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<document id="575E795405541B88FDE86B2DB27B6A08" ID-DOI="10.1080/00222933.2021.1931516" ID-ISSN="1464-5262" ID-Zenodo-Dep="5464831" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630917156140" checkinUser="plazi" docAuthor="Liu, Zhen-Hua &amp; Jia, Feng-Long" docDate="2021" docId="03F487FCF703762490B3FEDEFC9626A7" docLanguage="en" docName="JNATHIST.55.13-14.909-920.pdf" docOrigin="Journal of Natural History 55 (13 - 14)" docSource="http://dx.doi.org/10.1080/00222933.2021.1931516" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Mexico lawrencei Liu &amp; Jia 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="918" masterDocId="FFCDFF84F705762D9149FFD6F8152777" masterDocTitle="Descriptions of two new species of Mexico Spilman, 1972 from the Oriental region (Coleoptera: Limnichidae)" masterLastPageNumber="920" masterPageNumber="909" pageNumber="915" updateTime="1699282155100" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D7EE661191EBD177D8AC3A68E7ADB8B5">Descriptions of two new species of Mexico Spilman, 1972 from the Oriental region (Coleoptera: Limnichidae)</mods:title>
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<mods:namePart id="B3D670982376BF795C62534928BB6421">Liu, Zhen-Hua</mods:namePart>
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<mods:namePart id="69E0BD9D00F345F0B42BBBFAF55451B2">Jia, Feng-Long</mods:namePart>
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<mods:title id="F59673177BC57D5B9C4FA2A55D9F670A">Journal of Natural History</mods:title>
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<mods:number id="6E621341C641769583DB85FE62627179">13 - 14</mods:number>
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<subSubSection id="C3476561F703762B90B3FEDEFB3C2655" box="[506,809,264,290]" pageId="6" pageNumber="915" type="nomenclature">
<paragraph id="8BE236EAF703762B90B3FEDEFB3C2655" blockId="6.[506,809,264,324]" box="[506,809,264,290]" pageId="6" pageNumber="915">
<heading id="D0AA8186F703762B90B3FEDEFB3C2655" box="[506,809,264,290]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="915" reason="5">
<taxonomicName id="4C5D4D69F703762B90B3FEDEFADB2655" authority="Liu &amp; Jia, 2021" authorityName="Liu &amp; Jia" authorityYear="2021" box="[506,718,264,290]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="915" phylum="Arthropoda" rank="species" species="lawrencei" status="sp. nov.">
<emphasis id="B929EAF8F703762B90B3FEDEFADB2655" bold="true" box="[506,718,264,290]" italics="true" pageId="6" pageNumber="915">Mexico lawrencei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A5783F703762B9387FEDEFB3C2655" box="[718,809,264,290]" pageId="6" pageNumber="915" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C3476561F703762B9352FEFCFB1C2633" box="[539,777,298,324]" pageId="6" pageNumber="915" type="description">
<paragraph id="8BE236EAF703762B9352FEFCFB1C2633" blockId="6.[506,809,264,324]" box="[539,777,298,324]" pageId="6" pageNumber="915">
(
<figureCitation id="13662A6FF703762B936AFEFCFA9C2633" box="[547,649,298,324]" captionStart="Figure 1" captionStartId="2.[160,225,1261,1284]" captionTargetBox="[171,1151,145,1218]" captionTargetId="figure-105@2.[163,1153,142,1229]" captionTargetPageId="2" captionText="Figure 1. Habitus of Mexico spp. nov. (ab) Mexico xiei sp. nov. Holotype. (cd) Mexico lawrencei sp. nov. Paratype. (a, c) dorsal view; (b, d) ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5464835" httpUri="https://zenodo.org/record/5464835/files/figure.png" pageId="6" pageNumber="915">Figures 1</figureCitation>
(cd), 3, 4)
</paragraph>
</subSubSection>
<subSubSection id="C3476561F703762B91E9FE57FA0C2697" pageId="6" pageNumber="915" type="etymology">
<paragraph id="8BE236EAF703762B91E9FE57F90826EC" blockId="6.[160,1155,385,480]" box="[160,285,385,411]" pageId="6" pageNumber="915">
<emphasis id="B929EAF8F703762B91E9FE57F90826EC" bold="true" box="[160,285,385,411]" italics="true" pageId="6" pageNumber="915">Etymology</emphasis>
</paragraph>
<paragraph id="8BE236EAF703762B91E9FE72FA0C2697" blockId="6.[160,1155,385,480]" pageId="6" pageNumber="915">
The new species is named after John Lawrence, a coleopterist who collected these specimens from
<collectingCountry id="F34A767AF703762B9012FE11FA002697" box="[347,533,455,480]" name="Christmas Island" pageId="6" pageNumber="915">Christmas Island</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3476561F703762B91E9FDDAF96D2582" pageId="6" pageNumber="915" type="materials_examined">
<paragraph id="8BE236EAF703762B91E9FDDAF9232551" blockId="6.[160,461,524,584]" box="[160,310,524,550]" pageId="6" pageNumber="915">
<emphasis id="B929EAF8F703762B91E9FDDAF9232551" bold="true" box="[160,310,524,550]" italics="true" pageId="6" pageNumber="915">Type locality</emphasis>
</paragraph>
<paragraph id="8BE236EAF703762B91E9FDF9F9D8253F" blockId="6.[160,461,524,584]" box="[160,461,559,584]" pageId="6" pageNumber="915">
<collectingCountry id="F34A767AF703762B91E9FDF9F94E253F" box="[160,347,559,584]" name="Christmas Island" pageId="6" pageNumber="915">Christmas Island</collectingCountry>
,
<collectingCountry id="F34A767AF703762B902CFDF9F9DC253F" box="[357,457,559,584]" name="Australia" pageId="6" pageNumber="915">Australia</collectingCountry>
.
</paragraph>
<paragraph id="8BE236EAF703762B91E9FDA2F95125F9" blockId="6.[160,1156,628,757]" box="[160,324,628,654]" pageId="6" pageNumber="915">
<emphasis id="B929EAF8F703762B91E9FDA2F95125F9" bold="true" box="[160,324,628,654]" italics="true" pageId="6" pageNumber="915">Type material</emphasis>
</paragraph>
<paragraph id="8BE236EAF703762B91E9FD40F96D2582" blockId="6.[160,1156,628,757]" pageId="6" pageNumber="915">
<materialsCitation id="3B353CB7F703762B91E9FD40FAEE25A4" collectingDate="1989-04-12" collectingDateMax="1989-04-24" collectingDateMin="1989-04-12" collectionCode="ANIC" collectorName="J. F. Lawrence" country="Christmas Island" latitude="-10.31" location="Christmas Island" longLatPrecision="779" longitude="105.37" pageId="6" pageNumber="915" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B929EAF8F703762B91E9FD40F90725C7" bold="true" box="[160,274,662,688]" italics="true" pageId="6" pageNumber="915">
<typeStatus id="54E68848F703762B91E9FD40F91825C7" box="[160,269,662,688]" pageId="6" pageNumber="915" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D5BFD63F703762B9068FD41F92025D9" box="[289,309,663,686]" pageId="6" pageNumber="915" type="male"></specimenCount>
,
<collectingCountry id="F34A767AF703762B900DFD41FA1525C7" box="[324,512,663,688]" name="Christmas Island" pageId="6" pageNumber="915">Christmas Island</collectingCountry>
,
<geoCoordinate id="EE69502DF703762B9342FD41FA4125C7" box="[523,596,663,688]" degrees="10.31" direction="south" orientation="latitude" pageId="6" pageNumber="915" precision="555" value="-10.31">10.31S</geoCoordinate>
<geoCoordinate id="EE69502DF703762B9317FD41FAA125C7" box="[606,692,663,688]" degrees="105.37" direction="east" orientation="longitude" pageId="6" pageNumber="915" precision="555" value="105.37">105.37E</geoCoordinate>
nr. The Blowholes
<date id="FFE3102AF703762B92D8FD41FC4025C7" box="[913,1109,663,688]" pageId="6" pageNumber="915" value="1989-04-12" valueMax="1989-04-24" valueMin="1989-04-12">
<collectingDate id="EFA7E9C2F703762B92D8FD41FC4025C7" box="[913,1109,663,688]" pageId="6" pageNumber="915" value="1989-04-12" valueMax="1989-04-24" valueMin="1989-04-12">1224 April 1989</collectingDate>
</date>
.
<collectorName id="26A8533CF703762B9528FD41F90525A4" pageId="6" pageNumber="915">J.F. Lawrence</collectorName>
/mud beneath limestones pinnacles (
<collectionCode id="ED4CAE2FF703762B93FEFD6CFAE025A4" box="[695,757,698,723]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="6" pageNumber="915">ANIC</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54E68848F703762B9241FD6CFB6D25A4" box="[776,888,698,723]" pageId="6" pageNumber="915" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D5BFD63F703762B92CCFD6CFC0025A4" box="[901,1045,698,723]" pageId="6" pageNumber="915" type="generic">9 specimens</specimenCount>
, with the same labels (
<collectionCode id="ED4CAE2FF703762B9079FD0AF97B2582" box="[304,366,732,757]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="6" pageNumber="915">ANIC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3476561F703762B91E9FCF7FB55237C" pageId="6" pageNumber="915" type="diagnosis">
<paragraph id="8BE236EAF703762B91E9FCF7F901244C" blockId="6.[160,1156,801,1035]" box="[160,276,801,827]" pageId="6" pageNumber="915">
<emphasis id="B929EAF8F703762B91E9FCF7F901244C" bold="true" box="[160,276,801,827]" italics="true" pageId="6" pageNumber="915">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BE236EAF703762B91E9FC93FB55237C" blockId="6.[160,1156,801,1035]" pageId="6" pageNumber="915">
This species is similar to
<emphasis id="B929EAF8F703762B90E4FC93FB462429" box="[429,851,837,862]" italics="true" pageId="6" pageNumber="915">M. papuanus, M. litoralis, M. borneensis</emphasis>
and
<emphasis id="B929EAF8F703762B92C3FC93FC132429" box="[906,1030,837,862]" italics="true" pageId="6" pageNumber="915">M. baliensis</emphasis>
in terms of the very simple median lobe without divided subapical projections. It can be easily distinguished from those species by median lobe distinctly expanded subapically. It can be further distinguished from
<emphasis id="B929EAF8F703762B90B9FC7BFA6824B1" box="[496,637,941,966]" italics="true" pageId="6" pageNumber="915">M. papuanus</emphasis>
and
<emphasis id="B929EAF8F703762B93FDFC7BFB3124B1" box="[692,804,941,966]" italics="true" pageId="6" pageNumber="915">M. litoralis</emphasis>
by the long apical setae on the lateral lobes and lateral lobes being shorter than half of median lobe, and from
<emphasis id="B929EAF8F703762B91E9FC24F920237C" box="[160,309,1010,1035]" italics="true" pageId="6" pageNumber="915">M. borneensis</emphasis>
and
<emphasis id="B929EAF8F703762B9026FC24F9F9237C" box="[367,492,1010,1035]" italics="true" pageId="6" pageNumber="915">M. baliensis</emphasis>
by the more acuminate apex.
</paragraph>
</subSubSection>
<subSubSection id="C3476561F703762591E9FBE0FA36241A" lastPageId="8" lastPageNumber="917" pageId="6" pageNumber="915" type="description">
<paragraph id="8BE236EAF703762B91E9FBE0F9332327" blockId="6.[160,1156,1078,1728]" box="[160,294,1078,1104]" pageId="6" pageNumber="915">
<emphasis id="B929EAF8F703762B91E9FBE0F9332327" bold="true" box="[160,294,1078,1104]" italics="true" pageId="6" pageNumber="915">Description</emphasis>
</paragraph>
<paragraph id="8BE236EAF703762B91E9FB8FFA3F23CF" blockId="6.[160,1156,1078,1728]" pageId="6" pageNumber="915">
<emphasis id="B929EAF8F703762B91E9FB8FF97F2304" bold="true" box="[160,362,1113,1139]" italics="true" pageId="6" pageNumber="915">Colour and form.</emphasis>
Body (
<figureCitation id="13662A6FF703762B908BFB8CFA0D2304" box="[450,536,1114,1139]" captionStart="Figure 1" captionStartId="2.[160,225,1261,1284]" captionTargetBox="[171,1151,145,1218]" captionTargetId="figure-105@2.[163,1153,142,1229]" captionTargetPageId="2" captionText="Figure 1. Habitus of Mexico spp. nov. (ab) Mexico xiei sp. nov. Holotype. (cd) Mexico lawrencei sp. nov. Paratype. (a, c) dorsal view; (b, d) ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5464835" httpUri="https://zenodo.org/record/5464835/files/figure.png" pageId="6" pageNumber="915">Figure 1</figureCitation>
(c,d)) small-sized, broadly elliptical, length
<quantity id="4CA59B0FF703762B92B2FB8CFC952304" box="[1019,1152,1114,1139]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="6" pageNumber="915" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.92.1 mm</quantity>
, width 1.0
<quantity id="4CA59B0FF703762B905EFBABF97923E1" box="[279,364,1149,1174]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="915" unit="mm" value="1.1">1.1 mm</quantity>
. Dorsal side black, covered with short, dense golden and whitish setae; ventral side more or less brownish.
</paragraph>
<paragraph id="8BE236EAF703762A9189FB14F9782304" blockId="6.[160,1156,1078,1728]" lastBlockId="7.[160,1156,1079,1728]" lastPageId="7" lastPageNumber="916" pageId="6" pageNumber="915">
<emphasis id="B929EAF8F703762B9189FB14F8EB23AC" bold="true" box="[192,254,1218,1243]" pageId="6" pageNumber="915">Head</emphasis>
(
<figureCitation id="13662A6FF703762B9046FB14F96623AC" box="[271,371,1218,1243]" captionStart="Figure 1" captionStartId="2.[160,225,1261,1284]" captionTargetBox="[171,1151,145,1218]" captionTargetId="figure-105@2.[163,1153,142,1229]" captionTargetPageId="2" captionText="Figure 1. Habitus of Mexico spp. nov. (ab) Mexico xiei sp. nov. Holotype. (cd) Mexico lawrencei sp. nov. Paratype. (a, c) dorsal view; (b, d) ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5464835" httpUri="https://zenodo.org/record/5464835/files/figure.png" pageId="6" pageNumber="915">Figures 1</figureCitation>
(d), 4(g)) strongly hypognathous, with mouthparts mostly concealed by labrum and prosternum when retracted, surface with dense microsculptures, occipital incisions absent; genal areas with distinct antennal grooves, margined by distinct carinae both anteriorly and posteriorly, with anterior half glabrous and posterior half microsculptured; frontoclypeal suture well developed and straight, clypeus rectangular and slightly protruding. Eyes anterolaterally located, large and finely faceted, well separated. Antennal insertions exposed, well separated from eyes. Antennae (
<figureCitation id="13662A6FF703762B9268FA44FB6B22DC" box="[801,894,1426,1451]" captionStart="Figure 4" captionStartId="7.[160,225,872,895]" captionTargetBox="[178,1141,155,834]" captionTargetId="figure-229@7.[163,1153,142,840]" captionTargetPageId="7" captionText="Figure 4. Mexico lawrencei sp. nov., dissections. (a) pterothorax, ventral; (b) abdomen, ventral; (c) labium, ventral; (d) mandible, dorsal; (e) maxilla, ventral; (f) antenna; (g) head, front; (h) sternites IX, male; (i) aedeagus, dorsal; (j) ovipositor and spermatheca; (k) sternite VIII, female. Scale bars: a b = 0.5 mm; cf = 0.1 mm; gk = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5464849" httpUri="https://zenodo.org/record/5464849/files/figure.png" pageId="6" pageNumber="915">Figure 4</figureCitation>
(f)) 11-segmented, with 4-segmented club; scape and pedicel enlarged, antennomere 3 stout, antennomeres 47 narrow, last segment subulate. Labrum large and sub-trapezoid, wider than long. Mandibles (
<figureCitation id="13662A6FF703762B906EFA2CF99D2164" box="[295,392,1530,1555]" captionStart="Figure 4" captionStartId="7.[160,225,872,895]" captionTargetBox="[178,1141,155,834]" captionTargetId="figure-229@7.[163,1153,142,840]" captionTargetPageId="7" captionText="Figure 4. Mexico lawrencei sp. nov., dissections. (a) pterothorax, ventral; (b) abdomen, ventral; (c) labium, ventral; (d) mandible, dorsal; (e) maxilla, ventral; (f) antenna; (g) head, front; (h) sternites IX, male; (i) aedeagus, dorsal; (j) ovipositor and spermatheca; (k) sternite VIII, female. Scale bars: a b = 0.5 mm; cf = 0.1 mm; gk = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5464849" httpUri="https://zenodo.org/record/5464849/files/figure.png" pageId="6" pageNumber="915">Figure 4</figureCitation>
(d)) small and sub-triangular, dorsal side distinctly elevated, apex slender; inner edge membranous, with a membranous apical projection; mola well developed. Maxilla (
<figureCitation id="13662A6FF703762B90CBF9E9F9CA212F" box="[386,479,1599,1624]" captionStart="Figure 4" captionStartId="7.[160,225,872,895]" captionTargetBox="[178,1141,155,834]" captionTargetId="figure-229@7.[163,1153,142,840]" captionTargetPageId="7" captionText="Figure 4. Mexico lawrencei sp. nov., dissections. (a) pterothorax, ventral; (b) abdomen, ventral; (c) labium, ventral; (d) mandible, dorsal; (e) maxilla, ventral; (f) antenna; (g) head, front; (h) sternites IX, male; (i) aedeagus, dorsal; (j) ovipositor and spermatheca; (k) sternite VIII, female. Scale bars: a b = 0.5 mm; cf = 0.1 mm; gk = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5464849" httpUri="https://zenodo.org/record/5464849/files/figure.png" pageId="6" pageNumber="915">Figure 4</figureCitation>
(e)) slender, with 4-segmented palps, last segment enlarged and acuminate apically; lacinia elongated with apical hook and short subapical spine; galea subequal in length with lacinia, apex acuminate. Labium (
<figureCitation id="13662A6FF703762B9232F953FBCC21E9" box="[891,985,1669,1694]" captionStart="Figure 4" captionStartId="7.[160,225,872,895]" captionTargetBox="[178,1141,155,834]" captionTargetId="figure-229@7.[163,1153,142,840]" captionTargetPageId="7" captionText="Figure 4. Mexico lawrencei sp. nov., dissections. (a) pterothorax, ventral; (b) abdomen, ventral; (c) labium, ventral; (d) mandible, dorsal; (e) maxilla, ventral; (f) antenna; (g) head, front; (h) sternites IX, male; (i) aedeagus, dorsal; (j) ovipositor and spermatheca; (k) sternite VIII, female. Scale bars: a b = 0.5 mm; cf = 0.1 mm; gk = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5464849" httpUri="https://zenodo.org/record/5464849/files/figure.png" pageId="6" pageNumber="915">Figure 4</figureCitation>
(c)) with 3-segmented labial palps, second segment enlarged, last segment small; ligula well developed. Mentum rectangular, longer than wide. Gular area highly reduced. Cervical sclerites present and large.
</paragraph>
<caption id="DF226662F702762A91E9FCBEFA5D24A0" ID-DOI="http://doi.org/10.5281/zenodo.5464849" ID-Zenodo-Dep="5464849" httpUri="https://zenodo.org/record/5464849/files/figure.png" pageId="7" pageNumber="916" startId="7.[160,225,872,895]" targetBox="[178,1141,155,834]" targetPageId="7">
<paragraph id="8BE236EAF702762A91E9FCBEFA5D24A0" blockId="7.[160,1156,872,983]" pageId="7" pageNumber="916">
<emphasis id="B929EAF8F702762A91E9FCBEF8E82408" bold="true" box="[160,253,872,895]" pageId="7" pageNumber="916">Figure 4.</emphasis>
<taxonomicName id="4C5D4D69F702762A904EFCBEF9BB2408" authority="Liu &amp; Jia, 2021" authorityName="Liu &amp; Jia" authorityYear="2021" box="[263,430,872,895]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="916" phylum="Arthropoda" rank="species" species="lawrencei" status="sp. nov.">
<emphasis id="B929EAF8F702762A904EFCBEF9BB2408" box="[263,430,872,895]" italics="true" pageId="7" pageNumber="916">Mexico lawrencei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A5783F702762A90FEFCBEFA1C2408" box="[439,521,872,895]" pageId="7" pageNumber="916" rank="species">sp. nov.</taxonomicNameLabel>
, dissections. (a) pterothorax, ventral; (b) abdomen, ventral; (c) labium, ventral; (d) mandible, dorsal; (e) maxilla, ventral; (f) antenna; (g) head, front; (h) sternites IX, male; (i) aedeagus, dorsal; (j) ovipositor and spermatheca; (k) sternite VIII, female. Scale bars: a b = 0.5 mm; cf = 0.1 mm; gk = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BE236EAF70276259189FBABFB4E2622" blockId="7.[160,1156,1079,1728]" lastBlockId="8.[159,1156,142,757]" lastPageId="8" lastPageNumber="917" pageId="7" pageNumber="916">
<emphasis id="B929EAF8F702762A9189FBABF90123E1" bold="true" box="[192,276,1149,1174]" pageId="7" pageNumber="916">Thorax</emphasis>
. Pronotum transverse, about 0.4 times as long as wide, widest posteriorly; lateral margins slightly curved, posterior margin sinuate with distinct median lobe; anterior angles broadly acute and forwardly projecting; disc weakly convex, with dense microsculptures. Pronotal epipleuron broad and slightly depressed. Prosternum narrow, prosternal process broad and parallel-sided. Notosternal suture complete. Procoxae strongly transverse, protrochantins small and exposed; procoxal cavities well separated, with small anterolateral incisions, externally broadly opened and internally narrowly opened. Scutellum small and triangular. Elytra about 1.4 times as long as wide, with about 10 tiny teeth along anterolateral edges and one small apical projection on either side; disc slightly convex, with dense microsculptures; epipleuron broad at base, extending to apical projection. Hing wings well developed. Mesoventrite (
<figureCitation id="13662A6FF702762A92CEFA01FBF72287" box="[903,994,1495,1520]" captionStart="Figure 4" captionStartId="7.[160,225,872,895]" captionTargetBox="[178,1141,155,834]" captionTargetId="figure-229@7.[163,1153,142,840]" captionTargetPageId="7" captionText="Figure 4. Mexico lawrencei sp. nov., dissections. (a) pterothorax, ventral; (b) abdomen, ventral; (c) labium, ventral; (d) mandible, dorsal; (e) maxilla, ventral; (f) antenna; (g) head, front; (h) sternites IX, male; (i) aedeagus, dorsal; (j) ovipositor and spermatheca; (k) sternite VIII, female. Scale bars: a b = 0.5 mm; cf = 0.1 mm; gk = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5464849" httpUri="https://zenodo.org/record/5464849/files/figure.png" pageId="7" pageNumber="916">Figure 4</figureCitation>
(a)) shortened, with large median concavity fitting to prosternal process. Mesanepisternum and mesepimeron fused (
<figureCitation id="13662A6FF702762A900BF9CBF9B52141" box="[322,416,1565,1590]" captionStart="Figure 4" captionStartId="7.[160,225,872,895]" captionTargetBox="[178,1141,155,834]" captionTargetId="figure-229@7.[163,1153,142,840]" captionTargetPageId="7" captionText="Figure 4. Mexico lawrencei sp. nov., dissections. (a) pterothorax, ventral; (b) abdomen, ventral; (c) labium, ventral; (d) mandible, dorsal; (e) maxilla, ventral; (f) antenna; (g) head, front; (h) sternites IX, male; (i) aedeagus, dorsal; (j) ovipositor and spermatheca; (k) sternite VIII, female. Scale bars: a b = 0.5 mm; cf = 0.1 mm; gk = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5464849" httpUri="https://zenodo.org/record/5464849/files/figure.png" pageId="7" pageNumber="916">Figure 4</figureCitation>
(a)). Metaventrite subtriangular, strongly transverse at base, with deep and narrow apical incision; metendosternite with basal strut highly sclerotised, large and fusiform, fused with metaventrite by transverse carinae; lateral arms relatively long, with complicated ventral projections connected with apex of basal strut; metanepisternum highly reduced, partly exposed from ventral side which is triangular. Mesocoxae (
<figureCitation id="13662A6FF70D762591E1FF58F91727D0" box="[168,258,142,167]" captionStart="Figure 4" captionStartId="7.[160,225,872,895]" captionTargetBox="[178,1141,155,834]" captionTargetId="figure-229@7.[163,1153,142,840]" captionTargetPageId="7" captionText="Figure 4. Mexico lawrencei sp. nov., dissections. (a) pterothorax, ventral; (b) abdomen, ventral; (c) labium, ventral; (d) mandible, dorsal; (e) maxilla, ventral; (f) antenna; (g) head, front; (h) sternites IX, male; (i) aedeagus, dorsal; (j) ovipositor and spermatheca; (k) sternite VIII, female. Scale bars: a b = 0.5 mm; cf = 0.1 mm; gk = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5464849" httpUri="https://zenodo.org/record/5464849/files/figure.png" pageId="8" pageNumber="917">Figure 4</figureCitation>
(a)) slightly transverse, trochantins exposed; mesocoxal cavities widely separated, laterally open to mesepimeron. Metacoxae large and transverse, strongly oblique, narrowly separated; metacoxal plates large, sharply narrowed to lateral edges. Legs with femora slightly enlarged, tibiae slender, tarsi slightly shorter than tibiae, claws small; tarsal formula 4-4-4, basal three tarsomeres with pair of short apical spines. Hind leg distinctly longer than others, tibiae with long spines along outer edges.
</paragraph>
<paragraph id="8BE236EAF70D76259189FE88FC3E26A8" blockId="8.[159,1156,142,757]" pageId="8" pageNumber="917">
<emphasis id="B929EAF8F70D76259189FE88F9272600" bold="true" box="[192,306,350,375]" pageId="8" pageNumber="917">Abdomen</emphasis>
(
<figureCitation id="13662A6FF70D7625900DFE88F9B42600" box="[324,417,350,375]" captionStart="Figure 4" captionStartId="7.[160,225,872,895]" captionTargetBox="[178,1141,155,834]" captionTargetId="figure-229@7.[163,1153,142,840]" captionTargetPageId="7" captionText="Figure 4. Mexico lawrencei sp. nov., dissections. (a) pterothorax, ventral; (b) abdomen, ventral; (c) labium, ventral; (d) mandible, dorsal; (e) maxilla, ventral; (f) antenna; (g) head, front; (h) sternites IX, male; (i) aedeagus, dorsal; (j) ovipositor and spermatheca; (k) sternite VIII, female. Scale bars: a b = 0.5 mm; cf = 0.1 mm; gk = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5464849" httpUri="https://zenodo.org/record/5464849/files/figure.png" pageId="8" pageNumber="917">Figure 4</figureCitation>
(b)) 5-segmented with basal three ventrites connate, lateral edges expanded with narrow flanges.Ventrite 1nearly divided by metacoxae,with intercoxal process short and acute; ventrites 24 sub-equal in length; last ventrite elongate and subtriangular, with pair of small posterolateral projections, bearing clustered sub-lateral long setae.
</paragraph>
<paragraph id="8BE236EAF70D76259189FE3FF90B25D8" blockId="8.[159,1156,142,757]" pageId="8" pageNumber="917">
<emphasis id="B929EAF8F70D76259189FE3FF9782575" bold="true" box="[192,365,489,514]" pageId="8" pageNumber="917">Male genitalia</emphasis>
with tergite VIII as in
<figureCitation id="13662A6FF70D7625933BFE3FFAC72575" box="[626,722,489,514]" captionStart="Figure 4" captionStartId="7.[160,225,872,895]" captionTargetBox="[178,1141,155,834]" captionTargetId="figure-229@7.[163,1153,142,840]" captionTargetPageId="7" captionText="Figure 4. Mexico lawrencei sp. nov., dissections. (a) pterothorax, ventral; (b) abdomen, ventral; (c) labium, ventral; (d) mandible, dorsal; (e) maxilla, ventral; (f) antenna; (g) head, front; (h) sternites IX, male; (i) aedeagus, dorsal; (j) ovipositor and spermatheca; (k) sternite VIII, female. Scale bars: a b = 0.5 mm; cf = 0.1 mm; gk = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5464849" httpUri="https://zenodo.org/record/5464849/files/figure.png" pageId="8" pageNumber="917">Figure 4</figureCitation>
(h). Aedeagus (
<figureCitation id="13662A6FF70D76259237FE3FFBCA2575" box="[894,991,489,514]" captionStart="Figure 4" captionStartId="7.[160,225,872,895]" captionTargetBox="[178,1141,155,834]" captionTargetId="figure-229@7.[163,1153,142,840]" captionTargetPageId="7" captionText="Figure 4. Mexico lawrencei sp. nov., dissections. (a) pterothorax, ventral; (b) abdomen, ventral; (c) labium, ventral; (d) mandible, dorsal; (e) maxilla, ventral; (f) antenna; (g) head, front; (h) sternites IX, male; (i) aedeagus, dorsal; (j) ovipositor and spermatheca; (k) sternite VIII, female. Scale bars: a b = 0.5 mm; cf = 0.1 mm; gk = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5464849" httpUri="https://zenodo.org/record/5464849/files/figure.png" pageId="8" pageNumber="917">Figure 4</figureCitation>
(i)) with lateral lobes about half length of penis, bearing two long apical setae; basal piece sub-annular; median lobe elongate and simple, slightly constricted at about middle, apex acuminate. Female genitalia with spiculum ventrale (
<figureCitation id="13662A6FF70D76259339FD87FAD9251D" box="[624,716,593,618]" captionStart="Figure 4" captionStartId="7.[160,225,872,895]" captionTargetBox="[178,1141,155,834]" captionTargetId="figure-229@7.[163,1153,142,840]" captionTargetPageId="7" captionText="Figure 4. Mexico lawrencei sp. nov., dissections. (a) pterothorax, ventral; (b) abdomen, ventral; (c) labium, ventral; (d) mandible, dorsal; (e) maxilla, ventral; (f) antenna; (g) head, front; (h) sternites IX, male; (i) aedeagus, dorsal; (j) ovipositor and spermatheca; (k) sternite VIII, female. Scale bars: a b = 0.5 mm; cf = 0.1 mm; gk = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5464849" httpUri="https://zenodo.org/record/5464849/files/figure.png" pageId="8" pageNumber="917">Figure 4</figureCitation>
(k)) elongated and bifurcate; ovipositor (
<figureCitation id="13662A6FF70D762591E1FDA2F91025FA" box="[168,261,628,653]" captionStart="Figure 4" captionStartId="7.[160,225,872,895]" captionTargetBox="[178,1141,155,834]" captionTargetId="figure-229@7.[163,1153,142,840]" captionTargetPageId="7" captionText="Figure 4. Mexico lawrencei sp. nov., dissections. (a) pterothorax, ventral; (b) abdomen, ventral; (c) labium, ventral; (d) mandible, dorsal; (e) maxilla, ventral; (f) antenna; (g) head, front; (h) sternites IX, male; (i) aedeagus, dorsal; (j) ovipositor and spermatheca; (k) sternite VIII, female. Scale bars: a b = 0.5 mm; cf = 0.1 mm; gk = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5464849" httpUri="https://zenodo.org/record/5464849/files/figure.png" pageId="8" pageNumber="917">Figure 4</figureCitation>
(j)) heavily sclerotised, without stylus, baculus extremely long; spermatheca well sclerotised.
</paragraph>
<paragraph id="8BE236EAF70D76259189FD6FFA242582" blockId="8.[159,1156,142,757]" pageId="8" pageNumber="917">
<emphasis id="B929EAF8F70D76259189FD6FF9BD25A5" bold="true" box="[192,424,697,722]" pageId="8" pageNumber="917">Sexual dimorphism</emphasis>
not distinct, ventral side of apical tibiae and basal tarsomeres of fore legs with denser setae in male.
</paragraph>
<paragraph id="8BE236EAF70D762591E9FCE6F911243D" blockId="8.[160,1156,816,946]" box="[160,260,816,842]" pageId="8" pageNumber="917">
<emphasis id="B929EAF8F70D762591E9FCE6F911243D" bold="true" box="[160,260,816,842]" italics="true" pageId="8" pageNumber="917">Habitats</emphasis>
</paragraph>
<paragraph id="8BE236EAF70D762591E9FC82FA36241A" blockId="8.[160,1156,816,946]" box="[160,547,852,877]" pageId="8" pageNumber="917">Living in mud under seaside reefs.</paragraph>
</subSubSection>
<subSubSection id="C3476561F70D76249189FCA0FC9626A7" lastPageId="9" lastPageNumber="918" pageId="8" pageNumber="917" type="key">
<paragraph id="8BE236EAF70D76259189FCA0F94B24C5" blockId="8.[160,1156,816,946]" pageId="8" pageNumber="917">
<emphasis id="B929EAF8F70D76259189FCA0FC9124F8" bold="true" box="[192,1156,886,911]" pageId="8" pageNumber="917">
Key to species of
<taxonomicName id="4C5D4D69F70D762590FDFCA0FA6D24F8" ID-CoL="5R5P" authority="Spilman" authorityName="Spilman" authorityYear="1972" box="[436,632,886,911]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="917" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAF8F70D762590FDFCA0FA1324F8" bold="true" box="[436,518,886,911]" italics="true" pageId="8" pageNumber="917">Mexico</emphasis>
Spilman
</taxonomicName>
from the Oriental region, derived from
</emphasis>
<bibRefCitation id="EFCC4B1BF70D762591E9FC4FF94B24C5" author="Yoshitomi H" box="[160,350,921,946]" pageId="8" pageNumber="917" pagination="1 - 45" refId="ref4853" refString="Yoshitomi H. 2019. Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy. 583: 1 - 45." type="journal article" year="2019">Yoshitomi (2019)</bibRefCitation>
</paragraph>
<key id="FFCC61DBF70D762491F6FC08FC9626A7" lastPageId="9" lastPageNumber="918" pageId="8" pageNumber="917">
<keyStep id="30A92E4FF70D762591F6FC08FC962330" pageId="8" pageNumber="917">
<paragraph id="8BE236EAF70D762591F6FC08FC96236D" blockId="8.[191,1155,990,1050]" pageId="8" pageNumber="917">
<keyLead id="30AC95DFF70D762591F6FC08FC96236D" pageId="8" pageNumber="917">1. Median lobe with distinct subapical projections laterally, separated from ventral part....................... ......................................................................................................... 2</keyLead>
</paragraph>
<paragraph id="8BE236EAF70D762591F6FBF8FC962330" blockId="8.[191,1155,1070,1095]" box="[191,1155,1070,1095]" pageId="8" pageNumber="917">
<keyLead id="30AC95DFF70D762591F6FBF8FC962330" box="[191,1155,1070,1095]" pageId="8" pageNumber="917">- Median lobe simple, without subapical projections laterally.......................6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A92E4FF70D762591F6FB8AFC9623D5" pageId="8" pageNumber="917">
<paragraph id="8BE236EAF70D762591F6FB8AFC962302" blockId="8.[191,1155,1116,1141]" box="[191,1155,1116,1141]" pageId="8" pageNumber="917">
<keyLead id="30AC95DFF70D762591F6FB8AFC962302" box="[191,1155,1116,1141]" pageId="8" pageNumber="917">2. Subapical projections of median lobe distinctly curved outwardly.................. 3</keyLead>
</paragraph>
<paragraph id="8BE236EAF70D762591F6FB5FFC9623D5" blockId="8.[191,1155,1161,1186]" box="[191,1155,1161,1186]" pageId="8" pageNumber="917">
<keyLead id="30AC95DFF70D762591F6FB5FFC9623D5" box="[191,1155,1161,1186]" pageId="8" pageNumber="917">- Subapical projections of median lobe curved interiorly or ventrally................ 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A92E4FF70D762591F6FB60FC962235" pageId="8" pageNumber="917">
<paragraph id="8BE236EAF70D762591F6FB60FC962385" blockId="8.[191,1155,1206,1266]" pageId="8" pageNumber="917">
<keyLead id="30AC95DFF70D762591F6FB60FC962385" pageId="8" pageNumber="917">
3. Subapical projections of median lobe nearly equal in length, apex of median lobe slender and gradually acuminate;
<collectingCountry id="F34A767AF70D76259314FB0FFA8F2385" box="[605,666,1241,1266]" name="Palau" pageId="8" pageNumber="917">Palau</collectingCountry>
....................... .........
<emphasis id="B929EAF8F70D762592A6FB0FFC962385" box="[1007,1155,1241,1266]" italics="true" pageId="8" pageNumber="917">M. palauensis</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BE236EAF70D762591F6FAD0FC962235" blockId="8.[191,1155,1286,1346]" pageId="8" pageNumber="917">
<keyLead id="30AC95DFF70D762591F6FAD0FC962235" pageId="8" pageNumber="917">
- Subapical projections of median lobe with left part distinctly longer than right, apex of median lobe robust and sharply acuminate; Yongxing Island.......... ...
<taxonomicName id="4C5D4D69F70D76259574FAFFFC962235" authorityName="Liu &amp; Jia" authorityYear="2021" box="[1085,1155,1321,1346]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="917" phylum="Arthropoda" rank="species" species="xiei">
<emphasis id="B929EAF8F70D76259574FAFFFC962235" box="[1085,1155,1321,1346]" italics="true" pageId="8" pageNumber="917">M. xiei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A92E4FF70D762591F6FA80FC962295" pageId="8" pageNumber="917">
<paragraph id="8BE236EAF70D762591F6FA80FC9622E5" blockId="8.[191,1155,1366,1426]" pageId="8" pageNumber="917">
<keyLead id="30AC95DFF70D762591F6FA80FC9622E5" pageId="8" pageNumber="917">
4. Subapical projections of median lobe of same length, ventrally curved; apex of median lobe slender; Ogasawara Island.................. .........
<emphasis id="B929EAF8F70D762592FDFAAFFC9622E5" box="[948,1155,1401,1426]" italics="true" pageId="8" pageNumber="917">M. ogasawaraensis</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BE236EAF70D762591F6FA70FC962295" blockId="8.[191,1156,1446,1506]" pageId="8" pageNumber="917">
<keyLead id="30AC95DFF70D762591F6FA70FC962295" pageId="8" pageNumber="917">- Subapical projections of median lobe with left part distinctly longer than right, interiorly curved; apex of median lobe robust.......................................5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A92E4FF70D762591F6FA20FC9621F5" pageId="8" pageNumber="917">
<paragraph id="8BE236EAF70D762591F6FA20FC962145" blockId="8.[191,1155,1526,1586]" pageId="8" pageNumber="917">
<keyLead id="30AC95DFF70D762591F6FA20FC962145" pageId="8" pageNumber="917">
5. Subapical projections of median lobe short and small, widely separated from each other;
<collectingCountry id="F34A767AF70D76259061F9CFF96D2145" box="[296,376,1561,1586]" name="Taiwan" pageId="8" pageNumber="917">Taiwan</collectingCountry>
...... .....................................................................................................
<taxonomicName id="4C5D4D69F70D762592B7F9CFFC962145" box="[1022,1155,1561,1586]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="917" phylum="Arthropoda" rank="species" species="taiwanus">
<emphasis id="B929EAF8F70D762592B7F9CFFC962145" box="[1022,1155,1561,1586]" italics="true" pageId="8" pageNumber="917">M. taiwanus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE236EAF70D762591F6F990FC9621F5" blockId="8.[191,1156,1606,1666]" pageId="8" pageNumber="917">
<keyLead id="30AC95DFF70D762591F6F990FC9621F5" pageId="8" pageNumber="917">
- Subapical projections of median lobe large, nearly attached to each other at middle; Honshu to Ishigakijima.......................... .................... ...................
<taxonomicName id="4C5D4D69F70D7625954DF9BFFC9621F5" box="[1028,1155,1641,1666]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="917" phylum="Arthropoda" rank="species" species="masamii">
<emphasis id="B929EAF8F70D7625954DF9BFFC9621F5" box="[1028,1155,1641,1666]" italics="true" pageId="8" pageNumber="917">M. masamii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A92E4FF70C762491F6FF59FC96278F" pageId="9" pageNumber="918">
<paragraph id="8BE236EAF70C762491F6FF59FC9627BC" blockId="9.[191,1155,143,203]" pageId="9" pageNumber="918">
<keyLead id="30AC95DFF70C762491F6FF59FC9627BC" pageId="9" pageNumber="918">
6. Lateral lobes long, exceeding proximal half of median lobe, apex of lateral lobes with short setae;
<collectingCountry id="F34A767AF70C76249021FF64FA2B27BC" box="[360,574,178,203]" name="Papua New Guinea" pageId="9" pageNumber="918">Papua New Guinea</collectingCountry>
..................... ...............................
<emphasis id="B929EAF8F70C762492BCFF64FC9627BC" box="[1013,1155,178,203]" italics="true" pageId="9" pageNumber="918">M. papuanus</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BE236EAF70C762491F6FF09FC96278F" blockId="9.[191,1155,223,248]" box="[191,1155,223,248]" pageId="9" pageNumber="918">
<keyLead id="30AC95DFF70C762491F6FF09FC96278F" box="[191,1155,223,248]" pageId="9" pageNumber="918">- Lateral lobes shorter than half of median lobe, apex with long setae...............7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A92E4FF70C762491F6FEDAFC962602" pageId="9" pageNumber="918">
<paragraph id="8BE236EAF70C762491F6FEDAFC96263F" blockId="9.[191,1155,268,328]" pageId="9" pageNumber="918">
<keyLead id="30AC95DFF70C762491F6FEDAFC96263F" pageId="9" pageNumber="918">
7. Median lobe distinctly expanded subapically;
<collectingCountry id="F34A767AF70C76249392FEDAFB862652" box="[731,915,268,293]" name="Christmas Island" pageId="9" pageNumber="918">Christmas Island</collectingCountry>
........... .................. ..............................................................................................................................................
<taxonomicName id="4C5D4D69F70C762492B3FEF9FC96263F" authorityName="Liu &amp; Jia" authorityYear="2021" box="[1018,1155,303,328]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="918" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis id="B929EAF8F70C762492B3FEF9FC96263F" box="[1018,1155,303,328]" italics="true" pageId="9" pageNumber="918">M. lawrencei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE236EAF70C762491F6FE8AFC962602" blockId="9.[191,1155,348,373]" box="[191,1155,348,373]" pageId="9" pageNumber="918">
<keyLead id="30AC95DFF70C762491F6FE8AFC962602" box="[191,1155,348,373]" pageId="9" pageNumber="918">- Median lobe not distinctly expanded subapically...................................8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A92E4FF70C762491F6FE5CFC9626A7" pageId="9" pageNumber="918">
<paragraph id="8BE236EAF70C762491F6FE5CFC9626D4" blockId="9.[191,1155,394,419]" box="[191,1155,394,419]" pageId="9" pageNumber="918">
<keyLead id="30AC95DFF70C762491F6FE5CFC9626D4" box="[191,1155,394,419]" pageId="9" pageNumber="918">
8. Apex of median lobe nipple-like; Borneo...........................
<emphasis id="B929EAF8F70C762492A7FE5CFC9626D4" box="[1006,1155,394,419]" italics="true" pageId="9" pageNumber="918">M. borneensis</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BE236EAF70C762491F6FE61FC9626A7" blockId="9.[191,1155,439,464]" box="[191,1155,439,464]" pageId="9" pageNumber="918">
<keyLead id="30AC95DFF70C762491F6FE61FC9626A7" box="[191,1155,439,464]" pageId="9" pageNumber="918">
- Apex of median lobe simply pointed; Bali......................... ... ..
<emphasis id="B929EAF8F70C7624954CFE61FC9626A7" box="[1029,1155,439,464]" italics="true" pageId="9" pageNumber="918">M. baliensis</emphasis>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>