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<document id="FD492120316A11E47C818AEDAD98DCFD" ID-CLB-Dataset="30311" ID-DOI="10.11646/zootaxa.4407.2.5" ID-GBIF-Dataset="177c225c-e7db-4fc0-a099-f88eead488de" ID-ISSN="1175-5326" ID-Zenodo-Dep="1216378" ID-ZooBank="6B58A9F4-1A92-4E3A-92E3-1B80388AE534" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1523432164308" checkinUser="plazi" docAuthor="Yang, Yuxia, Liu, Haoyu &amp; Yang, Xingke" docDate="2018" docId="374387C2EA731237FF02B939D10D0442" docLanguage="en" docName="zootaxa.4407.2.5.pdf" docOrigin="Zootaxa 4407 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Themus (Haplothemus) elongatior Pic 1955" docType="treatment" docVersion="7" lastPageNumber="247" masterDocId="CB7AFFBAEA771231FF95BB1DD44A0622" masterDocTitle="A contribution to the knowledge of Themus (Haplothemus) Wittmer from China (Coleoptera: Cantharidae)" masterLastPageNumber="253" masterPageNumber="241" pageNumber="245" updateTime="1698535925069" updateUser="ExternalLinkService">
<mods:mods id="D0EA97F55CFADC27DF3D38F32A5A84F8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F1604F79A3B47A0109F76A48DDCCC036">
<mods:title id="7BA3CC8EF8DAB6796B61B676F72720AA">A contribution to the knowledge of Themus (Haplothemus) Wittmer from China (Coleoptera: Cantharidae)</mods:title>
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<mods:namePart id="E666AE88E38C0060E9A9335D2EB31F19">Yang, Yuxia</mods:namePart>
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<mods:namePart id="498968FD623799541B8165B65E8C4A52">Liu, Haoyu</mods:namePart>
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<mods:namePart id="18E074042DE4557055AA618BB2B836C7">Yang, Xingke</mods:namePart>
</mods:name>
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<mods:title id="D17409A3500F8A89D8A927562CEF56ED">Zootaxa</mods:title>
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<mods:part id="8E1F0E54AF8C98C24CE78C4C69A63D1E">
<mods:date id="096C1184D90E70C8D8E5BF6BFD58770D">2018</mods:date>
<mods:detail id="D4936081B3637B1BFF3B932B6CCABBB5" type="pubDate">
<mods:number id="CBAB5DB6F9DECFE05B9038067578C40B">2018-04-10</mods:number>
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<mods:detail id="9A84090556D10E75005839BDB927F733" type="volume">
<mods:number id="43628FA3D759F0B0465919F9ECF568FC">4407</mods:number>
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<mods:classification id="276DF013FDBE748689DBD70FEB6D1857">journal article</mods:classification>
<mods:identifier id="2AB7E9B85AB86A0D5B7BB9682A65C1A5" type="CLB-Dataset">30311</mods:identifier>
<mods:identifier id="CFDD9AA7490BE4378A22B094C3422AC8" type="DOI">10.11646/zootaxa.4407.2.5</mods:identifier>
<mods:identifier id="9A44233BB8B493C13303ADBC76CD127C" type="GBIF-Dataset">177c225c-e7db-4fc0-a099-f88eead488de</mods:identifier>
<mods:identifier id="FC17CC2A177F94AC4B405B9067CDF195" type="ISSN">1175-5326</mods:identifier>
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<treatment id="374387C2EA731237FF02B939D10D0442" ID-DOI="http://doi.org/10.5281/zenodo.5954981" ID-GBIF-Taxon="143536478" ID-Zenodo-Dep="5954981" LSID="urn:lsid:plazi:treatment:374387C2EA731237FF02B939D10D0442" httpUri="http://treatment.plazi.org/id/374387C2EA731237FF02B939D10D0442" lastPageId="6" lastPageNumber="247" pageId="4" pageNumber="245">
<subSubSection id="F7F0655FEA731235FF02B939D5CE047D" pageId="4" pageNumber="245" type="nomenclature">
<paragraph id="BF5536D4EA731235FF02B939D6FB041C" blockId="4.[151,689,548,607]" box="[151,689,548,574]" pageId="4" pageNumber="245">
<heading id="E41D81B8EA731235FF02B939D6FB041C" bold="true" box="[151,689,548,574]" fontSize="11" level="1" pageId="4" pageNumber="245" reason="1">
<taxonomicName id="78EA4D57EA731235FF02B939D6FB041C" authority="Pic, 1955" authorityName="Pic" authorityYear="1955" box="[151,689,548,574]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="elongatior" subGenus="Haplothemus">
<emphasis id="8D9EEAC6EA731235FF02B939D6FB041C" bold="true" box="[151,689,548,574]" pageId="4" pageNumber="245">
<emphasis id="8D9EEAC6EA731235FF02B939D4B3041C" bold="true" box="[151,249,548,574]" italics="true" pageId="4" pageNumber="245">Themus</emphasis>
(
<emphasis id="8D9EEAC6EA731235FE9FB939D5E7041C" bold="true" box="[266,429,548,574]" italics="true" pageId="4" pageNumber="245">Haplothemus</emphasis>
)
<emphasis id="8D9EEAC6EA731235FE28B939D673041C" bold="true" box="[445,569,548,574]" italics="true" pageId="4" pageNumber="245">elongatior</emphasis>
