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<document id="9568C6C6CAE97E4C16098204AE5E9C61" ID-DOI="10.11646/zootaxa.3949.4.1" ID-GBIF-Dataset="39ca09d7-bca6-4ac0-9dee-5bd38d66ff3c" ID-ISSN="1175-5326" ID-Zenodo-Dep="243464" ID-ZooBank="B3123099-882F-4C42-B83B-2BA1C2906F65" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461047353794" checkinUser="plazi" docAuthor="Hämäläinen, Matti, Dow, Rory A. &amp; Stokvis, Frank R." docDate="2015" docId="03F487E5FFC1FFFEFF780C454D219832" docLanguage="en" docName="zt03949p490.pdf" docOrigin="Zootaxa 3949 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dysphaea ulu Hämäläinen, Dow &amp; Stokvis, 2015, spec. nov." docType="treatment" docVersion="11" lastPageNumber="476" masterDocId="FFCDFF9DFFD4FFE7FFEF0B31484B9F1D" masterDocTitle="Revision of the Sundaland species of the genus Dysphaea Selys, 1853 using molecular and morphological methods, with notes on allied species (Odonata: Euphaeidae)" masterLastPageNumber="490" masterPageNumber="451" pageNumber="472" updateTime="1698577609733" updateUser="plazi">
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<mods:title id="BA9D1598E2FF3D7A0C5172090737E53F">Revision of the Sundaland species of the genus Dysphaea Selys, 1853 using molecular and morphological methods, with notes on allied species (Odonata: Euphaeidae)</mods:title>
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<taxonomicName id="4C5D4D70FFC1FFF2FF780C4549759893" ID-CoL="9WTWD" box="[151,318,1908,1934]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="ulu" status="sp. nov.">
<emphasis id="B929EAE1FFC1FFF2FF780C4549759893" bold="true" box="[151,318,1908,1934]" italics="true" pageId="21" pageNumber="472">Dysphaea ulu</emphasis>
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(
<figureCitation id="13662A76FFC1FFF2FF700CA648B098B2" box="[159,251,1943,1968]" captionStart="FIGURES 9 – 12" captionStartId="14.[151,264,1350,1373]" captionTargetBox="[151,1436,373,1328]" captionTargetId="figure@14.[151,1436,373,1330]" captionTargetPageId="14" captionText="FIGURES 9 – 12. Habitus of male Dysphaea: 9) D. dimidiata West Java; 10) D. lugens Danum Valley, Sabah; 11) D. ulu holotype (flipped horizontally); 12) D. vanida holotype (flipped horizontally)." httpUri="https://zenodo.org/record/243466/files/figure.png" pageId="21" pageNumber="472">Figs. 11</figureCitation>
,
<figureCitation id="13662A76FFC1FFF2FEE80CA6496E98AD" box="[263,293,1943,1968]" captionStart="FIGURES 13 – 18" captionStartId="15.[151,264,1951,1974]" captionTargetBox="[151,1436,517,1918]" captionTargetId="figure@15.[151,1436,517,1931]" captionTargetPageId="15" captionText="FIGURES 13 – 18. Synthorax male Dysphaea: 13) D. dimidiata West Java; 14) D. lugens Bakuan, Kalimantan Barat; 15) D. ulu holotype; 16) D. vanida holotype; 17) metaposternum, D. dimidiata Brunei; 18) metaposternum, D. ulu holotype." httpUri="https://zenodo.org/record/243467/files/figure.png" pageId="21" pageNumber="472">15</figureCitation>
,
<figureCitation id="13662A76FFC1FFF2FEDE0CA6490498B2" box="[305,335,1943,1967]" captionStart="FIGURES 25 – 30" captionStartId="18.[151,264,1692,1715]" captionTargetBox="[151,1436,193,1669]" captionTargetId="figure@18.[151,1436,193,1671]" captionTargetPageId="18" captionText="FIGURES 25 – 30. Wings of Dysphaea males: 25) D. lugens Pemantang, Kalimantan Tengah; 26) D. lugens Ampah, Kalimantan Tengah; 27) D. lugens Selekau River, Kalimantan Barat; 28) D. ulu paratype, Temburong, Brunei; 29) D. vanida spec. nov. paratype, Khlong Nakha, Ranong; 30) D. vanida paratype, Khlong Nakha, Ranong." httpUri="https://zenodo.org/record/243469/files/figure.png" pageId="21" pageNumber="472">28</figureCitation>
,
<figureCitation id="13662A76FFC1FFF2FEB40CA6493298B2" box="[347,377,1943,1967]" captionStart="FIGURES 31 – 36" captionStartId="19.[151,264,1705,1728]" captionTargetBox="[157,1436,185,1683]" captionTargetId="figure@19.[157,1442,185,1684]" captionTargetPageId="19" captionText="FIGURES 31 – 36. Male anal appendages. Dorsal view: 31) D. dimidiata West Java; 32) D. lugens Bakuan, Kalimantan Barat; 33) D. ulu holotype; 34) D. vanida holotype. Dorso-lateral views: 35) D. dimidiata Terengganu; 36) D. vanida paratype Lam Khlong Gnu, Kanchanaburi." httpUri="https://zenodo.org/record/243470/files/figure.png" pageId="21" pageNumber="472">33</figureCitation>
,
<figureCitation id="13662A76FFC1FFF2FE6A0CA649E898B2" box="[389,419,1943,1967]" captionStart="FIGURES 37 – 40" captionStartId="20.[151,264,1070,1093]" captionTargetBox="[162,1392,232,1036]" captionTargetId="figure@20.[151,1436,193,1050]" captionTargetPageId="20" captionText="FIGURES 37 – 40. Male anal appendages, lateral view (scale bars 1 mm): 37) D. dimidiata west Java; 38) D. lugens Bakuan, Kalimantan Barat; 39) D. ulu holotype; 40) D. vanida holotype." httpUri="https://zenodo.org/record/243471/files/figure.png" pageId="21" pageNumber="472">39</figureCitation>
,
<figureCitation id="13662A76FFC1FFF2FE400CA6498698B2" box="[431,461,1943,1967]" captionStart="FIGURES 41 – 44" captionStartId="21.[151,264,1790,1813]" captionTargetBox="[162,1384,617,1756]" captionTargetId="figure@21.[151,1436,555,1769]" captionTargetPageId="21" captionText="FIGURES 41 – 44. Penis, ventral view: 41) D. dimidiata west Java; 42) D. lugens Selekau River, Kalimantan Barat; 43) D. ulu holotype; 44) D. vanida paratype, Khlong Nakha, Ranong." httpUri="https://zenodo.org/record/243472/files/figure.png" pageId="21" pageNumber="472">43</figureCitation>
,
<figureCitation id="13662A76FFC1FFF2FE360CA649BC98B2" box="[473,503,1943,1967]" captionStart="FIGURES 45 – 48" captionStartId="23.[151,264,1735,1758]" captionTargetBox="[162,1365,783,1707]" captionTargetId="figure@23.[151,1436,721,1720]" captionTargetPageId="23" captionText="FIGURES 45 – 48. Penis, lateral view: 45) D. dimidiata west Java; 46) D. lugens Selekau River, Kalimantan Barat; 47) D. ulu holotype; 48) D. vanida paratype, Khlong Nakha, Ranong." httpUri="https://zenodo.org/record/243473/files/figure.png" pageId="21" pageNumber="472">47</figureCitation>
,
<figureCitation id="13662A76FFC1FFF2FDEC0CA64A6A98AD" box="[515,545,1943,1968]" captionStart="FIGURES 49 – 52" captionStartId="24.[151,264,1070,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@24.[151,1436,193,1049]" captionTargetPageId="24" captionText="FIGURES 49 – 52. Habitus of female Dysphaea: 49) D. dimidiata west Java; 50) D. lugens Mount Dulit, Sarawak, photograph by Benjamin Price; 51) D. ulu paratype; 52) D. vanida paratype Kanchanaburi." httpUri="https://zenodo.org/record/243474/files/figure.png" pageId="21" pageNumber="472">51</figureCitation>
,
<figureCitation id="13662A76FFC1FFF2FDC20CA64A0798AD" box="[557,588,1943,1968]" captionStart="FIGURES 53 – 56" captionStartId="25.[151,264,1609,1632]" captionTargetBox="[151,1436,625,1588]" captionTargetId="figure@25.[151,1436,625,1588]" captionTargetPageId="25" captionText="FIGURES 53 – 56. Female head: 53) D. dimidiata Sarawak, dorsal view; 54) D. lugens Gunung Sari, Kalimantan Timur, dorsal view; 55) D. ulu paratype, dorsal, slightly frontal view; 56) D. vanida paratype, Kanchanaburi, dorsal, slightly frontal view." httpUri="https://zenodo.org/record/243475/files/figure.png" pageId="21" pageNumber="472">55</figureCitation>
,
<figureCitation id="13662A76FFC1FFF2FDB80CA64A3E98AD" box="[599,629,1943,1968]" captionStart="FIGURES 57 – 62" captionStartId="27.[151,264,1913,1936]" captionTargetBox="[155,1436,401,1899]" captionTargetId="figure@27.[155,1440,401,1900]" captionTargetPageId="27" captionText="FIGURES 57 – 62. Female prothorax, dorsal view: 57) D. dimidiata west Java; 58) D. dimidiata Sarawak; 59) D. dimidiata Tandjong Pandan, Billiton; 60) D. lugens Gunung Sari, Kaliamantan Timur; 61) D. ulu paratype; 62) D. vanida paratype, Kanchanaburi." httpUri="https://zenodo.org/record/243476/files/figure.png" pageId="21" pageNumber="472">61</figureCitation>
,
<figureCitation id="13662A76FFC1FFF2FD6E0CA64AD498AD" box="[641,671,1943,1968]" captionStart="FIGURES 63 – 68" captionStartId="28.[151,264,1699,1722]" captionTargetBox="[151,1436,185,1683]" captionTargetId="figure@28.[151,1436,185,1684]" captionTargetPageId="28" captionText="FIGURES 63 – 68. Female synthorax, dorsal view: 63) D. dimidiata west Java; 64) D. dimidiata Sarawak; 65) D. dimidiata Johor; 66) D. lugens Gunung Sari, Kaliamantan Timur; 67) D. ulu paratype; 68) D. vanida paratype Kanchanaburi." httpUri="https://zenodo.org/record/243477/files/figure.png" pageId="21" pageNumber="472">67</figureCitation>
,
<figureCitation id="13662A76FFC1FFF2FD440CA64A8298B2" box="[683,713,1943,1967]" captionStart="FIGURES 69 – 74" captionStartId="29.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1797]" captionTargetId="figure@29.[151,1436,193,1799]" captionTargetPageId="29" captionText="FIGURES 69 – 74. Female synthorax, lateral view: 69) D. dimidiata west Java; 70) D. dimidiata Sarawak; 71) D. dimidiata Johor; 72) D. lugens Mount Dulit, Sarawak, photograph by Benjamin Price; 73) D. ulu paratype; 74) D. vanida paratype Kanchanaburi." httpUri="https://zenodo.org/record/243478/files/figure.png" pageId="21" pageNumber="472">73</figureCitation>
,
<figureCitation id="13662A76FFC1FFF2FD3A0CA64ABF98B2" box="[725,756,1943,1967]" captionStart="FIGURES 77 – 82" captionStartId="31.[151,264,1570,1593]" captionTargetBox="[151,1436,193,1548]" captionTargetId="figure@31.[151,1436,193,1549]" captionTargetPageId="31" captionText="FIGURES 77 – 82. Dysphaea males in life: 77) D. basitincta Hainan (photo by Mo Shanlian) 78) D. dimidiata Sarawak (photo by Graham T. Reels); 79) D. dimidiata Negeri Sembilan (photo by C. Y. Choong); 80) D. ulu Sarawak (photo by Graham T. Reels); 81) D. vanida Ranong (photo by Matti Hämäläinen); 82) D. vanida Surat Thani (photo by André Günther)." httpUri="https://zenodo.org/record/243480/files/figure.png" pageId="21" pageNumber="472">80</figureCitation>
,
<figureCitation id="13662A76FFC1FFF2FD100CA64B5798AD" box="[767,796,1943,1968]" captionStart="FIGURE 85" captionStartId="34.[151,250,2012,2034]" captionTargetBox="[159,1436,645,1988]" captionTargetId="figure@34.[159,1444,631,1989]" captionTargetPageId="34" captionText="FIGURE 85. Distribution map of Dysphaea ulu." httpUri="https://zenodo.org/record/243483/files/figure.png" pageId="21" pageNumber="472">85</figureCitation>
)
</paragraph>
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<taxonomicName id="4C5D4D70FFC1FFF2FF780CEE492D98E8" ID-CoL="6DSLB" box="[151,358,2015,2037]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B929EAE1FFC1FFF2FF780CEE492D98E8" box="[151,358,2015,2037]" italics="true" pageId="21" pageNumber="472">Dysphaea dimidiata</emphasis>
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[nec
