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<document id="66BACA2D1EEF248CD827635E2C240BA3" ID-CLB-Dataset="30311" ID-DOI="10.11646/zootaxa.4407.2.5" ID-GBIF-Dataset="177c225c-e7db-4fc0-a099-f88eead488de" ID-ISSN="1175-5326" ID-Zenodo-Dep="1216378" ID-ZooBank="6B58A9F4-1A92-4E3A-92E3-1B80388AE534" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1523432164308" checkinUser="plazi" docAuthor="Yang, Yuxia, Liu, Haoyu &amp; Yang, Xingke" docDate="2018" docId="374387C2EA761232FF02BA73D7AF06DA" docLanguage="en" docName="zootaxa.4407.2.5.pdf" docOrigin="Zootaxa 4407 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Themus (Haplothemus) quadratus Wittmer 1983" docType="treatment" docVersion="7" lastPageNumber="244" masterDocId="CB7AFFBAEA771231FF95BB1DD44A0622" masterDocTitle="A contribution to the knowledge of Themus (Haplothemus) Wittmer from China (Coleoptera: Cantharidae)" masterLastPageNumber="253" masterPageNumber="241" pageNumber="242" updateTime="1698535925069" updateUser="ExternalLinkService">
<mods:mods id="6EEE599C5EA846A9F3F3CE92DB606D6A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6A3DBFDDA7FB3D547DF00CCC86D65C75">
<mods:title id="9993933AE87CA6FF14D0403A6D7B3320">A contribution to the knowledge of Themus (Haplothemus) Wittmer from China (Coleoptera: Cantharidae)</mods:title>
</mods:titleInfo>
<mods:name id="7883CB667F2B0BE356351BE76A5FA766" type="personal">
<mods:role id="50190F8E9956BBB641836C2B938E403C">
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<mods:namePart id="3D96EA7D6B7334BD60E5C76F63897CAB">Yang, Yuxia</mods:namePart>
</mods:name>
<mods:name id="CE0BF5C30642B582B7D3D8B4D86B8726" type="personal">
<mods:role id="EF5EACA8D16AA8CF225515BA48C5129E">
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</mods:role>
<mods:namePart id="3B70DDDDFE1DD2011BB19D77C38A4017">Liu, Haoyu</mods:namePart>
</mods:name>
<mods:name id="8C2859CAD932B496D7CE7ABEDF1263C9" type="personal">
<mods:role id="5CDA486255C93FCB8876B6F84EE3F246">
<mods:roleTerm id="FF825558392FCA4CE99888990777FB66">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2B0DEE28751FE79C5F20C7C8A9C8239F">Yang, Xingke</mods:namePart>
</mods:name>
<mods:typeOfResource id="F0351574DFA2F64A901B35BC7998CBC4">text</mods:typeOfResource>
<mods:relatedItem id="1124F0E37B7A3CF1B1B3FB2600A035BD" type="host">
<mods:titleInfo id="F9CAAA29DA0E846B74969E6A3D2AC9C4">
<mods:title id="AA24EE08851EB9EC8E82489FB410BFFE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1EBD0E862BA007FA89184EA41F1A5A7A">
<mods:date id="27308FA5829B0170D62399CDBA278BF1">2018</mods:date>
<mods:detail id="2588B04A67EEEDFA28F4B9D762BF5FD3" type="pubDate">
<mods:number id="E4A6532CDA3503C8BB972040D6D4B879">2018-04-10</mods:number>
</mods:detail>
<mods:detail id="8F2385FCC6555E6275E0432F60E00084" type="volume">
<mods:number id="D342C7158064CEC9E3AD2C7F28B43CC4">4407</mods:number>
</mods:detail>
<mods:detail id="6BF91464FADE356A86D44799318A8439" type="issue">
<mods:number id="9A22C0687470F27C5893173BC73C43F7">2</mods:number>
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</mods:extent>
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<mods:classification id="203348D6C94A02B0B72FD1AE7A4BA70D">journal article</mods:classification>
<mods:identifier id="824F1F4A6F84E363918B9DDE2A6D280E" type="CLB-Dataset">30311</mods:identifier>
<mods:identifier id="B175273C7D023776BF13C00B9FA8A518" type="DOI">10.11646/zootaxa.4407.2.5</mods:identifier>
<mods:identifier id="6FBACEC7E7524DA2EC0C7F7A425B555D" type="GBIF-Dataset">177c225c-e7db-4fc0-a099-f88eead488de</mods:identifier>
<mods:identifier id="E1E8EBF92A223E63D4B0AB8538EE1F17" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="4ED09A434F749C8143B8600D0A34DC51" type="ZooBank">6B58A9F4-1A92-4E3A-92E3-1B80388AE534</mods:identifier>
</mods:mods>
<treatment id="374387C2EA761232FF02BA73D7AF06DA" ID-DOI="http://doi.org/10.5281/zenodo.5954975" ID-GBIF-Taxon="143536476" ID-Zenodo-Dep="5954975" LSID="urn:lsid:plazi:treatment:374387C2EA761232FF02BA73D7AF06DA" httpUri="http://treatment.plazi.org/id/374387C2EA761232FF02BA73D7AF06DA" lastPageId="3" lastPageNumber="244" pageId="1" pageNumber="242">
<subSubSection id="F7F0655FEA761230FF02BA73D522078B" pageId="1" pageNumber="242" type="nomenclature">
<paragraph id="BF5536D4EA761230FF02BA73D6BA07AA" blockId="1.[151,752,366,426]" box="[151,752,366,392]" pageId="1" pageNumber="242">
<heading id="E41D81B8EA761230FF02BA73D6BA07AA" bold="true" box="[151,752,366,392]" fontSize="11" level="1" pageId="1" pageNumber="242" reason="1">