<bibRefCitation id="DB7B4B25EA731235FDD5B939D6FB041C" box="[576,689,548,574]" pageId="4" pageNumber="245" refId="ref6267" refString="Pic, M. 1955. Descriptions diverses. Diversites Entomologiques, 14, 7 - 20." type="journal article">Pic, 1955</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="BF5536D4EA731235FF02B95BD5CE047D" blockId="4.[151,689,548,607]" box="[151,388,582,607]" pageId="4" pageNumber="245">
<figureCitation id="27D12A51EA731235FF02B95BD497047D" box="[151,221,582,607]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1947,1970]" captionTargetBox="[172,1394,461,1888]" captionTargetId="figure@2.[151,1436,447,1923]" captionTargetPageId="2" captionText="FIGURES 1–6. Male habitus, dorsal vieW: 1. Themus (Haplothemus) quadratus Wittmer, 1983; 2. T. (H.) quadratiformis sp. nov.; 3. T. (H.) elongatior Pic, 1955; 4. T. (H.) mishmiensis Wittmer, 1973; 5. weishanensis Kopetz, 2016; 6. T. (H.) longideverticulum sp. nov. Scale bars:5.0 mm." httpUri="https://zenodo.org/record/1216380/files/figure.png" pageId="4" pageNumber="245">Figs 3</figureCitation>
,
<figureCitation id="27D12A51EA731235FF7FB95BD578047D" box="[234,306,582,607]" captionStart="FIGURES7–15" captionStartId="5.[151,264,1972,1995]" captionTargetBox="[238,1365,233,1932]" captionTargetId="figure@5.[172,1420,193,1954]" captionTargetPageId="5" captionText="FIGURES7–15. Aedeagus(7,10, 13. ventral vieW;8,11, 14.dorsal vieW;9, 12, 15.lateral vieW):7‒9. Themus(Haplothemus) quadratus Wittmer, 1983; 10‒12. T. (H.) quadratiformis sp. nov.; 13‒15. T. (H.) elongatior Pic, 1955.Scale bars:1.0 mm." httpUri="https://zenodo.org/record/1216382/files/figure.png" pageId="4" pageNumber="245">13‒15</figureCitation>
,
<figureCitation id="27D12A51EA731235FEABB95AD516047D" box="[318,348,583,607]" captionStart="FIGURES25–29" captionStartId="8.[151,264,1912,1935]" captionTargetBox="[160,1412,274,1811]" captionTargetId="figure@8.[151,1436,241,1847]" captionTargetPageId="8" captionText="FIGURES25–29. Internalreproductive organ of female genitalia,lateral vieW:25.Themus (Haplothemus) quadratus Wittmer, 1983; 26. T. (H.) quadratiformis sp. nov.; 27. T. (H.) elongatior Pic, 1955; 28. T. (H.) weishanensis Kopetz, 2016; 29. T. (H.) longideverticulumsp.nov. Scalebars:1.0mm.(ag: accessorygland; di:diverticulum;sp: spermatheca; ov:medianoviduct; va: vagina; sr: sclerotized ring)" httpUri="https://zenodo.org/record/1216384/files/figure.png" pageId="4" pageNumber="245">27</figureCitation>
,
<figureCitation id="27D12A51EA731235FEFDB95AD5CE047D" box="[360,388,583,607]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1783,1806]" captionTargetBox="[202,1380,377,1733]" captionTargetId="figure@10.[151,1436,301,1762]" captionTargetPageId="10" captionText="FIGURES 30–34. Abdominal sternite VIII of female, ventral vieW: 30. Themus (Haplothemus) quadratus Wittmer, 1983; 31. T. (H.) quadratiformis sp. nov.; 32. T. (H.) elongatior Pic, 1955; 33. T. (H.) weishanensis Kopetz, 2016; 34. T. (H.) longideverticulum sp. nov. Scale bars:1.0 mm." httpUri="https://zenodo.org/record/1216386/files/figure.png" pageId="4" pageNumber="245">32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F7F0655FEA731235FF02B993D6F104C0" pageId="4" pageNumber="245" type="reference_group">
<paragraph id="BF5536D4EA731235FF02B993D5A20487" blockId="4.[151,1126,654,738]" box="[151,488,654,677]" pageId="4" pageNumber="245">
<treatmentCitationGroup id="9FFA11FAEA731235FF02B993D5A20487" box="[151,488,654,677]" pageId="4" pageNumber="245">
<taxonomicName id="78EA4D57EA731235FF02B993D5AE0487" authority="Pic, 1955: 16" authorityName="Pic" authorityPageNumber="16" authorityYear="1955" box="[151,484,654,677]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="elongatior">
<emphasis id="8D9EEAC6EA731235FF02B993D51C0486" box="[151,342,654,676]" italics="true" pageId="4" pageNumber="245">Themus elongatior</emphasis>
<treatmentCitation id="3E4B10C5EA731235FECEB992D5AE0487" author="" box="[347,484,655,677]" page="16" pageId="4" pageNumber="245">
<bibRefCitation id="DB7B4B25EA731235FECEB992D5F40487" box="[347,446,655,677]" pageId="4" pageNumber="245" refId="ref6267" refString="Pic, M. 1955. Descriptions diverses. Diversites Entomologiques, 14, 7 - 20." type="journal article">Pic, 1955</bibRefCitation>
: 16
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="BF5536D4EA731235FF02B9B0D6F104C0" blockId="4.[151,1126,654,738]" pageId="4" pageNumber="245">
<treatmentCitationGroup id="9FFA11FAEA731235FF02B9B0D6F104C0" pageId="4" pageNumber="245">
<taxonomicName id="78EA4D57EA731235FF02B9B0D62504E0" authority=": Wittmer, 1973: 191" authorityName="Wittmer" authorityPageNumber="191" authorityYear="1973" box="[151,623,685,707]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="elongatior">
<emphasis id="8D9EEAC6EA731235FF02B9B0D4AC04E1" box="[151,230,685,707]" italics="true" pageId="4" pageNumber="245">Themus</emphasis>
(s.str.)
