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<document id="96AE4494CEC82EA41DF66450CCB10396" ID-DOI="10.1649/0010-065x(2005)059[0127:obngas]2.0.co;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4924192" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623349059116" checkinUser="carolina" docAuthor="Ratcliffe, Brett C., Smith, Dena M. &amp; Erwin, Diane" docDate="2005" docId="03F4E93CF027FFE68B786DEF9ABAF9D1" docLanguage="en" docName="TheColeopBull.59.1.127-135.pdf.imf" docOrigin="The Coleopterists Bulletin 59 (1)" docStyle="DocumentStyle:07711681861DB98305E87147CAB6DAD5.4:TheColeopBull.2003-2009.journal_article.0cover" docStyleId="07711681861DB98305E87147CAB6DAD5" docStyleName="TheColeopBull.2003-2009.journal_article.0cover" docStyleVersion="4" docTitle="Oryctoantiquus Ratcliffe and Smith, new genus" docType="treatment" docVersion="3" lastPageNumber="129" masterDocId="FFCD9144F026FFE48A38685F9816FFF0" masterDocTitle="Oryctoantiquus borealis, New Genus And Species From The Eocene Of Oregon, U. S. A., The World S Oldest Fossil Dynastine And Largest Fossil Scarabaeid (Coleoptera: Scarabaeidae: Dynastinae)" masterLastPageNumber="135" masterPageNumber="127" pageNumber="128" updateTime="1698962260145" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E73523E742D16C9A2C8C4DE9053FB5E2">Oryctoantiquus borealis, New Genus And Species From The Eocene Of Oregon, U. S. A., The World S Oldest Fossil Dynastine And Largest Fossil Scarabaeid (Coleoptera: Scarabaeidae: Dynastinae)</mods:title>
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<mods:namePart id="B0C40332198B4942D1096DB3283C14D3">Ratcliffe, Brett C.</mods:namePart>
<mods:affiliation id="D73FE3ACA11BDE21AE15266526E63FE2">Systematics Research Collections W- 436 Nebraska Hall University of Nebraska Lincoln, NE 68588 - 0514, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="277B141555B559741AE77F46C25DEC07" type="email">bratcliffe1@unl.edu</mods:nameIdentifier>
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<mods:namePart id="1D62A7CFCB42FF3F10300F2D816A7F26">Smith, Dena M.</mods:namePart>
<mods:affiliation id="4E7FA3BC89F24890D3CDB02D76B4BDAA">University of Colorado Museum of Natural History UCB 265, University of Colorado Boulder, CO 80309, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="F41AE82F84A08CAE83F9A70D92CE18AD" type="email">Dena.Smith@colorado.edu</mods:nameIdentifier>
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<mods:namePart id="4C6082EE1F6370B938A3490BC40D4B37">Erwin, Diane</mods:namePart>
<mods:affiliation id="336A771B11095F08620D0E8E9B74097B">Museum of Paleontology 1101 Valley Life Sciences Bldg. University of California Berkeley, CA 94720, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="7AE517D52BE80FD8A80CC42AB2E63275" type="email">dmerwin@uclink4.berkeley.edu</mods:nameIdentifier>
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<mods:title id="5FC4EA396257C4DAFA7B275AFC0195CD">The Coleopterists Bulletin</mods:title>
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<mods:date id="55EC6AF8B316FE193897520B4AE59BB1">2005</mods:date>
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<mods:number id="8EE0AF97D256AA8A96DDC814FF80EDD2">2005-03-01</mods:number>
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<mods:number id="6A8799327CDD1BFB8C806A29F915D5D1">59</mods:number>
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<mods:identifier id="F816D2ECA332E02234E7E04C39A6F265" type="DOI">10.1649/0010-065x(2005)059[0127:obngas]2.0.co;2</mods:identifier>
<mods:identifier id="95C288C017008226640442786B3D6BF1" type="ISSN">1938-4394</mods:identifier>
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<treatment id="03F4E93CF027FFE68B786DEF9ABAF9D1" ID-DOI="http://doi.org/10.5281/zenodo.4924194" ID-Zenodo-Dep="4924194" LSID="urn:lsid:plazi:treatment:03F4E93CF027FFE68B786DEF9ABAF9D1" httpUri="http://treatment.plazi.org/id/03F4E93CF027FFE68B786DEF9ABAF9D1" lastPageId="2" lastPageNumber="129" pageId="1" pageNumber="128">
<subSubSection id="C3470BA1F027FFE58B786DEF9B33FA35" box="[320,805,1456,1477]" pageId="1" pageNumber="128" type="nomenclature">
<paragraph id="8BE2582AF027FFE58B786DEF9B33FA35" blockId="1.[320,805,1456,1477]" box="[320,805,1456,1477]" pageId="1" pageNumber="128">
<heading id="D0AAEF46F027FFE58B786DEF9B33FA35" box="[320,805,1456,1477]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="128" reason="2">
<emphasis id="B9298438F027FFE58B786DEF9B33FA35" bold="true" box="[320,805,1456,1477]" pageId="1" pageNumber="128">
<taxonomicName id="4C5D23A9F027FFE58B786DEF9ABAFA35" authority="Ratcliffe and Smith" authorityName="Ratcliffe and Smith" box="[320,684,1456,1477]" class="Insecta" family="Dynastidae" genus="Oryctoantiquus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="128" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9298438F027FFE58B786DEF99CFFA35" bold="true" box="[320,473,1456,1477]" italics="true" pageId="1" pageNumber="128">Oryctoantiquus</emphasis>
Ratcliffe and Smith
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,
<taxonomicNameLabel id="A21A3943F027FFE588816DEF9B33FA35" box="[697,805,1456,1477]" pageId="1" pageNumber="128" rank="genus">new genus</taxonomicNameLabel>
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<subSubSection id="C3470BA1F027FFE58A946D879AEFFA1D" box="[172,761,1496,1517]" pageId="1" pageNumber="128" type="type_taxon">
<paragraph id="8BE2582AF027FFE58A946D879AEFFA1D" blockId="1.[148,977,1496,1570]" box="[172,761,1496,1517]" pageId="1" pageNumber="128">
<emphasis id="B9298438F027FFE58A946D87992EFA1D" bold="true" box="[172,312,1496,1517]" pageId="1" pageNumber="128">
<typeStatus id="54E6E688F027FFE58A946D8798F6FA1D" box="[172,224,1496,1517]" pageId="1" pageNumber="128">Type</typeStatus>
Species.
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<taxonomicName id="4C5D23A9F027FFE58B7C6D879AEFFA1D" authority="Ratcliffe and Smith." authorityName="Ratcliffe and Smith." box="[324,761,1496,1517]" class="Insecta" family="Dynastidae" genus="Oryctoantiquus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="128" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B9298438F027FFE58B7C6D879A3BFA1D" box="[324,557,1496,1517]" italics="true" pageId="1" pageNumber="128">Oryctoantiquus borealis</emphasis>
Ratcliffe and Smith.
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<subSubSection id="C3470BA1F027FFE68A946DAD9913FA47" lastPageId="2" lastPageNumber="129" pageId="1" pageNumber="128" type="diagnosis">
<paragraph id="8BE2582AF027FFE68A946DAD9913FA47" blockId="1.[148,977,1496,1570]" lastBlockId="2.[175,1005,1415,1569]" lastPageId="2" lastPageNumber="129" pageId="1" pageNumber="128">
<emphasis id="B9298438F027FFE58A946DAD9902F9F7" bold="true" box="[172,276,1522,1543]" pageId="1" pageNumber="128">Diagnosis.</emphasis>
The same as for the monotypic species,
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<emphasis id="B9298438F027FFE5888A6DAC9B8BF9F8" box="[690,925,1523,1544]" italics="true" pageId="1" pageNumber="128">Oryctoantiquus borealis</emphasis>
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. The pattern of punctures on the metasternum, combined with the large body size and form of the posterior legs, distinguishes
<taxonomicName id="4C5D23A9F024FFE688206DD89ABAFA6C" box="[536,684,1415,1436]" class="Insecta" family="Dynastidae" genus="Oryctoantiquus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B9298438F024FFE688206DD89ABAFA6C" box="[536,684,1415,1436]" italics="true" pageId="2" pageNumber="129">Oryctoantiquus</emphasis>
</taxonomicName>
from other genera in the tribe
<taxonomicName id="4C5D23A9F024FFE68A976DFD9917FA47" box="[175,257,1442,1463]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="tribe" tribe="Oryctini">Oryctini</taxonomicName>
.
</paragraph>
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<caption id="DF2208A2F024FFE68AFD6D679AD0FABB" ID-DOI="http://doi.org/10.5281/zenodo.4924196" ID-Zenodo-Dep="4924196" box="[197,710,1336,1355]" httpUri="https://zenodo.org/record/4924196/files/figure.png" pageId="2" pageNumber="129" startId="2.[197,228,1336,1355]" targetBox="[177,978,194,1314]" targetPageId="2">
<paragraph id="8BE2582AF024FFE68AFD6D679AD0FABB" blockId="2.[197,710,1336,1355]" box="[197,710,1336,1355]" pageId="2" pageNumber="129">
<emphasis id="B9298438F024FFE68AFD6D6798E9FABB" bold="true" box="[197,255,1336,1355]" pageId="2" pageNumber="129">Fig. 1.</emphasis>
<taxonomicName id="4C5D23A9F024FFE68B2C6D6799F5FABB" box="[276,483,1336,1355]" class="Insecta" family="Dynastidae" genus="Oryctoantiquus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B9298438F024FFE68B2C6D6799F5FABB" box="[276,483,1336,1355]" italics="true" pageId="2" pageNumber="129">Oryctoantiquus borealis</emphasis>
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. Scale bar equals 10 mm.
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<subSubSection id="C3470BA1F024FFE68AFF6DE39ABAF9D1" pageId="2" pageNumber="129" type="etymology">
<paragraph id="8BE2582AF024FFE68AFF6DE39ABAF9D1" blockId="2.[175,1005,1415,1569]" pageId="2" pageNumber="129">
<emphasis id="B9298438F024FFE68AFF6DE3992DFA21" bold="true" box="[199,315,1468,1489]" pageId="2" pageNumber="129">Etymology.</emphasis>
The generic name is derived from the Greek
<emphasis id="B9298438F024FFE6893B6DE39B51FA21" box="[771,839,1468,1489]" italics="true" pageId="2" pageNumber="129">oryktes</emphasis>
, meaning digger (also the stem of the dynastine tribal name
<taxonomicName id="4C5D23A9F024FFE688696D889AB5FA1C" box="[593,675,1495,1516]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="tribe" tribe="Oryctini">Oryctini</taxonomicName>
), and the Latin
<emphasis id="B9298438F024FFE689036D889B9BFA1C" box="[827,909,1495,1516]" italics="true" pageId="2" pageNumber="129">antiquus</emphasis>
, meaning old; hence the old digger or old oryctine in reference to the 45 million year old age of the fossil and the tribe to which we assign it.
</paragraph>
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