2573 lines
387 KiB
XML
2573 lines
387 KiB
XML
<document id="0C0C2905086A0B94AE73C23C8A8F4552" ID-CLB-Dataset="29472" ID-DOI="10.11646/zootaxa.4471.3.1" ID-GBIF-Dataset="51adf720-0d97-45dd-ac98-c3c79ba4d3ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="1439790" ID-ZooBank="294764A6-5EE6-40DC-B05C-BE23398A4B29" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538365037966" checkinUser="plazi" docAuthor="Li, Fengyuan, Liu, Chang, Wongprom, Prasit & Li, Shuqiang" docDate="2018" docId="03F487D1B63AFFFCFF4CF9EF0550F91D" docLanguage="en" docName="zootaxa.4471.3.1.pdf" docOrigin="Zootaxa 4471 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Althepus Thorell 1898" docType="treatment" docVersion="7" lastPageNumber="412" masterDocId="FFCDFFA9B63BFFF7FFDBFFBB0012FFC2" masterDocTitle="Sixteen new species of the spider genus Althepus Thorell, 1898 (Araneae: Ochyroceratidae) from Southeast Asia" masterLastPageNumber="445" masterPageNumber="401" pageNumber="402" updateTime="1698558335820" updateUser="ExternalLinkService">
|
||
<mods:mods id="4109641520F9A46D64292D3CFE181C1D" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="243F58D165EC1224CE051EE181217582">
|
||
<mods:title id="6B85ED957B6F19ABD1BD8DF166E2948D">Sixteen new species of the spider genus Althepus Thorell, 1898 (Araneae: Ochyroceratidae) from Southeast Asia</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="CD9F003310EF9D0B8E1A46868926D101" type="personal">
|
||
<mods:role id="73EF230370FFF9957DDC16104211FABB">
|
||
<mods:roleTerm id="53A2E68DD91AFBDBAC044BDAA1F36B6C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="54053526700DF2A34FCAC79329A77108">Li, Fengyuan</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="72ECD0340EFD9E50E5D9BD9642F0F36B" type="personal">
|
||
<mods:role id="8B4D90E1D6453662438BCEFE89CD3819">
|
||
<mods:roleTerm id="FD82A879B49D8309303D11C0BACE6F14">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="3BDCBEC59C551507C7F52B53C3CC4435">Liu, Chang</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="2968770B1F8CFA3849F3AE9D05920884" type="personal">
|
||
<mods:role id="4C19969291ABE98DFCF084354E64276A">
|
||
<mods:roleTerm id="0EBEC1CC9998C56182134235766D83CC">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B26C879155547F5046EEFF1FD073096D">Wongprom, Prasit</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="1A493BFCCD13DD1CC5FF598ED2733711" type="personal">
|
||
<mods:role id="4FE2F6DC78E95A2FEC9424CF72EEA64D">
|
||
<mods:roleTerm id="7972E3D209A865D44C03687DB91A9E00">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="FEC8F4807A8D8A31B39827CBE7318CDB">Li, Shuqiang</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="C4B8EF71AEC79C936FB1B31F7798E17A">text</mods:typeOfResource>
|
||
<mods:relatedItem id="BA37C88C8309F3AEB1D71D0FDEA015E2" type="host">
|
||
<mods:titleInfo id="6D9B2A4948A6C4C994EBC9FDA40AEC89">
|
||
<mods:title id="039718FC14F13501C796F7DAF0A0CD17">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="B4F3A3D82CD3602026A489C81D7CE79A">
|
||
<mods:date id="EB12A99219E292B377FE59A62BE144DA">2018</mods:date>
|
||
<mods:detail id="2EC3D0C9A35C394D0656CFC34DB632FB" type="pubDate">
|
||
<mods:number id="5FFEF687FAD7933514CE873DCFFD8DB1">2018-09-06</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="187A7F87E98E0AE6012653CAC9EB3998" type="volume">
|
||
<mods:number id="AE3F2B29B08E3318723713F4EC6ACA8A">4471</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="06AC41229B40CB421FC55902AF32B4FF" type="issue">
|
||
<mods:number id="4D8F355BF8FEDACC97F81412E1C29AC5">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="AF3B27A14FBBE656E5FE262C57DA3BAE" unit="page">
|
||
<mods:start id="67CBF6F61EEA7EDE24A6E74A025B52FC">401</mods:start>
|
||
<mods:end id="D4F9FABD772FEE98D6FA7697468D329C">445</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="BB2CD611007F93C35829EC6BCF3D3D90">journal article</mods:classification>
|
||
<mods:identifier id="99B943F3C5C08DE7D33B54C064467B07" type="CLB-Dataset">29472</mods:identifier>
|
||
<mods:identifier id="BD6E2D2B420BBE1EFA70AF2A8E595C00" type="DOI">10.11646/zootaxa.4471.3.1</mods:identifier>
|
||
<mods:identifier id="BBD8D304B925BF7725420F8BEE24758D" type="GBIF-Dataset">51adf720-0d97-45dd-ac98-c3c79ba4d3ce</mods:identifier>
|
||
<mods:identifier id="24F75C3DD9C9688B9358610F372D61FD" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="84389BD805DE9BC151F93B26B0063385" type="Zenodo-Dep">1439790</mods:identifier>
|
||
<mods:identifier id="3CED7BA861BBA7F7A8C5AE8DF2A5E343" type="ZooBank">294764A6-5EE6-40DC-B05C-BE23398A4B29</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03F487D1B63AFFFCFF4CF9EF0550F91D" ID-DOI="http://doi.org/10.5281/zenodo.5997333" ID-GBIF-Taxon="148415768" ID-Zenodo-Dep="5997333" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D" httpUri="http://treatment.plazi.org/id/03F487D1B63AFFFCFF4CF9EF0550F91D" lastPageId="11" lastPageNumber="412" pageId="1" pageNumber="402">
|
||
<subSubSection id="C347654CB63AFFF6FF4CF9EF0215F9AC" box="[151,519,1620,1646]" pageId="1" pageNumber="402" type="nomenclature">
|
||
<paragraph id="8BE236C7B63AFFF6FF4CF9EF0215F9AC" blockId="1.[151,519,1620,1646]" box="[151,519,1620,1646]" pageId="1" pageNumber="402">
|
||
<heading id="D0AA81ABB63AFFF6FF4CF9EF0215F9AC" bold="true" box="[151,519,1620,1646]" fontSize="11" level="1" pageId="1" pageNumber="402" reason="1">
|
||
<emphasis id="B929EAD5B63AFFF6FF4CF9EF0215F9AC" bold="true" box="[151,519,1620,1646]" pageId="1" pageNumber="402">
|
||
Genus
|
||
<taxonomicName id="4C5D4D44B63AFFF6FF35F9EF0215F9AC" authority="Thorell, 1898" authorityName="Thorell" authorityYear="1898" box="[238,519,1620,1646]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="402" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B929EAD5B63AFFF6FF35F9EF0148F9AC" bold="true" box="[238,346,1620,1646]" italics="true" pageId="1" pageNumber="402">Althepus</emphasis>
|
||
Thorell, 1898
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63AFFF6FF4CF9200243F972" box="[151,593,1690,1712]" pageId="1" pageNumber="402" type="materials_examined">
|
||
<paragraph id="8BE236C7B63AFFF6FF4CF9200243F972" blockId="1.[151,593,1690,1712]" box="[151,593,1690,1712]" pageId="1" pageNumber="402">
|
||
<typeStatus id="54E68865B63AFFF6FF4CF92000DBF972" box="[151,201,1691,1712]" pageId="1" pageNumber="402">Type</typeStatus>
|
||
species.