<emphasis id="8D9EEAC6EA731235FEA4B9B0D5D004E1" box="[305,410,685,707]" italics="true" pageId="4" pageNumber="245">elongatior</emphasis>
: Wittmer, 1973: 191
</taxonomicName>
, 210, figs 23, 24;
<treatmentCitation id="3E4B10C5EA731235FCB0B9B0D7BC04E1" author="" box="[805,1014,685,707]" page="57" pageId="4" pageNumber="245">
<bibRefCitation id="DB7B4B25EA731235FCB0B9B0D79A04E0" box="[805,976,685,707]" pageId="4" pageNumber="245" refId="ref6223" refString="Okushima, Y. (1999) Cantharidae collected by the Hokkaido University Expeditions to Nepal Himalaya (Coleoptera). Insecta Matsumurana, 56, 51 - 68. [http: // hdl. handle. net / 2115 / 9897]" type="journal article">Okushima, 1999</bibRefCitation>
: 57
</treatmentCitation>
, figs 5, 30.
<taxonomicName id="78EA4D57EA731235FF02B9D1D6FD04C0" authority=": Svihla, 2008: 185" authorityName=": Svihla" authorityPageNumber="185" authorityYear="2008" box="[151,695,715,738]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="elongatior" subGenus="Haplothemus">
<emphasis id="8D9EEAC6EA731235FF02B9D1D4AC04C0" box="[151,230,716,738]" italics="true" pageId="4" pageNumber="245">Themus</emphasis>
(
<emphasis id="8D9EEAC6EA731235FF61B9D1D53104C0" box="[244,379,716,738]" italics="true" pageId="4" pageNumber="245">Haplothemus</emphasis>
)
<emphasis id="8D9EEAC6EA731235FE1FB9D1D5BE04C0" box="[394,500,716,738]" italics="true" pageId="4" pageNumber="245">elongatior</emphasis>
: Švihla, 2008: 185
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F7F0655FEA731235FF02B813D5DD0201" pageId="4" pageNumber="245" type="materials_examined">
<paragraph id="BF5536D4EA731235FF02B813D514054D" blockId="4.[151,1437,782,2031]" pageId="4" pageNumber="245">
<emphasis id="8D9EEAC6EA731235FF02B813D59B0505" bold="true" box="[151,465,782,807]" pageId="4" pageNumber="245">
<typeStatus id="60518876EA731235FF02B813D4980505" box="[151,210,782,807]" pageId="4" pageNumber="245">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="60518876EA731235FE74B813D6040505" box="[481,590,782,807]" pageId="4" pageNumber="245" type="holotype">Holotype</typeStatus>
:
<specimenCount id="A9ECFD5DEA731235FDF4B812D6CE0505" box="[609,644,783,807]" pageId="4" pageNumber="245" type="male">1♂</specimenCount>
(
<collectionCode id="D9FBAE11EA731235FD09B812D6BD0505" box="[668,759,783,807]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="245">MNHN</collectionCode>
), [p] “
<collectingCountry id="C7FD7644EA731235FCDBB812D0560505" box="[846,1052,783,807]" name="India" pageId="4" pageNumber="245">BRITISH INDIA</collectingCountry>
\
<collectionCode id="D9FBAE11EA731235FBD1B812D0D40505" box="[1092,1182,783,807]" pageId="4" pageNumber="245">SKKIM</collectionCode>
\ Lachen-Lachung \
<date id="CB541014EA731235FF02B82ED5400569" box="[151,266,819,843]" pageId="4" pageNumber="245" value="1933-08">VIII.1933</date>
”, [h] “
<taxonomicName id="78EA4D57EA731235FEF1B82FD58B0569" authorityName="Motschulsky" authorityYear="1858" box="[356,449,818,843]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="genus">Themus</taxonomicName>
\
<taxonomicName id="78EA4D57EA731235FE7CB82FD6110569" authorityName=": Svihla" authorityYear="2008" box="[489,603,818,843]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="elongatior">elongatior</taxonomicName>
\
<emphasis id="8D9EEAC6EA731235FD16B82FD6890569" bold="true" box="[643,707,818,843]" pageId="4" pageNumber="245">n. sp</emphasis>
.”, [p] “
<taxonomicName id="78EA4D57EA731235FCBCB82FD7CC0569" authorityName="Motschulsky" authorityYear="1858" box="[809,902,818,843]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="genus">Themus</taxonomicName>
\
<taxonomicName id="78EA4D57EA731235FC38B82FD0550569" authorityName=": Svihla" authorityYear="2008" box="[941,1055,818,843]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="elongatior">elongatior</taxonomicName>
\ Pic \ det. W. Wittmer”, [p] “
<typeStatus id="60518876EA731235FF34B84AD51A054D" box="[161,336,855,879]" pageId="4" pageNumber="245" type="holotype">HOLOTYPUS</typeStatus>
”.