<taxonomicName id="78EA4D57EA761230FF02BA73D6BA07AA" authority="Wittmer, 1983" authorityName="Wittmer" authorityYear="1983" box="[151,752,366,392]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="242" phylum="Arthropoda" rank="species" species="quadratus" subGenus="Haplothemus">
<emphasis id="8D9EEAC6EA761230FF02BA73D6BA07AA" bold="true" box="[151,752,366,392]" pageId="1" pageNumber="242">
<emphasis id="8D9EEAC6EA761230FF02BA73D4B307AA" bold="true" box="[151,249,366,392]" italics="true" pageId="1" pageNumber="242">Themus</emphasis>
(
<emphasis id="8D9EEAC6EA761230FE9FBA73D5E707AA" bold="true" box="[266,429,366,392]" italics="true" pageId="1" pageNumber="242">Haplothemus</emphasis>
)
<emphasis id="8D9EEAC6EA761230FE28BA73D67D07AA" bold="true" box="[445,567,366,392]" italics="true" pageId="1" pageNumber="242">quadratus</emphasis>
Wittmer, 1983
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="BF5536D4EA761230FF02BA8CD522078B" blockId="1.[151,752,366,426]" box="[151,360,401,426]" pageId="1" pageNumber="242">
<figureCitation id="27D12A51EA761230FF02BA8CD496078B" box="[151,220,401,426]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1947,1970]" captionTargetBox="[172,1394,461,1888]" captionTargetId="figure@2.[151,1436,447,1923]" captionTargetPageId="2" captionText="FIGURES 1–6. Male habitus, dorsal vieW: 1. Themus (Haplothemus) quadratus Wittmer, 1983; 2. T. (H.) quadratiformis sp. nov.; 3. T. (H.) elongatior Pic, 1955; 4. T. (H.) mishmiensis Wittmer, 1973; 5. weishanensis Kopetz, 2016; 6. T. (H.) longideverticulum sp. nov. Scale bars:5.0 mm." httpUri="https://zenodo.org/record/1216380/files/figure.png" pageId="1" pageNumber="242">Figs 1</figureCitation>
,
<figureCitation id="27D12A51EA761230FF7FBA8CD55C078B" box="[234,278,401,425]" captionStart="FIGURES7–15" captionStartId="5.[151,264,1972,1995]" captionTargetBox="[238,1365,233,1932]" captionTargetId="figure@5.[172,1420,193,1954]" captionTargetPageId="5" captionText="FIGURES7–15. Aedeagus(7,10, 13. ventral vieW;8,11, 14.dorsal vieW;9, 12, 15.lateral vieW):7‒9. Themus(Haplothemus) quadratus Wittmer, 1983; 10‒12. T. (H.) quadratiformis sp. nov.; 13‒15. T. (H.) elongatior Pic, 1955.Scale bars:1.0 mm." httpUri="https://zenodo.org/record/1216382/files/figure.png" pageId="1" pageNumber="242">7‒9</figureCitation>
,
<figureCitation id="27D12A51EA761230FEB7BA8CD50B0788" box="[290,321,401,426]" captionStart="FIGURES25–29" captionStartId="8.[151,264,1912,1935]" captionTargetBox="[160,1412,274,1811]" captionTargetId="figure@8.[151,1436,241,1847]" captionTargetPageId="8" captionText="FIGURES25–29. Internalreproductive organ of female genitalia,lateral vieW:25.Themus (Haplothemus) quadratus Wittmer, 1983; 26. T. (H.) quadratiformis sp. nov.; 27. T. (H.) elongatior Pic, 1955; 28. T. (H.) weishanensis Kopetz, 2016; 29. T. (H.) longideverticulumsp.nov. Scalebars:1.0mm.(ag: accessorygland; di:diverticulum;sp: spermatheca; ov:medianoviduct; va: vagina; sr: sclerotized ring)" httpUri="https://zenodo.org/record/1216384/files/figure.png" pageId="1" pageNumber="242">25</figureCitation>
,
<figureCitation id="27D12A51EA761230FED9BA8CD522078B" box="[332,360,401,425]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1783,1806]" captionTargetBox="[202,1380,377,1733]" captionTargetId="figure@10.[151,1436,301,1762]" captionTargetPageId="10" captionText="FIGURES 30–34. Abdominal sternite VIII of female, ventral vieW: 30. Themus (Haplothemus) quadratus Wittmer, 1983; 31. T. (H.) quadratiformis sp. nov.; 32. T. (H.) elongatior Pic, 1955; 33. T. (H.) weishanensis Kopetz, 2016; 34. T. (H.) longideverticulum sp. nov. Scale bars:1.0 mm." httpUri="https://zenodo.org/record/1216386/files/figure.png" pageId="1" pageNumber="242">30</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F7F0655FEA761230FF02BAC4D6F2042C" pageId="1" pageNumber="242" type="reference_group">
<paragraph id="BF5536D4EA761230FF02BAC4D6F2042C" blockId="1.[151,709,473,526]" pageId="1" pageNumber="242">
<taxonomicName id="78EA4D57EA761230FF02BAC4D62F07CC" authority="Wittmer, 1983: 224" authorityName="Wittmer" authorityPageNumber="224" authorityYear="1983" box="[151,613,473,495]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="242" phylum="Arthropoda" rank="species" species="quadratus">
<emphasis id="8D9EEAC6EA761230FF02BAC4D4AC07CD" box="[151,230,473,495]" italics="true" pageId="1" pageNumber="242">Themus</emphasis>
(s.str.)