|
||
<taxonomicName id="4C5D4D44B63AFFF6FEFEF9210243F972" authority="Thorell, 1898" authorityName="Thorell" authorityYear="1898" box="[293,593,1690,1712]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="pictus">
|
||
<emphasis id="B929EAD5B63AFFF6FEFEF92101D2F972" box="[293,448,1690,1712]" italics="true" pageId="1" pageNumber="402">Althepus pictus</emphasis>
|
||
Thorell, 1898
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63AFFF6FF4CF966054EF8A0" pageId="1" pageNumber="402" type="diagnosis">
|
||
<paragraph id="8BE236C7B63AFFF6FF4CF966054EF8A0" blockId="1.[151,1436,1757,1998]" pageId="1" pageNumber="402">
|
||
<emphasis id="B929EAD5B63AFFF6FF4CF9660194F934" bold="true" box="[151,390,1757,1782]" pageId="1" pageNumber="402">Emended diagnosis.</emphasis>
|
||
The genus
|
||
<taxonomicName id="4C5D4D44B63AFFF6FDD0F966027FF937" authorityName="Thorell" authorityYear="1898" box="[523,621,1757,1781]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="402" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B929EAD5B63AFFF6FDD0F966027FF937" box="[523,621,1757,1781]" italics="true" pageId="1" pageNumber="402">Althepus</emphasis>
|
||
</taxonomicName>
|
||
belongs to Psilodercinae and can be distinguished from other genera of the subfamily by the following combination of characters: cheliceral promargin with lamina and 1–2 teeth, retromargin with 1–2 small teeth; tarsus of male palp with lateral protrusion bearing a hook-shaped spine; short bulb with embolus; and female internal genitalia often with paired spermathecae (Deeleman-Reinhold 1995).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63AFFF5FF1CF8D6015EFF11" lastPageId="2" lastPageNumber="403" pageId="1" pageNumber="402" type="discussion">
|
||
<paragraph id="8BE236C7B63AFFF6FF1CF8D600FBF80C" blockId="1.[151,1436,1757,1998]" pageId="1" pageNumber="402">
|
||
<emphasis id="B929EAD5B63AFFF6FF1CF8D60129F844" bold="true" box="[199,315,1901,1926]" pageId="1" pageNumber="402">Remarks.</emphasis>
|
||
According to our observations, we used hook-shaped spine instead the "lanceolate apophysis" used by Deeleman-Reinhold (1995), and used lamina and 1–2 teeth instead the "3 teeth" used by Deeleman-Reinhold (1995).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B639FFF5FF4CFF2C022FFF73" blockId="2.[151,573,151,211]" box="[151,573,151,178]" pageId="2" pageNumber="403">
|
||
<taxonomicName id="4C5D4D44B639FFF5FF4CFF2C01CFFF70" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[151,477,151,178]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="bamensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B639FFF5FF4CFF2C01CFFF70" bold="true" box="[151,477,151,178]" pageId="2" pageNumber="403">
|
||
<emphasis id="B929EAD5B639FFF5FF4CFF2C0168FF73" bold="true" box="[151,378,151,177]" italics="true" pageId="2" pageNumber="403">Althepus bamensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B639FFF5FE7FFF2C01ABFF73" bold="true" box="[420,441,151,177]" italics="true" pageId="2" pageNumber="403">et</emphasis>
|
||
Li
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB639FFF5FE3FFF23022FFF73" box="[484,573,152,177]" pageId="2" pageNumber="403" rank="species">
|
||
<emphasis id="B929EAD5B639FFF5FE3FFF23022FFF73" bold="true" box="[484,573,152,177]" pageId="2" pageNumber="403">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B639FFF5FF4CFF01015EFF11" blockId="2.[151,573,151,211]" box="[151,332,186,211]" pageId="2" pageNumber="403">
|
||
<figureCitation id="13662A42B639FFF5FF4CFF0100E8FF11" box="[151,250,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="13.[151,250,1933,1955]" captionStartId-1="14.[151,250,1934,1956]" captionTargetBox-0="[188,1343,193,1908]" captionTargetBox-1="[207,1365,193,1896]" captionTargetId-0="figure@13.[188,1356,190,1908]" captionTargetId-1="figure@14.[207,1365,193,1896]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURE 1. Althepus bamensis Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 1A and 1C as in figure 1D." captionText-1="FIGURE 2. Althepus bamensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 2D as in figure 2E." httpUri-0="https://zenodo.org/record/1439792/files/figure.png" httpUri-1="https://zenodo.org/record/1439794/files/figure.png" pageId="2" pageNumber="403">Figs 1–2</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B639FFF5FEDDFF000136FF11" box="[262,292,187,211]" captionStart="FIGURE 30" captionStartId="42.[151,250,1917,1939]" captionTargetBox="[177,1410,193,1896]" captionTargetId="figure@42.[177,1410,193,1896]" captionTargetPageId="42" captionText="FIGURE 30. Left cheliceral retromargin of male holotype. A, Althepus bamensis Li et Li sp. nov.; B, A. changmao Li et Li sp. nov.; C, A. dongnaiensis Li et Li sp. nov.; D, A. guan Li et Li sp. nov.; E, A. maechamensis Li et Li sp. nov.; F, A. menglaensis Li et Li sp. nov.; G, A. naphongensis Li et Li sp. nov.; H, A. natmataungensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439850/files/figure.png" pageId="2" pageNumber="403">30</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B639FFF5FEEBFF00015EFF11" box="[304,332,187,211]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="2" pageNumber="403">32</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB639FFF5FF4CFEB903ABFEA1" pageId="2" pageNumber="403" type="materials_examined">
|
||
<paragraph id="8BE236C7B639FFF5FF4CFEB903ABFEA1" blockId="2.[151,1437,258,1255]" pageId="2" pageNumber="403">
|
||
<materialsCitation id="3B353C9AB639FFF5FF4CFEB904EAFEFD" ID-GBIF-Occurrence="1913888973" collectingDate="2016-11-19" country="Thailand" elevation="169" latitude="17.276716" location="Bam Thung Tham Sub District" longLatPrecision="1" longitude="98.19162" municipality="Tha Song Yang District" pageId="2" pageNumber="403" specimenCount="2" stateProvince="Tak Province" typeStatus="holotype">
|
||
<emphasis id="B929EAD5B639FFF5FF4CFEB901D0FED9" bold="true" box="[151,450,258,283]" pageId="2" pageNumber="403">
|
||
<typeStatus id="54E68865B639FFF5FF4CFEB900C0FED9" box="[151,210,258,283]" pageId="2" pageNumber="403">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68865B639FFF5FE97FEB901AFFED9" box="[332,445,258,283]" pageId="2" pageNumber="403" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male (IZCAS-Ar 33986),
|
||
<collectingCountry id="F34A7657B639FFF5FD37FEB90340FED9" box="[748,850,258,283]" name="Thailand" pageId="2" pageNumber="403">Thailand</collectingCountry>
|
||
,
|
||
<collectingRegion id="4999F825B639FFF5FC85FEB903E4FED9" box="[862,1014,258,283]" country="Thailand" name="Tak" pageId="2" pageNumber="403">Tak Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B86ACBDB639FFF5FBDBFEB9051EFED9" box="[1024,1292,258,283]" pageId="2" pageNumber="403">Tha Song Yang District</collectingMunicipality>
|
||
,
|
||
<location id="8E82601CB639FFF5FACCFEB8017EFEFD" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB639FFF5FACCFEB8017EFEFD" country="Thailand" latitude="17.276716" longLatPrecision="1" longitude="98.19162" municipality="Tha Song Yang District" name="Bam Thung Tham Sub District" pageId="2" pageNumber="403" stateProvince="Tak Province">Bam Thung Tham Sub District</location>
|
||
, unnamed cave,
|
||
<geoCoordinate id="EE695000B639FFF5FDF7FE9D02AEFEFD" box="[556,700,294,319]" direction="north" orientation="latitude" pageId="2" pageNumber="403" precision="1" value="17.276716">17°16.603'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695000B639FFF5FD12FE9C0347FEFD" box="[713,853,295,319]" direction="east" orientation="longitude" pageId="2" pageNumber="403" precision="1" value="98.19162">98°11.497'E</geoCoordinate>
|
||
,
|
||
<elevation id="0070D1F4B639FFF5FCB8FE9D03BEFEFC" box="[867,940,294,319]" metricMagnitude="2" metricUnit="m" metricValue="1.69" pageId="2" pageNumber="403" unit="m" value="169.0">
|
||
<quantity id="4CA59B22B639FFF5FCB8FE9D03BEFEFC" box="[867,940,294,319]" metricMagnitude="2" metricUnit="m" metricValue="1.69" pageId="2" pageNumber="403" unit="m" value="169.0">169 m</quantity>
|
||
</elevation>
|
||
altitude,
|
||
<collectingDate id="EFA7E9EFB639FFF5FBC3FE9C04EAFEFD" box="[1048,1272,294,319]" pageId="2" pageNumber="403" value="2016-11-19">
|
||
<date id="FFE31007B639FFF5FBC3FE9C04EAFEFD" box="[1048,1272,294,319]" pageId="2" pageNumber="403" value="2016-11-19">19 November 2016</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B353C9AB639FFF5FADFFE9D03ABFEA1" ID-GBIF-Occurrence="1913888979" collectingDate="2016-11-19" country="Thailand" elevation="169" latitude="17.276716" location="Bam Thung Tham Sub District" longLatPrecision="1" longitude="98.19162" municipality="Tha Song Yang District" pageId="2" pageNumber="403" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Tak Province" typeStatus="paratype">
|
||
<emphasis id="B929EAD5B639FFF5FADFFE9D0597FEFD" bold="true" box="[1284,1413,294,319]" pageId="2" pageNumber="403">
|
||