</paragraph>
<paragraph id="BF5536D4EA731235FF52B867D5DD0201" blockId="4.[151,1437,782,2031]" pageId="4" pageNumber="245">
<materialsCitation id="0F823C89EA731235FF52B867D7980595" ID-GBIF-Occurrence="1837803957" collectingDate="2014-07-27" collectionCode="MHBU" collectorName="G. D. Ren &amp; Bai" country="China" county="Other" elevation="3492" location="Rongxia" municipality="Tingri" pageId="4" pageNumber="245" specimenCount="1" stateProvince="Xizang">
<emphasis id="8D9EEAC6EA731235FF52B867D64C05B1" bold="true" box="[199,518,890,915]" pageId="4" pageNumber="245">
<collectingCounty id="56344E58EA731235FF52B867D54505B1" box="[199,271,890,915]" pageId="4" pageNumber="245">Other</collectingCounty>
material examined.
</emphasis>
<collectingCountry id="C7FD7644EA731235FD84B866D62705B1" box="[529,621,891,915]" name="China" pageId="4" pageNumber="245">CHINA</collectingCountry>
,
<collectingRegion id="7D2EF836EA731235FDE8B867D69805B1" box="[637,722,890,915]" country="China" name="Xizang" pageId="4" pageNumber="245">Xizang</collectingRegion>
: one male, two females (
<collectionCode id="D9FBAE11EA731235FB95B866D01305B1" box="[1024,1113,891,915]" pageId="4" pageNumber="245">MHBU</collectionCode>
),
<collectingMunicipality id="5F31ACAEEA731235FBE7B867D0F005B1" box="[1138,1210,890,915]" pageId="4" pageNumber="245">Tingri</collectingMunicipality>
,
<location id="BA35600FEA731235FB5CB867D16105B1" LSID="urn:lsid:plazi:treatment:374387C2EA731237FF02B939D10D0442:BA35600FEA731235FB5CB867D16105B1" box="[1225,1323,890,915]" country="China" county="Other" municipality="Tingri" name="Rongxia" pageId="4" pageNumber="245" stateProvince="Xizang">Rongxia</location>
,
<elevation id="34C7D1E7EA731235FAAEB866D1DD05B0" box="[1339,1431,891,915]" metricMagnitude="3" metricUnit="m" metricValue="3.492" pageId="4" pageNumber="245" unit="m" value="3492.0">
<quantity id="78129B31EA731235FAAEB866D1DD05B0" box="[1339,1431,891,915]" metricMagnitude="3" metricUnit="m" metricValue="3.492" pageId="4" pageNumber="245" unit="m" value="3492.0">3492 m</quantity>
</elevation>
, 28°03.673ʹN, 86°20.919ʹE,
<collectingDate id="DB10E9FCEA731235FE4CB883D62E0595" box="[473,612,926,951]" pageId="4" pageNumber="245" value="2014-07-27">
<date id="CB541014EA731235FE4CB883D62E0595" box="[473,612,926,951]" pageId="4" pageNumber="245" value="2014-07-27">27.VII.2014</date>
</collectingDate>
, leg.
<collectorName id="121F5302EA731235FD39B883D7560594" box="[684,796,926,951]" pageId="4" pageNumber="245">G.D. Ren</collectorName>
, X.L.
<collectorName id="121F5302EA731235FCFBB883D7DF0595" box="[878,917,926,951]" pageId="4" pageNumber="245">Bai</collectorName>
&amp; J
</materialsCitation>
.