<emphasis id="8D9EEAC6EA761230FEA4BAC4D5DD07CD" box="[305,407,473,495]" italics="true" pageId="1" pageNumber="242">quadratus</emphasis>
Wittmer, 1983: 224
</taxonomicName>
, fig. 112.
<taxonomicName id="78EA4D57EA761230FF02BAE5D6FF042C" authority=": Svihla, 2008: 185" authorityName="Svihla" authorityPageNumber="185" authorityYear="2008" box="[151,693,503,526]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="242" phylum="Arthropoda" rank="species" species="quadratus" subGenus="Haplothemus">
<emphasis id="8D9EEAC6EA761230FF02BAE5D4AC042C" box="[151,230,504,526]" italics="true" pageId="1" pageNumber="242">Themus</emphasis>
(
<emphasis id="8D9EEAC6EA761230FF61BAE5D531042C" box="[244,379,504,526]" italics="true" pageId="1" pageNumber="242">Haplothemus</emphasis>
)
<emphasis id="8D9EEAC6EA761230FE1FBAE5D5BB042C" box="[394,497,504,526]" italics="true" pageId="1" pageNumber="242">quadratus</emphasis>
: Švihla, 2008: 185
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F7F0655FEA761230FF02B923D62604C8" pageId="1" pageNumber="242" type="materials_examined">
<paragraph id="BF5536D4EA761230FF02B923D7B50481" blockId="1.[151,1436,573,2007]" pageId="1" pageNumber="242">
<materialsCitation id="0F823C89EA761230FF02B923D7B60481" ID-GBIF-Occurrence="1837803962" collectingDate="1928-08" collectingDateMax="1957-08-28" collectingDateMin="1928-08" collectionCode="IZAS" collectorName="F. X. Zhu" country="China" elevation="1900" location="Xixiangchi" municipality="Emei Shan" pageId="1" pageNumber="242" specimenCount="3" stateProvince="Sichuan" typeStatus="holotype">
<emphasis id="8D9EEAC6EA761230FF02B923D5F70475" bold="true" box="[151,445,574,599]" pageId="1" pageNumber="242">
<typeStatus id="60518876EA761230FF02B923D4980475" box="[151,210,574,599]" pageId="1" pageNumber="242">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="60518876EA761230FE51B923D67A0475" box="[452,560,574,599]" pageId="1" pageNumber="242" type="holotype">Holotype</typeStatus>
: one female (
<collectionCode id="D9FBAE11EA761230FD56B922D7420475" box="[707,776,575,599]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="1" pageNumber="242">IZAS</collectionCode>
): [p] “洗象池 \ 1800-2000公尺 \ 采集者: 朱复兴”, [p] “四 川: 峨 眉山 \ 1957.