<bibRefCitation id="EFCC4B02FFC1FFF2FE4B0CEE4A5798E8" author="Selys" box="[420,540,2015,2037]" pageId="21" pageNumber="472" refString="Selys Longchamps, E. de (1853) Synopsis des Calopterygines. Bulletins de l'Academie royale des sciences, des lettres et des beaux-arts de Belgique, 20 (Annexe), 1 - 73." type="journal article" year="1853">Selys, 1853</bibRefCitation>
];—
<bibRefCitation id="EFCC4B02FFC1FFF2FDAE0CEE4CCA98E8" author="Schmidt" box="[577,1153,2015,2037]" pageId="21" pageNumber="472" refString="Schmidt, E. (1934) Odonata der Deutschen Limnologischen Sunda-Expedition. 1. Imagines. Mit Beschreibungen zweier neuer Ictinus aus Celebes und Neu - Guinea. Archiv fur Hydrobiologie, Supplement - Band, 13, 316 - 397, plates 14 - 17, excl." type="journal article" year="1934">Schmidt (1934: 330–331, part: Baram, plate 16, fig. 8)</bibRefCitation>
;—
<bibRefCitation id="EFCC4B02FFC1FFF1FB700CEE491B9FB3" author="Lieftinck" lastPageId="22" lastPageNumber="473" pageId="21" pageNumber="472" refString="Lieftinck, M. A. (1954) Handlist of Malaysian Odonata. A catalogue of the dragonflies of the Malay Peninsula, Sumatra, Java and Borneo, including the adjacent small islands. Treubia, 22 (Supplement), i - xiii + 1 - 202, 1 folded map excl." type="journal article" year="1954">Lieftinck (1954: 19–20, part: Borneo)</bibRefCitation>
;—
<bibRefCitation id="EFCC4B02FFC2FFF1FE810BA949929FB3" author="Orr" box="[366,473,152,174]" pageId="22" pageNumber="473" refString="Orr, A. G. (2001) An annotated checklist of the Odonata of Brunei with ecological notes and descriptions of hitherto unknown males and larvae. International Journal of Odonatology, 4, 167 - 220. http: // dx. doi. org / 10.1080 / 13887890.2001.9748168" type="journal article" year="2001">Orr (2001</bibRefCitation>
: 178, specimens from Kuala Belalong Field Studies Centre, Sites 1,2; except one male which was
<taxonomicName id="4C5D4D70FFC2FFF1FF1B0B8949A59FD1" ID-CoL="6DSLB" authority="Selys, 1853" authorityName="Selys" authorityYear="1853" box="[244,494,182,205]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B929EAE1FFC2FFF1FF1B0B89493F9FD1" box="[244,372,182,205]" italics="true" pageId="22" pageNumber="473">D. dimidiata</emphasis>
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<bibRefCitation id="EFCC4B02FFC2FFF1FDFA0B874ACB9FD1" author="Orr" box="[533,640,182,204]" pageId="22" pageNumber="473" refString="Orr, A. G. (2003) A guide to the dragonflies of Borneo: their identification and biology. Natural History Publications (Borneo), Kota Kinabalu, x + 195 pp." type="book" year="2003">Orr (2003</bibRefCitation>
: 37, part; 58 part; p. 59, fig. 72; plate 3h);—
<bibRefCitation id="EFCC4B02FFC2FFF1FBD50B874D079FD1" author="Orr" box="[1082,1356,182,204]" pageId="22" pageNumber="473" refString="Orr, A. G. (2005) A pocket guide: Dragonflies of Peninsular Malaysia and Singapore. Natural History Publications (Borneo), Kota Kinabalu, vi + 127 pp." type="book" year="2005">Orr (2005: 26, illustration)</bibRefCitation>
;—
<bibRefCitation id="EFCC4B02FFC2FFF1FA850B874AEF9FF6" author="Dow" pageId="22" pageNumber="473" refString="Dow, R. A. (2005) Odonata, burglary and ballistic cicadas in South-east Asia. Agrion, 9 (1), 10 - 12." type="journal article" year="2005">Dow (2005: 11, Gunung Mulu National Park, part)</bibRefCitation>
;—
<bibRefCitation id="EFCC4B02FFC2FFF1FD2C0BE74D659FF6" author="Dow" box="[707,1326,213,235]" pageId="22" pageNumber="473" refString="Dow, R. A. &amp; Reels, G. T. (2008) List of species recorded at Gunung Mulu National Park, Sarawak, Malaysian Borneo in 2005 - 2006. Echo, 5, 2 - 3 [in Agrion, 12 (1)]." type="journal article" year="2008">Dow &amp; Reels (2008: 3, Gunung Mulu National Park, part)</bibRefCitation>
;—
<bibRefCitation id="EFCC4B02FFC2FFF1FAA20BE749B59E17" author="Dow" pageId="22" pageNumber="473" refString="Dow, R. A. &amp; Reels, G. T. (2009) Expedition to Mount Dulit, Sarawak, August - September 2008 - Odonata. International Dragonfly Fund Report, 19, 1 - 16." type="journal article" year="2009">Dow &amp; Reels (2009: 13, Mount Dulit)</bibRefCitation>
;—
<bibRefCitation id="EFCC4B02FFC2FFF1FDF20BC54C7C9E17" author="Dow" box="[541,1079,244,266]" pageId="22" pageNumber="473" refString="Dow, R. A. &amp; Reels, G. T. (2010) The Odonata of three National Parks in Sarawak. Agrion, 14 (1), 14 - 19." type="journal article" year="2010">Dow &amp; Reels (2010: 15, Kubah National Park, part)</bibRefCitation>
;—
<bibRefCitation id="EFCC4B02FFC2FFF1FBB90BC44D359E17" author="Dow" box="[1110,1406,244,266]" pageId="22" pageNumber="473" refString="Dow, R. A., Reels, G. T. &amp; Butler, S. G. (2013 a) Odonata of the Dulit Range in Sarawak, Malaysian Borneo. Notulae odonatologicae, 8 (1), 8 - 14." type="journal article" year="2013" yearSuffix="a">
Dow,
<bibRefCitation id="EFCC4B02FFC2FFF1FB7C0BC54D359E17" author="Reels" box="[1171,1406,244,266]" pageId="22" pageNumber="473" refString="Dow, R. A., Reels, G. T. &amp; Butler, S. G. (2013 a) Odonata of the Dulit Range in Sarawak, Malaysian Borneo. Notulae odonatologicae, 8 (1), 8 - 14." type="journal article" year="2013" yearSuffix="a">Reels &amp; Butler (2013a</bibRefCitation>
</bibRefCitation>
: 9, Dulit range);—
<bibRefCitation id="EFCC4B02FFC2FFF1FE8E0A224ACC9E35" author="Dow" box="[353,647,274,296]" pageId="22" pageNumber="473" refString="Dow, R. A., Reels, G. T. &amp; Butler, S. G. (2013 b) Previously unpublished Odonata records from Sarawak, Borneo. Part II. Kubah National Park. Faunistic Studies in South-East Asian and Pacific Island Odonata, 6, 1 - 21." type="journal article" year="2013" yearSuffix="b">
Dow,
<bibRefCitation id="EFCC4B02FFC2FFF1FE710A234ACC9E35" author="Reels" box="[414,647,274,296]" pageId="22" pageNumber="473" refString="Dow, R. A., Reels, G. T. &amp; Butler, S. G. (2013 b) Previously unpublished Odonata records from Sarawak, Borneo. Part II. Kubah National Park. Faunistic Studies in South-East Asian and Pacific Island Odonata, 6, 1 - 21." type="journal article" year="2013" yearSuffix="b">Reels &amp; Butler (2013b</bibRefCitation>
</bibRefCitation>
: 11, Kubah National Park, part).
</paragraph>
<paragraph id="8BE236F3FFC2FFF1FF780A004C2C9E5A" blockId="22.[151,1436,152,358]" box="[151,1127,305,327]" pageId="22" pageNumber="473">
<treatmentCitationGroup id="AB4D11DDFFC2FFF1FF780A004C2C9E5A" box="[151,1127,305,327]" pageId="22" pageNumber="473">
<taxonomicName id="4C5D4D70FFC2FFF1FF780A0049999E5A" ID-CoL="6DSL8" box="[151,466,305,327]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="473" phylum="Arthropoda" rank="subSpecies" species="limbata" subSpecies="semilimbata">
<emphasis id="B929EAE1FFC2FFF1FF780A0049999E5A" box="[151,466,305,327]" italics="true" pageId="22" pageNumber="473">Dysphaea limbata semilimbata</emphasis>
</taxonomicName>
[nec
<bibRefCitation id="EFCC4B02FFC2FFF1FDE50A004A349E5A" author="Selys" box="[522,639,305,327]" pageId="22" pageNumber="473" refString="Selys Longchamps, E. de (1873) Troisiemes additions au synopsis de Calopterygines. Bulletins de l'Academie royale des sciences, des lettres et des beaux-arts de Belgique, Series 2, 35, 469 - 519." type="journal article" year="1873">Selys, 1873</bibRefCitation>
];—
<treatmentCitation id="0AFC10E2FFC2FFF1FD4B0A004B239E5A" author="Laidlaw" box="[676,872,305,327]" page="328" pageId="22" pageNumber="473" year="1920">
<bibRefCitation id="EFCC4B02FFC2FFF1FD4B0A004B739E5A" author="Laidlaw" box="[676,824,305,327]" pageId="22" pageNumber="473" refString="Laidlaw, F. F. (1920) Contributions to the study of the dragonfly fauna of Borneo. Part IV. A list of species known to occur in the island. Proceedings of the Zoological Society of London, 1920, 311 - 342. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1920. tb 07074. x" type="proceedings paper" year="1920">Laidlaw 1920</bibRefCitation>
: 328
</treatmentCitation>
(specimen from Baram).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE236F3FFC2FFF1FF780A614B709E7B" blockId="22.[151,1436,152,358]" box="[151,827,336,358]" pageId="22" pageNumber="473">
<treatmentCitationGroup id="AB4D11DDFFC2FFF1FF780A614B709E7B" box="[151,827,336,358]" pageId="22" pageNumber="473">
<taxonomicName id="4C5D4D70FFC2FFF1FF780A6148B79E7B" ID-CoL="496W" box="[151,252,336,358]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="473" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAE1FFC2FFF1FF780A6148B79E7B" box="[151,252,336,358]" italics="true" pageId="22" pageNumber="473">Dysphaea</emphasis>
</taxonomicName>
species;—
<treatmentCitation id="0AFC10E2FFC2FFF1FE840A614AE59E7B" author="Dow" box="[363,686,336,358]" page="11" pageId="22" pageNumber="473" year="2015">
<bibRefCitation id="EFCC4B02FFC2FFF1FE840A614AC19E7B" author="Dow" box="[363,650,336,358]" pageId="22" pageNumber="473" refString="Dow, R. A., Reels, G. T. &amp; Ngiam, R. W. J. (2015) Odonata collected at Usun Apau National Park, Miri Division, Sarawak, Malaysia in April and May 2012. International Dragonfly Fund Report, 79, 1 - 17." type="journal article" year="2015">
Dow,
<bibRefCitation id="EFCC4B02FFC2FFF1FE470A614AC19E7B" author="Reels" box="[424,650,336,358]" pageId="22" pageNumber="473" refString="Dow, R. A., Reels, G. T. &amp; Ngiam, R. W. J. (2015) Odonata collected at Usun Apau National Park, Miri Division, Sarawak, Malaysia in April and May 2012. International Dragonfly Fund Report, 79, 1 - 17." type="journal article" year="2015">Reels &amp; Ngiam (2015</bibRefCitation>
</bibRefCitation>
: 11
</treatmentCitation>
, Usun Apau).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3476578FFC2FFF1FF780AA34B5A99A6" pageId="22" pageNumber="473" type="materials_examined">
<paragraph id="8BE236F3FFC2FFF1FF780AA34B5A99A6" blockId="22.[151,1437,402,2011]" pageId="22" pageNumber="473">
<emphasis id="B929EAE1FFC2FFF1FF780AA349289EB6" bold="true" box="[151,355,402,427]" pageId="22" pageNumber="473">Material studied</emphasis>
:
<emphasis id="B929EAE1FFC2FFF1FE980AA349AF9EB6" bold="true" box="[375,484,402,427]" pageId="22" pageNumber="473">
<typeStatus id="54E68851FFC2FFF1FE980AA349AF9EB6" box="[375,484,402,427]" pageId="22" pageNumber="473" type="holotype">Holotype</typeStatus>
</emphasis>
♂ (SAR13_14_EUP15): Borneo,
<collectingCountry id="F34A7663FFC2FFF1FC930AA34B949EB6" box="[892,991,402,427]" name="Malaysia" pageId="22" pageNumber="473">Sarawak</collectingCountry>
, Miri division, Upper Baram, Sungai Pejelai,
<taxonomicName id="4C5D4D70FFC2FFF1FF1B0A8749559ED2" box="[244,286,438,463]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="486" phylum="Arthropoda" rank="species" species="ulu">Ulu</taxonomicName>
Moh,
<date id="FFE31033FFC2FFF1FE810A8649B59ED2" box="[366,510,438,463]" pageId="22" pageNumber="473" value="2014-08-24">24 viii 2014</date>
, leg. M. Tegong. Deposited at
<collectionCode id="ED4CAE36FFC2FFF1FC880A864BF59ED3" LSID="urn:lsid:biocol.org:col:34992" box="[871,958,439,462]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
, Leiden.
<emphasis id="B929EAE1FFC2FFF1FBC50A874CE99ED2" bold="true" box="[1066,1186,438,463]" pageId="22" pageNumber="473">
<typeStatus id="54E68851FFC2FFF1FBC50A874CE99ED2" box="[1066,1186,438,463]" pageId="22" pageNumber="473" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D5BFD7AFFC2FFF1FB5B0A864CBB9ED3" box="[1204,1264,439,463]" pageId="22" pageNumber="473" type="generic">47 ♂</specimenCount>
, 2 ♀; all from Borneo; deposited in Coll. Dow unless otherwise stated).—From
<emphasis id="B929EAE1FFC2FFF1FC690AEB4BA49EEE" bold="true" box="[902,1007,474,499]" pageId="22" pageNumber="473">
<collectingCountry id="F34A7663FFC2FFF1FC690AEB4BA49EEE" box="[902,1007,474,499]" name="Malaysia" pageId="22" pageNumber="473">Sarawak</collectingCountry>
</emphasis>
: 1 ♀ (SAR13_14_EUP15, in tandem with
<typeStatus id="54E68851FFC2FFF1FF3F0ACF497D9D0A" box="[208,310,510,535]" pageId="22" pageNumber="473" type="holotype">holotype</typeStatus>
),
<collectingCountry id="F34A7663FFC2FFF1FEA60ACF49E79D0A" box="[329,428,510,535]" name="Malaysia" pageId="22" pageNumber="473">Sarawak</collectingCountry>
, Miri division, Upper Baram, Sungai Pejelai,
<taxonomicName id="4C5D4D70FFC2FFF1FC410ACF4B939D0A" box="[942,984,510,535]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="486" phylum="Arthropoda" rank="species" species="ulu">Ulu</taxonomicName>
Moh,
<date id="FFE31033FFC2FFF1FBCC0ACE4CE69D0A" box="[1059,1197,510,535]" pageId="22" pageNumber="473" value="2014-08-24">24 viii 2014</date>
, leg. M. Tegong;
<specimenCount id="9D5BFD7AFFC2FFF1FA9D0ACE4DD79D0B" box="[1394,1436,511,535]" pageId="22" pageNumber="473" type="generic">3 ♂</specimenCount>
(SAR13_14_EUP24–26), same location and date as above, leg. N. Sigau;
<specimenCount id="9D5BFD7AFFC2FFF1FC3B09124BB59D27" box="[980,1022,547,571]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR13_14_EUP23), same location as above,
<date id="FFE31033FFC2FFF1FEE9097749DA9D42" box="[262,401,582,607]" pageId="22" pageNumber="473" value="2014-08-26">26 viii 2014</date>
, leg. M. Tegong, E. Aran &amp; N. Sigau;
<specimenCount id="9D5BFD7AFFC2FFF1FCAE09764B209D43" box="[833,875,583,607]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR13_14_EUP29), same location as above,
<date id="FFE31033FFC2FFF1FA6F097648B49D9E" pageId="22" pageNumber="473" value="2014-08-27">27 viii 2014</date>
, leg. E. Aran &amp; O. Udau;
<specimenCount id="9D5BFD7AFFC2FFF1FDC1095A4A129D9F" box="[558,601,619,643]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR13_14_EUP19),
<collectingCountry id="F34A7663FFC2FFF1FCB0095B4B889D9E" box="[863,963,618,643]" name="Malaysia" pageId="22" pageNumber="473">Sarawak</collectingCountry>
, Miri division, Upper Baram, tributary of Sungai Pejelai,
<taxonomicName id="4C5D4D70FFC2FFF1FEAA09BF49249DBA" box="[325,367,654,679]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="486" phylum="Arthropoda" rank="species" species="ulu">Ulu</taxonomicName>
Moh,
<date id="FFE31033FFC2FFF1FE5709BE4A099DBA" box="[440,578,654,679]" pageId="22" pageNumber="473" value="2014-08-23">23 viii 2014</date>
, leg. Y. Tegong;
<specimenCount id="9D5BFD7AFFC2FFF1FD1309BE4B6D9DBB" box="[764,806,655,679]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR13_14_EUP35), same area as above, Sungai Moh,
<date id="FFE31033FFC2FFF1FF780982496B9DD6" box="[151,288,690,715]" pageId="22" pageNumber="473" value="2014-08-24">24 viii 2014</date>
, leg. R.A. Dow;
<specimenCount id="9D5BFD7AFFC2FFF1FE3609824A489DD7" box="[473,515,691,715]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR13_14_EUP27), same area as above, Sungai Sii,
<date id="FFE31033FFC2FFF1FB8709824CA29DD6" box="[1128,1257,690,715]" pageId="22" pageNumber="473" value="2014-07-10">10 vii 2014</date>
, leg. R.A. Dow;
<specimenCount id="9D5BFD7AFFC2FFF1FF7809E6488B9DF3" box="[151,192,727,751]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR13_14_EUP34), same location as above,
<date id="FFE31033FFC2FFF1FD3E09E64B199DF2" box="[721,850,726,751]" pageId="22" pageNumber="473" value="2014-07-12">12 vii 2014</date>
, leg. R.W.J. Ngiam;
<specimenCount id="9D5BFD7AFFC2FFF1FBD809E64C2A9DF3" box="[1079,1121,727,751]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR13_14_EUP36), same location as above,
<date id="FFE31033FFC2FFF1FE8909CA49A39C0E" box="[358,488,762,787]" pageId="22" pageNumber="473" value="2014-07-13">13 vii 2014</date>
, leg. N. Pawi;
<specimenCount id="9D5BFD7AFFC2FFF1FD6509CA4AFF9C0F" box="[650,692,763,787]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR13_14_EUP31), same location as above,
<date id="FFE31033FFC2FFF1FB2A09CB4D0C9C0E" box="[1221,1351,762,787]" pageId="22" pageNumber="473" value="2014-07-15">15 vii 2014</date>
, leg. O. Udau;
<specimenCount id="9D5BFD7AFFC2FFF1FF0C082E49469C2B" box="[227,269,799,823]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR13_14_EUP17), same location as above,
<date id="FFE31033FFC2FFF1FCCB082E4BE39C2A" box="[804,936,798,823]" pageId="22" pageNumber="473" value="2014-07-17">17 vii 2014</date>
, leg. O. Udau;
<specimenCount id="9D5BFD7AFFC2FFF1FBBC082E4C359C2B" box="[1107,1150,799,823]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR09_10_EUP142, in
<collectionCode id="ED4CAE36FFC2FFF1FF78087248BB9C47" LSID="urn:lsid:biocol.org:col:34992" box="[151,240,835,858]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
:
<collectionCode id="ED4CAE36FFC2FFF1FEED087349179C46" LSID="urn:lsid:biocol.org:col:34992" box="[258,348,834,859]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
.INS.503424),
<collectingCountry id="F34A7663FFC2FFF1FDEA08734A239C46" box="[517,616,834,859]" name="Malaysia" pageId="22" pageNumber="473">Sarawak</collectingCountry>
, Miri division, Upper Baram, Sungai Pawan,
<date id="FFE31033FFC2FFF1FB4B08724D739C46" box="[1188,1336,834,859]" pageId="22" pageNumber="473" value="2010-07-19">19 vii 2010</date>
, leg. L. Southwell;
<specimenCount id="9D5BFD7AFFC2FFF1FEF8085649099C63" box="[279,322,871,895]" pageId="22" pageNumber="473" type="generic">2 ♂</specimenCount>
(SAR06_EUP18–19),
<collectingCountry id="F34A7663FFC2FFF1FDAA08574AE39C62" box="[581,680,870,895]" name="Malaysia" pageId="22" pageNumber="473">Sarawak</collectingCountry>
, Miri division, foot of Mount Dulit, Sungai Nuam,
<date id="FFE31033FFC2FFF1FB0408564D239C62" box="[1259,1384,870,895]" pageId="22" pageNumber="473" value="2006-03-30">30 iii 2006</date>
, leg. R.A. Dow;
<specimenCount id="9D5BFD7AFFC2FFF1FEF508BA490D9CBF" box="[282,326,907,931]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR06_EUP17, in
<collectionCode id="ED4CAE36FFC2FFF1FDD908BA4AC49CBF" LSID="urn:lsid:biocol.org:col:34992" box="[566,655,907,930]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
:
<collectionCode id="ED4CAE36FFC2FFF1FD7508BB4ABF9CBE" LSID="urn:lsid:biocol.org:col:34992" box="[666,756,906,931]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
.INS.509905), same area as above, Sungai Long Aton,
<date id="FFE31033FFC2FFF1FA8F08BA48999CDA" pageId="22" pageNumber="473" value="2006-03-31">31 iii 2006</date>
, leg. G.T. Reels;
<specimenCount id="9D5BFD7AFFC2FFF1FE7B089E498B9CDB" box="[404,448,943,967]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR09_10_EUP100; Coll. Hämäläinen),
<collectingCountry id="F34A7663FFC2FFF1FC45089F4C469CDA" box="[938,1037,942,967]" name="Malaysia" pageId="22" pageNumber="473">Sarawak</collectingCountry>
, Miri division, Upper Tinjar River, Sungai Lobang,
<date id="FFE31033FFC2FFF1FEBA08E349909CF6" box="[341,475,978,1003]" pageId="22" pageNumber="473" value="2010-05-04" valueMax="2010-05-05" valueMin="2010-05-04">4–5 v 2010</date>
, leg. L. Southwell;
<specimenCount id="9D5BFD7AFFC2FFF1FD2E08E24AA69CF7" box="[705,749,979,1003]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR11_12_EUP159),
<collectingCountry id="F34A7663FFC2FFF1FBEC08E34C2D9CF6" box="[1027,1126,978,1003]" name="Malaysia" pageId="22" pageNumber="473">Sarawak</collectingCountry>
, Miri division, Usun Apau National Park, Sungai Julan,
<date id="FFE31033FFC2FFF1FE0D08C64A029B12" box="[482,585,1015,1039]" pageId="22" pageNumber="473" value="2012-05-01">1 v 2012</date>
, leg. C. Wilfred;
<specimenCount id="9D5BFD7AFFC2FFF1FCE108C64B729B13" box="[782,825,1015,1039]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR07_8_EUP48, in
<collectionCode id="ED4CAE36FFC2FFF1FBAA08C64CD59B13" LSID="urn:lsid:biocol.org:col:34992" box="[1093,1182,1015,1038]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
:
<collectionCode id="ED4CAE36FFC2FFF1FB4608C74D489B12" LSID="urn:lsid:biocol.org:col:34992" box="[1193,1283,1014,1039]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
.INS.509906), Miri division, Gunung Mulu National Park, foot of Gunung Mulu, stream on old trail to
<collectingCountry id="F34A7663FFC2FFF1FB6A0F2B4CAE9B2E" box="[1157,1253,1050,1075]" name="Malaysia" pageId="22" pageNumber="473">Sarawak</collectingCountry>
Chamber,
<date id="FFE31033FFC2FFF1FA8D0F2A48999B4A" pageId="22" pageNumber="473" value="2008-09-10">10 ix 2008</date>
, leg. R.A. Dow;
<specimenCount id="9D5BFD7AFFC2FFF1FE760F0E498D9B4B" box="[409,454,1087,1111]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR05-EUP57; Coll. Hämäläinen),
<collectingCountry id="F34A7663FFC2FFF1FC960F0F4B969B4A" box="[889,989,1086,1111]" name="Malaysia" pageId="22" pageNumber="473">Sarawak</collectingCountry>
, Miri division, Gunung Mulu National Park, Long Lansat,
<date id="FFE31033FFC2FFF1FE990F5249B99B66" box="[374,498,1122,1147]" pageId="22" pageNumber="473" value="2005-04-19">19 iv 2005</date>
, leg. R.A. Dow;
<specimenCount id="9D5BFD7AFFC2FFF1FD5C0F524A959B67" box="[691,734,1123,1147]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR05-EUP31),
<collectingCountry id="F34A7663FFC2FFF1FC5A0F534C539B66" box="[949,1048,1122,1147]" name="Malaysia" pageId="22" pageNumber="473">Sarawak</collectingCountry>
, Miri division, Kelabit Highlands, Bario area, Pa’Dapur river,
<date id="FFE31033FFC2FFF1FE3F0FB74A749B82" box="[464,575,1158,1183]" pageId="22" pageNumber="473" value="2005-04-05">5 iv 2005</date>
, leg. G.T. Reels;
<specimenCount id="9D5BFD7AFFC2FFF1FCEF0FB64B609B83" box="[768,811,1159,1183]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR06_EUP33),
<collectingCountry id="F34A7663FFC2FFF1FBE90FB74C229B82" box="[1030,1129,1158,1183]" name="Malaysia" pageId="22" pageNumber="473">Sarawak</collectingCountry>
, Kuching division, Matang Range, Kubah National Park, Sungai Rayu at Matang Wildlife Centre,
<date id="FFE31033FFC2FFF1FBE50F9B4CC29BDE" box="[1034,1161,1194,1219]" pageId="22" pageNumber="473" value="2006-01-25">25 i 2006</date>
, leg. G.T. Reels;
<specimenCount id="9D5BFD7AFFC2FFF1FA860F9A4DD79BDF" box="[1385,1436,1195,1219]" pageId="22" pageNumber="473" type="generic">2 ♂</specimenCount>
(SAR07_8_EUP37; SAR07_8_EUP
<quantity id="4CA59B16FFC2FFF1FDDF0FFE4A209BFA" box="[560,619,1230,1255]" metricMagnitude="0" metricUnit="m" metricValue="1.2446" pageId="22" pageNumber="473" unit="in" value="49.0">49 in</quantity>
<collectionCode id="ED4CAE36FFC2FFF1FD9B0FFE4A879BFB" LSID="urn:lsid:biocol.org:col:34992" box="[628,716,1231,1254]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
:
<collectionCode id="ED4CAE36FFC2FFF1FD360FFF4B789BFA" LSID="urn:lsid:biocol.org:col:34992" box="[729,819,1230,1255]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
.INS.509907),
<collectingCountry id="F34A7663FFC2FFF1FC3A0FFF4C739BFA" box="[981,1080,1230,1255]" name="Malaysia" pageId="22" pageNumber="473">Sarawak</collectingCountry>
, Sri
<collectingCountry id="F34A7663FFC2FFF1FB9F0FFE4CFE9BFA" box="[1136,1205,1231,1255]" name="Oman" pageId="22" pageNumber="473">Aman</collectingCountry>
division, Batang Ai National Park, Sungai Bebiong Besar,
<date id="FFE31033FFC2FFF1FDA70FC24AF59A16" box="[584,702,1266,1291]" pageId="22" pageNumber="473" value="2007-12-04">4 xii 2007</date>
, leg. R.A. Dow;
<specimenCount id="9D5BFD7AFFC2FFF1FC940FC24BEE9A17" box="[891,933,1267,1291]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR07_8_EUP12), same location and date as above, leg. G.T. Reels;
<specimenCount id="9D5BFD7AFFC2FFF1FE580E2649AB9A33" box="[439,480,1303,1327]" pageId="22" pageNumber="473" type="generic">3 ♂</specimenCount>
(SAR13_14_EUP7–8, SAR13_14_EUP
<quantity id="4CA59B16FFC2FFF1FC470E264B999A32" box="[936,978,1302,1327]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="22" pageNumber="473" unit="in" value="2.0">2 in</quantity>
<collectionCode id="ED4CAE36FFC2FFF1FC370E264C7B9A33" LSID="urn:lsid:biocol.org:col:34992" box="[984,1072,1303,1326]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
:
<collectionCode id="ED4CAE36FFC2FFF1FBD50E274CDF9A32" LSID="urn:lsid:biocol.org:col:34992" box="[1082,1172,1302,1327]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
.INS.557687),
<collectingCountry id="F34A7663FFC2FFF1FADA0E274DD39A32" box="[1333,1432,1302,1327]" name="Malaysia" pageId="22" pageNumber="473">Sarawak</collectingCountry>
, Kapit division, Lanjak Entimau Wildlife Sanctuary,
<taxonomicName id="4C5D4D70FFC2FFF1FD080E0B4B5A9A4E" box="[743,785,1338,1363]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="486" phylum="Arthropoda" rank="species" species="ulu">Ulu</taxonomicName>
Katibas, Sungai Datai,
<date id="FFE31033FFC2FFF1FBCE0E0A4CE49A4E" box="[1057,1199,1338,1363]" pageId="22" pageNumber="473" value="2013-08-22">22 viii 2013</date>
, leg. R.A. Dow;
<specimenCount id="9D5BFD7AFFC2FFF1FA9E0E0A4DD79A4F" box="[1393,1436,1339,1363]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(SAR13_14_EUP12), same location and date as above, leg. J. ak Awan &amp; M. ak Adau.—From
<emphasis id="B929EAE1FFC2FFF1FAE80E6F4D119A6A" bold="true" box="[1287,1370,1374,1399]" pageId="22" pageNumber="473">
<collectingCountry id="F34A7663FFC2FFF1FAE80E6F4D119A6A" box="[1287,1370,1374,1399]" name="Brunei" pageId="22" pageNumber="473">Brunei</collectingCountry>
</emphasis>
:
<specimenCount id="9D5BFD7AFFC2FFF1FA820E6E4DD09A6B" box="[1389,1435,1375,1399]" pageId="22" pageNumber="473" type="generic">2 ♂</specimenCount>
(
<collectionCode id="ED4CAE36FFC2FFF1FF710EB248BC9A86" LSID="urn:lsid:biocol.org:col:34992" box="[158,247,1411,1435]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
, ex. collection P. Mill), (Temburong district),
<taxonomicName id="4C5D4D70FFC2FFF1FD100EB34BB79A86" authority="Temburong, 1978" authorityName="Temburong" authorityYear="1978" box="[767,1020,1410,1435]" class="Insecta" family="Euphaeidae" genus="Ulu" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="486" phylum="Arthropoda" rank="genus">Ulu Temburong, 1978</taxonomicName>
, leg. S.L. Sutton;
<specimenCount id="9D5BFD7AFFC2FFF1FB260EB24CB89A87" box="[1225,1267,1411,1435]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED4CAE36FFC2FFF1FAEC0EB24D169A86" LSID="urn:lsid:biocol.org:col:34992" box="[1283,1373,1411,1435]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
), 1 ♀ (Coll. A.G. Orr)
<collectingCountry id="F34A7663FFC2FFF1FEB80E9749E89AA2" box="[343,419,1446,1471]" name="Brunei" pageId="22" pageNumber="473">Brunei</collectingCountry>
(Temburong district), Kuala Belalong Field Studies Centre,
<date id="FFE31033FFC2FFF1FBB10E974CF29AA2" box="[1118,1209,1446,1471]" pageId="22" pageNumber="473" value="1991-03">iii 1991</date>
, leg. A.G. Orr;
<specimenCount id="9D5BFD7AFFC2FFF1FA800E964DD79AA3" box="[1391,1436,1447,1471]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED4CAE36FFC2FFF1FF700EFA48B29AFE" LSID="urn:lsid:biocol.org:col:34992" box="[159,249,1483,1507]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
), same data,
<date id="FFE31033FFC2FFF1FE7A0EFA49A29AFE" box="[405,489,1483,1507]" pageId="22" pageNumber="473" value="1991-10">x 1991</date>
;
<specimenCount id="9D5BFD7AFFC2FFF1FE160EFA4A6C9AFF" box="[505,551,1483,1507]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED4CAE36FFC2FFF1FDD50EFA4ADF9AFE" LSID="urn:lsid:biocol.org:col:34992" box="[570,660,1483,1507]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
), same data,
<date id="FFE31033FFC2FFF1FCDF0EFB4BDE9AFE" box="[816,917,1482,1507]" pageId="22" pageNumber="473" value="1992-07">vii 1992</date>
;
<specimenCount id="9D5BFD7AFFC2FFF1FC4B0EFA4B999AFF" box="[932,978,1483,1507]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED4CAE36FFC2FFF1FC0A0EFA4C749AFE" LSID="urn:lsid:biocol.org:col:34992" box="[997,1087,1483,1507]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
), same data,
<date id="FFE31033FFC2FFF1FB340EFA4D159AFE" box="[1243,1374,1482,1507]" pageId="22" pageNumber="473" value="1992-09-29">29 ix 1992</date>
;
<specimenCount id="9D5BFD7AFFC2FFF1FA810EFA4DD79AFF" box="[1390,1436,1483,1507]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED4CAE36FFC2FFF1FF700EDE48B2991A" LSID="urn:lsid:biocol.org:col:34992" box="[159,249,1519,1543]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
), same data,
<date id="FFE31033FFC2FFF1FE650EDE49B3991A" box="[394,504,1518,1543]" pageId="22" pageNumber="473" value="1993-01-23">23 i 1993</date>
;
<specimenCount id="9D5BFD7AFFC2FFF1FDEB0EDE4A64991B" box="[516,559,1519,1543]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED4CAE36FFC2FFF1FDD10EDE4AD3991A" LSID="urn:lsid:biocol.org:col:34992" box="[574,664,1519,1543]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
), same data,
<date id="FFE31033FFC2FFF1FCC50EDF4BC1991A" box="[810,906,1518,1543]" pageId="22" pageNumber="473" value="1993-07">vii 1993</date>
;
<specimenCount id="9D5BFD7AFFC2FFF1FC790EDE4B8A991B" box="[918,961,1519,1543]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED4CAE36FFC2FFF1FC3F0EDE4C6C991A" box="[976,1063,1519,1543]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="22" pageNumber="473">CUMZ</collectionCode>
),
<collectingCountry id="F34A7663FFC2FFF1FBD50EDF4CC2991A" box="[1082,1161,1518,1543]" name="Brunei" pageId="22" pageNumber="473">Brunei</collectingCountry>
, Temburong,
<date id="FFE31033FFC2FFF1FACB0EDE4DD3991A" box="[1316,1432,1519,1543]" pageId="22" pageNumber="473" value="1978-10-10">10 x 1978</date>
, leg. T. Harman (ex coll. D.A.L. Davies);
<specimenCount id="9D5BFD7AFFC2FFF1FD840D224ADD9937" box="[619,662,1555,1579]" pageId="22" pageNumber="473" type="generic">3 ♂</specimenCount>
(
<collectionCode id="ED4CAE36FFC2FFF1FD490D224AB69936" box="[678,765,1555,1579]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="22" pageNumber="473">CUMZ</collectionCode>
),
<collectingCountry id="F34A7663FFC2FFF1FCFE0D234B2B9936" box="[785,864,1554,1579]" name="Brunei" pageId="22" pageNumber="473">Brunei</collectingCountry>
, Snake Creek,
<date id="FFE31033FFC2FFF1FBE40D234CCA9936" box="[1035,1153,1554,1579]" pageId="22" pageNumber="473" value="1982-02-16">16 ii 1982</date>
, leg. T. Harman (ex coll. D.A.L. Davies).—From
<emphasis id="B929EAE1FFC2FFF1FE440D0749BD9952" bold="true" box="[427,502,1590,1615]" pageId="22" pageNumber="473">Sabah</emphasis>
:
<specimenCount id="9D5BFD7AFFC2FFF1FDEB0D064A649953" box="[516,559,1591,1615]" pageId="22" pageNumber="473" type="generic">3 ♂</specimenCount>
(
<collectionCode id="ED4CAE36FFC2FFF1FDD10D064AD39952" LSID="urn:lsid:biocol.org:col:34992" box="[574,664,1591,1615]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
), Sabah,
<quantity id="4CA59B16FFC2FFF1FD110D074B189952" box="[766,851,1590,1615]" metricMagnitude="5" metricUnit="m" metricValue="1.05" pageId="22" pageNumber="473" unit="km" value="105.0">105 km</quantity>
S of Beaufort, Long Pasia area, Sungai Padas, S. of Kampong Long Pasia,
<quantity id="4CA59B16FFC2FFF1FE730D6A49B9996F" box="[412,498,1627,1651]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="22" pageNumber="473" unit="m" value="1030.0">1030 m</quantity>
asl.,
<geoCoordinate id="EE695034FFC2FFF1FDDF0D6A4AC9996E" box="[560,642,1627,1651]" direction="north" orientation="latitude" pageId="22" pageNumber="473" precision="925" value="4.4">4 24’N</geoCoordinate>
,
<geoCoordinate id="EE695034FFC2FFF1FD600D6B4AB1996E" box="[655,762,1626,1651]" direction="east" orientation="longitude" pageId="22" pageNumber="473" precision="925" value="115.73333">115 44’E</geoCoordinate>
,
<date id="FFE31033FFC2FFF1FCE80D6A4BE4996E" box="[775,943,1626,1651]" pageId="22" pageNumber="473" value="1987-04-12" valueMax="1987-04-13" valueMin="1987-04-12">12–13 iv 1987</date>
, leg. J. van Tol;
<specimenCount id="9D5BFD7AFFC2FFF1FB820D6A4CD2996F" box="[1133,1177,1627,1651]" pageId="22" pageNumber="473" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED4CAE36FFC2FFF1FB460D6A4D48996E" LSID="urn:lsid:biocol.org:col:34992" box="[1193,1283,1627,1651]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
), Sabah, surr. Long Pasia, Sg. Matang, leg. J. van Tol.;
<specimenCount id="9D5BFD7AFFC2FFF1FD810D4E4AD2998B" box="[622,665,1663,1687]" pageId="22" pageNumber="473" type="generic">2 ♂</specimenCount>
(
<collectionCode id="ED4CAE36FFC2FFF1FD460D4E4B4B998A" box="[681,768,1663,1687]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="22" pageNumber="473">CUMZ</collectionCode>
), Borneo, Sabah, Maliau Basin, tributary to Sungai Maliau near Maliau Basin Field Centre,
<date id="FFE31033FFC2FFF1FDEB0D924A3C99A6" box="[516,631,1699,1723]" pageId="22" pageNumber="473" value="2013-05-14">14 v 2013</date>
, leg. S. Luke.
</paragraph>
</subSubSection>
<subSubSection id="C3476578FFC2FFF1FF280DF74ADC983A" pageId="22" pageNumber="473" type="etymology">
<paragraph id="8BE236F3FFC2FFF1FF280DF74ADC983A" blockId="22.[151,1437,402,2011]" pageId="22" pageNumber="473">
<emphasis id="B929EAE1FFC2FFF1FF280DF7490599C2" bold="true" box="[199,334,1734,1759]" pageId="22" pageNumber="473">Etymology.</emphasis>
The species epithet is based on the word ‘ulu’, the form generally in use in Borneo of the Bahasa Melayu/
<collectingCountry id="F34A7663FFC2FFF1FF1B0DDB492A981E" box="[244,353,1770,1795]" name="Indonesia" pageId="22" pageNumber="473">Indonesia</collectingCountry>
word ‘hulu’, which means upstream. The species epithet is used as a noun in apposition. The species typically inhabits ‘upstream’ habitats.
</paragraph>
</subSubSection>
<subSubSection id="C3476578FFC2FFF1FF280C034AB39872" pageId="22" pageNumber="473" type="diagnosis">
<paragraph id="8BE236F3FFC2FFF1FF280C034AB39872" blockId="22.[151,1437,402,2011]" pageId="22" pageNumber="473">
<emphasis id="B929EAE1FFC2FFF1FF280C03490A9856" bold="true" box="[199,321,1842,1867]" pageId="22" pageNumber="473">Diagnosis.</emphasis>
A narrow-winged
<taxonomicName id="4C5D4D70FFC2FFF1FDFB0C024ACF9856" box="[532,644,1843,1867]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="473" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAE1FFC2FFF1FDFB0C024ACF9856" box="[532,644,1843,1867]" italics="true" pageId="22" pageNumber="473">Dysphaea</emphasis>
</taxonomicName>
species with male wings broadly opaque at basal half and at wing tips. Cerci with lower border nearly straight in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C3476578FFC2FFFEFF280C4B48889D20" lastPageId="25" lastPageNumber="476" pageId="22" pageNumber="473" type="description">
<paragraph id="8BE236F3FFC2FFF1FF280C4B499A98AA" blockId="22.[151,1437,402,2011]" pageId="22" pageNumber="473">
<emphasis id="B929EAE1FFC2FFF1FF280C4B4A55988E" bold="true" box="[199,542,1914,1939]" pageId="22" pageNumber="473">
Description of
<typeStatus id="54E68851FFC2FFF1FE980C4B4995988E" box="[375,478,1914,1939]" pageId="22" pageNumber="473" type="holotype">holotype</typeStatus>
male
</emphasis>
(
<figureCitation id="13662A76FFC2FFF1FDC20C4B4A36988E" box="[557,637,1914,1939]" captionStart="FIGURES 9 – 12" captionStartId="14.[151,264,1350,1373]" captionTargetBox="[151,1436,373,1328]" captionTargetId="figure@14.[151,1436,373,1330]" captionTargetPageId="14" captionText="FIGURES 9 – 12. Habitus of male Dysphaea: 9) D. dimidiata West Java; 10) D. lugens Danum Valley, Sabah; 11) D. ulu holotype (flipped horizontally); 12) D. vanida holotype (flipped horizontally)." httpUri="https://zenodo.org/record/243466/files/figure.png" pageId="22" pageNumber="473">Fig. 11</figureCitation>
). Head: Labium, labrum, base of mandibles and clypeus shining black, frons and vertex matt black.
</paragraph>
<paragraph id="8BE236F3FFC2FFF0FF280CF349269FC8" blockId="22.[151,1437,402,2011]" lastBlockId="23.[151,1437,151,680]" lastPageId="23" lastPageNumber="474" pageId="22" pageNumber="473">
Thorax: Matt black, with very faint obscure brown marking on metepisternum above second lateral suture and similar marking on metepimeron (
<figureCitation id="13662A76FFC3FFF0FDCF0BA64A3D9FAD" box="[544,630,151,176]" captionStart="FIGURES 13 – 18" captionStartId="15.[151,264,1951,1974]" captionTargetBox="[151,1436,517,1918]" captionTargetId="figure@15.[151,1436,517,1931]" captionTargetPageId="15" captionText="FIGURES 13 – 18. Synthorax male Dysphaea: 13) D. dimidiata West Java; 14) D. lugens Bakuan, Kalimantan Barat; 15) D. ulu holotype; 16) D. vanida holotype; 17) metaposternum, D. dimidiata Brunei; 18) metaposternum, D. ulu holotype." httpUri="https://zenodo.org/record/243467/files/figure.png" pageId="23" pageNumber="474">Fig. 15</figureCitation>
). Venter of thorax without tiny tubercles on metaposternum (
<figureCitation id="13662A76FFC3FFF0FAD60BA64DC49FAD" box="[1337,1423,151,176]" captionStart="FIGURES 13 – 18" captionStartId="15.[151,264,1951,1974]" captionTargetBox="[151,1436,517,1918]" captionTargetId="figure@15.[151,1436,517,1931]" captionTargetPageId="15" captionText="FIGURES 13 – 18. Synthorax male Dysphaea: 13) D. dimidiata West Java; 14) D. lugens Bakuan, Kalimantan Barat; 15) D. ulu holotype; 16) D. vanida holotype; 17) metaposternum, D. dimidiata Brunei; 18) metaposternum, D. ulu holotype." httpUri="https://zenodo.org/record/243467/files/figure.png" pageId="23" pageNumber="474">Fig. 18</figureCitation>
). Legs wholly black.
</paragraph>
<paragraph id="8BE236F3FFC3FFF0FF280BEE49199EE8" blockId="23.[151,1437,151,680]" pageId="23" pageNumber="474">
Wings: Basal half of wings opaque black with strong metallic blue reflections. In Fw opaque area extends to level of 3–4 cells before nodus, except in costal field where the opaque stripe extends 1 or 2 cells beyond the nodus. Otherwise costal field between nodus and pterostigma hyaline. In Hw basal opaque area extends more apicad, not quite reaching half way between nodus and proximal border of pterostigma; costal field between nodus and pterostigma opaque throughout. Tips of wings narrowly opaque, in Hw slightly more extensively than in Fw (
<figureCitation id="13662A76FFC3FFF0FA800A5E48FF9EB1" captionStart="FIGURES 9 – 12" captionStartId="14.[151,264,1350,1373]" captionTargetBox="[151,1436,373,1328]" captionTargetId="figure@14.[151,1436,373,1330]" captionTargetPageId="14" captionText="FIGURES 9 – 12. Habitus of male Dysphaea: 9) D. dimidiata West Java; 10) D. lugens Danum Valley, Sabah; 11) D. ulu holotype (flipped horizontally); 12) D. vanida holotype (flipped horizontally)." httpUri="https://zenodo.org/record/243466/files/figure.png" pageId="23" pageNumber="474">Fig. 11</figureCitation>
). Venation typical of genus. Fw with 37 antenodals in first row; Hw correspondingly with 27 antenodals. Quadrangle with 2 crossveins in Fw,
<quantity id="4CA59B16FFC3FFF0FDA80A894ADA9ECD" box="[583,657,439,464]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="23" pageNumber="474" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">2–3 in</quantity>
Hw. Pterostigma long and narrow, broadest in middle; covering 14 underlying cells.
</paragraph>
<paragraph id="8BE236F3FFC3FFF0FF280ACE490D9D7D" blockId="23.[151,1437,151,680]" pageId="23" pageNumber="474">
Abdomen: Matt black throughout. Appendages black; cerci in dorsal (
<figureCitation id="13662A76FFC3FFF0FC160ACE4C059D05" box="[1017,1102,511,536]" captionStart="FIGURES 31 – 36" captionStartId="19.[151,264,1705,1728]" captionTargetBox="[157,1436,185,1683]" captionTargetId="figure@19.[157,1442,185,1684]" captionTargetPageId="19" captionText="FIGURES 31 – 36. Male anal appendages. Dorsal view: 31) D. dimidiata West Java; 32) D. lugens Bakuan, Kalimantan Barat; 33) D. ulu holotype; 34) D. vanida holotype. Dorso-lateral views: 35) D. dimidiata Terengganu; 36) D. vanida paratype Lam Khlong Gnu, Kanchanaburi." httpUri="https://zenodo.org/record/243470/files/figure.png" pageId="23" pageNumber="474">Fig. 33</figureCitation>
) and ventral view of typical shape for genus; in lateral view ventral margin of cercus almost straight (
<figureCitation id="13662A76FFC3FFF0FC3809154C679D21" box="[983,1068,548,573]" captionStart="FIGURES 37 – 40" captionStartId="20.[151,264,1070,1093]" captionTargetBox="[162,1392,232,1036]" captionTargetId="figure@20.[151,1436,193,1050]" captionTargetPageId="20" captionText="FIGURES 37 – 40. Male anal appendages, lateral view (scale bars 1 mm): 37) D. dimidiata west Java; 38) D. lugens Bakuan, Kalimantan Barat; 39) D. ulu holotype; 40) D. vanida holotype." httpUri="https://zenodo.org/record/243471/files/figure.png" pageId="23" pageNumber="474">Fig. 39</figureCitation>
). Paraprocts very short, rounded and featureless.
</paragraph>
<paragraph id="8BE236F3FFC3FFF0FF28095D4BEC9DB5" blockId="23.[151,1437,151,680]" pageId="23" pageNumber="474">
Penis: Terminal segment with two apical arms directed upwards on either side of shaft, turning out and down for short distance at ends, slightly expanded in this part (
<figureCitation id="13662A76FFC3FFF0FCFE09BE4B259DB5" box="[785,878,655,680]" captionStart="FIGURES 41 – 44" captionStartId="21.[151,264,1790,1813]" captionTargetBox="[162,1384,617,1756]" captionTargetId="figure@21.[151,1436,555,1769]" captionTargetPageId="21" captionText="FIGURES 41 – 44. Penis, ventral view: 41) D. dimidiata west Java; 42) D. lugens Selekau River, Kalimantan Barat; 43) D. ulu holotype; 44) D. vanida paratype, Khlong Nakha, Ranong." httpUri="https://zenodo.org/record/243472/files/figure.png" pageId="23" pageNumber="474">Figs. 43</figureCitation>
,
<figureCitation id="13662A76FFC3FFF0FC9509A14BD19DB5" box="[890,922,656,680]" captionStart="FIGURES 45 – 48" captionStartId="23.[151,264,1735,1758]" captionTargetBox="[162,1365,783,1707]" captionTargetId="figure@23.[151,1436,721,1720]" captionTargetPageId="23" captionText="FIGURES 45 – 48. Penis, lateral view: 45) D. dimidiata west Java; 46) D. lugens Selekau River, Kalimantan Barat; 47) D. ulu holotype; 48) D. vanida paratype, Khlong Nakha, Ranong." httpUri="https://zenodo.org/record/243473/files/figure.png" pageId="23" pageNumber="474">47</figureCitation>
).
</paragraph>
<caption id="DF22667BFFC3FFF0FF780DF64AA199E1" httpUri="https://zenodo.org/record/243473/files/figure.png" pageId="23" pageNumber="474" targetBox="[162,1365,783,1707]" targetPageId="23">
<paragraph id="8BE236F3FFC3FFF0FF780DF64AA199E1" blockId="23.[151,1436,1735,1789]" pageId="23" pageNumber="474">
<emphasis id="B929EAE1FFC3FFF0FF780DF6491F99C0" bold="true" box="[151,340,1735,1758]" pageId="23" pageNumber="474">FIGURES 45–48.</emphasis>
Penis, lateral view:
<emphasis id="B929EAE1FFC3FFF0FDCB0DF64A0D99C3" bold="true" box="[548,582,1735,1758]" pageId="23" pageNumber="474">45)</emphasis>
<taxonomicName id="4C5D4D70FFC3FFF0FDA20DF84A8699C3" box="[589,717,1736,1758]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B929EAE1FFC3FFF0FDA20DF84A8699C3" box="[589,717,1736,1758]" italics="true" pageId="23" pageNumber="474">D. dimidiata</emphasis>
</taxonomicName>
west Java;
<emphasis id="B929EAE1FFC3FFF0FCAA0DF64B2D99C3" bold="true" box="[837,870,1735,1758]" pageId="23" pageNumber="474">46)</emphasis>
<taxonomicName id="4C5D4D70FFC3FFF0FC820DF84C2C99C3" authority="Selekau River" authorityName="Selekau River" box="[877,1127,1736,1758]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="lugens">
<emphasis id="B929EAE1FFC3FFF0FC820DF84B8499C3" box="[877,975,1736,1758]" italics="true" pageId="23" pageNumber="474">D. lugens</emphasis>
Selekau River
</taxonomicName>
, Kalimantan Barat;
<emphasis id="B929EAE1FFC3FFF0FADC0DF64D1E99C3" bold="true" box="[1331,1365,1735,1758]" pageId="23" pageNumber="474">47)</emphasis>
<taxonomicName id="4C5D4D70FFC3FFF0FAB30DF84DD799C3" box="[1372,1436,1736,1758]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="ulu">
<emphasis id="B929EAE1FFC3FFF0FAB30DF84DD799C3" box="[1372,1436,1736,1758]" italics="true" pageId="23" pageNumber="474">D. ulu</emphasis>
</taxonomicName>
holotype;
<emphasis id="B929EAE1FFC3FFF0FF120DD7495599E0" bold="true" box="[253,286,1766,1789]" pageId="23" pageNumber="474">48)</emphasis>
<taxonomicName id="4C5D4D70FFC3FFF0FECA0DD949C299E1" box="[293,393,1766,1789]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="vanida">
<emphasis id="B929EAE1FFC3FFF0FECA0DD949C299E1" box="[293,393,1766,1789]" italics="true" pageId="23" pageNumber="474">D. vanida</emphasis>
</taxonomicName>
paratype, Khlong Nakha, Ranong.
</paragraph>
</caption>
<paragraph id="8BE236F3FFC3FFF0FF280C154C4C9821" blockId="23.[151,1437,1828,2033]" box="[199,1031,1828,1853]" pageId="23" pageNumber="474">
<emphasis id="B929EAE1FFC3FFF0FF280C15493D9820" bold="true" box="[199,374,1828,1853]" pageId="23" pageNumber="474">Measurements</emphasis>
(mm). Fw 33, Hw 31, abdomen (apps. excl.) 35.5, cerci 2.
</paragraph>
<paragraph id="8BE236F3FFC3FFF0FF280C794BEA98EC" blockId="23.[151,1437,1828,2033]" pageId="23" pageNumber="474">
<emphasis id="B929EAE1FFC3FFF0FF280C794987987C" bold="true" box="[199,460,1864,1889]" pageId="23" pageNumber="474">Description of female</emphasis>
(
<figureCitation id="13662A76FFC3FFF0FE300C794A7E987C" box="[479,565,1864,1889]" captionStart="FIGURES 49 – 52" captionStartId="24.[151,264,1070,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@24.[151,1436,193,1049]" captionTargetPageId="24" captionText="FIGURES 49 – 52. Habitus of female Dysphaea: 49) D. dimidiata west Java; 50) D. lugens Mount Dulit, Sarawak, photograph by Benjamin Price; 51) D. ulu paratype; 52) D. vanida paratype Kanchanaburi." httpUri="https://zenodo.org/record/243474/files/figure.png" pageId="23" pageNumber="474">Fig. 51</figureCitation>
). Head: Labium shining black with lateral lobes largely yellow, with black hooks. Labrum shining black, with broadly yellow centre, indented by black basally. Base of mandibles largely yellow, with black incomplete stripe medially. Clypeus shining black with narrow yellow stripe along ridge. Frons matt black with sides below antennae yellow, yellow colour continuing over genae (
<figureCitation id="13662A76FFC3FFF0FB9C0C854C8198D0" box="[1139,1226,1972,1997]" captionStart="FIGURES 53 – 56" captionStartId="25.[151,264,1609,1632]" captionTargetBox="[151,1436,625,1588]" captionTargetId="figure@25.[151,1436,625,1588]" captionTargetPageId="25" captionText="FIGURES 53 – 56. Female head: 53) D. dimidiata Sarawak, dorsal view; 54) D. lugens Gunung Sari, Kalimantan Timur, dorsal view; 55) D. ulu paratype, dorsal, slightly frontal view; 56) D. vanida paratype, Kanchanaburi, dorsal, slightly frontal view." httpUri="https://zenodo.org/record/243475/files/figure.png" pageId="23" pageNumber="474">Fig. 55</figureCitation>
). Antennae black. Vertex and occiput matt black, with two tiny yellow spots on occiput.
</paragraph>
<caption id="DF22667BFFCCFFFFFF780F1F4BFC9B7E" httpUri="https://zenodo.org/record/243474/files/figure.png" pageId="24" pageNumber="475" targetBox="[151,1436,193,1049]" targetPageId="24">
<paragraph id="8BE236F3FFCCFFFFFF780F1F4BFC9B7E" blockId="24.[151,1436,1070,1124]" pageId="24" pageNumber="475">
<emphasis id="B929EAE1FFCCFFFFFF780F1F49189B59" bold="true" box="[151,339,1070,1093]" pageId="24" pageNumber="475">FIGURES 49–52.</emphasis>
Habitus of female
<taxonomicName id="4C5D4D70FFCCFFFFFDF80F1E4A379B58" box="[535,636,1071,1093]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAE1FFCCFFFFFDF80F1E4A379B58" box="[535,636,1071,1093]" italics="true" pageId="24" pageNumber="475">Dysphaea</emphasis>
</taxonomicName>
:
<emphasis id="B929EAE1FFCCFFFFFD650F1F4AE09B58" bold="true" box="[650,683,1070,1093]" pageId="24" pageNumber="475">49)</emphasis>
<taxonomicName id="4C5D4D70FFCCFFFFFD5D0F014B789B58" box="[690,819,1071,1093]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B929EAE1FFCCFFFFFD5D0F014B789B58" box="[690,819,1071,1093]" italics="true" pageId="24" pageNumber="475">D. dimidiata</emphasis>
</taxonomicName>
west Java;
<emphasis id="B929EAE1FFCCFFFFFC460F1F4B819B58" bold="true" box="[937,970,1070,1093]" pageId="24" pageNumber="475">50)</emphasis>
<taxonomicName id="4C5D4D70FFCCFFFFFC3E0F014CF19B58" authority="Mount Dulit" authorityName="Mount Dulit" box="[977,1210,1071,1093]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="lugens">
<emphasis id="B929EAE1FFCCFFFFFC3E0F014C789B58" box="[977,1075,1071,1093]" italics="true" pageId="24" pageNumber="475">D. lugens</emphasis>
Mount Dulit
</taxonomicName>
, Sarawak, photograph by Benjamin Price;
<emphasis id="B929EAE1FFCCFFFFFE8E0F7C49C99B79" bold="true" box="[353,386,1101,1124]" pageId="24" pageNumber="475">51)</emphasis>
<taxonomicName id="4C5D4D70FFCCFFFFFE660F7E49839B7E" box="[393,456,1101,1124]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="ulu">
<emphasis id="B929EAE1FFCCFFFFFE660F7E49839B7E" box="[393,456,1101,1124]" italics="true" pageId="24" pageNumber="475">D. ulu</emphasis>
</taxonomicName>
paratype;
<emphasis id="B929EAE1FFCCFFFFFDDC0F7C4A1F9B79" bold="true" box="[563,596,1101,1124]" pageId="24" pageNumber="475">52)</emphasis>
<taxonomicName id="4C5D4D70FFCCFFFFFDB40F7E4AF49B7E" box="[603,703,1101,1124]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="vanida">
<emphasis id="B929EAE1FFCCFFFFFDB40F7E4AF49B7E" box="[603,703,1101,1124]" italics="true" pageId="24" pageNumber="475">D. vanida</emphasis>
</taxonomicName>
paratype Kanchanaburi.
</paragraph>
</caption>
<paragraph id="8BE236F3FFCCFFFFFF280FBA4D469A4A" blockId="24.[151,1437,1163,2016]" pageId="24" pageNumber="475">
Thorax: Prothorax black with rounded large yellow spots on either side of dorsum of middle lobe. Posterior part of hind lobe raised obliquely upwards to form an elongate rectangular flap, which is narrowly bordered by yellow, more broadly so at sides (
<figureCitation id="13662A76FFCCFFFFFDF60FE24A259BF1" box="[537,622,1235,1260]" captionStart="FIGURES 57 – 62" captionStartId="27.[151,264,1913,1936]" captionTargetBox="[155,1436,401,1899]" captionTargetId="figure@27.[155,1440,401,1900]" captionTargetPageId="27" captionText="FIGURES 57 – 62. Female prothorax, dorsal view: 57) D. dimidiata west Java; 58) D. dimidiata Sarawak; 59) D. dimidiata Tandjong Pandan, Billiton; 60) D. lugens Gunung Sari, Kaliamantan Timur; 61) D. ulu paratype; 62) D. vanida paratype, Kanchanaburi." httpUri="https://zenodo.org/record/243476/files/figure.png" pageId="24" pageNumber="475">Fig. 61</figureCitation>
). Lateral parts of flap not prominent, lying in ca same plane as median part. Synthorax matt black, with moderately narrow yellow stripes as in
<figureCitation id="13662A76FFCCFFFFFBF20FC74CC99A12" box="[1053,1154,1270,1295]" captionStart="FIGURES 63 – 68" captionStartId="28.[151,264,1699,1722]" captionTargetBox="[151,1436,185,1683]" captionTargetId="figure@28.[151,1436,185,1684]" captionTargetPageId="28" captionText="FIGURES 63 – 68. Female synthorax, dorsal view: 63) D. dimidiata west Java; 64) D. dimidiata Sarawak; 65) D. dimidiata Johor; 66) D. lugens Gunung Sari, Kaliamantan Timur; 67) D. ulu paratype; 68) D. vanida paratype Kanchanaburi." httpUri="https://zenodo.org/record/243477/files/figure.png" pageId="24" pageNumber="475">Figs. 67</figureCitation>
,
<figureCitation id="13662A76FFCCFFFFFB790FC64CFE9A12" box="[1174,1205,1271,1295]" captionStart="FIGURES 69 – 74" captionStartId="29.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1797]" captionTargetId="figure@29.[151,1436,193,1799]" captionTargetPageId="29" captionText="FIGURES 69 – 74. Female synthorax, lateral view: 69) D. dimidiata west Java; 70) D. dimidiata Sarawak; 71) D. dimidiata Johor; 72) D. lugens Mount Dulit, Sarawak, photograph by Benjamin Price; 73) D. ulu paratype; 74) D. vanida paratype Kanchanaburi." httpUri="https://zenodo.org/record/243478/files/figure.png" pageId="24" pageNumber="475">73</figureCitation>
. Yellow stripes on mesepisternum not connected at wing base, but those on metepimeron forming a loop at wing base. Legs black, with yellow markings on both apical and posterior sides of coxae and obscure streaks on hind femora.
</paragraph>
<paragraph id="8BE236F3FFCCFFFFFF280E524C1B9911" blockId="24.[151,1437,1163,2016]" pageId="24" pageNumber="475">
Wings: Hyaline with broad blackish opaque streaks in middle section of wings, at base from subcostal field to MA, costal field and much of lower part of wing hyaline (
<figureCitation id="13662A76FFCCFFFFFCF00EB74B399A82" box="[799,882,1414,1439]" captionStart="FIGURES 49 – 52" captionStartId="24.[151,264,1070,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@24.[151,1436,193,1049]" captionTargetPageId="24" captionText="FIGURES 49 – 52. Habitus of female Dysphaea: 49) D. dimidiata west Java; 50) D. lugens Mount Dulit, Sarawak, photograph by Benjamin Price; 51) D. ulu paratype; 52) D. vanida paratype Kanchanaburi." httpUri="https://zenodo.org/record/243474/files/figure.png" pageId="24" pageNumber="475">Fig. 51</figureCitation>
). In Fw opaque streak extends to level of 12 cells before nodus, in Hw it extends gradually narrowing to proximal end of pterostigma. Apex of Fw narrowly darkened, slightly more extensively on Hw. Fw with 37 antenodals in first row; hindwings with 26–28. Quadrangle with 3 crossveins in Fw,
<quantity id="4CA59B16FFCCFFFFFE420EC249939911" box="[429,472,1523,1548]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="24" pageNumber="475" unit="in" value="5.0">5 in</quantity>
Hw. Pterostigma long, covering 11–13 underlying cells.
</paragraph>
<paragraph id="8BE236F3FFCCFFFFFF280D2749F29981" blockId="24.[151,1437,1163,2016]" pageId="24" pageNumber="475">Abdomen: Matt black, with yellow markings as follows: S1 with lateral spot, extending upwards and downwards at apex of segment. S2–7 with lateral stripe, broadest on S2–3 and gradually narrowing towards apical segments. Separate small lateral spots at base of S3–7. On S2–5 stripe occupies almost whole segment length, on S6–7 stripe is interrupted.</paragraph>
<paragraph id="8BE236F3FFCCFFFFFF280D974C5099A2" blockId="24.[151,1437,1163,2016]" box="[199,1051,1702,1727]" pageId="24" pageNumber="475">
<emphasis id="B929EAE1FFCCFFFFFF280D97493D99A2" bold="true" box="[199,374,1702,1727]" pageId="24" pageNumber="475">Measurements</emphasis>
(mm). Hw 31.5–33, abdomen (apps. excl.) 30.5–32, cerci 1.
</paragraph>
<paragraph id="8BE236F3FFCCFFFFFF280DFA4CCF9869" blockId="24.[151,1437,1163,2016]" pageId="24" pageNumber="475">
<emphasis id="B929EAE1FFCCFFFFFF280DFA4A6899F9" bold="true" box="[199,547,1739,1764]" pageId="24" pageNumber="475">
Variation in male
<typeStatus id="54E68851FFCCFFFFFE420DFA4A6899F9" box="[429,547,1739,1764]" pageId="24" pageNumber="475" type="paratype">paratypes</typeStatus>
</emphasis>
. In some specimens the pale markings on the synthorax are slightly more distinct, in others they are entirely absent; clearly this is an age dependent character. The extent of the opaque area in wings is somewhat variable. In some specimens the opaque basal area in the Fw extends to the level of the nodus or even 1–4 cells beyond the nodus (up to
<quantity id="4CA59B16FFCCFFFFFD6A0C064AF89852" box="[645,691,1846,1871]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="24" pageNumber="475" unit="in" value="8.0">8 in</quantity>
costal field) and in Hw beyond half way between the nodus and proximal border of pterostigma. There is also some slight variability in venational details.
</paragraph>
<paragraph id="8BE236F3FFCCFFFFFF280C4F4B93988A" blockId="24.[151,1437,1163,2016]" box="[199,984,1918,1943]" pageId="24" pageNumber="475">
<emphasis id="B929EAE1FFCCFFFFFF280C4F493D988A" bold="true" box="[199,374,1918,1943]" pageId="24" pageNumber="475">Measurements</emphasis>
(mm): Hw 27.5–32.5, abdomen (apps. excl.) 32.5–37.
</paragraph>
<paragraph id="8BE236F3FFCCFFFEFF280C9248889D20" blockId="24.[151,1437,1163,2016]" lastBlockId="25.[151,1436,151,573]" lastPageId="25" lastPageNumber="476" pageId="24" pageNumber="475">
<emphasis id="B929EAE1FFCCFFFFFF280C9249B098A1" bold="true" box="[199,507,1955,1980]" pageId="24" pageNumber="475">Distinguishing characters</emphasis>
.
<emphasis id="B929EAE1FFCCFFFFFDE30C924A0298A1" bold="true" box="[524,585,1955,1980]" pageId="24" pageNumber="475">Male</emphasis>
: Superficially
<taxonomicName id="4C5D4D70FFCCFFFFFD1B0C954B7698A6" box="[756,829,1955,1979]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="ulu">
<emphasis id="B929EAE1FFCCFFFFFD1B0C954B7698A6" box="[756,829,1955,1979]" italics="true" pageId="24" pageNumber="475">D. ulu</emphasis>
</taxonomicName>
male closely resembles
<taxonomicName id="4C5D4D70FFCCFFFFFBB60C954CA298A6" box="[1113,1257,1955,1979]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B929EAE1FFCCFFFFFBB60C954CA298A6" box="[1113,1257,1955,1979]" italics="true" pageId="24" pageNumber="475">D. dimidiata</emphasis>
</taxonomicName>
, a species with which it co-occurs in many locations in northern Borneo. However, these species are easy to separate by the shape of cercus as seen in lateral view; in
<taxonomicName id="4C5D4D70FFCDFFFEFDCD0BA84A239FAD" box="[546,616,152,176]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="ulu">
<emphasis id="B929EAE1FFCDFFFEFDCD0BA84A239FAD" box="[546,616,152,176]" italics="true" pageId="25" pageNumber="476">D. ulu</emphasis>
</taxonomicName>
(
<figureCitation id="13662A76FFCDFFFEFD980BA64A839FAD" box="[631,712,151,176]" captionStart="FIGURES 37 – 40" captionStartId="20.[151,264,1070,1093]" captionTargetBox="[162,1392,232,1036]" captionTargetId="figure@20.[151,1436,193,1050]" captionTargetPageId="20" captionText="FIGURES 37 – 40. Male anal appendages, lateral view (scale bars 1 mm): 37) D. dimidiata west Java; 38) D. lugens Bakuan, Kalimantan Barat; 39) D. ulu holotype; 40) D. vanida holotype." httpUri="https://zenodo.org/record/243471/files/figure.png" pageId="25" pageNumber="476">Fig. 39</figureCitation>
) the ventral margin of the cercus is almost straight, but distinctly arched in
<taxonomicName id="4C5D4D70FFCDFFFEFEE60B8C49D39FC9" box="[265,408,188,212]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B929EAE1FFCDFFFEFEE60B8C49D39FC9" box="[265,408,188,212]" italics="true" pageId="25" pageNumber="476">D. dimidiata</emphasis>
</taxonomicName>
(
<figureCitation id="13662A76FFCDFFFEFE450B8D4A4B9FC9" box="[426,512,188,213]" captionStart="FIGURES 37 – 40" captionStartId="20.[151,264,1070,1093]" captionTargetBox="[162,1392,232,1036]" captionTargetId="figure@20.[151,1436,193,1050]" captionTargetPageId="20" captionText="FIGURES 37 – 40. Male anal appendages, lateral view (scale bars 1 mm): 37) D. dimidiata west Java; 38) D. lugens Bakuan, Kalimantan Barat; 39) D. ulu holotype; 40) D. vanida holotype." httpUri="https://zenodo.org/record/243471/files/figure.png" pageId="25" pageNumber="476">Fig. 37</figureCitation>
). Other characters, although less consistent, include the colour of the costal field between nodus and pterostigma in the Fw; in
<taxonomicName id="4C5D4D70FFCDFFFEFD7E0BD04A9C9FE5" box="[657,727,224,248]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="ulu">
<emphasis id="B929EAE1FFCDFFFEFD7E0BD04A9C9FE5" box="[657,727,224,248]" italics="true" pageId="25" pageNumber="476">D. ulu</emphasis>
</taxonomicName>
(
<figureCitation id="13662A76FFCDFFFEFD0A0BEE4B7D9FE5" box="[741,822,223,248]" captionStart="FIGURES 25 – 30" captionStartId="18.[151,264,1692,1715]" captionTargetBox="[151,1436,193,1669]" captionTargetId="figure@18.[151,1436,193,1671]" captionTargetPageId="18" captionText="FIGURES 25 – 30. Wings of Dysphaea males: 25) D. lugens Pemantang, Kalimantan Tengah; 26) D. lugens Ampah, Kalimantan Tengah; 27) D. lugens Selekau River, Kalimantan Barat; 28) D. ulu paratype, Temburong, Brunei; 29) D. vanida spec. nov. paratype, Khlong Nakha, Ranong; 30) D. vanida paratype, Khlong Nakha, Ranong." httpUri="https://zenodo.org/record/243469/files/figure.png" pageId="25" pageNumber="476">Fig. 28</figureCitation>
) the field is hyaline, but in most specimens of Bornean
<taxonomicName id="4C5D4D70FFCDFFFEFF780A34496F9E01" box="[151,292,260,284]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B929EAE1FFCDFFFEFF780A34496F9E01" box="[151,292,260,284]" italics="true" pageId="25" pageNumber="476">D. dimidiata</emphasis>
</taxonomicName>
the costal field in the Fw is opaque (
<figureCitation id="13662A76FFCDFFFEFD2B0A354B539E01" box="[708,792,260,285]" captionStart="FIGURES 19 – 24" captionStartId="16.[151,264,1692,1715]" captionTargetBox="[151,1436,204,1669]" captionTargetId="figure@16.[151,1436,193,1671]" captionTargetPageId="16" captionText="FIGURES 19 – 24. Wings of D. dimidiata males: 19) West Java; 20) West Java; 21) Billiton; 22) Babi Djuton, Kalimantan Timur; 23) Perak; 24) Labuan." httpUri="https://zenodo.org/record/243468/files/figure.png" pageId="25" pageNumber="476">Fig. 22</figureCitation>
). In
<taxonomicName id="4C5D4D70FFCDFFFEFCA50A344BDB9E01" box="[842,912,260,284]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="ulu">
<emphasis id="B929EAE1FFCDFFFEFCA50A344BDB9E01" box="[842,912,260,284]" italics="true" pageId="25" pageNumber="476">D. ulu</emphasis>
</taxonomicName>
the venter of the thorax is always without tiny tubercles on metaposternum (
<figureCitation id="13662A76FFCDFFFEFE0A0A164A739E5D" box="[485,568,295,320]" captionStart="FIGURES 13 – 18" captionStartId="15.[151,264,1951,1974]" captionTargetBox="[151,1436,517,1918]" captionTargetId="figure@15.[151,1436,517,1931]" captionTargetPageId="15" captionText="FIGURES 13 – 18. Synthorax male Dysphaea: 13) D. dimidiata West Java; 14) D. lugens Bakuan, Kalimantan Barat; 15) D. ulu holotype; 16) D. vanida holotype; 17) metaposternum, D. dimidiata Brunei; 18) metaposternum, D. ulu holotype." httpUri="https://zenodo.org/record/243467/files/figure.png" pageId="25" pageNumber="476">Fig. 18</figureCitation>
), whereas in Bornean
<taxonomicName id="4C5D4D70FFCDFFFEFCD80A184B8D9E5D" box="[823,966,296,320]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B929EAE1FFCDFFFEFCD80A184B8D9E5D" box="[823,966,296,320]" italics="true" pageId="25" pageNumber="476">D. dimidiata</emphasis>
</taxonomicName>
they are almost always present (
<figureCitation id="13662A76FFCDFFFEFAD50A164DC49E5D" box="[1338,1423,295,320]" captionStart="FIGURES 13 – 18" captionStartId="15.[151,264,1951,1974]" captionTargetBox="[151,1436,517,1918]" captionTargetId="figure@15.[151,1436,517,1931]" captionTargetPageId="15" captionText="FIGURES 13 – 18. Synthorax male Dysphaea: 13) D. dimidiata West Java; 14) D. lugens Bakuan, Kalimantan Barat; 15) D. ulu holotype; 16) D. vanida holotype; 17) metaposternum, D. dimidiata Brunei; 18) metaposternum, D. ulu holotype." httpUri="https://zenodo.org/record/243467/files/figure.png" pageId="25" pageNumber="476">Fig. 17</figureCitation>
). There are also differences in the shape of the apical arms of the penis: the terminal, upward directed part is distinctly shorter in
<taxonomicName id="4C5D4D70FFCDFFFEFE980A4049F69E95" box="[375,445,368,392]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="ulu">
<emphasis id="B929EAE1FFCDFFFEFE980A4049F69E95" box="[375,445,368,392]" italics="true" pageId="25" pageNumber="476">D. ulu</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C5D4D70FFCDFFFEFDF70A404AEE9E95" box="[536,677,368,392]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B929EAE1FFCDFFFEFDF70A404AEE9E95" box="[536,677,368,392]" italics="true" pageId="25" pageNumber="476">D. dimidiata</emphasis>
</taxonomicName>
, best seen in lateral view (cf.
<figureCitation id="13662A76FFCDFFFEFC1B0A5E4CE89E95" box="[1012,1187,367,392]" captionStart="FIGURES 45 – 48" captionStartId="23.[151,264,1735,1758]" captionTargetBox="[162,1365,783,1707]" captionTargetId="figure@23.[151,1436,721,1720]" captionTargetPageId="23" captionText="FIGURES 45 – 48. Penis, lateral view: 45) D. dimidiata west Java; 46) D. lugens Selekau River, Kalimantan Barat; 47) D. ulu holotype; 48) D. vanida paratype, Khlong Nakha, Ranong." httpUri="https://zenodo.org/record/243473/files/figure.png" pageId="25" pageNumber="476">Figs. 45 and 47</figureCitation>
), and is more rounded in ventral view (cf.
<figureCitation id="13662A76FFCDFFFEFE9C0AA54A6F9EB1" box="[371,548,404,429]" captionStart="FIGURES 41 – 44" captionStartId="21.[151,264,1790,1813]" captionTargetBox="[162,1384,617,1756]" captionTargetId="figure@21.[151,1436,555,1769]" captionTargetPageId="21" captionText="FIGURES 41 – 44. Penis, ventral view: 41) D. dimidiata west Java; 42) D. lugens Selekau River, Kalimantan Barat; 43) D. ulu holotype; 44) D. vanida paratype, Khlong Nakha, Ranong." httpUri="https://zenodo.org/record/243472/files/figure.png" pageId="25" pageNumber="476">Figs. 41 and 43</figureCitation>
).
<emphasis id="B929EAE1FFCDFFFEFDD80AA54AC59EB0" bold="true" box="[567,654,404,429]" pageId="25" pageNumber="476">Female</emphasis>
:
<taxonomicName id="4C5D4D70FFCDFFFEFD720AA44AA89EB1" box="[669,739,404,428]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="ulu">
<emphasis id="B929EAE1FFCDFFFEFD720AA44AA89EB1" box="[669,739,404,428]" italics="true" pageId="25" pageNumber="476">D. ulu</emphasis>
</taxonomicName>
is easy to separate by the distinct opaque streak in both wings (
<figureCitation id="13662A76FFCDFFFEFF700A8648B89ECD" box="[159,243,439,464]" captionStart="FIGURES 49 – 52" captionStartId="24.[151,264,1070,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@24.[151,1436,193,1049]" captionTargetPageId="24" captionText="FIGURES 49 – 52. Habitus of female Dysphaea: 49) D. dimidiata west Java; 50) D. lugens Mount Dulit, Sarawak, photograph by Benjamin Price; 51) D. ulu paratype; 52) D. vanida paratype Kanchanaburi." httpUri="https://zenodo.org/record/243474/files/figure.png" pageId="25" pageNumber="476">Fig. 51</figureCitation>
). In
<taxonomicName id="4C5D4D70FFCDFFFEFEC80A8849FD9ECD" box="[295,438,440,464]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B929EAE1FFCDFFFEFEC80A8849FD9ECD" box="[295,438,440,464]" italics="true" pageId="25" pageNumber="476">D. dimidiata</emphasis>
</taxonomicName>
the wings are largely hyaline or semihyaline with brownish tinge, the tips being slightly darkened (
<figureCitation id="13662A76FFCDFFFEFEE20AED49299EE9" box="[269,354,476,501]" captionStart="FIGURES 49 – 52" captionStartId="24.[151,264,1070,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@24.[151,1436,193,1049]" captionTargetPageId="24" captionText="FIGURES 49 – 52. Habitus of female Dysphaea: 49) D. dimidiata west Java; 50) D. lugens Mount Dulit, Sarawak, photograph by Benjamin Price; 51) D. ulu paratype; 52) D. vanida paratype Kanchanaburi." httpUri="https://zenodo.org/record/243474/files/figure.png" pageId="25" pageNumber="476">Fig. 49</figureCitation>
). In
<taxonomicName id="4C5D4D70FFCDFFFEFE7B0AEC49919EE9" box="[404,474,476,500]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="ulu">
<emphasis id="B929EAE1FFCDFFFEFE7B0AEC49919EE9" box="[404,474,476,500]" italics="true" pageId="25" pageNumber="476">D. ulu</emphasis>
</taxonomicName>
the yellowish stripes on thorax are narrower (
<figureCitation id="13662A76FFCDFFFEFC300AED4C7A9EE8" box="[991,1073,476,501]" captionStart="FIGURES 63 – 68" captionStartId="28.[151,264,1699,1722]" captionTargetBox="[151,1436,185,1683]" captionTargetId="figure@28.[151,1436,185,1684]" captionTargetPageId="28" captionText="FIGURES 63 – 68. Female synthorax, dorsal view: 63) D. dimidiata west Java; 64) D. dimidiata Sarawak; 65) D. dimidiata Johor; 66) D. lugens Gunung Sari, Kaliamantan Timur; 67) D. ulu paratype; 68) D. vanida paratype Kanchanaburi." httpUri="https://zenodo.org/record/243477/files/figure.png" pageId="25" pageNumber="476">Fig. 67</figureCitation>
) than in
<taxonomicName id="4C5D4D70FFCDFFFEFB7A0AEC4D689EE9" box="[1173,1315,476,500]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B929EAE1FFCDFFFEFB7A0AEC4D689EE9" box="[1173,1315,476,500]" italics="true" pageId="25" pageNumber="476">D. dimidiata</emphasis>
</taxonomicName>
(
<figureCitation id="13662A76FFCDFFFEFADD0AED48FC9D05" captionStart="FIGURES 63 – 68" captionStartId="28.[151,264,1699,1722]" captionTargetBox="[151,1436,185,1683]" captionTargetId="figure@28.[151,1436,185,1684]" captionTargetPageId="28" captionText="FIGURES 63 – 68. Female synthorax, dorsal view: 63) D. dimidiata west Java; 64) D. dimidiata Sarawak; 65) D. dimidiata Johor; 66) D. lugens Gunung Sari, Kaliamantan Timur; 67) D. ulu paratype; 68) D. vanida paratype Kanchanaburi." httpUri="https://zenodo.org/record/243477/files/figure.png" pageId="25" pageNumber="476">Figs. 63– 65</figureCitation>
).
<taxonomicName id="4C5D4D70FFCDFFFEFF240930495A9D05" box="[203,273,512,536]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="ulu">
<emphasis id="B929EAE1FFCDFFFEFF240930495A9D05" box="[203,273,512,536]" italics="true" pageId="25" pageNumber="476">D. ulu</emphasis>
</taxonomicName>
lacks the conspicuous yellow stripe on antefrons (
<figureCitation id="13662A76FFCDFFFEFCA40ACE4BD49D05" box="[843,927,511,536]" captionStart="FIGURES 53 – 56" captionStartId="25.[151,264,1609,1632]" captionTargetBox="[151,1436,625,1588]" captionTargetId="figure@25.[151,1436,625,1588]" captionTargetPageId="25" captionText="FIGURES 53 – 56. Female head: 53) D. dimidiata Sarawak, dorsal view; 54) D. lugens Gunung Sari, Kalimantan Timur, dorsal view; 55) D. ulu paratype, dorsal, slightly frontal view; 56) D. vanida paratype, Kanchanaburi, dorsal, slightly frontal view." httpUri="https://zenodo.org/record/243475/files/figure.png" pageId="25" pageNumber="476">Fig. 55</figureCitation>
), which is often present in
<taxonomicName id="4C5D4D70FFCDFFFEFB3E09304D149D05" box="[1233,1375,512,536]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B929EAE1FFCDFFFEFB3E09304D149D05" box="[1233,1375,512,536]" italics="true" pageId="25" pageNumber="476">D. dimidiata</emphasis>
</taxonomicName>
(
<figureCitation id="13662A76FFCDFFFEFA810ACE48FC9D20" captionStart="FIGURES 53 – 56" captionStartId="25.[151,264,1609,1632]" captionTargetBox="[151,1436,625,1588]" captionTargetId="figure@25.[151,1436,625,1588]" captionTargetPageId="25" captionText="FIGURES 53 – 56. Female head: 53) D. dimidiata Sarawak, dorsal view; 54) D. lugens Gunung Sari, Kalimantan Timur, dorsal view; 55) D. ulu paratype, dorsal, slightly frontal view; 56) D. vanida paratype, Kanchanaburi, dorsal, slightly frontal view." httpUri="https://zenodo.org/record/243475/files/figure.png" pageId="25" pageNumber="476">Fig. 53</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF22667BFFCDFFFEFF780D784D319963" httpUri="https://zenodo.org/record/243475/files/figure.png" pageId="25" pageNumber="476" targetBox="[151,1436,625,1588]" targetPageId="25">
<paragraph id="8BE236F3FFCDFFFEFF780D784D319963" blockId="25.[151,1436,1609,1663]" pageId="25" pageNumber="476">
<emphasis id="B929EAE1FFCDFFFEFF780D78491F9942" bold="true" box="[151,340,1609,1632]" pageId="25" pageNumber="476">FIGURES 53–56.</emphasis>
Female head:
<emphasis id="B929EAE1FFCDFFFEFE080D784A43997D" bold="true" box="[487,520,1609,1632]" pageId="25" pageNumber="476">53)</emphasis>
<taxonomicName id="4C5D4D70FFCDFFFEFDE00D7A4AC4997D" box="[527,655,1610,1632]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B929EAE1FFCDFFFEFDE00D7A4AC4997D" box="[527,655,1610,1632]" italics="true" pageId="25" pageNumber="476">D. dimidiata</emphasis>
</taxonomicName>
Sarawak, dorsal view;
<emphasis id="B929EAE1FFCDFFFEFC930D784BD6997D" bold="true" box="[892,925,1609,1632]" pageId="25" pageNumber="476">54)</emphasis>
<taxonomicName id="4C5D4D70FFCDFFFEFC4B0D7A4CC5997D" authority="Gunung Sari" authorityName="Gunung Sari" box="[932,1166,1610,1632]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="lugens">
<emphasis id="B929EAE1FFCDFFFEFC4B0D7A4C4F997D" box="[932,1028,1610,1632]" italics="true" pageId="25" pageNumber="476">D. lugens</emphasis>
Gunung Sari
</taxonomicName>
, Kalimantan Timur, dorsal view;
<emphasis id="B929EAE1FFCDFFFEFF3A0D5948BC9962" bold="true" box="[213,247,1640,1663]" pageId="25" pageNumber="476">55)</emphasis>
<taxonomicName id="4C5D4D70FFCDFFFEFF120D5B49779963" box="[253,316,1640,1663]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="ulu">
<emphasis id="B929EAE1FFCDFFFEFF120D5B49779963" box="[253,316,1640,1663]" italics="true" pageId="25" pageNumber="476">D. ulu</emphasis>
</taxonomicName>
paratype, dorsal, slightly frontal view;
<emphasis id="B929EAE1FFCDFFFEFD230D594AA59962" bold="true" box="[716,750,1640,1663]" pageId="25" pageNumber="476">56)</emphasis>
<taxonomicName id="4C5D4D70FFCDFFFEFD1B0D5B4B139963" box="[756,856,1640,1663]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="vanida">
<emphasis id="B929EAE1FFCDFFFEFD1B0D5B4B139963" box="[756,856,1640,1663]" italics="true" pageId="25" pageNumber="476">D. vanida</emphasis>
</taxonomicName>
paratype, Kanchanaburi, dorsal, slightly frontal view.
</paragraph>
</caption>
<subSubSection id="C3476578FFCDFFFEFF280D9A4D219832" pageId="25" pageNumber="476" type="discussion">
<paragraph id="8BE236F3FFCDFFFEFF280D9A4D219832" blockId="25.[151,1437,1707,1840]" pageId="25" pageNumber="476">
<emphasis id="B929EAE1FFCDFFFEFF280D9A497099D9" bold="true" box="[199,315,1707,1732]" pageId="25" pageNumber="476">Remarks.</emphasis>
Some published records on Bornean
<taxonomicName id="4C5D4D70FFCDFFFEFD1C0D9A4B2899DE" box="[755,867,1707,1731]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAE1FFCDFFFEFD1C0D9A4B2899DE" box="[755,867,1707,1731]" italics="true" pageId="25" pageNumber="476">Dysphaea</emphasis>
</taxonomicName>
species still remain uncertain as regards the real identity of the species. The
<taxonomicName id="4C5D4D70FFCDFFFEFE3A0DE14A2E99FA" box="[469,613,1743,1767]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B929EAE1FFCDFFFEFE3A0DE14A2E99FA" box="[469,613,1743,1767]" italics="true" pageId="25" pageNumber="476">D. dimidiata</emphasis>
</taxonomicName>
record from Lanjak Entimau Wildlife Sanctuary in
<collectingCountry id="F34A7663FFCDFFFEFB580DFE4D5C99F5" box="[1207,1303,1743,1768]" name="Malaysia" pageId="25" pageNumber="476">Sarawak</collectingCountry>
by Norma- Rashid &amp; al. (2010, p. 326) could refer to either to
<taxonomicName id="4C5D4D70FFCDFFFEFD330DC54B209816" box="[732,875,1779,1803]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B929EAE1FFCDFFFEFD330DC54B209816" box="[732,875,1779,1803]" italics="true" pageId="25" pageNumber="476">D. dimidiata</emphasis>
</taxonomicName>
or
<taxonomicName id="4C5D4D70FFCDFFFEFC7D0DC54B929816" box="[914,985,1779,1803]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="ulu">
<emphasis id="B929EAE1FFCDFFFEFC7D0DC54B929816" box="[914,985,1779,1803]" italics="true" pageId="25" pageNumber="476">D. ulu</emphasis>
</taxonomicName>
; both species are known to occur there. The
<taxonomicName id="4C5D4D70FFCDFFFEFF260C2949139832" box="[201,344,1815,1839]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B929EAE1FFCDFFFEFF260C2949139832" box="[201,344,1815,1839]" italics="true" pageId="25" pageNumber="476">D. dimidiata</emphasis>
</taxonomicName>
record from ‘
<collectingCountry id="F34A7663FFCDFFFEFE010C264A1D982D" box="[494,598,1815,1840]" name="Malaysia" pageId="25" pageNumber="476">Sarawak</collectingCountry>
, Kampong Seku’ by
<bibRefCitation id="EFCC4B02FFCDFFFEFCAD0C264BBB982D" author="Asahina" box="[834,1008,1815,1840]" pageId="25" pageNumber="476" refString="Asahina, S. (1966) Sarawak Odonata taken by Mr. H. Hayashi. Tombo - Acta Odonatologica, 9 (1 - 4), 28 - 30." type="journal article" year="1966">Asahina (1966)</bibRefCitation>
might just as well refer to
<taxonomicName id="4C5D4D70FFCDFFFEFAF30C294D289832" box="[1308,1379,1815,1839]" class="Insecta" family="Euphaeidae" genus="Dysphaea" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="ulu">
<emphasis id="B929EAE1FFCDFFFEFAF30C294D289832" box="[1308,1379,1815,1839]" italics="true" pageId="25" pageNumber="476">D. ulu</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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