<materialsCitation id="0F823C89EA731235FC42B883D12205F9" ID-GBIF-Occurrence="1837803964" collectingDate="2014-08-03" collectionCode="MHBU" collectorName="S. Shan &amp; G. D. Ren &amp; Bai &amp; Shan" country="China" elevation="3234" location="Chentang" municipality="Tingri" pageId="4" pageNumber="245" specimenCount="1" stateProvince="Xizang">
<collectorName id="121F5302EA731235FC42B883D07C0595" box="[983,1078,926,951]" pageId="4" pageNumber="245">S. Shan</collectorName>
; one female (
<collectionCode id="D9FBAE11EA731235FB48B883D17D0595" box="[1245,1335,926,951]" pageId="4" pageNumber="245">MHBU</collectionCode>
),
<collectingMunicipality id="5F31ACAEEA731235FAC5B883D1D20595" box="[1360,1432,926,951]" pageId="4" pageNumber="245">Tingri</collectingMunicipality>
,
<location id="BA35600FEA731235FF02B8DFD54905F9" LSID="urn:lsid:plazi:treatment:374387C2EA731237FF02B939D10D0442:BA35600FEA731235FF02B8DFD54905F9" box="[151,259,962,987]" country="China" municipality="Tingri" name="Chentang" pageId="4" pageNumber="245" stateProvince="Xizang">Chentang</location>
power station,
<elevation id="34C7D1E7EA731235FE3BB8DFD64E05F8" box="[430,516,962,987]" metricMagnitude="3" metricUnit="m" metricValue="3.2339999999999995" pageId="4" pageNumber="245" unit="m" value="3234.0">
<quantity id="78129B31EA731235FE3BB8DFD64E05F8" box="[430,516,962,987]" metricMagnitude="3" metricUnit="m" metricValue="3.2339999999999995" pageId="4" pageNumber="245" unit="m" value="3234.0">3234 m</quantity>
</elevation>
, 27°55.069ʹN, 87°28.171ʹE,
<collectingDate id="DB10E9FCEA731235FCD0B8DFD78105F9" box="[837,971,962,987]" pageId="4" pageNumber="245" value="2014-08-03">
<date id="CB541014EA731235FCD0B8DFD78105F9" box="[837,971,962,987]" pageId="4" pageNumber="245" value="2014-08-03">3.VIII.2014</date>
</collectingDate>
, leg.
<collectorName id="121F5302EA731235FB93B8DFD03A05F8" box="[1030,1136,962,987]" pageId="4" pageNumber="245">G.D. Ren</collectorName>
, X.L.
<collectorName id="121F5302EA731235FB21B8DFD09105F9" box="[1204,1243,962,987]" pageId="4" pageNumber="245">Bai</collectorName>
&amp; J.S.
<collectorName id="121F5302EA731235FAB8B8DFD12205F9" box="[1325,1384,962,987]" pageId="4" pageNumber="245">Shan</collectorName>
</materialsCitation>
;
<materialsCitation id="0F823C89EA731235FAE1B8DED5DD0201" ID-GBIF-Occurrence="1837803955" collectionCode="IZAS" collectorName="Baibung, E &amp; Liang H. B." country="China" elevation="3210" latitude="29.46751" location="Medog" longLatPrecision="1" longitude="95.00232" pageId="4" pageNumber="245" specimenCount="1" stateProvince="Tibet">
one male (
<collectionCode id="D9FBAE11EA731235FF48B8FAD56A05DD" box="[221,288,999,1023]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="4" pageNumber="245">IZAS</collectionCode>
), “
<collectingRegion id="7D2EF836EA731235FEABB8FBD53705DD" box="[318,381,998,1023]" country="China" name="Xizang" pageId="4" pageNumber="245">Tibet</collectingRegion>
,
<location id="BA35600FEA731235FE1DB8FBD59005DD" LSID="urn:lsid:plazi:treatment:374387C2EA731237FF02B939D10D0442:BA35600FEA731235FE1DB8FBD59005DD" box="[392,474,998,1023]" country="China" latitude="29.46751" longLatPrecision="1" longitude="95.00232" name="Medog" pageId="4" pageNumber="245" stateProvince="Tibet">Mêdog</location>
,
<collectorName id="121F5302EA731235FE73B8FBD62D05DC" box="[486,615,998,1023]" pageId="4" pageNumber="245">Baibung, E</collectorName>
. Doxong Pass Lage hotel,
<collectingMethod id="66AB4EC3EA731235FC07B8FBD04005DC" box="[914,1034,998,1023]" pageId="4" pageNumber="245">pitfall trap</collectingMethod>
,
<geoCoordinate id="DADE5013EA731235FB83B8FBD0EB05DD" box="[1046,1185,998,1023]" direction="north" orientation="latitude" pageId="4" pageNumber="245" precision="1" value="29.46751">29.46751°N</geoCoordinate>
,
<geoCoordinate id="DADE5013EA731235FB38B8FBD17E05DD" box="[1197,1332,998,1023]" direction="east" orientation="longitude" pageId="4" pageNumber="245" precision="1" value="95.00232">95.00232°E</geoCoordinate>
,
<elevation id="34C7D1E7EA731235FAD5B8FAD1DD05DC" box="[1344,1431,999,1023]" metricMagnitude="3" metricUnit="m" metricValue="3.21" pageId="4" pageNumber="245" unit="m" value="3210.0">