<collectingDate id="DB10E9FCEA761230FE1CB978D594045C" box="[393,478,613,638]" pageId="1" pageNumber="242" value="1928-08">
<date id="CB541014EA761230FE1CB978D594045C" box="[393,478,613,638]" pageId="1" pageNumber="242" value="1928-08">VIII.28</date>
</collectingDate>
\ 中 国科学院 ”, [p] “
<typeStatus id="60518876EA761230FD7BB97BD7D6045C" box="[750,924,614,638]" pageId="1" pageNumber="242" type="holotype">HOLOTYPUS</typeStatus>
”, [p] “IOZ (E) 217899”. [one female,
<collectingCountry id="C7FD7644EA761230FAC7B978D1D2045C" box="[1362,1432,613,638]" name="China" pageId="1" pageNumber="242">China</collectingCountry>
,
<collectingRegion id="7D2EF836EA761230FF02B997D4BE0481" box="[151,244,650,675]" country="China" name="Sichuan" pageId="1" pageNumber="242">Sichuan</collectingRegion>
,
<collectingMunicipality id="5F31ACAEEA761230FF6AB997D5360481" box="[255,380,650,675]" pageId="1" pageNumber="242">Emei Shan</collectingMunicipality>
,
<location id="BA35600FEA761230FE12B997D64C0481" LSID="urn:lsid:plazi:treatment:374387C2EA761232FF02BA73D7AF06DA:BA35600FEA761230FE12B997D64C0481" box="[391,518,650,675]" country="China" municipality="Emei Shan" name="Xixiangchi" pageId="1" pageNumber="242" stateProvince="Sichuan">Xixiangchi</location>
,
<elevation id="34C7D1E7EA761230FD87B997D6E50480" box="[530,687,650,674]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="1" pageNumber="242" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">
<quantity id="78129B31EA761230FD87B997D6E50480" box="[530,687,650,674]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="1" pageNumber="242" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">1800‒2000 m</quantity>
</elevation>
,
<collectingDate id="DB10E9FCEA761230FD2FB997D7050481" box="[698,847,650,675]" pageId="1" pageNumber="242" value="1957-08-28">
<date id="CB541014EA761230FD2FB997D7050481" box="[698,847,650,675]" pageId="1" pageNumber="242" value="1957-08-28">28.VIII.1957</date>
</collectingDate>
, leg.
<collectorName id="121F5302EA761230FC1EB996D7BE0481" box="[907,1012,650,675]" pageId="1" pageNumber="242">F.X. Zhu</collectorName>
]
</materialsCitation>
.
</paragraph>
<paragraph id="BF5536D4EA761230FF52B9B0D62604C8" blockId="1.[151,1436,573,2007]" pageId="1" pageNumber="242">
<materialsCitation id="0F823C89EA761230FF52B9B0D62604C8" ID-GBIF-Occurrence="1837803965" collectingDate="2003-07-08" collectionCode="MHBU" collectorName="Y. B. Ba &amp; Yu." country="China" county="Other" elevation="2350" location="Huangtuling" municipality="Wenxian" pageId="1" pageNumber="242" specimenCount="1" stateProvince="Gansu">
<emphasis id="8D9EEAC6EA761230FF52B9B0D64C04E4" bold="true" box="[199,518,685,710]" pageId="1" pageNumber="242">
<collectingCounty id="56344E58EA761230FF52B9B0D54504E4" box="[199,271,685,710]" pageId="1" pageNumber="242">Other</collectingCounty>
material examined.
</emphasis>
<collectingCountry id="C7FD7644EA761230FD84B9B3D62704E4" box="[529,621,686,710]" name="China" pageId="1" pageNumber="242">CHINA</collectingCountry>
,
<collectingRegion id="7D2EF836EA761230FDE8B9B3D68304E4" box="[637,713,686,710]" country="China" name="Gansu" pageId="1" pageNumber="242">Gansu</collectingRegion>
: three males, three females (
<collectionCode id="D9FBAE11EA761230FBB5B9B3D03104E4" box="[1056,1147,686,710]" pageId="1" pageNumber="242">MHBU</collectionCode>
):
<collectingMunicipality id="5F31ACAEEA761230FB06B9B0D0B304E4" box="[1171,1273,685,710]" pageId="1" pageNumber="242">Wenxian</collectingMunicipality>
,
<location id="BA35600FEA761230FA9CB9B0D1D204E4" LSID="urn:lsid:plazi:treatment:374387C2EA761232FF02BA73D7AF06DA:BA35600FEA761230FA9CB9B0D1D204E4" box="[1289,1432,685,710]" country="China" county="Other" municipality="Wenxian" name="Huangtuling" pageId="1" pageNumber="242" stateProvince="Gansu">Huangtuling</location>
,
<elevation id="34C7D1E7EA761230FF02B9CFD4A204C9" box="[151,232,722,747]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="1" pageNumber="242" unit="m" value="2350.0">
<quantity id="78129B31EA761230FF02B9CFD4A204C9" box="[151,232,722,747]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="1" pageNumber="242" unit="m" value="2350.0">2350m</quantity>
</elevation>
,
<collectingDate id="DB10E9FCEA761230FF66B9CFD53A04C8" box="[243,368,722,746]" pageId="1" pageNumber="242" value="2003-07-08">
<date id="CB541014EA761230FF66B9CFD53A04C8" box="[243,368,722,746]" pageId="1" pageNumber="242" value="2003-07-08">8.VII.2003</date>
</collectingDate>
, leg.
<collectorName id="121F5302EA761230FE39B9CED64904C8" box="[428,515,723,746]" pageId="1" pageNumber="242">Y.B. Ba</collectorName>
&amp; Y.