<typeStatus id="54E68865B639FFF5FADFFE9D0592FEFD" box="[1284,1408,294,319]" pageId="2" pageNumber="403" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D5BFD4EB639FFF5FA56FE9C00CAFEA1" pageId="2" pageNumber="403" type="male">2 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D5BFD4EB639FFF5FED5FEF0016CFEA1" box="[270,382,330,355]" pageId="2" pageNumber="403" type="female">2 females</specimenCount>
|
||
, same data as holotype (IZCAS-Ar 33987–33990).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB639FFF5FF1CFED5040FFE45" box="[199,1053,366,391]" pageId="2" pageNumber="403" type="etymology">
|
||
<paragraph id="8BE236C7B639FFF5FF1CFED5040FFE45" blockId="2.[151,1437,258,1255]" box="[199,1053,366,391]" pageId="2" pageNumber="403">
|
||
<emphasis id="B929EAD5B639FFF5FF1CFED5015CFE45" bold="true" box="[199,334,366,391]" pageId="2" pageNumber="403">Etymology.</emphasis>
|
||
The specific name is an adjective and refers to the
|
||
<typeStatus id="54E68865B639FFF5FC57FED403AEFE45" box="[908,956,367,391]" pageId="2" pageNumber="403">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB639FFF5FF1CFE290333FE31" pageId="2" pageNumber="403" type="diagnosis">
|
||
<paragraph id="8BE236C7B639FFF5FF1CFE290333FE31" blockId="2.[151,1437,258,1255]" pageId="2" pageNumber="403">
|
||
<emphasis id="B929EAD5B639FFF5FF1CFE290153FE69" bold="true" box="[199,321,402,427]" pageId="2" pageNumber="403">Diagnosis.</emphasis>
|
||
This new species can be easily distinguished from all the other species by their short, slightly sigmoid embolus (
|
||
<figureCitation id="13662A42B639FFF5FEABFE0D01DEFE0D" box="[368,460,438,463]" captionStart="FIGURE 1" captionStartId="13.[151,250,1933,1955]" captionTargetBox="[188,1343,193,1908]" captionTargetId="figure@13.[188,1356,190,1908]" captionTargetPageId="13" captionText="FIGURE 1. Althepus bamensis Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 1A and 1C as in figure 1D." httpUri="https://zenodo.org/record/1439792/files/figure.png" pageId="2" pageNumber="403">Fig. 1A</figureCitation>
|
||
) and their horn-shaped distal part of the conductor in males; and by two pairs of spermathecae with irregular margins in females (
|
||
<figureCitation id="13662A42B639FFF5FD61FE610306FE31" box="[698,788,474,499]" captionStart="FIGURE 2" captionStartId="14.[151,250,1934,1956]" captionTargetBox="[207,1365,193,1896]" captionTargetId="figure@14.[207,1365,193,1896]" captionTargetPageId="14" captionText="FIGURE 2. Althepus bamensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 2D as in figure 2E." httpUri="https://zenodo.org/record/1439794/files/figure.png" pageId="2" pageNumber="403">Fig. 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB639FFF5FF1CFE45040AFBB9" pageId="2" pageNumber="403" type="description">
|
||
<paragraph id="8BE236C7B639FFF5FF1CFE450227FC61" blockId="2.[151,1437,258,1255]" pageId="2" pageNumber="403">
|
||
<emphasis id="B929EAD5B639FFF5FF1CFE450237FDD5" bold="true" box="[199,549,510,535]" pageId="2" pageNumber="403">
|
||
Description. Male (
|
||
<typeStatus id="54E68865B639FFF5FE71FE450205FDD5" box="[426,535,510,535]" pageId="2" pageNumber="403" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 4.48; carapace 1.48 length, 1.52 width; abdomen 2.69 length, 1.62 width. Carapace round (
|
||
<figureCitation id="13662A42B639FFF5FE73FD990213FDF9" box="[424,513,546,571]" captionStart="FIGURE 2" captionStartId="14.[151,250,1934,1956]" captionTargetBox="[207,1365,193,1896]" captionTargetId="figure@14.[207,1365,193,1896]" captionTargetPageId="14" captionText="FIGURE 2. Althepus bamensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 2D as in figure 2E." httpUri="https://zenodo.org/record/1439794/files/figure.png" pageId="2" pageNumber="403">Fig. 2C</figureCitation>
|
||
), yellow, with three longitudinal brown bands of different widths, the middle one shortened to form a triangular band around eyes. Anterior margin of thoracic region distinctly elevated. Clypeus brown. Cheliceral promargin with two teeth, followed by a lamina, retromargin with two small teeth, posterior surface of fang with 20 small denticles (
|
||
<figureCitation id="13662A42B639FFF5FDA3FD3502F4FD65" box="[632,742,654,679]" captionStart="FIGURE 30" captionStartId="42.[151,250,1917,1939]" captionTargetBox="[177,1410,193,1896]" captionTargetId="figure@42.[177,1410,193,1896]" captionTargetPageId="42" captionText="FIGURE 30. Left cheliceral retromargin of male holotype. A, Althepus bamensis Li et Li sp. nov.; B, A. changmao Li et Li sp. nov.; C, A. dongnaiensis Li et Li sp. nov.; D, A. guan Li et Li sp. nov.; E, A. maechamensis Li et Li sp. nov.; F, A. menglaensis Li et Li sp. nov.; G, A. naphongensis Li et Li sp. nov.; H, A. natmataungensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439850/files/figure.png" pageId="2" pageNumber="403">Fig. 30A</figureCitation>
|
||
). Labium brown. Sternum brown. Abdomen elongate, with complex pattern of white and dark-brown dorsally and ventrally. Legs brown, femur and tibia with white annulations. Leg measurements: I - (11.86, 0.63, 11.79, 21.15, -), II missing, III 17.32 (5.00, 0.57, 4.25, 6.28, 1.22), IV 24.76 (6.99, 0.59, 6.54, 9.04, 1.60). Male palp (
|
||
<figureCitation id="13662A42B639FFF5FD11FD41035EFCD1" box="[714,844,762,787]" captionStart="FIGURE 1" captionStartId="13.[151,250,1933,1955]" captionTargetBox="[188,1343,193,1908]" captionTargetId="figure@13.[188,1356,190,1908]" captionTargetPageId="13" captionText="FIGURE 1. Althepus bamensis Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 1A and 1C as in figure 1D." httpUri="https://zenodo.org/record/1439792/files/figure.png" pageId="2" pageNumber="403">Figs 1A–D</figureCitation>
|
||
): tarsus with three slightly curved, serrated bristles at the top of cymbial protrusion (two of them missing,
|
||
<figureCitation id="13662A42B639FFF5FD33FCA5032DFCF5" box="[744,831,798,823]" captionStart="FIGURE 1" captionStartId="13.[151,250,1933,1955]" captionTargetBox="[188,1343,193,1908]" captionTargetId="figure@13.[188,1356,190,1908]" captionTargetPageId="13" captionText="FIGURE 1. Althepus bamensis Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 1A and 1C as in figure 1D." httpUri="https://zenodo.org/record/1439792/files/figure.png" pageId="2" pageNumber="403">Fig. 1D</figureCitation>
|
||
), one hooked spine with tip directed proximally (
|
||
<figureCitation id="13662A42B639FFF5FAB4FCA500A9FC99" captionStart="FIGURE 1" captionStartId="13.[151,250,1933,1955]" captionTargetBox="[188,1343,193,1908]" captionTargetId="figure@13.[188,1356,190,1908]" captionTargetPageId="13" captionText="FIGURE 1. Althepus bamensis Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 1A and 1C as in figure 1D." httpUri="https://zenodo.org/record/1439792/files/figure.png" pageId="2" pageNumber="403">Fig. 1A</figureCitation>
|
||
); bulb bright yellow, ovate; embolus arising distally from bulb, short, slightly sigmoid; conductor arising proximally from bulb, with two horn-shaped protrusions distally; embolus and conductor widely separated (distance less than diameter of bulb).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B639FFF5FF1CFC150476FBF1" blockId="2.[151,1437,258,1255]" pageId="2" pageNumber="403">
|
||
<emphasis id="B929EAD5B639FFF5FF1CFC1501BFFC05" bold="true" box="[199,429,942,967]" pageId="2" pageNumber="403">
|
||
Female (
|
||
<typeStatus id="54E68865B639FFF5FEF5FC1501B2FC05" box="[302,416,942,967]" pageId="2" pageNumber="403" type="paratype">paratype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Similar to male in color and general features (
|
||
<figureCitation id="13662A42B639FFF5FC65FC15042EFC05" box="[958,1084,942,967]" captionStart="FIGURE 2" captionStartId="14.[151,250,1934,1956]" captionTargetBox="[207,1365,193,1896]" captionTargetId="figure@14.[207,1365,193,1896]" captionTargetPageId="14" captionText="FIGURE 2. Althepus bamensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 2D as in figure 2E." httpUri="https://zenodo.org/record/1439794/files/figure.png" pageId="2" pageNumber="403">Figs 2D–E</figureCitation>
|
||
), but smaller. Internal genitalia with two pairs of spermathecae with irregular margins (
|
||
<figureCitation id="13662A42B639FFF5FCC9FC69037FFC29" box="[786,877,978,1003]" captionStart="FIGURE 2" captionStartId="14.[151,250,1934,1956]" captionTargetBox="[207,1365,193,1896]" captionTargetId="figure@14.[207,1365,193,1896]" captionTargetPageId="14" captionText="FIGURE 2. Althepus bamensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 2D as in figure 2E." httpUri="https://zenodo.org/record/1439794/files/figure.png" pageId="2" pageNumber="403">Fig. 2A</figureCitation>
|
||
). Measurements: total length 3.90; carapace 1.25 length, 1.34 width; abdomen 2.44 length, 1.25 width. Leg measurements: I 40.95 (9.61, 0.56, 9.94, 17.63, 3.21), II 21.67 (5.44, 0.53, 5.77, 8.33, 1.60), III 14.14 (4.08, 0.51, 3.41, 5.00, 1.14), IV missing.
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B639FFF5FF1CFB85040AFBB9" blockId="2.[151,1437,258,1255]" pageId="2" pageNumber="403">
|
||
<emphasis id="B929EAD5B639FFF5FF1CFB85012EFB95" bold="true" box="[199,316,1086,1111]" pageId="2" pageNumber="403">Variation.</emphasis>
|
||
Males: carapace 1.25–1.48 length, 1.30–1.52 width; femur I: 11.86, 12.31 (n = 3; leg I lost in one specimen). Females: carapace 1.10, 1.25 length, 1.02, 1.34 width; femur I: 9.61.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB639FFF5FF1CFB3D01EEFB25" pageId="2" pageNumber="403" type="distribution">
|
||
<paragraph id="8BE236C7B639FFF5FF1CFB3D034EFB5D" blockId="2.[151,1437,258,1255]" box="[199,860,1158,1183]" pageId="2" pageNumber="403">
|
||
<emphasis id="B929EAD5B639FFF5FF1CFB3D0172FB5D" bold="true" box="[199,352,1158,1183]" pageId="2" pageNumber="403">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54E68865B639FFF5FDBBFB3C0282FB5D" box="[608,656,1159,1183]" pageId="2" pageNumber="403">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13662A42B639FFF5FD21FB3D035DFB5D" box="[762,847,1158,1183]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="2" pageNumber="403">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B639FFF5FF1CFB1101EEFB25" blockId="2.[151,1437,258,1255]" pageId="2" pageNumber="403">
|
||
<emphasis id="B929EAD5B639FFF5FF1CFB110199FB01" bold="true" box="[199,395,1194,1219]" pageId="2" pageNumber="403">Natural history.</emphasis>
|
||
Collected on a depression of the cave wall. The cave is located in the bamboo plant community of deciduous forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB639FFF5FF4CFA81015EFAB4" pageId="2" pageNumber="403" type="description">
|
||
<paragraph id="8BE236C7B639FFF5FF4CFA81025EFA96" blockId="2.[151,588,1338,1398]" box="[151,588,1338,1365]" pageId="2" pageNumber="403">
|
||
<taxonomicName id="4C5D4D44B639FFF5FF4CFA8101F9FA97" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[151,491,1338,1365]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="changmao" status="sp. nov.">
|
||
<emphasis id="B929EAD5B639FFF5FF4CFA8101F9FA97" bold="true" box="[151,491,1338,1365]" pageId="2" pageNumber="403">
|
||
<emphasis id="B929EAD5B639FFF5FF4CFA81019BFA96" bold="true" box="[151,393,1338,1364]" italics="true" pageId="2" pageNumber="403">Althepus changmao</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B639FFF5FE68FA8101DAFA96" bold="true" box="[435,456,1338,1364]" italics="true" pageId="2" pageNumber="403">et</emphasis>
|
||
Li
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB639FFF5FE29FA80025EFA96" box="[498,588,1339,1364]" pageId="2" pageNumber="403" rank="species">
|
||
<emphasis id="B929EAD5B639FFF5FE29FA80025EFA96" bold="true" box="[498,588,1339,1364]" pageId="2" pageNumber="403">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B639FFF5FF4CFAE6015EFAB4" blockId="2.[151,588,1338,1398]" box="[151,332,1373,1398]" pageId="2" pageNumber="403">
|
||
<figureCitation id="13662A42B639FFF5FF4CFAE600E8FAB4" box="[151,250,1373,1398]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="15.[151,250,1933,1955]" captionStartId-1="16.[151,250,1938,1960]" captionTargetBox-0="[209,1376,193,1911]" captionTargetBox-1="[200,1368,193,1911]" captionTargetId-0="figure@15.[209,1378,193,1911]" captionTargetId-1="figure@16.[200,1368,193,1911]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURE 3. Althepus changmao Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, retrolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 3A and 3C as in figure 3D." captionText-1="FIGURE 4. Althepus changmao Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 4D as in figure 4E." httpUri-0="https://zenodo.org/record/1439796/files/figure.png" httpUri-1="https://zenodo.org/record/1439798/files/figure.png" pageId="2" pageNumber="403">Figs 3–4</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B639FFF5FEDDFAE50136FAB4" box="[262,292,1374,1398]" captionStart="FIGURE 30" captionStartId="42.[151,250,1917,1939]" captionTargetBox="[177,1410,193,1896]" captionTargetId="figure@42.[177,1410,193,1896]" captionTargetPageId="42" captionText="FIGURE 30. Left cheliceral retromargin of male holotype. A, Althepus bamensis Li et Li sp. nov.; B, A. changmao Li et Li sp. nov.; C, A. dongnaiensis Li et Li sp. nov.; D, A. guan Li et Li sp. nov.; E, A. maechamensis Li et Li sp. nov.; F, A. menglaensis Li et Li sp. nov.; G, A. naphongensis Li et Li sp. nov.; H, A. natmataungensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439850/files/figure.png" pageId="2" pageNumber="403">30</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B639FFF5FEEBFAE5015EFAB4" box="[304,332,1374,1398]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="2" pageNumber="403">32</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB639FFF5FF4CFA1E0150F9C4" pageId="2" pageNumber="403" type="materials_examined">
|
||
<paragraph id="8BE236C7B639FFF5FF4CFA1E0150F9C4" blockId="2.[151,1437,1445,2010]" pageId="2" pageNumber="403">
|
||
<materialsCitation id="3B353C9AB639FFF5FF4CFA1E0302FA20" ID-GBIF-Occurrence="1913888971" collectingDate="2014-06-27" country="Thailand" county="Pha Daeng Cave" elevation="577" latitude="19.503117" location="Chiangmai" longLatPrecision="1" longitude="99.1473" municipality="Tham" pageId="2" pageNumber="403" specimenCount="2" typeStatus="holotype">
|
||
<emphasis id="B929EAD5B639FFF5FF4CFA1E01C0FA7C" bold="true" box="[151,466,1445,1470]" pageId="2" pageNumber="403">
|
||
<typeStatus id="54E68865B639FFF5FF4CFA1E00C0FA7C" box="[151,210,1445,1470]" pageId="2" pageNumber="403">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68865B639FFF5FE80FA1E01DEFA7C" box="[347,460,1445,1470]" pageId="2" pageNumber="403" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male (IZCAS-Ar 33991),
|
||
<collectingCountry id="F34A7657B639FFF5FCC0FA1E0393FA7C" box="[795,897,1445,1470]" name="Thailand" pageId="2" pageNumber="403">Thailand</collectingCountry>
|
||
,
|
||
<location id="8E82601CB639FFF5FC4FFA1E0400FA7C" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB639FFF5FC4FFA1E0400FA7C" box="[916,1042,1445,1470]" country="Thailand" county="Pha Daeng Cave" latitude="19.503117" longLatPrecision="1" longitude="99.1473" municipality="Tham" name="Chiangmai" pageId="2" pageNumber="403">Chiangmai</location>
|
||
,
|
||
<collectingMunicipality id="6B86ACBDB639FFF5FBFEFA1E0474FA7C" box="[1061,1126,1445,1470]" pageId="2" pageNumber="403">Tham</collectingMunicipality>
|
||
(cave)
|
||
<collectingCounty id="62834E4BB639FFF5FB10FA1E0585FA7C" box="[1227,1431,1445,1470]" pageId="2" pageNumber="403">Pha Daeng Cave</collectingCounty>
|
||
,
|
||
<geoCoordinate id="EE695000B639FFF5FF4CFA710137FA20" box="[151,293,1482,1506]" direction="north" orientation="latitude" pageId="2" pageNumber="403" precision="1" value="19.503117">19°30.187'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695000B639FFF5FEEBFA7101A9FA20" box="[304,443,1482,1506]" direction="east" orientation="longitude" pageId="2" pageNumber="403" precision="1" value="99.1473">99°08.838'E</geoCoordinate>
|
||
,
|
||
<elevation id="0070D1F4B639FFF5FE1DFA72021FFA23" box="[454,525,1481,1506]" metricMagnitude="2" metricUnit="m" metricValue="5.77" pageId="2" pageNumber="403" unit="m" value="577.0">
|
||
<quantity id="4CA59B22B639FFF5FE1DFA72021FFA23" box="[454,525,1481,1506]" metricMagnitude="2" metricUnit="m" metricValue="5.77" pageId="2" pageNumber="403" unit="m" value="577.0">577 m</quantity>
|
||
</elevation>
|
||
altitude,
|
||
<collectingDate id="EFA7E9EFB639FFF5FDADFA710302FA20" box="[630,784,1482,1506]" pageId="2" pageNumber="403" value="2014-06-27">
|
||
<date id="FFE31007B639FFF5FDADFA710302FA20" box="[630,784,1482,1506]" pageId="2" pageNumber="403" value="2014-06-27">27 June 2014</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B353C9AB639FFF5FCC1FA720150F9C4" ID-GBIF-Occurrence="1913888978" collectingDate="2014-06-27" country="Thailand" county="Pha Daeng Cave" elevation="577" latitude="19.503117" location="Chiangmai" longLatPrecision="1" longitude="99.1473" municipality="Tham" pageId="2" pageNumber="403" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
|
||
<emphasis id="B929EAD5B639FFF5FCC1FA720389FA20" bold="true" box="[794,923,1481,1506]" pageId="2" pageNumber="403">
|
||
<typeStatus id="54E68865B639FFF5FCC1FA720384FA20" box="[794,918,1481,1506]" pageId="2" pageNumber="403" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D5BFD4EB639FFF5FC79FA710403FA20" box="[930,1041,1481,1506]" pageId="2" pageNumber="403" type="female">2 females</specimenCount>
|
||
, same data as holotype (IZCAS-Ar 33992–33993).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB639FFF5FF1CF9AA0436F98C" pageId="2" pageNumber="403" type="etymology">
|
||
<paragraph id="8BE236C7B639FFF5FF1CF9AA0436F98C" blockId="2.[151,1437,1445,2010]" pageId="2" pageNumber="403">
|
||
<emphasis id="B929EAD5B639FFF5FF1CF9AA015CF9E8" bold="true" box="[199,334,1553,1578]" pageId="2" pageNumber="403">Etymology.</emphasis>
|
||
The specific name is derived from the Chinese pinyin 'cháng máo', which means 'long bristle', referring to the long bristle of the male palpal tibia (
|
||
<figureCitation id="13662A42B639FFF5FD06F98E0325F98C" box="[733,823,1589,1614]" captionStart="FIGURE 3" captionStartId="15.[151,250,1933,1955]" captionTargetBox="[209,1376,193,1911]" captionTargetId="figure@15.[209,1378,193,1911]" captionTargetPageId="15" captionText="FIGURE 3. Althepus changmao Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, retrolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 3A and 3C as in figure 3D." httpUri="https://zenodo.org/record/1439796/files/figure.png" pageId="2" pageNumber="403">Fig. 3A</figureCitation>
|
||
); noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB639FFF5FF1CF9E202C0F8E7" pageId="2" pageNumber="403" type="diagnosis">
|
||
<paragraph id="8BE236C7B639FFF5FF1CF9E202C0F8E7" blockId="2.[151,1437,1445,2010]" pageId="2" pageNumber="403">
|
||
<emphasis id="B929EAD5B639FFF5FF1CF9E20153F9B0" bold="true" box="[199,321,1625,1650]" pageId="2" pageNumber="403">Diagnosis.</emphasis>
|
||
The species resembles
|
||
<taxonomicName id="4C5D4D44B639FFF5FD97F9E103FDF9B0" authority="Deeleman-Reinhold, 1985" authorityName="Deeleman-Reinhold" authorityYear="1985" box="[588,1007,1625,1650]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="tibiatus">
|
||
<emphasis id="B929EAD5B639FFF5FD97F9E102D2F9B3" box="[588,704,1625,1649]" italics="true" pageId="2" pageNumber="403">A. tibiatus</emphasis>
|
||
Deeleman-Reinhold, 1985
|
||
</taxonomicName>
|
||
(Deeleman-Reinhold 1985) in having the upward curling part of the distal conductor and one distinctively long bristle on the male palpal tibia (
|
||
<figureCitation id="13662A42B639FFF5FAECF9C6059CF954" box="[1335,1422,1661,1686]" captionStart="FIGURE 3" captionStartId="15.[151,250,1933,1955]" captionTargetBox="[209,1376,193,1911]" captionTargetId="figure@15.[209,1378,193,1911]" captionTargetPageId="15" captionText="FIGURE 3. Althepus changmao Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, retrolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 3A and 3C as in figure 3D." httpUri="https://zenodo.org/record/1439796/files/figure.png" pageId="2" pageNumber="403">Fig. 3C</figureCitation>
|
||
), and the spermathecae with many bifurcations in females (
|
||
<figureCitation id="13662A42B639FFF5FCF8F91A036CF978" box="[803,894,1697,1722]" captionStart="FIGURE 4" captionStartId="16.[151,250,1938,1960]" captionTargetBox="[200,1368,193,1911]" captionTargetId="figure@16.[200,1368,193,1911]" captionTargetPageId="16" captionText="FIGURE 4. Althepus changmao Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 4D as in figure 4E." httpUri="https://zenodo.org/record/1439798/files/figure.png" pageId="2" pageNumber="403">Fig. 4A</figureCitation>
|
||
). Males of the new species can be distinguished by the jagged distal part of the conductor (
|
||
<figureCitation id="13662A42B639FFF5FD52F97E02F1F91C" box="[649,739,1733,1758]" captionStart="FIGURE 3" captionStartId="15.[151,250,1933,1955]" captionTargetBox="[209,1376,193,1911]" captionTargetId="figure@15.[209,1378,193,1911]" captionTargetPageId="15" captionText="FIGURE 3. Althepus changmao Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, retrolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 3A and 3C as in figure 3D." httpUri="https://zenodo.org/record/1439796/files/figure.png" pageId="2" pageNumber="403">Fig. 3B</figureCitation>
|
||
), compared with the sharp distal part of the conductor in
|
||
<taxonomicName id="4C5D4D44B639FFF5FA5FF97D00F9F8C3" authorityName="Deeleman-Reinhold" authorityYear="1985" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="tibiatus">
|
||
<emphasis id="B929EAD5B639FFF5FA5FF97D00F9F8C3" italics="true" pageId="2" pageNumber="403">A. tibiatus</emphasis>
|
||
</taxonomicName>
|
||
. Females can be distinguished by two branches of the medial spermathecae (
|
||
<figureCitation id="13662A42B639FFF5FB88F95204BCF8C0" box="[1107,1198,1769,1794]" captionStart="FIGURE 4" captionStartId="16.[151,250,1938,1960]" captionTargetBox="[200,1368,193,1911]" captionTargetId="figure@16.[200,1368,193,1911]" captionTargetPageId="16" captionText="FIGURE 4. Althepus changmao Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 4D as in figure 4E." httpUri="https://zenodo.org/record/1439798/files/figure.png" pageId="2" pageNumber="403">Fig. 4A</figureCitation>
|
||
), compared with 3–4 branches of the medial spermathecae in
|
||
<taxonomicName id="4C5D4D44B639FFF5FD83F8B502D9F8E7" authorityName="Deeleman-Reinhold" authorityYear="1985" box="[600,715,1805,1829]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="tibiatus">
|
||
<emphasis id="B929EAD5B639FFF5FD83F8B502D9F8E7" box="[600,715,1805,1829]" italics="true" pageId="2" pageNumber="403">A. tibiatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB639FFF4FF1CF88A044CFDA2" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403" type="description">
|
||
<paragraph id="8BE236C7B639FFF4FF1CF88A0351FE4A" blockId="2.[151,1437,1445,2010]" lastBlockId="3.[151,1436,151,717]" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403">
|
||
<emphasis id="B929EAD5B639FFF5FF1CF88A0237F888" bold="true" box="[199,549,1841,1866]" pageId="2" pageNumber="403">
|
||
Description. Male (
|
||
<typeStatus id="54E68865B639FFF5FE71F88A0205F888" box="[426,535,1841,1866]" pageId="2" pageNumber="403" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 4.42; carapace 1.44 length, 1.62 width; abdomen 2.03 length, 1.10 width. Carapace round (
|
||
<figureCitation id="13662A42B639FFF5FE69F8EE021DF8AC" box="[434,527,1877,1902]" captionStart="FIGURE 4" captionStartId="16.[151,250,1938,1960]" captionTargetBox="[200,1368,193,1911]" captionTargetId="figure@16.[200,1368,193,1911]" captionTargetPageId="16" captionText="FIGURE 4. Althepus changmao Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 4D as in figure 4E." httpUri="https://zenodo.org/record/1439798/files/figure.png" pageId="2" pageNumber="403">Fig. 4C</figureCitation>
|
||
), yellowish, with three longitudinal brown bands of different width. Anterior margin of thoracic region distinctly elevated. Clypeus brown. Cheliceral promargin with two teeth, followed by a lamina, retromargin with two small teeth, posterior surface of fang provided with 21 small denticles (
|
||
<figureCitation id="13662A42B639FFF5FAFDF826059DF874" box="[1318,1423,1949,1974]" captionStart="FIGURE 30" captionStartId="42.[151,250,1917,1939]" captionTargetBox="[177,1410,193,1896]" captionTargetId="figure@42.[177,1410,193,1896]" captionTargetPageId="42" captionText="FIGURE 30. Left cheliceral retromargin of male holotype. A, Althepus bamensis Li et Li sp. nov.; B, A. changmao Li et Li sp. nov.; C, A. dongnaiensis Li et Li sp. nov.; D, A. guan Li et Li sp. nov.; E, A. maechamensis Li et Li sp. nov.; F, A. menglaensis Li et Li sp. nov.; G, A. naphongensis Li et Li sp. nov.; H, A. natmataungensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439850/files/figure.png" pageId="2" pageNumber="403">Fig. 30B</figureCitation>
|
||
). Labium brown. Sternum brown. Abdomen elongate, with complex patterns of white and brown dorsally and ventrally. Legs brown, femur and tibia with white annulations. Leg measurements: I 44.60 (10.83, 0.66, 11.22, 19.23, 2.66), II missing, III 26.58 (7.69, 0.64, 7.37, 9.28, 1.60), IV missing. Male palp (
|
||
<figureCitation id="13662A42B638FFF4FBBDFF0704FAFF16" box="[1126,1256,188,213]" captionStart="FIGURE 3" captionStartId="15.[151,250,1933,1955]" captionTargetBox="[209,1376,193,1911]" captionTargetId="figure@15.[209,1378,193,1911]" captionTargetPageId="15" captionText="FIGURE 3. Althepus changmao Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, retrolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 3A and 3C as in figure 3D." httpUri="https://zenodo.org/record/1439796/files/figure.png" pageId="3" pageNumber="404">Figs 3A–D</figureCitation>
|
||
): tarsus distinct, extends distally, with three slightly curved, serrated bristles at the top of the cymbial protrusion (one of them missing,
|
||
<figureCitation id="13662A42B638FFF4FF27FEBF0146FEDE" box="[252,340,260,285]" captionStart="FIGURE 3" captionStartId="15.[151,250,1933,1955]" captionTargetBox="[209,1376,193,1911]" captionTargetId="figure@15.[209,1378,193,1911]" captionTargetPageId="15" captionText="FIGURE 3. Althepus changmao Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, retrolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 3A and 3C as in figure 3D." httpUri="https://zenodo.org/record/1439796/files/figure.png" pageId="3" pageNumber="404">Fig. 3A</figureCitation>
|
||
), one hooked spine with tip directed proximally (
|
||
<figureCitation id="13662A42B638FFF4FCA3FEBF03C3FEDE" box="[888,977,260,285]" captionStart="FIGURE 3" captionStartId="15.[151,250,1933,1955]" captionTargetBox="[209,1376,193,1911]" captionTargetId="figure@15.[209,1378,193,1911]" captionTargetPageId="15" captionText="FIGURE 3. Althepus changmao Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, retrolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 3A and 3C as in figure 3D." httpUri="https://zenodo.org/record/1439796/files/figure.png" pageId="3" pageNumber="404">Fig. 3A</figureCitation>
|
||
); tibia with one distinctively long bristle; bulb yellow, ovate; embolus arising prolatero-distally from bulb, slender and proximal part curved; conductor arising proximally from bulb, curved, with the distal part curling upward forming a groove; embolus and conductor widely separated (distance almost equal to diameter of bulb).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B638FFF4FF1CFE2F0173FDFF" blockId="3.[151,1436,151,717]" pageId="3" pageNumber="404">
|
||
<emphasis id="B929EAD5B638FFF4FF1CFE2F01BFFE6F" bold="true" box="[199,429,404,429]" pageId="3" pageNumber="404">
|
||
Female (
|
||
<typeStatus id="54E68865B638FFF4FEF5FE2F01B2FE6F" box="[302,416,404,429]" pageId="3" pageNumber="404" type="paratype">paratype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Similar to male in color and general features (
|
||
<figureCitation id="13662A42B638FFF4FC65FE2F042EFE6E" box="[958,1084,404,429]" captionStart="FIGURE 4" captionStartId="16.[151,250,1938,1960]" captionTargetBox="[200,1368,193,1911]" captionTargetId="figure@16.[200,1368,193,1911]" captionTargetPageId="16" captionText="FIGURE 4. Althepus changmao Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 4D as in figure 4E." httpUri="https://zenodo.org/record/1439798/files/figure.png" pageId="3" pageNumber="404">Figs 4D–E</figureCitation>
|
||
), but smaller. Internal genitalia with two pairs of spermathecae, curved, bifurcate, with a swollen distal part; the spermachecae left and right showing different number of branches (
|
||
<figureCitation id="13662A42B638FFF4FDB3FE6702D5FE36" box="[616,711,476,501]" captionStart="FIGURE 4" captionStartId="16.[151,250,1938,1960]" captionTargetBox="[200,1368,193,1911]" captionTargetId="figure@16.[200,1368,193,1911]" captionTargetPageId="16" captionText="FIGURE 4. Althepus changmao Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 4D as in figure 4E." httpUri="https://zenodo.org/record/1439798/files/figure.png" pageId="3" pageNumber="404">Fig. 4A</figureCitation>
|
||
). Measurements: total length 3.36; carapace 1.05 length, 1.20 width; abdomen 2.13 length, 1.34 width. Leg measurements: I missing, II missing, III 11.52 (3.24, 0.49, 2.88, 3.80, 1.11), IV missing.
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B638FFF4FF1CFDFC044CFDA2" blockId="3.[151,1436,151,717]" box="[199,1118,583,608]" pageId="3" pageNumber="404">
|
||
<emphasis id="B929EAD5B638FFF4FF1CFDFC012EFDA2" bold="true" box="[199,316,583,608]" pageId="3" pageNumber="404">Variation.</emphasis>
|
||
Females: carapace 1.05, 1.15 length, 1.19, 1.20 wide; leg I lost (n = 2).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB638FFF4FF1CFDD703E6FD6A" pageId="3" pageNumber="404" type="distribution">
|
||
<paragraph id="8BE236C7B638FFF4FF1CFDD7034EFD46" blockId="3.[151,1436,151,717]" box="[199,860,620,645]" pageId="3" pageNumber="404">
|
||
<emphasis id="B929EAD5B638FFF4FF1CFDD70172FD47" bold="true" box="[199,352,620,645]" pageId="3" pageNumber="404">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54E68865B638FFF4FDBBFDD60282FD47" box="[608,656,621,645]" pageId="3" pageNumber="404">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13662A42B638FFF4FD21FDD7035DFD46" box="[762,847,620,645]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="3" pageNumber="404">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B638FFF4FF1CFD3403E6FD6A" blockId="3.[151,1436,151,717]" box="[199,1012,655,680]" pageId="3" pageNumber="404">
|
||
<emphasis id="B929EAD5B638FFF4FF1CFD340197FD6A" bold="true" box="[199,389,655,680]" pageId="3" pageNumber="404">Natural history.</emphasis>
|
||
Collected on the cave wall (high humidity in the cave).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB638FFF4FF1CFD0F015EFC99" pageId="3" pageNumber="404" type="discussion">
|
||
<paragraph id="8BE236C7B638FFF4FF1CFD0F0522FD0E" blockId="3.[151,1436,151,717]" box="[199,1328,692,717]" pageId="3" pageNumber="404">
|
||
<emphasis id="B929EAD5B638FFF4FF1CFD0F0122FD0F" bold="true" box="[199,304,692,717]" pageId="3" pageNumber="404">Remark.</emphasis>
|
||
<taxonomicName id="4C5D4D44B638FFF4FEECFD0F0230FD0E" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[311,546,692,716]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="changmao" status="sp. nov.">
|
||
<emphasis id="B929EAD5B638FFF4FEECFD0F01D9FD0E" box="[311,459,692,716]" italics="true" pageId="3" pageNumber="404">A. changmao</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B638FFF4FE2BFD0E0216FD0E" box="[496,516,693,716]" italics="true" pageId="3" pageNumber="404">et</emphasis>
|
||
Li
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB638FFF4FDF3FD0F0290FD0F" box="[552,642,692,717]" pageId="3" pageNumber="404" rank="species">
|
||
<emphasis id="B929EAD5B638FFF4FDF3FD0F0290FD0F" bold="true" box="[552,642,692,717]" pageId="3" pageNumber="404">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
was labeled as “sp. 101 ” in the analysis of Li and Li (2018).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B638FFF4FF4CFCA4027FFCFB" blockId="3.[151,621,799,859]" box="[151,621,799,826]" pageId="3" pageNumber="404">
|
||
<taxonomicName id="4C5D4D44B638FFF4FF4CFCA4021EFCF8" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[151,524,799,826]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="dongnaiensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B638FFF4FF4CFCA4021EFCF8" bold="true" box="[151,524,799,826]" pageId="3" pageNumber="404">
|
||
<emphasis id="B929EAD5B638FFF4FF4CFCA401BBFCFB" bold="true" box="[151,425,799,825]" italics="true" pageId="3" pageNumber="404">Althepus dongnaiensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B638FFF4FE0FFCA401FBFCFB" bold="true" box="[468,489,799,825]" italics="true" pageId="3" pageNumber="404">et</emphasis>
|
||
Li
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB638FFF4FDC8FC9B027FFCFB" box="[531,621,800,825]" pageId="3" pageNumber="404" rank="species">
|
||
<emphasis id="B929EAD5B638FFF4FDC8FC9B027FFCFB" bold="true" box="[531,621,800,825]" pageId="3" pageNumber="404">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B638FFF4FF4CFCF9015EFC99" blockId="3.[151,621,799,859]" box="[151,332,834,859]" pageId="3" pageNumber="404">
|
||
<figureCitation id="13662A42B638FFF4FF4CFCF900E8FC99" box="[151,250,834,859]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="17.[151,250,1933,1955]" captionStartId-1="18.[151,250,1933,1955]" captionTargetBox-0="[209,1367,202,1911]" captionTargetBox-1="[209,1378,193,1911]" captionTargetId-0="figure@17.[209,1378,193,1911]" captionTargetId-1="figure@18.[209,1378,193,1911]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="FIGURE 5. Althepus dongnaiensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 5A and 5C as in figure 5D." captionText-1="FIGURE 6. Althepus dongnaiensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 6D as in figure 6E." httpUri-0="https://zenodo.org/record/1439800/files/figure.png" httpUri-1="https://zenodo.org/record/1439802/files/figure.png" pageId="3" pageNumber="404">Figs 5–6</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B638FFF4FEDDFCF80136FC99" box="[262,292,835,859]" captionStart="FIGURE 30" captionStartId="42.[151,250,1917,1939]" captionTargetBox="[177,1410,193,1896]" captionTargetId="figure@42.[177,1410,193,1896]" captionTargetPageId="42" captionText="FIGURE 30. Left cheliceral retromargin of male holotype. A, Althepus bamensis Li et Li sp. nov.; B, A. changmao Li et Li sp. nov.; C, A. dongnaiensis Li et Li sp. nov.; D, A. guan Li et Li sp. nov.; E, A. maechamensis Li et Li sp. nov.; F, A. menglaensis Li et Li sp. nov.; G, A. naphongensis Li et Li sp. nov.; H, A. natmataungensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439850/files/figure.png" pageId="3" pageNumber="404">30</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B638FFF4FEEBFCF8015EFC99" box="[304,332,835,859]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="3" pageNumber="404">32</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB638FFF4FF4CFC310164FC29" pageId="3" pageNumber="404" type="materials_examined">
|
||
<paragraph id="8BE236C7B638FFF4FF4CFC310164FC29" blockId="3.[151,1437,906,1867]" pageId="3" pageNumber="404">
|
||
<materialsCitation id="3B353C9AB638FFF4FF4CFC3103A0FC05" ID-GBIF-Occurrence="1913888970" collectingDate="2015-09-05" country="Vietnam" elevation="173" latitude="11.450084" location="Natural Forest" longLatPrecision="1" longitude="107.36438" municipality="Cat Tien National Park" pageId="3" pageNumber="404" specimenCount="2" stateProvince="Dong Nai Province" typeStatus="holotype">
|
||
<emphasis id="B929EAD5B638FFF4FF4CFC3101D3FC61" bold="true" box="[151,449,906,931]" pageId="3" pageNumber="404">
|
||
<typeStatus id="54E68865B638FFF4FF4CFC3100C0FC61" box="[151,210,906,931]" pageId="3" pageNumber="404">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68865B638FFF4FE90FC3101AEFC61" box="[331,444,906,931]" pageId="3" pageNumber="404" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male (IZCAS-Ar 33994),
|
||
<collectingCountry id="F34A7657B638FFF4FD32FC310359FC61" box="[745,843,906,931]" name="Vietnam" pageId="3" pageNumber="404">Vietnam</collectingCountry>
|
||
,
|
||
<collectingRegion id="4999F825B638FFF4FC8EFC30043DFC61" box="[853,1071,906,931]" country="Vietnam" name="Dong Nai" pageId="3" pageNumber="404">
|
||
Dong
|
||
<collectingRegion id="4999F825B638FFF4FC41FC31043DFC61" box="[922,1071,906,931]" inferredBySuffix="true" pageId="3" pageNumber="404">Nai Province</collectingRegion>
|
||
</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B86ACBDB638FFF4FBE1FC30052FFC61" box="[1082,1341,906,931]" pageId="3" pageNumber="404">Cat Tien National Park</collectingMunicipality>
|
||
,
|
||
<location id="8E82601CB638FFF4FA93FC3100F2FC04" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB638FFF4FA93FC3100F2FC04" country="Vietnam" latitude="11.450084" longLatPrecision="1" longitude="107.36438" municipality="Cat Tien National Park" name="Natural Forest" pageId="3" pageNumber="404" stateProvince="Dong Nai Province">Natural Forest</location>
|
||
,
|
||
<geoCoordinate id="EE695000B638FFF4FF30FC150165FC05" box="[235,375,942,967]" direction="north" orientation="latitude" pageId="3" pageNumber="404" precision="1" value="11.450084">11°27.005'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695000B638FFF4FE59FC150208FC05" box="[386,538,942,967]" direction="east" orientation="longitude" pageId="3" pageNumber="404" precision="1" value="107.36438">107°21.863'E</geoCoordinate>
|
||
,
|
||
<elevation id="0070D1F4B638FFF4FDFDFC14027EFC04" box="[550,620,943,967]" metricMagnitude="2" metricUnit="m" metricValue="1.73" pageId="3" pageNumber="404" unit="m" value="173.0">
|
||
<quantity id="4CA59B22B638FFF4FDFDFC14027EFC04" box="[550,620,943,967]" metricMagnitude="2" metricUnit="m" metricValue="1.73" pageId="3" pageNumber="404" unit="m" value="173.0">173 m</quantity>
|
||
</elevation>
|
||
altitude, 0
|
||
<collectingDate id="EFA7E9EFB638FFF4FD38FC1503A0FC05" box="[739,946,942,967]" pageId="3" pageNumber="404" value="2015-09-05">
|
||
<date id="FFE31007B638FFF4FD38FC1503A0FC05" box="[739,946,942,967]" pageId="3" pageNumber="404" value="2015-09-05">5 September 2015</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B353C9AB638FFF4FC60FC150164FC29" ID-GBIF-Occurrence="1913888977" collectingDate="2015-09-05" country="Vietnam" elevation="173" latitude="11.450084" location="Natural Forest" longLatPrecision="1" longitude="107.36438" municipality="Cat Tien National Park" pageId="3" pageNumber="404" specimenCount="1" specimenCount-female="1" stateProvince="Dong Nai Province" typeStatus="paratype">
|
||
<emphasis id="B929EAD5B638FFF4FC60FC150423FC05" bold="true" box="[955,1073,942,967]" pageId="3" pageNumber="404">
|
||
<typeStatus id="54E68865B638FFF4FC60FC15043EFC05" box="[955,1068,942,967]" pageId="3" pageNumber="404" type="paratype">Paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D5BFD4EB638FFF4FBE3FC140489FC05" box="[1080,1179,942,967]" pageId="3" pageNumber="404" type="female">1 female</specimenCount>
|
||
, same data as holotype (IZCAS-Ar 33995).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB638FFF4FF1CFC4D040FFBCD" box="[199,1053,1014,1039]" pageId="3" pageNumber="404" type="etymology">
|
||
<paragraph id="8BE236C7B638FFF4FF1CFC4D040FFBCD" blockId="3.[151,1437,906,1867]" box="[199,1053,1014,1039]" pageId="3" pageNumber="404">
|
||
<emphasis id="B929EAD5B638FFF4FF1CFC4D015CFBCD" bold="true" box="[199,334,1014,1039]" pageId="3" pageNumber="404">Etymology.</emphasis>
|
||
The specific name is an adjective and refers to the
|
||
<typeStatus id="54E68865B638FFF4FC57FC4C03AEFBCD" box="[908,956,1015,1039]" pageId="3" pageNumber="404">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB638FFF4FF1CFBA10266FBB9" pageId="3" pageNumber="404" type="diagnosis">
|
||
<paragraph id="8BE236C7B638FFF4FF1CFBA10266FBB9" blockId="3.[151,1437,906,1867]" pageId="3" pageNumber="404">
|
||
<emphasis id="B929EAD5B638FFF4FF1CFBA10153FBF1" bold="true" box="[199,321,1050,1075]" pageId="3" pageNumber="404">Diagnosis.</emphasis>
|
||
This new species can be easily distinguished from all the other species by the wide, curved conductor and the arc-shaped margin of the distal conductor in males (
|
||
<figureCitation id="13662A42B638FFF4FC74FB85043CFB95" box="[943,1070,1086,1111]" captionStart="FIGURE 5" captionStartId="17.[151,250,1933,1955]" captionTargetBox="[209,1367,202,1911]" captionTargetId="figure@17.[209,1378,193,1911]" captionTargetPageId="17" captionText="FIGURE 5. Althepus dongnaiensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 5A and 5C as in figure 5D." httpUri="https://zenodo.org/record/1439800/files/figure.png" pageId="3" pageNumber="404">Figs 5A–D</figureCitation>
|
||
) and by one pair of distinctively curved spermathecae in females (
|
||
<figureCitation id="13662A42B638FFF4FDD6FBD90275FBB9" box="[525,615,1122,1147]" captionStart="FIGURE 6" captionStartId="18.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@18.[209,1378,193,1911]" captionTargetPageId="18" captionText="FIGURE 6. Althepus dongnaiensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 6D as in figure 6E." httpUri="https://zenodo.org/record/1439802/files/figure.png" pageId="3" pageNumber="404">Fig. 6A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB638FFF4FF1CFB3D031DF91D" pageId="3" pageNumber="404" type="description">
|
||
<paragraph id="8BE236C7B638FFF4FF1CFB3D0417F9E9" blockId="3.[151,1437,906,1867]" pageId="3" pageNumber="404">
|
||
<emphasis id="B929EAD5B638FFF4FF1CFB3D0237FB5D" bold="true" box="[199,549,1158,1183]" pageId="3" pageNumber="404">
|
||
Description. Male (
|
||