<quantity id="78129B31EA731235FAD5B8FAD1DD05DC" box="[1344,1431,999,1023]" metricMagnitude="3" metricUnit="m" metricValue="3.21" pageId="4" pageNumber="245" unit="m" value="3210.0">3210 m</quantity>
</elevation>
, 2006.8.8,
<collectorName id="121F5302EA731235FE92BF17D5CC0201" box="[263,390,1034,1059]" pageId="4" pageNumber="245">Liang H.B.</collectorName>
”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="F7F0655FEA731235FF52BF33D6FF0265" box="[199,693,1070,1095]" pageId="4" pageNumber="245" type="distribution">
<paragraph id="BF5536D4EA731235FF52BF33D6FF0265" blockId="4.[151,1437,782,2031]" box="[199,693,1070,1095]" pageId="4" pageNumber="245">
<emphasis id="8D9EEAC6EA731235FF52BF33D52A0265" bold="true" box="[199,352,1070,1095]" pageId="4" pageNumber="245">Distribution.</emphasis>
<collectingCountry id="C7FD7644EA731235FEF2BF33D5E00265" box="[359,426,1070,1095]" name="China" pageId="4" pageNumber="245">China</collectingCountry>
(
<collectingRegion id="7D2EF836EA731235FE2CBF33D65A0265" box="[441,528,1070,1095]" country="China" name="Xizang" pageId="4" pageNumber="245">Xizang</collectingRegion>
);
<collectingCountry id="C7FD7644EA731235FDB6BF33D6150265" box="[547,607,1070,1095]" name="India" pageId="4" pageNumber="245">India</collectingCountry>
,
<collectingCountry id="C7FD7644EA731235FDFEBF33D6F80265" box="[619,690,1070,1095]" name="Nepal" pageId="4" pageNumber="245">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="F7F0655FEA731237FF52BF4FD481043A" lastPageId="6" lastPageNumber="247" pageId="4" pageNumber="245" type="description">
<paragraph id="BF5536D4EA731235FF52BF4FD4BE02D9" blockId="4.[151,1437,782,2031]" pageId="4" pageNumber="245">
<emphasis id="8D9EEAC6EA731235FF52BF4FD53E0249" bold="true" box="[199,372,1106,1131]" pageId="4" pageNumber="245">Redescription.</emphasis>
Male (
<figureCitation id="27D12A51EA731235FE59BF4FD65E0249" box="[460,532,1106,1131]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1947,1970]" captionTargetBox="[172,1394,461,1888]" captionTargetId="figure@2.[151,1436,447,1923]" captionTargetPageId="2" captionText="FIGURES 1–6. Male habitus, dorsal vieW: 1. Themus (Haplothemus) quadratus Wittmer, 1983; 2. T. (H.) quadratiformis sp. nov.; 3. T. (H.) elongatior Pic, 1955; 4. T. (H.) mishmiensis Wittmer, 1973; 5. weishanensis Kopetz, 2016; 6. T. (H.) longideverticulum sp. nov. Scale bars:5.0 mm." httpUri="https://zenodo.org/record/1216380/files/figure.png" pageId="4" pageNumber="245">Fig. 3</figureCitation>
). Body black, mouthparts orange brown, darkened at mandibular apices, legs orange brown at coxae, trochanters and basal parts of femora, ventral surface orange brown, abdominal ventrites except the terminal one each side with a small round black marking. Body densely covered with decumbent black pubescence on dorsal surface but brown on ventral surface, mixed with a few long hairs along anterior margin of clypeus.
</paragraph>
<paragraph id="BF5536D4EA731235FF52BE1BD62C03D5" blockId="4.[151,1437,782,2031]" pageId="4" pageNumber="245">Head rounded, sparsely and finely punctate, surface lustrous; eyes moderately protruding, head width across eyes slightly wider than anterior margin of pronotum; terminal maxillary palpomeres nearly long-triangular, widest at basal two-fifths length, with outer margins slightly arcuate, inner apical margins slightly sharp and apices acute; antennae slightly thickened, almost extending to elytral mid-length, antennomeres II about four times as long as wide, III distinctly shorter than II, IV about twice as long as III, V longest, V‒X gradually shortened and narrowed one by one, XI slightly shorter than X and pointed at apices, IV‒VIII each with a short longitudinal or oblong impression at apical part of outer margin.</paragraph>
<paragraph id="BF5536D4EA731235FF52BD1ED73F0040" blockId="4.[151,1437,782,2031]" pageId="4" pageNumber="245">Pronotum subquadrate, about 1.5 times as wide as long, anterior and lateral margins arcuate, posterior margin nearly straight and narrowly bordered, anterior and posterior angles widely rounded, disc convex at postero-lateral parts, sparsely and finely punctate like that on head, surface matt.</paragraph>
<paragraph id="BF5536D4EA731235FF52BD73D7DB0088" blockId="4.[151,1437,782,2031]" pageId="4" pageNumber="245">Elytra about 6.0 times longer than pronotum, 3.3 times as long as humeral width, with lateral margins nearly parallel, disc rugulose-lacunose and coarsely punctate, surface matt.</paragraph>
<paragraph id="BF5536D4EA731235FF52BDABD75D0185" blockId="4.[151,1437,782,2031]" pageId="4" pageNumber="245">
Aedeagus (
<figureCitation id="27D12A51EA731235FED3BDABD58600ED" box="[326,460,1718,1743]" captionStart="FIGURES7–15" captionStartId="5.[151,264,1972,1995]" captionTargetBox="[238,1365,233,1932]" captionTargetId="figure@5.[172,1420,193,1954]" captionTargetPageId="5" captionText="FIGURES7–15. Aedeagus(7,10, 13. ventral vieW;8,11, 14.dorsal vieW;9, 12, 15.lateral vieW):7‒9. Themus(Haplothemus) quadratus Wittmer, 1983; 10‒12. T. (H.) quadratiformis sp. nov.; 13‒15. T. (H.) elongatior Pic, 1955.Scale bars:1.0 mm." httpUri="https://zenodo.org/record/1216382/files/figure.png" pageId="4" pageNumber="245">Figs 13‒15</figureCitation>
): ventral process of each paramere moderately bent inwards in ventral view, abruptly narrowed at base, distinctly protuberant at basal part of outer margin, longitudinally depressed along the whole length of ventral side, rounded and slightly turned ventrally at apex; conjoint dorsal plate of parameres distinctly shorter than ventral processes, widely and roundly emarginated in middle of apical margin, lateral angles rectangular and around with long hairs; laterophyses slightly shorter than conjoint dorsal plate, situated on dorsal side of median lobe, conjoint and depressed, widely and triangularly emarginated in middle of apical part, with latero-apical part acute at apex and slightly bent dorsally.
</paragraph>
<paragraph id="BF5536D4EA731235FF52BCAFD55301CD" blockId="4.[151,1437,782,2031]" pageId="4" pageNumber="245">Female. Similar to male, but antennae slightly shorter and thinner, antennomeres IV‒VIII without any impression.</paragraph>
<caption id="EB95665CEA721234FF02BCA9D12401CB" httpUri="https://zenodo.org/record/1216382/files/figure.png" pageId="5" pageNumber="246" startId="5.[151,264,1972,1995]" targetBox="[238,1365,233,1932]" targetPageId="5">
<paragraph id="BF5536D4EA721234FF02BCA9D12401CB" blockId="5.[151,1436,1972,2025]" pageId="5" pageNumber="246">
<emphasis id="8D9EEAC6EA721234FF02BCA9D50C01E8" bold="true" box="[151,326,1972,1995]" pageId="5" pageNumber="246">FIGURES 7–15.</emphasis>
Aedeagus (7, 10, 13. ventral vieW; 8, 11, 14. dorsal vieW; 9, 12, 15. lateral vieW): 7‒9.
<taxonomicName id="78EA4D57EA721234FB25BCA9D5D301CB" authority="Wittmer, 1983" authorityName="Wittmer" authorityYear="1983" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="quadratus" subGenus="Haplothemus">
<emphasis id="8D9EEAC6EA721234FB25BCA9D0B501E8" box="[1200,1279,1972,1994]" italics="true" pageId="5" pageNumber="246">Themus</emphasis>
(
<emphasis id="8D9EEAC6EA721234FA99BCA9D1D901E8" box="[1292,1427,1972,1994]" italics="true" pageId="5" pageNumber="246">Haplothemus</emphasis>
)
<emphasis id="8D9EEAC6EA721234FF02BCCED4B401CB" box="[151,254,2003,2025]" italics="true" pageId="5" pageNumber="246">quadratus</emphasis>
Wittmer, 1983
</taxonomicName>
; 10‒12.
<taxonomicName id="78EA4D57EA721234FE7ABCC9D68501CB" authority="Yang &amp; Liu &amp; Yang, 2018" authorityName="Yang &amp; Liu &amp; Yang" authorityYear="2018" box="[495,719,2003,2025]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="quadratiformis" status="sp. nov." subGenus="Haplothemus">
<emphasis id="8D9EEAC6EA721234FE7ABCC9D5B101CB" box="[495,507,2004,2025]" italics="true" pageId="5" pageNumber="246">T</emphasis>
. (
<emphasis id="8D9EEAC6EA721234FD85BCC9D66801CB" box="[528,546,2004,2025]" italics="true" pageId="5" pageNumber="246">H</emphasis>
.)
<emphasis id="8D9EEAC6EA721234FDA2BCCED68501CB" box="[567,719,2003,2025]" italics="true" pageId="5" pageNumber="246">quadratiformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="96AD57BDEA721234FD40BCC9D76C01CB" box="[725,806,2004,2025]" pageId="5" pageNumber="246" rank="species">
<emphasis id="8D9EEAC6EA721234FD40BCC9D76C01CB" bold="true" box="[725,806,2004,2025]" pageId="5" pageNumber="246">sp. nov.</emphasis>
</taxonomicNameLabel>
; 13‒15.