<collectorName id="121F5302EA761230FDD3B9CED62604C8" box="[582,620,723,746]" pageId="1" pageNumber="242">Yu.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="F7F0655FEA761230FF52B9E8D631052C" box="[199,635,757,782]" pageId="1" pageNumber="242" type="distribution">
<paragraph id="BF5536D4EA761230FF52B9E8D631052C" blockId="1.[151,1436,573,2007]" box="[199,635,757,782]" pageId="1" pageNumber="242">
<emphasis id="8D9EEAC6EA761230FF52B9E8D513052C" bold="true" box="[199,345,757,782]" pageId="1" pageNumber="242">Distribution</emphasis>
.
<collectingCountry id="C7FD7644EA761230FEF2B9E8D5E0052C" box="[359,426,757,782]" name="China" pageId="1" pageNumber="242">China</collectingCountry>
(
<collectingRegion id="7D2EF836EA761230FE2CB9E8D652052C" box="[441,536,757,782]" country="China" name="Sichuan" pageId="1" pageNumber="242">Sichuan</collectingRegion>
,
<collectingRegion id="7D2EF836EA761230FDB6B9EBD625052C" box="[547,623,758,782]" country="China" name="Gansu" pageId="1" pageNumber="242">Gansu</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="F7F0655FEA761233FF52B807D73707A5" lastPageId="2" lastPageNumber="243" pageId="1" pageNumber="242" type="description">
<paragraph id="BF5536D4EA761230FF52B807D79E020C" blockId="1.[151,1436,573,2007]" pageId="1" pageNumber="242">
<emphasis id="8D9EEAC6EA761230FF52B807D53E0511" bold="true" box="[199,372,794,819]" pageId="1" pageNumber="242">Redescription.</emphasis>
Male (
<figureCitation id="27D12A51EA761230FE5BB807D6590510" box="[462,531,794,819]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1947,1970]" captionTargetBox="[172,1394,461,1888]" captionTargetId="figure@2.[151,1436,447,1923]" captionTargetPageId="2" captionText="FIGURES 1–6. Male habitus, dorsal vieW: 1. Themus (Haplothemus) quadratus Wittmer, 1983; 2. T. (H.) quadratiformis sp. nov.; 3. T. (H.) elongatior Pic, 1955; 4. T. (H.) mishmiensis Wittmer, 1973; 5. weishanensis Kopetz, 2016; 6. T. (H.) longideverticulum sp. nov. Scale bars:5.0 mm." httpUri="https://zenodo.org/record/1216380/files/figure.png" pageId="1" pageNumber="242">Fig. 1</figureCitation>
). Head dark brown, black in middle of clypeus and both sides of vertex on dorsum, mouthparts dark brown, darkened at apices of mandibles, antennae black, prothorax dark brown, pronotum with two large irregular black markings on disc, markings more or less conjoint together in middle, sometimes extending to anterior margin but never to posterior or lateral margins, scutellum dark brown, elytra black, legs dark brown, slightly darkened at apical parts of femora and tibiae and tarsi, ventral surface dark brown, black at both sides of each abdominal ventrite except the terminal one, sometimes more or less darkened at meso- and metasterna. Body densely covered with decumbent brown pubescence, mixed with slightly long, semi-erected pubescence on elytra and few long hairs along anterior margin of clypeus.
</paragraph>
<paragraph id="BF5536D4EA761230FF52BF27D6100309" blockId="1.[151,1436,573,2007]" pageId="1" pageNumber="242">Head rounded, sparsely and finely punctate, surface lustrous; eyes moderately protruding, head width across eyes slightly wider than anterior margin of pronotum; terminal maxillary palpomeres nearly long-triangular, widest at basal two-fifths length, with outer margins slightly arcuate, inner apical margins slightly sharp and apices acute; antennae slightly thickened, almost extending to basal two-thirds length of elytra, antennomeres II about twice as long as wide, III slightly shorter than II, IV about 1.5 times as long as III, V longest, V‒X gradually shortened and narrowed one by one, XI slightly shorter than X and pointed at apices, IV‒VIII each with a small round or oblong impression near middle of outer margin.</paragraph>
<paragraph id="BF5536D4EA761230FF52BE2AD16403B4" blockId="1.[151,1436,573,2007]" pageId="1" pageNumber="242">Pronotum subquadrate, about 1.2 times as wide as long, anterior margin arcuate, lateral margins nearly parallel, posterior margin nearly straight and narrowly bordered, anterior angles nearly rectangular, posterior angles rounded, disc convex at postero-lateral parts, sparsely and finely punctate like that on head, surface matt.</paragraph>
<paragraph id="BF5536D4EA761230FF52BEBFD7DB03FC" blockId="1.[151,1436,573,2007]" pageId="1" pageNumber="242">Elytra about 5.5 times longer than pronotum, 3.0 times as long as humeral width, with lateral margins nearly parallel, disc rugulose-lacunose and coarsely punctate, surface matt.</paragraph>
<paragraph id="BF5536D4EA761230FF52BEF7D7970095" blockId="1.[151,1436,573,2007]" pageId="1" pageNumber="242">
Aedeagus (
<figureCitation id="27D12A51EA761230FEDDBEF7D5F80020" box="[328,434,1514,1539]" captionStart="FIGURES7–15" captionStartId="5.[151,264,1972,1995]" captionTargetBox="[238,1365,233,1932]" captionTargetId="figure@5.[172,1420,193,1954]" captionTargetPageId="5" captionText="FIGURES7–15. Aedeagus(7,10, 13. ventral vieW;8,11, 14.dorsal vieW;9, 12, 15.lateral vieW):7‒9. Themus(Haplothemus) quadratus Wittmer, 1983; 10‒12. T. (H.) quadratiformis sp. nov.; 13‒15. T. (H.) elongatior Pic, 1955.Scale bars:1.0 mm." httpUri="https://zenodo.org/record/1216382/files/figure.png" pageId="1" pageNumber="242">Figs 7‒9</figureCitation>
): ventral process of each paramere narrow, nearly straight and nearly pointed at apex; conjoint dorsal plate of parameres slightly shorter than ventral processes, with lateral margins nearly parallel, widely and triangularly emarginated in middle and slightly sinuate on both sides of apical margin, latero-apical angles slightly protuberant and sub-rounded; laterophyses depressed and slightly shorter than conjoint dorsal plate, separated on both sides of median lobe and distinctly diverging from each other, acute and slightly hooked at apices, which pointed towards latero-apical angles of conjoint dorsal plate.
</paragraph>
<paragraph id="BF5536D4EA761230FF52BDDFD5A90101" blockId="1.[151,1436,573,2007]" pageId="1" pageNumber="242">Female. Similar to male, but antennae slightly thinner and shorter, nearly extending to elytral mid-length, IV‒VIII without impressions; pronotum slightly wider, about 1.3 times as wide as long; elytra with lateral margins slightly diverging posteriorly.</paragraph>
<paragraph id="BF5536D4EA761230FF52BC33D0A80190" blockId="1.[151,1436,573,2007]" pageId="1" pageNumber="242">
Abdominal sternite VIII (
<figureCitation id="27D12A51EA761230FE76BC33D67E0165" box="[483,564,1838,1863]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1783,1806]" captionTargetBox="[202,1380,377,1733]" captionTargetId="figure@10.[151,1436,301,1762]" captionTargetPageId="10" captionText="FIGURES 30–34. Abdominal sternite VIII of female, ventral vieW: 30. Themus (Haplothemus) quadratus Wittmer, 1983; 31. T. (H.) quadratiformis sp. nov.; 32. T. (H.) elongatior Pic, 1955; 33. T. (H.) weishanensis Kopetz, 2016; 34. T. (H.) longideverticulum sp. nov. Scale bars:1.0 mm." httpUri="https://zenodo.org/record/1216386/files/figure.png" pageId="1" pageNumber="242">Fig. 30</figureCitation>
) triangularly emarginated in middle and bi-sinuately emarginate on both sides of posterior margin, the lateral emarginations both slightly shallower than the middle one, the anterior lateral emargination slightly wider than the posterior one, the portion between the two lateral emarginations nearly rectangular at apex, and that between the middle and posterior lateral emarginations acute at apex.
</paragraph>
<paragraph id="BF5536D4EA761233FF52BCA3D0640747" blockId="1.[151,1436,573,2007]" lastBlockId="2.[151,1436,151,392]" lastPageId="2" lastPageNumber="243" pageId="1" pageNumber="242">
Internal genitalia (
<figureCitation id="27D12A51EA761230FE02BCA3D5A601F5" box="[407,492,1982,2007]" captionStart="FIGURES25–29" captionStartId="8.[151,264,1912,1935]" captionTargetBox="[160,1412,274,1811]" captionTargetId="figure@8.[151,1436,241,1847]" captionTargetPageId="8" captionText="FIGURES25–29. Internalreproductive organ of female genitalia,lateral vieW:25.Themus (Haplothemus) quadratus Wittmer, 1983; 26. T. (H.) quadratiformis sp. nov.; 27. T. (H.) elongatior Pic, 1955; 28. T. (H.) weishanensis Kopetz, 2016; 29. T. (H.) longideverticulumsp.nov. Scalebars:1.0mm.(ag: accessorygland; di:diverticulum;sp: spermatheca; ov:medianoviduct; va: vagina; sr: sclerotized ring)" httpUri="https://zenodo.org/record/1216384/files/figure.png" pageId="1" pageNumber="242">Fig. 25</figureCitation>
): vagina stout, with diverticulum and spermatheca situated at apex, median oviduct at basal one-third part of ventral side; diverticulum moderately long and thick sac-shaped, slightly shorter than vagina, distinctly expanded at apical half part, about twice as long as maximal width, rounded at apex; spermatheca short sac-shaped, distinctly shorter than diverticulum, distinctly expanded apically and rounded at apex, and provided with a moderately long and thin accessory gland at base; diverticulum and spermatheca surrounded with a strongly sclerotized ring at base, which is confluent in middle and extending to median oviduct on ventral side, the opening of spermatheca arising from middle of the sclerotized ring on dorsal side.
