi-American assemblage’ but is located on the disto-lateral extension of the dorsal marginal cartilage (J. McEachran, pers comm.). The position of the spiracles relative to the eyeball in
|
||
<taxonomicName id="0FF04D71402B5258FB1DF4441C1FFEE0" box="[1190,1290,332,356]" class="Elasmobranchii" family="Rajidae" genus="Orbiraja" kingdom="Animalia" order="Rajiformes" pageId="2" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="FA84EAE0402B5258FB1DF4441C1FFEE0" box="[1190,1290,332,356]" italics="true" pageId="2">Orbiraja</emphasis>
|
||
</taxonomicName>
|
||
is somewhat unusual within skates (placed more posteriorly) and needs further exploration. The orbit + spiracle length is 1.7– 2.0 times the orbit diameter in
|
||
<taxonomicName id="0FF04D71402B5258FDB4F49C1BF4FE28" box="[527,737,404,428]" class="Elasmobranchii" family="Rajidae" genus="Orbiraja" kingdom="Animalia" order="Rajiformes" pageId="2" phylum="Chordata" rank="species" species="jensenae">
|
||
<emphasis id="FA84EAE0402B5258FDB4F49C1BF4FE28" box="[527,737,404,428]" italics="true" pageId="2">Orbiraja jensenae</emphasis>
|
||
</taxonomicName>
|
||
versus 1.2–l.6 times in several rajin skates examined (PL, unpubl.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |