treatments-xml/data/37/00/8F/37008F9045DF5615852D618C7DD10215.xml

<document id="4CC0D380EDC9179BE0192B40A6E386B7" ID-CLB-Dataset="298082" ID-DOI="10.3897/zookeys.1204.111842" ID-GBIF-Dataset="fb1e044e-9149-4837-8df1-c9ea3e5fdbc7" ID-ZooBank="41A41142-ED13-4322-8B86-3681C2FAE4F3" ID-publisher-id="111842" URI-arpha="39D1C612-7089-580C-B242-71FD3D97E4EA" URI-zoobank="http://zoobank.org/41A41142-ED13-4322-8B86-3681C2FAE4F3" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1717518108510" checkinUser="pensoft" docAuthor="Hou, Yanmeng &amp; Zhang, Feng" docDate="2024" docId="37008F9045DF5615852D618C7DD10215" docLanguage="en" docName="ZooKeys 1204: 135-154" docOrigin="ZooKeys 1204" docSource="https://zookeys.pensoft.net/article/111842/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Spelaeochthonius huanglaoensis Hou &amp; Zhang 2024, sp. nov." docType="treatment" docUuid="AFD5997B-116A-44E4-91B8-F682E488F620" docUuidSource="ZooBank" docVersion="5" dtd-version="3.0" lastPageNumber="154" masterDocId="39D1C6127089580CB24271FD3D97E4EA" masterDocTitle="Three new species of dragon pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from China" masterLastPageNumber="154" masterPageNumber="135" pageNumber="135" updateTime="1717536209949" updateUser="ExternalLinkService">
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<mods:title id="0CD7239BAB5EDFABCBF95257C932B63B">Three new species of dragon pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from China</mods:title>
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<mods:namePart id="1B0A20435923E6748DBDD39AD54043B3">Hou, Yanmeng</mods:namePart>
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<mods:affiliation id="8BA9A88C6EA0F800FBAF50567D0B68D3">College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China &amp; The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="ED4B866C0B4B865AE79E041483F52CF0">Zhang, Feng</mods:namePart>
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<mods:affiliation id="C1F5FC351D8553278FDCC71EE5AB861A">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<treatment id="37008F9045DF5615852D618C7DD10215" ID-DOI="http://doi.org/10.5281/zenodo.11476706" ID-GBIF-Taxon="232137082" ID-Zenodo-Dep="11476706" ID-arpha="37008F90-45DF-5615-852D-618C7DD10215" ID-zoobank="https://zoobank.org/AFD5997B-116A-44E4-91B8-F682E488F620" LSID="urn:lsid:zoobank.org:act:AFD5997B-116A-44E4-91B8-F682E488F620" LSID-ZBK="urn:lsid:zoobank.org:act:AFD5997B-116A-44E4-91B8-F682E488F620" httpUri="http://treatment.plazi.org/id/37008F9045DF5615852D618C7DD10215">
<subSubSection id="5FD87FA88C29BFFD36E175FA8B8DE8D6" type="nomenclature">
<paragraph id="90B01820FF3AC8A7A8F60CBB3828C789">
<taxonomicName id="9970AFF7E450C8B2800ED74A78D78FD8" authority="Hou &amp; Zhang, 2024" authorityName="Hou &amp; Zhang" authorityYear="2024" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="huanglaoensis" status="sp. nov.">
<emphasis id="6126424F2CE8127BE07CF24AE46A4900" italics="true">Spelaeochthonius huanglaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="78E963FB834C67FB4C12455C3FD2B171" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="227ABEF6FE6A31F5CBC07677834B2B29" type="description">
<paragraph id="770F4B23A5A815692358BB2F41C238AB">
<figureCitation id="288314466B919629EFED0040D1908001" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of all Pseudotyrannochthoniidae species in China. 1 Allochthonius liaoningensis Hu &amp; Zhang, 2012 2 A. exornatus Gao &amp; Zhang, 2013 3 A. wui Hu &amp; Zhang, 2011 4 A. jingyuanus Zhang &amp; Zhang, 2014 5 A. brevitus 6 A. sichuanensis (Schawaller, 1995) 7 A. hispidus sp. nov. 8 A. lini Li, 2023 9 A. xuae Li, 2023 10 A. bainiensis 11 A. pandus 12 A. xinqiaoensis 13 A. fanjingshan Gao, Zhang &amp; Zhang, 2016 14 A. trigonus Hu &amp; Zhang, 2011 15 A. fuscus Hu &amp; Zhang, 2011 16 Centrochthonius kozlovi (Redikorzev, 1918) 17 C. cheni (Gao, Zhang &amp; Zhang, 2016) 18 Spelaeochthonius huanglaoensis sp. nov. 19 S. tuoliangensis sp. nov. 20 S. yinae 21 S. wulibeiensis." figureDoi="10.3897/zookeys.1204.111842.figure1" httpUri="https://binary.pensoft.net/fig/1062043">Figs 1</figureCitation>
, 
<figureCitation id="73B946E20AA1FBF7A53C35FFB853C926" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Spelaeochthonius huanglaoensis sp. nov. A holotype male, habitus (dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1204.111842.figure5" httpUri="https://binary.pensoft.net/fig/1062047">5</figureCitation>
, 
<figureCitation id="06C9E08615CE4F4B0994EBCFE5F71C5B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Spelaeochthonius huanglaoensis sp. nov. A carapace (dorsal view) B left chelicera (dorsal view) C coxae (ventral view) D left chela (lateral view) E male genital area (ventral view) F female genital area (ventral view). Scale bars: 0.20 mm (C, D); 0.10 mm (A, B, E, F)." figureDoi="10.3897/zookeys.1204.111842.figure6" httpUri="https://binary.pensoft.net/fig/1062048">6</figureCitation>
, 
<figureCitation id="6570C09274E3D65DCA14944C16FBE0C9" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Spelaeochthonius huanglaoensis sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C rallum D left chela (lateral view), with details of trichobothrial pattern E left pedipalp (minus chela, dorsal view) F left chela (dorsal view) G leg I (lateral view) H leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1204.111842.figure7" httpUri="https://binary.pensoft.net/fig/1062049">7</figureCitation>
, 
<figureCitation id="2CED29BC2171223AEACC2B26B6C2A709" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Spelaeochthonius huanglaoensis sp. nov. scanning electron micrographs, paratype female A coxal spines in overview, with details of tips B serrula exterior C manducatory process D rallum. Scale bars: 50 μm (C, D); 20 μm (B); 10 μm (A)." figureDoi="10.3897/zookeys.1204.111842.figure8" httpUri="https://binary.pensoft.net/fig/1062050">8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1VAE" type="Chinese name">
<paragraph id="116131E06CF5A8DAAA11BB1E125528F6">
<heading id="3D0EC385D8010157D1186BD085EA2ED9" reason="title">Chinese name.</heading>
</paragraph>
<paragraph id="7661331BAB3EEC1BC57C7177506509E7">黄老穴伪蝎.</paragraph>
</subSubSection>
<subSubSection id="SECID0E6VAE" type="type material">
<paragraph id="52EF63FE8D93357502DF58BE57FA4585">
<heading id="46AEF3615FA0DF6D39830CFC033740D1" reason="title">Type material.</heading>
</paragraph>
<paragraph id="99D5E600AB856E3899F3CB69B65C5579">
<materialsCitation id="7C484BAC39ACEAF3073FB2879A3D9665" ID-GBIF-Occurrence="4873230308" collectedFrom="under a stone in the deep zone" collectingDate="2021-10-19" collectionCode="MHBU" collectorName="Nana Zhan" country="China" county="Fangshan District" elevation="495" latitude="39.681934" location="Huanglao Cave" longLatPrecision="1" longitude="115.65068" municipality="Shidu Town" specimenCode="BJFS-21-10-19-02-01" specimenCount="1" specimenCount-male="1" stateProvince="Beijing City" typeStatus="holotype">
<emphasis id="5A4B98185BFE77AAF0B5D477BC261968" bold="true" italics="true">
<typeStatus id="772E8C07D6DFE59F4D0FB73E7EAC99FC" type="holotype">Holotype</typeStatus>
</emphasis>
: 
<collectingCountry id="7E4F3557C51DDE42294381B81AB4AA10" name="China">China</collectingCountry>
• 
<specimenCount id="0ECBA060A6250972E5BB926860E68406" count="1" type="male">♂</specimenCount>
; 
<collectingRegion id="1554BFD684271FEA4843A15E1A3049AE">Beijing City</collectingRegion>
, 
<collectingCounty id="BFE6681DDAA67E5297EED3795BA6CBB7">Fangshan District</collectingCounty>
, 
<collectingMunicipality id="4D85A40517DBA86AC6651FD1516748AC">Shidu Town</collectingMunicipality>
, 
<collectingMunicipality id="4A208CAEE002DEF7C0DD051F4B3E8433">Wanglaopu Village</collectingMunicipality>
, 
<location id="4967537DC755EA9A6982994B9FBE68F9" LSID="urn:lsid:plazi:treatment:37008F9045DF5615852D618C7DD10215:4967537DC755EA9A6982994B9FBE68F9" country="China" county="Fangshan District" latitude="39.681934" longLatPrecision="1" longitude="115.65068" municipality="Shidu Town" name="Huanglao Cave" stateProvince="Beijing City">Huanglao Cave</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENWAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[115.650683,39.681933]}">
<geoCoordinate id="1DEEDE7455D8D8387A4F03CD5E2246FB" degrees="39" direction="north" minutes="40.916" orientation="latitude" precision="1" value="39.681934">39 ° 40.916 ′ N</geoCoordinate>
, 
<geoCoordinate id="8C4C2F5677EDDF3E3F7A38BE9BEDB8D1" degrees="115" direction="east" minutes="39.041" orientation="longitude" precision="1" value="115.65068">115 ° 39.041 ′ E</geoCoordinate>
</named-content>
; 
<elevation id="BB1A82317131389FE44144EF50785CC4" metricMagnitude="2" metricUnit="m" metricValue="4.95" unit="m" value="495.0">
<quantity id="9CB2EA9FF6AD92DFB205499CAAC1C5AF" metricMagnitude="2" metricUnit="m" metricValue="4.95" unit="m" value="495.0">495 m</quantity>
a. s. l.
</elevation>
; 
<date id="C7289D6EDD3542903C2A24EC48FB0F7E" value="2021-10-19">
<collectingDate id="9045458F155B0FF37C016C286CF25927" value="2021-10-19">19 Oct. 2021</collectingDate>
</date>
; 
<collectorName id="4C9CEEE0857C02D74569E83CDE07C844">Nana Zhan</collectorName>
leg.; 
<collectedFrom id="5051EB0CDCA1A692AA6DE72272681CA6">under a stone in the deep zone</collectedFrom>
(Fig. 
<figureCitation id="0D434A831CE0565B63E49A524D37A93F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of all Pseudotyrannochthoniidae species in China. 1 Allochthonius liaoningensis Hu &amp; Zhang, 2012 2 A. exornatus Gao &amp; Zhang, 2013 3 A. wui Hu &amp; Zhang, 2011 4 A. jingyuanus Zhang &amp; Zhang, 2014 5 A. brevitus 6 A. sichuanensis (Schawaller, 1995) 7 A. hispidus sp. nov. 8 A. lini Li, 2023 9 A. xuae Li, 2023 10 A. bainiensis 11 A. pandus 12 A. xinqiaoensis 13 A. fanjingshan Gao, Zhang &amp; Zhang, 2016 14 A. trigonus Hu &amp; Zhang, 2011 15 A. fuscus Hu &amp; Zhang, 2011 16 Centrochthonius kozlovi (Redikorzev, 1918) 17 C. cheni (Gao, Zhang &amp; Zhang, 2016) 18 Spelaeochthonius huanglaoensis sp. nov. 19 S. tuoliangensis sp. nov. 20 S. yinae 21 S. wulibeiensis." figureDoi="10.3897/zookeys.1204.111842.figure1" httpUri="https://binary.pensoft.net/fig/1062043">1</figureCitation>
); Ps. - 
<abbrev id="ABBRID0EWWAE" xlink_title="Museum of Hebei University">
<collectionCode id="6B0E987677F04689B0247A8889FAB82C">MHBU</collectionCode>
</abbrev>
- 
<specimenCode id="637ACA5F29A67BA13D78115E6B9971E5">BJFS-21-10-19-02-01</specimenCode>
</materialsCitation>
. 
<materialsCitation id="A8F797C936200DF17C2D00F9D5EA89DA" ID-GBIF-Occurrence="4873230310" collectedFrom="under a stone in the deep zone" collectingDate="2021-10-19" collectionCode="MHBU" collectorName="Nana Zhan &amp; Ps." country="China" county="City" elevation="495" latitude="39.681934" location="Shidu Town" longLatPrecision="1" longitude="115.65068" municipality="Fangshan District" specimenCode="BJFS-21-10-19-02-02" specimenCount="1" specimenCount-female="1" stateProvince="Beijing" typeStatus="paratype">
<emphasis id="5D97A21DB4769BCF6759757E1C52B1C8" bold="true" italics="true">
<typeStatus id="7D06E35F6B4E519EFF77B52BA33D8981" type="paratype">Paratype</typeStatus>
</emphasis>
: • 
<specimenCount id="48C68F60E8683F17B66D91F8A406A336" count="1" type="female">1 ♀</specimenCount>
; same data as for holotype; Ps. - 
<abbrev id="ABBRID0E4WAE" xlink_title="Museum of Hebei University">
<collectionCode id="9CE29A060FFD9B8FC6BE222F0FC21662">MHBU</collectionCode>
</abbrev>
- 
<specimenCode id="30930CD9326C625550124B00A4B32938">BJFS-21-10-19-02-02</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBXAE" type="diagnosis">
<paragraph id="3799B2BF1B36C6D9257E070B5700BD85">
<heading id="6BD317CD5B3723D4CF0215BB8D277E94" reason="title">Diagnosis</heading>
</paragraph>
<paragraph id="A62E1685EEFDEAF93E70558854798EC3">
<emphasis id="70ADD843D39603EAF856993FD6BB5751" bold="true">
(
<specimenCount id="D0264A37CFEDB6BF31A32D50387ECCA4" count="1" type="male">♂</specimenCount>
<specimenCount id="F031B6D0AAF49A82A9A5E503073AB915" count="1" type="female">♀</specimenCount>
).
</emphasis>
<taxonomicName id="D48A1EA9107B130B06C7344E913D9802" authority="Hou &amp; Zhang, 2024" authorityName="Hou &amp; Zhang" authorityYear="2024" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="huanglaoensis" status="sp. nov.">
<emphasis id="944AD9351548FDC73486B0B5399BE16B" italics="true">Spelaeochthonius huanglaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1C3294E9FDF46A8494CD4DDD85B1E52" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to 
<taxonomicName id="F6864268F31F8E027E662E933B80308D" authority="Gao, Hou &amp; Zhang, 2023" authorityName="Gao, Hou &amp; Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="wulibeiensis">
<emphasis id="214F81D062A4202557F88272127A4C42" italics="true">S. wulibeiensis</emphasis>
<bibRefCitation id="3CB1AF4C71C70102E99CACF52EB080F0" DOI="10.3897/zookeys.1139.96639" author="Gao ZZ &amp; Hou YM &amp; Zhang F" journalOrPublisher="ZooKeys" pagination="33-69" refId="B6" refString="Gao ZZ, Hou YM, Zhang F (2023) Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33 - 69. https: // doi. org / 10.3897 / zookeys. 1139.96639" title="Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China." volume="1139" year="2023">Gao, Hou &amp; Zhang, 2023</bibRefCitation>
</taxonomicName>
, but differs from it in having shorter pedipalps (e. g. chela 7.94 (
<specimenCount id="932404F4142F5C80AA1FAC0C4A545917" count="1" type="male">♂</specimenCount>
), 6.14 (
<specimenCount id="C98F16BC748011F3566955BB15F55601" count="1" type="female">♀</specimenCount>
) × vs 6.21–6.22 (
<specimenCount id="048E1AC08E24E668C71A964E7228129C" count="1" type="male">♂</specimenCount>
), 5.68 (
<specimenCount id="6F185BD6EA128C2FBC259943E24C84FC" count="1" type="female">♀</specimenCount>
) × longer than broad, length 1.43 (
<specimenCount id="3AC32F10391AE8CE6D8F263F0DBB0599" count="1" type="male">♂</specimenCount>
), 1.72 (
<specimenCount id="6273E98DE402F5D76A0358F19C5003F1" count="1" type="female">♀</specimenCount>
) mm vs 1.68–1.74 (
<specimenCount id="413C0FA7028082596E8B1F1FF04502B0" count="1" type="male">♂</specimenCount>
), 1.76 (
<specimenCount id="A5891E8AD28FE1CBD00BF9F7312CEEE1" count="1" type="female">♀</specimenCount>
) mm), 1 additional cheliceral seta (seven vs six), and more numerous fixed chelal finger teeth (29 vs 22–24). It differs from 
<taxonomicName id="BCADCAC2A972B493EE8CB510AD9F729C" authority="Li, 2023" authorityName="Li" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="yinae">
<emphasis id="CAD3BA6EC7337E3CB245D4C7EFDC94FC" italics="true">S. yinae</emphasis>
Li, 
<quantity id="4625A9D31B240B33210FB01B006F6E2B" metricMagnitude="1" metricUnit="m" metricValue="5.13842" unit="in" value="2023.0">2023 in</quantity>
</taxonomicName>
the number of setae on tergite II (four vs two), smaller body size (e. g. chela 7.94 (
<specimenCount id="75E363BC2AA68A3072D20745D8FEEFE6" count="1" type="male">♂</specimenCount>
), 6.14 (
<specimenCount id="85939F6C4CF57D003A10F16C10B492BE" count="1" type="female">♀</specimenCount>
) × vs 5.93 (
<specimenCount id="253918607E175441924107B5621F3DA1" count="1" type="male">♂</specimenCount>
), 6.30 (
<specimenCount id="B03B45D99D62A81B7DF209F4A6F4E7CA" count="1" type="female">♀</specimenCount>
) × longer than broad, length 1.43 (
<specimenCount id="D37445AA81D04601E06EB6BBA806EA76" count="1" type="male">♂</specimenCount>
), 1.72 (
<specimenCount id="AE72EE807C3872F083344D9429A87CB0" count="1" type="female">♀</specimenCount>
) mm vs 1.72 (
<specimenCount id="157CC143F5CBB4DD6FC99834D9FAA0E2" count="1" type="male">♂</specimenCount>
), 1.89 (
<specimenCount id="E8119E7D1BE3F6C7F74392EFF1EC8F0C" count="1" type="female">♀</specimenCount>
) mm), and more numerous fixed chelal finger teeth (
<specimenCount id="B7B2217EFA82EDE5C1E1D93518F8F95E" count="1" type="male">♂</specimenCount>
) (29 vs 23).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKYAE" type="etymology">
<paragraph id="78AC4B513A90708F3C89DE3697DB23E3">
<heading id="90FA595F6858EE4A7589D8CDB1F5EEF3" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="763C7F4A889DE8226064C101F901D8D4">The species is named after its type locality, Huanglao Cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0EPYAE" type="description">
<paragraph id="1221BA53EBD7C65D3594A059BB9B783D">
<heading id="33FA498249F7F7A667314DBB1638349C" reason="title">Description.</heading>
</paragraph>
<paragraph id="6D7EC8B5CAFE1E82FD81AA015641A8BD">
<emphasis id="44CDA41546CDA61D66F0AF6816F17870" bold="true">Adult male</emphasis>
(Figs 
<figureCitation id="F43D7C189F31D32FA3D3A623F9BB1CD0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Spelaeochthonius huanglaoensis sp. nov. A holotype male, habitus (dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1204.111842.figure5" httpUri="https://binary.pensoft.net/fig/1062047">5 A</figureCitation>
, 
<figureCitation id="20BDB1E95DE18E51CB03F0F9D8C539B8" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Spelaeochthonius huanglaoensis sp. nov. A carapace (dorsal view) B left chelicera (dorsal view) C coxae (ventral view) D left chela (lateral view) E male genital area (ventral view) F female genital area (ventral view). Scale bars: 0.20 mm (C, D); 0.10 mm (A, B, E, F)." figureDoi="10.3897/zookeys.1204.111842.figure6" httpUri="https://binary.pensoft.net/fig/1062048">6 A – E</figureCitation>
, 
<figureCitation id="56E59699F3B223C99C74164AEB2527A8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Spelaeochthonius huanglaoensis sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C rallum D left chela (lateral view), with details of trichobothrial pattern E left pedipalp (minus chela, dorsal view) F left chela (dorsal view) G leg I (lateral view) H leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1204.111842.figure7" httpUri="https://binary.pensoft.net/fig/1062049">7</figureCitation>
). 
<emphasis id="46813AF72ADE7053BBF565C9316A9432" bold="true" italics="true">Colour</emphasis>
: generally pale yellow; chelicerae, pedipalps and tergites slightly darker; soft parts pale. 
<emphasis id="F9A4FB0DE13A943B25B27F90D8427D57" bold="true" italics="true">Cephalothorax</emphasis>
(Figs 
<figureCitation id="9D12E8AE52C67E2F51EF275CB4607EA7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Spelaeochthonius huanglaoensis sp. nov. A carapace (dorsal view) B left chelicera (dorsal view) C coxae (ventral view) D left chela (lateral view) E male genital area (ventral view) F female genital area (ventral view). Scale bars: 0.20 mm (C, D); 0.10 mm (A, B, E, F)." figureDoi="10.3897/zookeys.1204.111842.figure6" httpUri="https://binary.pensoft.net/fig/1062048">6 A, C</figureCitation>
, 
<figureCitation id="95B613CEB93174936F35C17075D55BD6" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Spelaeochthonius huanglaoensis sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C rallum D left chela (lateral view), with details of trichobothrial pattern E left pedipalp (minus chela, dorsal view) F left chela (dorsal view) G leg I (lateral view) H leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1204.111842.figure7" httpUri="https://binary.pensoft.net/fig/1062049">7 A</figureCitation>
): carapace inverted-trapezoid, 1.04 × longer than broad, gently narrowed posteriorly; surface mostly with fine reticulations; with four anterior lyrifissures and two posterior lyrifissures; no traces of eyes but eye region bulging and convex in dorsal view; epistome present and with some tiny spinules; with 16 setae arranged s 4 s: 4: 2: 2: 2, most setae heavy, long, and gently curved. Chaetotaxy of coxae: P 3, I 6, II 5, III 4–5, IV 4; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta (refer to female, Fig. 
<figureCitation id="A4D565EA72CEF7FCC6B6D066135E7CEE" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Spelaeochthonius huanglaoensis sp. nov. scanning electron micrographs, paratype female A coxal spines in overview, with details of tips B serrula exterior C manducatory process D rallum. Scale bars: 50 μm (C, D); 20 μm (B); 10 μm (A)." figureDoi="10.3897/zookeys.1204.111842.figure8" httpUri="https://binary.pensoft.net/fig/1062050">8 C</figureCitation>
); coxal spines present on coxa I only, comprising a transverse, contiguous series of six or seven tridentate blades, which arise from a lightly sclerotized or translucent hillock, the central ramus of each blade (except the basal one) sharply acumino-spatulate and extending beyond the lateral rami (refer to female, Fig. 
<figureCitation id="BF4E3E2C076DE32F14B39CF48FDDF3A2" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Spelaeochthonius huanglaoensis sp. nov. scanning electron micrographs, paratype female A coxal spines in overview, with details of tips B serrula exterior C manducatory process D rallum. Scale bars: 50 μm (C, D); 20 μm (B); 10 μm (A)." figureDoi="10.3897/zookeys.1204.111842.figure8" httpUri="https://binary.pensoft.net/fig/1062050">8 A</figureCitation>
); a small, bisetose intercoxal tubercle present between coxae III and IV (Fig. 
<figureCitation id="DC7F50964913E9388C6BF9599C885689" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Spelaeochthonius huanglaoensis sp. nov. A carapace (dorsal view) B left chelicera (dorsal view) C coxae (ventral view) D left chela (lateral view) E male genital area (ventral view) F female genital area (ventral view). Scale bars: 0.20 mm (C, D); 0.10 mm (A, B, E, F)." figureDoi="10.3897/zookeys.1204.111842.figure6" httpUri="https://binary.pensoft.net/fig/1062048">6 C</figureCitation>
). 
<emphasis id="614D598B56191EFBE3BBDBE886872761" bold="true" italics="true">Chelicera</emphasis>
(Figs 
<figureCitation id="165C0483E94FD107C73D7DDA4DFBEE51" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Spelaeochthonius huanglaoensis sp. nov. A carapace (dorsal view) B left chelicera (dorsal view) C coxae (ventral view) D left chela (lateral view) E male genital area (ventral view) F female genital area (ventral view). Scale bars: 0.20 mm (C, D); 0.10 mm (A, B, E, F)." figureDoi="10.3897/zookeys.1204.111842.figure6" httpUri="https://binary.pensoft.net/fig/1062048">6 B</figureCitation>
, 
<figureCitation id="6D842577814517600B8F0F3221CE6107" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Spelaeochthonius huanglaoensis sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C rallum D left chela (lateral view), with details of trichobothrial pattern E left pedipalp (minus chela, dorsal view) F left chela (dorsal view) G leg I (lateral view) H leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1204.111842.figure7" httpUri="https://binary.pensoft.net/fig/1062049">7 B, C</figureCitation>
): large, approximately as long as carapace, 2.50 × longer than broad; six setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, movable finger with one medial seta, all setae acuminate, ventrobasal seta shorter than others. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers with well-developed teeth, fixed finger with 14 acute teeth, distal one largest; movable finger with 11 retrorse contiguous teeth of equal length; galea absent. Serrula exterior with 19 blades (refer to female, Fig. 
<figureCitation id="B013CE93FF6763086CF55FFEB3B2BC95" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Spelaeochthonius huanglaoensis sp. nov. scanning electron micrographs, paratype female A coxal spines in overview, with details of tips B serrula exterior C manducatory process D rallum. Scale bars: 50 μm (C, D); 20 μm (B); 10 μm (A)." figureDoi="10.3897/zookeys.1204.111842.figure8" httpUri="https://binary.pensoft.net/fig/1062050">8 B</figureCitation>
) and serrula interior with 15 blades. Rallum in two rows and composed of ten finely pinnate blades (11 blades in female), of which the basal-most blade shorter than the others (Figs 
<figureCitation id="2CD5F68F36C2A9A53F37DD00856141D8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Spelaeochthonius huanglaoensis sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C rallum D left chela (lateral view), with details of trichobothrial pattern E left pedipalp (minus chela, dorsal view) F left chela (dorsal view) G leg I (lateral view) H leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1204.111842.figure7" httpUri="https://binary.pensoft.net/fig/1062049">7 C</figureCitation>
, 
<figureCitation id="9EF524DE62CE94D07821D9B3EB28A85B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Spelaeochthonius huanglaoensis sp. nov. scanning electron micrographs, paratype female A coxal spines in overview, with details of tips B serrula exterior C manducatory process D rallum. Scale bars: 50 μm (C, D); 20 μm (B); 10 μm (A)." figureDoi="10.3897/zookeys.1204.111842.figure8" httpUri="https://binary.pensoft.net/fig/1062050">8 D</figureCitation>
). 
<emphasis id="322CE341BBCFD78F4E67134C56617D0C" bold="true" italics="true">Pedipalp</emphasis>
(Figs 
<figureCitation id="25CA2C1FA289A4220D5EA6D05027C251" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Spelaeochthonius huanglaoensis sp. nov. A carapace (dorsal view) B left chelicera (dorsal view) C coxae (ventral view) D left chela (lateral view) E male genital area (ventral view) F female genital area (ventral view). Scale bars: 0.20 mm (C, D); 0.10 mm (A, B, E, F)." figureDoi="10.3897/zookeys.1204.111842.figure6" httpUri="https://binary.pensoft.net/fig/1062048">6 D</figureCitation>
, 
<figureCitation id="0F7803BC477430DD4C0191C1B1C7A593" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Spelaeochthonius huanglaoensis sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C rallum D left chela (lateral view), with details of trichobothrial pattern E left pedipalp (minus chela, dorsal view) F left chela (dorsal view) G leg I (lateral view) H leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1204.111842.figure7" httpUri="https://binary.pensoft.net/fig/1062049">7 D – F</figureCitation>
): surfaces mostly with fine reticulations; long and slender, trochanter 1.87, femur 6.38, patella 2.69, chela 7.94, hand 3.00 × longer than broad; femur 2.37 × longer than patella; movable chelal finger 1.69 × longer than hand and 0.64 × longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (Fig. 
<figureCitation id="EF404A18CDE0C7796101449DF5D64410" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Spelaeochthonius huanglaoensis sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C rallum D left chela (lateral view), with details of trichobothrial pattern E left pedipalp (minus chela, dorsal view) F left chela (dorsal view) G leg I (lateral view) H leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1204.111842.figure7" httpUri="https://binary.pensoft.net/fig/1062049">7 E</figureCitation>
). Chelal palm slightly constricted towards fingers. Fixed chelal finger and hand with eight trichobothria plus duplex trichobothrium (
<emphasis id="ED41CA52483137445D4376C1AE53B98B" italics="true">dt</emphasis>
), movable chelal finger with four trichobothria, 
<emphasis id="72F909566DEE1A642095260A2D9027D3" italics="true">ib</emphasis>
, 
<emphasis id="017DC3EB8454D89FC48790E4146DA43D" italics="true">isb</emphasis>
, 
<emphasis id="A19628654DB5AAB27540ED48420A52BB" italics="true">eb</emphasis>
, 
<emphasis id="0AE2ABCEE18CA9A0A5DFEEFBDEF8F1E9" italics="true">esb</emphasis>
, and 
<emphasis id="3453BCB7D5294332C3D045957E015E1B" italics="true">ist</emphasis>
clustered at the base of fixed finger, 
<emphasis id="066134E48DAF7B883291C8A777AF21C3" italics="true">esb</emphasis>
slightly distal to 
<emphasis id="C618D7ABDD9ABDE04080BFE2E79E996D" italics="true">ist</emphasis>
; 
<emphasis id="E6E289AD04D5AF5013AEA75F6E4E0D84" italics="true">it</emphasis>
slightly distal to 
<emphasis id="1765F37331E10AC64E4FE89B08E352D5" italics="true">est</emphasis>
, situated subdistally and forming a pair; 
<emphasis id="ABFEF75C859B0C81AA8B6A14AFEB271E" italics="true">et</emphasis>
situated subdistally, very close to chelal teeth; 
<emphasis id="063A4B954D32A7DD4145BA9F35D64DFF" italics="true">dt</emphasis>
situated distal to 
<emphasis id="6885FDEC24A9846FB13E89FF4D638800" italics="true">et</emphasis>
, near the tip of fixed finger; 
<emphasis id="7E5AA869AC60816A16618435A687EAEF" italics="true">sb</emphasis>
distinctly closer to 
<emphasis id="F4A0B2F89A40EAA37A8D6B94BC3FDD18" italics="true">b</emphasis>
than to 
<emphasis id="AD868592B1DEA88C7B042A973E70E940" italics="true">st</emphasis>
(Fig. 
<figureCitation id="7363E51705AAD80DB6B9B916F053FE8F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Spelaeochthonius huanglaoensis sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C rallum D left chela (lateral view), with details of trichobothrial pattern E left pedipalp (minus chela, dorsal view) F left chela (dorsal view) G leg I (lateral view) H leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1204.111842.figure7" httpUri="https://binary.pensoft.net/fig/1062049">7 D</figureCitation>
). Microsetae (chemosensory setae) absent on hand and both palpal fingers. Sensilla absent. Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 29 teeth, slightly retrorse and pointed; movable chelal finger with 19 teeth (slightly smaller than teeth on fixed chelal finger) (Figs 
<figureCitation id="8212B8E4AEA6447A4A9FCCCCE8901BDD" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Spelaeochthonius huanglaoensis sp. nov. A carapace (dorsal view) B left chelicera (dorsal view) C coxae (ventral view) D left chela (lateral view) E male genital area (ventral view) F female genital area (ventral view). Scale bars: 0.20 mm (C, D); 0.10 mm (A, B, E, F)." figureDoi="10.3897/zookeys.1204.111842.figure6" httpUri="https://binary.pensoft.net/fig/1062048">6 D</figureCitation>
, 
<figureCitation id="D44E4989877D43BD1110526D30E4A94A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Spelaeochthonius huanglaoensis sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C rallum D left chela (lateral view), with details of trichobothrial pattern E left pedipalp (minus chela, dorsal view) F left chela (dorsal view) G leg I (lateral view) H leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1204.111842.figure7" httpUri="https://binary.pensoft.net/fig/1062049">7 D</figureCitation>
). Chelal fingers slightly curved in dorsal view (Fig. 
<figureCitation id="8AE684A6C893C78D436FB8004FAB8D15" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Spelaeochthonius huanglaoensis sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C rallum D left chela (lateral view), with details of trichobothrial pattern E left pedipalp (minus chela, dorsal view) F left chela (dorsal view) G leg I (lateral view) H leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1204.111842.figure7" httpUri="https://binary.pensoft.net/fig/1062049">7 F</figureCitation>
). 
<emphasis id="189040611D1592D7BB90B031A5BD9FD7" bold="true" italics="true">Opisthosoma</emphasis>
: generally typical, ovate, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 2: 4: 4: 5: 7: 7: 7: 6: 5: 4: TT: 0. Sternal chaetotaxy III – XII: 9: 8: 10: 9: 10: 9: 7: 8: 0: 2. Anterior genital operculum with nine setae, genital opening pit-like, with seven marginal setae on each side, 
<quantity id="0DB3946C54CB8065FD2ADA70153763BF" metricMagnitude="-1" metricUnit="m" metricValue="5.842" unit="in" value="23.0">23 in</quantity>
total (Fig. 
<figureCitation id="361B61CC72B98E4F0A2C2466F339456B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Spelaeochthonius huanglaoensis sp. nov. A carapace (dorsal view) B left chelicera (dorsal view) C coxae (ventral view) D left chela (lateral view) E male genital area (ventral view) F female genital area (ventral view). Scale bars: 0.20 mm (C, D); 0.10 mm (A, B, E, F)." figureDoi="10.3897/zookeys.1204.111842.figure6" httpUri="https://binary.pensoft.net/fig/1062048">6 E</figureCitation>
). 
<emphasis id="A175487951E751EA877A9066B9A81932" bold="true" italics="true">Legs</emphasis>
(Fig. 
<figureCitation id="4E4DAFED7E2FDBE5E45DF9CD7FDB50C3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Spelaeochthonius huanglaoensis sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C rallum D left chela (lateral view), with details of trichobothrial pattern E left pedipalp (minus chela, dorsal view) F left chela (dorsal view) G leg I (lateral view) H leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1204.111842.figure7" httpUri="https://binary.pensoft.net/fig/1062049">7 G, H</figureCitation>
): generally typical, long, and slender. Fine granulation present on anterodorsal faces of patella IV. Femur of leg I 1.73 × longer than patella and with one lyrifissure at the base of femur; tarsus 2.24 × longer than tibia. Femoropatella of leg IV 3.04 × longer than deep and with one lyrifissure at the base of femur; tibia 5.80 × longer than deep; with a long tactile seta on both tarsal segments: basitarsus 3.86 × longer than deep (TS = 0.37), telotarsus 11.40 × longer than deep and 2.11 × longer than basitarsus (TS = 0.35). Arolium slightly shorter than the claws, not divided; claws simple. 
<emphasis id="E30DF6916CCAD5DA3BF2CFED66B62224" bold="true" italics="true">Dimensions of adult male</emphasis>
(length / breadth or, in the case of the legs, length / depth in mm). Body length 1.80. Pedipalps: trochanter 0.28 / 0.15, femur 1.02 / 0.16, patella 0.43 / 0.16, chela 1.43 / 0.18, hand 0.54 / 0.18, movable finger length 0.91. Chelicera 0.55 / 0.22, movable finger length 0.28. Carapace 0.53 / 0.51. Leg I: trochanter 0.21 / 0.15, femur 0.52 / 0.09, patella 0.30 / 0.08, tibia 0.25 / 0.06, tarsus 0.56 / 0.05. Leg IV: trochanter 0.27 / 0.15, femoropatella 0.70 / 0.23, tibia 0.58 / 0.10, basitarsus 0.27 / 0.07, telotarsus 0.57 / 0.05.
</paragraph>
<caption id="130DC3971A104C170A52868450E3DE8A" ID-DOI="10.3897/zookeys.1204.111842.figure5" ID-arpha="6CDB10BE-062B-5825-9B80-FBB3E3D90F41" httpUri="https://binary.pensoft.net/fig/1062047" startId="F5">
<paragraph id="F0970D06647E64202AFD567C4380E1FD">
<label id="CF4E23982DAE67CAC9D330C5EBE73063">Figure 5.</label>
</paragraph>
<paragraph id="C9638A2E00D085F654C2D034655ADF5C">
<taxonomicName id="5F4924402FC49A2E2008AAAF385D6C76" authority="Hou &amp; Zhang, 2024" authorityName="Hou &amp; Zhang" authorityYear="2024" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="huanglaoensis" status="sp. nov.">
<emphasis id="23E69A58A307D68B929DFA2A4ABA6F3A" italics="true">Spelaeochthonius huanglaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AB951A381FB816B642EC1E27EC2A90C3" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="5069D7CEA29BB822BD2C68DD4C6AEE06" bold="true">A</emphasis>
holotype male, habitus (dorsal view) 
<emphasis id="F1984347554DF18456B2281E879895BD" bold="true">B</emphasis>
paratype female, habitus (dorsal view). Scale bars: 0.50 mm.
</paragraph>
</caption>
<caption id="80EC3E409CEBC2EC714D07BBA1661FAA" ID-DOI="10.3897/zookeys.1204.111842.figure6" ID-arpha="467A76CF-C0A2-5C9D-8AE2-564F5E9B7CB0" httpUri="https://binary.pensoft.net/fig/1062048" startId="F6">
<paragraph id="1164DA7A625A2C0A43EBE3780BD62CBF">
<label id="16EF99D89DC200AFC84493D4F07BF258">Figure 6.</label>
</paragraph>
<paragraph id="7C2F9BA2893C5D03684F1800B0C46AF8">
<taxonomicName id="A7F9F95D58F631D501A523DA9056A997" authority="Hou &amp; Zhang, 2024" authorityName="Hou &amp; Zhang" authorityYear="2024" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="huanglaoensis" status="sp. nov.">
<emphasis id="E55DA547D34380BF70BE32CB168133CE" italics="true">Spelaeochthonius huanglaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7CFB599CC89AA651F1425B0D3C4B5630" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="E9E01B1E8A959A0D5995FAF02A377DAD" bold="true">A</emphasis>
carapace (dorsal view) 
<emphasis id="21C621A994312AFBC4F307FABA375CD0" bold="true">B</emphasis>
left chelicera (dorsal view) 
<emphasis id="0097D3C6AB2C61F2EE54510EF2AFC06C" bold="true">C</emphasis>
coxae (ventral view) 
<emphasis id="DEF065FB554EC57D9FCBB0A69CF8E2EA" bold="true">D</emphasis>
left chela (lateral view) 
<emphasis id="65C0ED00A45AF7E0A1C039772930EDFA" bold="true">E</emphasis>
male genital area (ventral view) 
<emphasis id="00668C54424A50DA025F0C7396A5D4C6" bold="true">F</emphasis>
female genital area (ventral view). Scale bars: 0.20 mm (
<emphasis id="F9E3EC69914CF7BCE915B5DD4B20044F" bold="true">C, D</emphasis>
); 0.10 mm (
<emphasis id="9B54E93CB938C59AD26D8553D74E012F" bold="true">A, B, E, F</emphasis>
).
</paragraph>
</caption>
<caption id="045609DE2007EC14F4078E3077B661B5" ID-DOI="10.3897/zookeys.1204.111842.figure7" ID-arpha="B0AC2C9D-EE10-505F-9E24-484F98FC5E8A" httpUri="https://binary.pensoft.net/fig/1062049" startId="F7">
<paragraph id="B65DD8B28A04C97CD23DFAA3A1F21504">
<label id="862D444B7CDD0C1225A7707700585EE1">Figure 7.</label>
</paragraph>
<paragraph id="FD4EAD6586C62390BC4A9C1C827BDE55">
<taxonomicName id="D3CC87BA98D729BA9CBF08A4D601E4F3" authority="Hou &amp; Zhang, 2024" authorityName="Hou &amp; Zhang" authorityYear="2024" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="huanglaoensis" status="sp. nov.">
<emphasis id="DCE785DDB108C4B8A179A512669A87D5" italics="true">Spelaeochthonius huanglaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="88F4DD7483DF73DF36E5A0A5BFBF3B19" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male 
<emphasis id="C98725A8B64A9B9A520930B2246D5210" bold="true">A</emphasis>
carapace (dorsal view) 
<emphasis id="055B4FC70E927519488542B1BFD25A00" bold="true">B</emphasis>
left chelicera (dorsal view), with details of teeth 
<emphasis id="F7DCF9192853E244D838944B8141C9CF" bold="true">C</emphasis>
rallum 
<emphasis id="08FD8E367C232033B152CDDACEA4E338" bold="true">D</emphasis>
left chela (lateral view), with details of trichobothrial pattern 
<emphasis id="6B10AC2ED11DF58FC6F489FB79693185" bold="true">E</emphasis>
left pedipalp (minus chela, dorsal view) 
<emphasis id="C48F038440A6B415F8D951AFE8CF8397" bold="true">F</emphasis>
left chela (dorsal view) 
<emphasis id="DF17BB8F2BC47D115390BAD56049B2DD" bold="true">G</emphasis>
leg I (lateral view) 
<emphasis id="5D04F354B424BB307CC122712E168E0F" bold="true">H</emphasis>
leg IV (lateral view). Scale bars: 0.20 mm.
</paragraph>
</caption>
<paragraph id="9D6842F4FCA63288A2AEA8A6FBE0264A">
<emphasis id="F00AC48C4C70595F9CBE7E1E00A8690E" bold="true">Adult female</emphasis>
(Figs 
<figureCitation id="307DC3DF7D2CC4B6A96CEF8E940BC6B1" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Spelaeochthonius huanglaoensis sp. nov. A holotype male, habitus (dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1204.111842.figure5" httpUri="https://binary.pensoft.net/fig/1062047">5 B</figureCitation>
, 
<figureCitation id="9AB29FA9C633E8E4E5D56C7249DD02F6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Spelaeochthonius huanglaoensis sp. nov. A carapace (dorsal view) B left chelicera (dorsal view) C coxae (ventral view) D left chela (lateral view) E male genital area (ventral view) F female genital area (ventral view). Scale bars: 0.20 mm (C, D); 0.10 mm (A, B, E, F)." figureDoi="10.3897/zookeys.1204.111842.figure6" httpUri="https://binary.pensoft.net/fig/1062048">6 F</figureCitation>
, 
<figureCitation id="79E7FF58E2A7EF91DD2EAE62776DDA94" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Spelaeochthonius huanglaoensis sp. nov. scanning electron micrographs, paratype female A coxal spines in overview, with details of tips B serrula exterior C manducatory process D rallum. Scale bars: 50 μm (C, D); 20 μm (B); 10 μm (A)." figureDoi="10.3897/zookeys.1204.111842.figure8" httpUri="https://binary.pensoft.net/fig/1062050">8</figureCitation>
). Mostly same as male; tergal chaetotaxy I – XII: 2: 4: 4: 5: 6: 6: 6: 6: 5: 4: TT: 0; sternal chaetotaxy IV – XII: 5: 6: 8: 8: 9: 9: 8: 0: 2; anterior genital operculum with five setae, posterior margin with six marginal setae, 
<quantity id="3665CCD0EBA7E56BD25B072ABB865F63" metricMagnitude="-1" metricUnit="m" metricValue="2.794" unit="in" value="11.0">11 in</quantity>
total; leg IV with a long tactile seta on both tarsal segments: basitarsus 3.44 × longer than deep (TS = 0.35), telotarsus 9.86 × longer than deep and 2.23 × longer than basitarsus (TS = 0.36). Body length 1.86. Pedipalps: trochanter 0.35 / 0.19 (1.84 ×), femur 1.20 / 0.20 (6.00 ×), patella 0.52 / 0.21 (2.48 ×), chela 1.72 / 0.28 (6.14 ×), hand 0.62 / 0.28 (2.21 ×), movable chelal finger length 1.09. Chelicera 0.81 / 0.33 (2.45 ×), movable finger length 0.41. Carapace 0.69 / 0.74 (0.93 ×). Leg I: trochanter 0.22 / 0.14 (1.57 ×), femur 0.56 / 0.08 (7.00 ×), patella 0.37 / 0.08 (4.63 ×), tibia 0.32 / 0.07 (4.57 ×), tarsus 0.68 / 0.06 (11.33 ×). Leg IV: trochanter 0.31 / 0.18 (1.72 ×), femoropatella 0.82 / 0.28 (2.93 ×), tibia 0.68 / 0.11 (6.18 ×), basitarsus 0.31 / 0.09 (3.44 ×), telotarsus 0.69 / 0.07 (9.86 ×).
</paragraph>
<caption id="B7B9AD43C3596228AFDDDDCFF6BD3A4C" ID-DOI="10.3897/zookeys.1204.111842.figure8" ID-arpha="27AD253E-2A16-5074-BE45-2FCD00BF32F0" httpUri="https://binary.pensoft.net/fig/1062050" startId="F8">
<paragraph id="8FF98B42CD7159E933DE6E73E32116FB">
<label id="83430550EA663260CE0E96C91714CC61">Figure 8.</label>
</paragraph>
<paragraph id="F35B653BDF4A91F0E369F64928B11FDE">
<taxonomicName id="8F2D7C776C8593A7FE70717864485A9E" authority="Hou &amp; Zhang, 2024" authorityName="Hou &amp; Zhang" authorityYear="2024" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="huanglaoensis" status="sp. nov.">
<emphasis id="B94F94A4267F345CCDD0E1E463EEB46C" italics="true">Spelaeochthonius huanglaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="039F797AC5FE4948D29D4DD5F0DF0DF3" rank="species">sp. nov.</taxonomicNameLabel>
scanning electron micrographs, paratype female 
<emphasis id="B8B90C99E621D07810AF6C6F591F6885" bold="true">A</emphasis>
coxal spines in overview, with details of tips 
<emphasis id="383A769648E4D8FB12C185AD5817AA08" bold="true">B</emphasis>
serrula exterior 
<emphasis id="A1C22D955D161C269A587D92AF037114" bold="true">C</emphasis>
manducatory process 
<emphasis id="B881DAB618D2DCE13DCB30C754E028F9" bold="true">D</emphasis>
rallum. Scale bars: 50 μm (
<emphasis id="C7E190B8F3C3EFF802E2E8AB40CD62DE" bold="true">C, D</emphasis>
); 20 μm (
<emphasis id="FBFE7109B3485B67733938DC7A891814" bold="true">B</emphasis>
); 10 μm (
<emphasis id="A32333DA095406D987010DC0B299A8B8" bold="true">A</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EHCAG" type="distribution">
<paragraph id="320D6888F158B6C42219C6947FDACE99">
<heading id="F94E969089D6F84BFA7EA94DD1971746" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="347F495FF141C5B28E5CF89E609F6B88">
<collectingCountry id="A7BD5AFE6868E57A638881B073ADB49C" name="China">China</collectingCountry>
(
<collectingRegion id="F013DE1A11649AF4B67C1026B9267529" country="China" name="Beijing">Beijing</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>