</paragraph>
<paragraph id="BF5536D4EA751233FF52BA72D73707A5" blockId="2.[151,1436,151,392]" box="[199,893,367,392]" pageId="2" pageNumber="243">
Body length (both sexes): 17.0‒18.0 mm; width:
<quantity id="78129B31EA751233FD7BBA72D73207A5" box="[750,888,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="2" pageNumber="243" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.5‒4.5 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="EB95665CEA751233FF02BC86D63C01CD" httpUri="https://zenodo.org/record/1216380/files/figure.png" pageId="2" pageNumber="243" startId="2.[151,264,1947,1970]" targetBox="[172,1394,461,1888]" targetPageId="2">
<paragraph id="BF5536D4EA751233FF02BC86D63C01CD" blockId="2.[151,1436,1947,2031]" pageId="2" pageNumber="243">
<emphasis id="8D9EEAC6EA751233FF02BC86D5710193" bold="true" box="[151,315,1947,1970]" pageId="2" pageNumber="243">FIGURES 1–6.</emphasis>
Male habitus, dorsal vieW: 1.
<taxonomicName id="78EA4D57EA751233FDE1BC81D0240193" authority="Wittmer, 1983" authorityName="Wittmer" authorityYear="1983" box="[628,1134,1948,1970]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="243" phylum="Arthropoda" rank="species" species="quadratus" subGenus="Haplothemus">
<emphasis id="8D9EEAC6EA751233FDE1BC81D6890190" box="[628,707,1948,1970]" italics="true" pageId="2" pageNumber="243">Themus</emphasis>
(
<emphasis id="8D9EEAC6EA751233FD46BC81D7100190" box="[723,858,1948,1970]" italics="true" pageId="2" pageNumber="243">Haplothemus</emphasis>
)
<emphasis id="8D9EEAC6EA751233FCFFBC81D79B0190" box="[874,977,1948,1970]" italics="true" pageId="2" pageNumber="243">quadratus</emphasis>
Wittmer, 1983
</taxonomicName>
; 2.
<taxonomicName id="78EA4D57EA751233FB01BC80D13C0190" authority="Yang &amp; Liu &amp; Yang, 2018" authorityName="Yang &amp; Liu &amp; Yang" authorityYear="2018" box="[1172,1398,1948,1970]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="243" phylum="Arthropoda" rank="species" species="quadratiformis" status="sp. nov." subGenus="Haplothemus">
<emphasis id="8D9EEAC6EA751233FB01BC80D0EA0190" box="[1172,1184,1949,1970]" italics="true" pageId="2" pageNumber="243">T</emphasis>
. (
<emphasis id="8D9EEAC6EA751233FB23BC80D0820190" box="[1206,1224,1949,1970]" italics="true" pageId="2" pageNumber="243">H</emphasis>
.)
<emphasis id="8D9EEAC6EA751233FB4BBC81D13C0190" box="[1246,1398,1948,1970]" italics="true" pageId="2" pageNumber="243">quadratiformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="96AD57BDEA751233FAE8BC81D48901F2" pageId="2" pageNumber="243" rank="species">
<emphasis id="8D9EEAC6EA751233FAE8BC81D48901F2" bold="true" pageId="2" pageNumber="243">sp. nov.</emphasis>
</taxonomicNameLabel>
; 3.
<taxonomicName id="78EA4D57EA751233FF63BCA6D66301F2" authority="Pic, 1955" authorityName="Pic" authorityYear="1955" box="[246,553,1978,2001]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="243" phylum="Arthropoda" rank="species" species="elongatior" subGenus="Haplothemus">
<emphasis id="8D9EEAC6EA751233FF63BCA6D54801F2" box="[246,258,1979,2000]" italics="true" pageId="2" pageNumber="243">T</emphasis>
. (
<emphasis id="8D9EEAC6EA751233FE88BCA6D56501F2" box="[285,303,1979,2000]" italics="true" pageId="2" pageNumber="243">H</emphasis>
.)
<emphasis id="8D9EEAC6EA751233FEDEBCA7D5FF01F2" box="[331,437,1978,2000]" italics="true" pageId="2" pageNumber="243">elongatior</emphasis>
Pic, 1955
</taxonomicName>
; 4.
<taxonomicName id="78EA4D57EA751233FDCCBCA6D79B01F2" authority="Wittmer, 1973" authorityName="Wittmer" authorityYear="1973" box="[601,977,1978,2001]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="243" phylum="Arthropoda" rank="species" species="mishmiensis" subGenus="Haplothemus">
<emphasis id="8D9EEAC6EA751233FDCCBCA6D62F01F2" box="[601,613,1979,2000]" italics="true" pageId="2" pageNumber="243">T</emphasis>
. (
<emphasis id="8D9EEAC6EA751233FD15BCA6D6D801F2" box="[640,658,1979,2000]" italics="true" pageId="2" pageNumber="243">H</emphasis>
.)
<emphasis id="8D9EEAC6EA751233FD3BBCA7D76001F2" box="[686,810,1978,2000]" italics="true" pageId="2" pageNumber="243">mishmiensis</emphasis>
Wittmer, 1973
</taxonomicName>
; 5.
<taxonomicName id="78EA4D57EA751233FB95BCA7D16E01F2" authority="Kopetz, 2016" authorityName="Kopetz" authorityYear="2016" box="[1024,1316,1978,2000]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="243" phylum="Arthropoda" rank="species" species="weishanensis">
<emphasis id="8D9EEAC6EA751233FB95BCA7D0CC01F2" box="[1024,1158,1978,2000]" italics="true" pageId="2" pageNumber="243">weishanensis</emphasis>
Kopetz, 2016
</taxonomicName>
; 6.
<taxonomicName id="78EA4D57EA751233FAC1BCA6D50501CD" authority="Yang &amp; Liu &amp; Yang, 2018" authorityName="Yang &amp; Liu &amp; Yang" authorityYear="2018" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="243" phylum="Arthropoda" rank="species" species="longideverticulum" status="sp. nov." subGenus="Haplothemus">
<emphasis id="8D9EEAC6EA751233FAC1BCA6D12A01F2" box="[1364,1376,1979,2000]" italics="true" pageId="2" pageNumber="243">T</emphasis>
. (
<emphasis id="8D9EEAC6EA751233FAEEBCA6D1C701F2" box="[1403,1421,1979,2000]" italics="true" pageId="2" pageNumber="243">H</emphasis>
.)
<emphasis id="8D9EEAC6EA751233FF02BCC4D50501CD" box="[151,335,2009,2031]" italics="true" pageId="2" pageNumber="243">longideverticulum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="96AD57BDEA751233FEC0BCC4D5EF01CC" box="[341,421,2009,2030]" pageId="2" pageNumber="243" rank="species">
<emphasis id="8D9EEAC6EA751233FEC0BCC4D5EF01CC" bold="true" box="[341,421,2009,2030]" pageId="2" pageNumber="243">sp. nov.</emphasis>
</taxonomicNameLabel>
Scale bars: 5.0 mm.
</paragraph>
</caption>
<subSubSection id="F7F0655FEA741232FF52BB8AD7AF06DA" pageId="3" pageNumber="244" type="discussion">
<paragraph id="BF5536D4EA741232FF52BB8AD7AF06DA" blockId="3.[151,1437,151,248]" pageId="3" pageNumber="244">
<emphasis id="8D9EEAC6EA741232FF52BB8AD5710692" bold="true" box="[199,315,151,176]" pageId="3" pageNumber="244">Remarks.</emphasis>
The male of
<taxonomicName id="78EA4D57EA741232FE47BB84D70B0692" authority="Wittmer, 1983" authorityName="Wittmer" authorityYear="1983" box="[466,833,151,176]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="244" phylum="Arthropoda" rank="species" species="quadratus" subGenus="Haplothemus">
<emphasis id="8D9EEAC6EA741232FE47BB84D5950692" box="[466,479,153,176]" italics="true" pageId="3" pageNumber="244">T</emphasis>
. (
<emphasis id="8D9EEAC6EA741232FE6DBB84D6460692" box="[504,524,153,176]" italics="true" pageId="3" pageNumber="244">H</emphasis>
.)
<emphasis id="8D9EEAC6EA741232FDB1BB8AD6DF0692" box="[548,661,151,176]" italics="true" pageId="3" pageNumber="244">quadratus</emphasis>
Wittmer, 1983
</taxonomicName>
, which was described on a single female specimen in the original manuscript, is discovered and described here for the first time. At the same time, a related species is recognized and described under the name of
<taxonomicName id="78EA4D57EA741232FD19BBFCD7CE06DA" authority="Yang &amp; Liu &amp; Yang, 2018" authorityName="Yang &amp; Liu &amp; Yang" authorityYear="2018" box="[652,900,223,248]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="244" phylum="Arthropoda" rank="species" species="quadratiformis" status="sp. nov." subGenus="Haplothemus">
<emphasis id="8D9EEAC6EA741232FD19BBFCD6D306DA" box="[652,665,225,248]" italics="true" pageId="3" pageNumber="244">T</emphasis>
. (
<emphasis id="8D9EEAC6EA741232FD24BBFCD68F06DA" box="[689,709,225,248]" italics="true" pageId="3" pageNumber="244">H</emphasis>
.)
<emphasis id="8D9EEAC6EA741232FD49BBC2D7CE06DA" box="[732,900,223,248]" italics="true" pageId="3" pageNumber="244">quadratiformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="96AD57BDEA741232FC19BBC2D7AF06DA" box="[908,997,223,248]" pageId="3" pageNumber="244" rank="species">
<emphasis id="8D9EEAC6EA741232FC19BBC2D7AF06DA" bold="true" box="[908,997,223,248]" pageId="3" pageNumber="244">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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