<taxonomicName id="78EA4D57EA721234FC15BCC9D0D101CB" authority="Pic, 1955" authorityName="Pic" authorityYear="1955" box="[896,1179,2003,2025]" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="elongatior" subGenus="Haplothemus">
<emphasis id="8D9EEAC6EA721234FC15BCC9D7C601CB" box="[896,908,2004,2025]" italics="true" pageId="5" pageNumber="246">T</emphasis>
. (
<emphasis id="8D9EEAC6EA721234FC34BCC9D7F901CB" box="[929,947,2004,2025]" italics="true" pageId="5" pageNumber="246">H</emphasis>
.)
<emphasis id="8D9EEAC6EA721234FC5DBCCED07801CB" box="[968,1074,2003,2025]" italics="true" pageId="5" pageNumber="246">elongatior</emphasis>
Pic, 1955
</taxonomicName>
. Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="BF5536D4EA711237FF52BB8AD7CE06D5" blockId="6.[151,1436,151,608]" pageId="6" pageNumber="247">
Abdominal sternite VIII (
<figureCitation id="27D12A51EA711237FE7EBB8AD6740692" box="[491,574,151,176]" captionStart="FIGURES 30–34" captionStartId="10.[151,264,1783,1806]" captionTargetBox="[202,1380,377,1733]" captionTargetId="figure@10.[151,1436,301,1762]" captionTargetPageId="10" captionText="FIGURES 30–34. Abdominal sternite VIII of female, ventral vieW: 30. Themus (Haplothemus) quadratus Wittmer, 1983; 31. T. (H.) quadratiformis sp. nov.; 32. T. (H.) elongatior Pic, 1955; 33. T. (H.) weishanensis Kopetz, 2016; 34. T. (H.) longideverticulum sp. nov. Scale bars:1.0 mm." httpUri="https://zenodo.org/record/1216386/files/figure.png" pageId="6" pageNumber="247">Fig. 32</figureCitation>
) triangularly emarginated in middle and rectangularly emarginated on both sides of posterior margin, the lateral emargination nearly as deep as and slightly wider than the middle one, the portion between middle and lateral emarginations rounded at apex.
</paragraph>
<paragraph id="BF5536D4EA711237FF52BA19D481043A" blockId="6.[151,1436,151,608]" pageId="6" pageNumber="247">
Internal reproductive organ of genitalia (
<figureCitation id="27D12A51EA711237FD07BA19D6AD073E" box="[658,743,260,285]" captionStart="FIGURES25–29" captionStartId="8.[151,264,1912,1935]" captionTargetBox="[160,1412,274,1811]" captionTargetId="figure@8.[151,1436,241,1847]" captionTargetPageId="8" captionText="FIGURES25–29. Internalreproductive organ of female genitalia,lateral vieW:25.Themus (Haplothemus) quadratus Wittmer, 1983; 26. T. (H.) quadratiformis sp. nov.; 27. T. (H.) elongatior Pic, 1955; 28. T. (H.) weishanensis Kopetz, 2016; 29. T. (H.) longideverticulumsp.nov. Scalebars:1.0mm.(ag: accessorygland; di:diverticulum;sp: spermatheca; ov:medianoviduct; va: vagina; sr: sclerotized ring)" httpUri="https://zenodo.org/record/1216384/files/figure.png" pageId="6" pageNumber="247">Fig. 27</figureCitation>
): vagina stout, with diverticulum and spermatheca situated at apical one-third part, median oviduct at basal one-third part of ventral side; diverticulum slightly long tube-shaped, distinctly shorter than vagina, evenly thick in the whole length, about twice as long as wide, rounded at apex; spermatheca slightly long sac-shaped, distinctly expanded apically and rounded at apex, which is slightly longer than diverticulum, and provided with a long and thin accessory gland at basal one-third part; diverticulum and spermatheca surrounded with a strongly sclerotized ring at base, which is confluent in middle and extending to median oviduct on ventral side, the opening of spermatheca arising from middle of the sclerotized ring on dorsal side.
</paragraph>
</subSubSection>
<subSubSection id="F7F0655FEA711237FF52B939D10D0442" pageId="6" pageNumber="247" type="discussion">
<paragraph id="BF5536D4EA711237FF52B939D10D0442" blockId="6.[151,1436,151,608]" pageId="6" pageNumber="247">
<emphasis id="8D9EEAC6EA711237FF52B939D571041F" bold="true" box="[199,315,548,573]" pageId="6" pageNumber="247">Remarks.</emphasis>
The abdominal sternite VIII of female was illustrated by
<bibRefCitation id="DB7B4B25EA711237FC46B939D0D6041E" box="[979,1180,548,573]" pageId="6" pageNumber="247" refId="ref6223" refString="Okushima, Y. (1999) Cantharidae collected by the Hokkaido University Expeditions to Nepal Himalaya (Coleoptera). Insecta Matsumurana, 56, 51 - 68. [http: // hdl. handle. net / 2115 / 9897]" type="journal article">Okushima (1999)</bibRefCitation>
, but it seems different from the photo provided in the present study probably because of different preparation way of the material.
</paragraph>
</subSubSection>
</treatment>
</document>
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