<typeStatus id="54E68865B638FFF4FE71FB3D0205FB5D" box="[426,535,1158,1183]" pageId="3" pageNumber="404" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 3.68; carapace 1.16 length, 1.13 width; abdomen 2.15 length, 0.90 width. Carapace round (
|
||
<figureCitation id="13662A42B638FFF4FE70FB110217FB01" box="[427,517,1194,1219]" captionStart="FIGURE 6" captionStartId="18.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@18.[209,1378,193,1911]" captionTargetPageId="18" captionText="FIGURE 6. Althepus dongnaiensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 6D as in figure 6E." httpUri="https://zenodo.org/record/1439802/files/figure.png" pageId="3" pageNumber="404">Fig. 6C</figureCitation>
|
||
), yellowish, with three longitudinal brown bands of different widths, the middle one cleaved and shortened to form a triangular band around eyes. Anterior margin of thoracic region distinctly elevated. Clypeus brown. Cheliceral promargin with two teeth, followed by a lamina, retromargin with two small teeth, posterior surface of fang with 20 small denticles (
|
||
<figureCitation id="13662A42B638FFF4FCF4FAAD0388FAED" box="[815,922,1302,1327]" captionStart="FIGURE 30" captionStartId="42.[151,250,1917,1939]" captionTargetBox="[177,1410,193,1896]" captionTargetId="figure@42.[177,1410,193,1896]" captionTargetPageId="42" captionText="FIGURE 30. Left cheliceral retromargin of male holotype. A, Althepus bamensis Li et Li sp. nov.; B, A. changmao Li et Li sp. nov.; C, A. dongnaiensis Li et Li sp. nov.; D, A. guan Li et Li sp. nov.; E, A. maechamensis Li et Li sp. nov.; F, A. menglaensis Li et Li sp. nov.; G, A. naphongensis Li et Li sp. nov.; H, A. natmataungensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439850/files/figure.png" pageId="3" pageNumber="404">Fig. 30C</figureCitation>
|
||
). Labium brown. Sternum yellow, with two irregular brown lines each side. Abdomen elongate, with complex patterns of white and dark-brown dorsally and ventrally. Legs brown. Leg measurements: I 33.00 (8.00, 0.40, 8.00, 14.15, 2.45), II 21.45 (5.45, 0.40, 5.20, 8.70, 1.70), III 16.68 (5.00, 0.40, 4.49, 5.51, 1.28), IV 23.16 (6.86, 0.40, 6.41, 7.82, 1.67). Male palp (
|
||
<figureCitation id="13662A42B638FFF4FB1DFA39055AFA59" box="[1222,1352,1410,1435]" captionStart="FIGURE 5" captionStartId="17.[151,250,1933,1955]" captionTargetBox="[209,1367,202,1911]" captionTargetId="figure@17.[209,1378,193,1911]" captionTargetPageId="17" captionText="FIGURE 5. Althepus dongnaiensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 5A and 5C as in figure 5D." httpUri="https://zenodo.org/record/1439800/files/figure.png" pageId="3" pageNumber="404">Figs 5A–D</figureCitation>
|
||
): tarsus distinct, extends distally, with three slightly curved, serrated bristles at the top of cymbial protrusion (
|
||
<figureCitation id="13662A42B638FFF4FADDFA1D054DFA7D" box="[1286,1375,1446,1471]" captionStart="FIGURE 5" captionStartId="17.[151,250,1933,1955]" captionTargetBox="[209,1367,202,1911]" captionTargetId="figure@17.[209,1378,193,1911]" captionTargetPageId="17" captionText="FIGURE 5. Althepus dongnaiensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 5A and 5C as in figure 5D." httpUri="https://zenodo.org/record/1439800/files/figure.png" pageId="3" pageNumber="404">Fig. 5A</figureCitation>
|
||
), one hooked spine with tip directed proximally (
|
||
<figureCitation id="13662A42B638FFF4FDA1FA7102C6FA21" box="[634,724,1482,1507]" captionStart="FIGURE 5" captionStartId="17.[151,250,1933,1955]" captionTargetBox="[209,1367,202,1911]" captionTargetId="figure@17.[209,1378,193,1911]" captionTargetPageId="17" captionText="FIGURE 5. Althepus dongnaiensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 5A and 5C as in figure 5D." httpUri="https://zenodo.org/record/1439800/files/figure.png" pageId="3" pageNumber="404">Fig. 5A</figureCitation>
|
||
); bulb yellow, pyriform; embolus arising proximally from bulb, slender and curved (
|
||
<figureCitation id="13662A42B638FFF4FE5EFA55021BF9C5" box="[389,521,1518,1543]" captionStart="FIGURE 5" captionStartId="17.[151,250,1933,1955]" captionTargetBox="[209,1367,202,1911]" captionTargetId="figure@17.[209,1378,193,1911]" captionTargetPageId="17" captionText="FIGURE 5. Althepus dongnaiensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 5A and 5C as in figure 5D." httpUri="https://zenodo.org/record/1439800/files/figure.png" pageId="3" pageNumber="404">Figs 5A–B</figureCitation>
|
||
); conductor arising distally from bulb, curved, distal part with a large lamina; embolus and conductor widely separated (distance less than diameter of bulb).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B638FFF4FF1CF98D031DF91D" blockId="3.[151,1437,906,1867]" pageId="3" pageNumber="404">
|
||
<emphasis id="B929EAD5B638FFF4FF1CF98D01BCF98D" bold="true" box="[199,430,1590,1615]" pageId="3" pageNumber="404">
|
||
Female (
|
||
<typeStatus id="54E68865B638FFF4FEF4F98D01B3F98D" box="[303,417,1590,1615]" pageId="3" pageNumber="404" type="paratype">paratype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Similar to male in color and general features (
|
||
<figureCitation id="13662A42B638FFF4FC12F98D0458F98D" box="[969,1098,1590,1615]" captionStart="FIGURE 6" captionStartId="18.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@18.[209,1378,193,1911]" captionTargetPageId="18" captionText="FIGURE 6. Althepus dongnaiensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 6D as in figure 6E." httpUri="https://zenodo.org/record/1439802/files/figure.png" pageId="3" pageNumber="404">Figs 6D–E</figureCitation>
|
||
), but larger. Internal genitalia with one pair of spermathecae, distinctively curved at almost a right-angle, distal part swollen (
|
||
<figureCitation id="13662A42B638FFF4FAF6F9E1059DF9B1" box="[1325,1423,1626,1651]" captionStart="FIGURE 6" captionStartId="18.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@18.[209,1378,193,1911]" captionTargetPageId="18" captionText="FIGURE 6. Althepus dongnaiensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 6D as in figure 6E." httpUri="https://zenodo.org/record/1439802/files/figure.png" pageId="3" pageNumber="404">Fig. 6A</figureCitation>
|
||
). Measurements: total length 3.84; carapace 1.19 length, 1.14 width; abdomen 2.33 length, 1.10 width. Leg measurements: I 16.40 (4.35, 0.40, 4.10, 6.05, 1.50), II 16.00 (4.25, 0.40, 4.00, 6.00, 1.35), III 10.75 (3.10, 0.40, 2.75, 3.50, 1.00), IV 19.57 (5.71, 0.40, 5.38, 6.41, 1.67).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB638FFF4FF1CF95103EEF8E4" pageId="3" pageNumber="404" type="distribution">
|
||
<paragraph id="8BE236C7B638FFF4FF1CF951034EF8C1" blockId="3.[151,1437,906,1867]" box="[199,860,1770,1795]" pageId="3" pageNumber="404">
|
||
<emphasis id="B929EAD5B638FFF4FF1CF9510172F8C1" bold="true" box="[199,352,1770,1795]" pageId="3" pageNumber="404">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54E68865B638FFF4FDBBF9500282F8C1" box="[608,656,1771,1795]" pageId="3" pageNumber="404">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13662A42B638FFF4FD21F951035DF8C1" box="[762,847,1770,1795]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="3" pageNumber="404">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B638FFF4FF1CF8B503EEF8E4" blockId="3.[151,1437,906,1867]" box="[199,1020,1806,1831]" pageId="3" pageNumber="404">
|
||
<emphasis id="B929EAD5B638FFF4FF1CF8B50197F8E5" bold="true" box="[199,389,1806,1831]" pageId="3" pageNumber="404">Natural history.</emphasis>
|
||
Collected among tree buttresses at an altitude of
|
||
<specimenCount id="9D5BFD4EB638FFF4FC75F8B403EEF8E4" box="[942,1020,1807,1831]" pageId="3" pageNumber="404" type="generic">
|
||
<quantity id="4CA59B22B638FFF4FC75F8B403EAF8E4" box="[942,1016,1807,1831]" metricMagnitude="2" metricUnit="m" metricValue="1.73" pageId="3" pageNumber="404" unit="m" value="173.0">173 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB638FFF3FF1CF889015EFF11" lastPageId="4" lastPageNumber="405" pageId="3" pageNumber="404" type="discussion">
|
||
<paragraph id="8BE236C7B638FFF4FF1CF889055BF889" blockId="3.[151,1437,906,1867]" box="[199,1353,1842,1867]" pageId="3" pageNumber="404">
|
||
<emphasis id="B929EAD5B638FFF4FF1CF8890122F889" bold="true" box="[199,304,1842,1867]" pageId="3" pageNumber="404">Remark.</emphasis>
|
||
<taxonomicName id="4C5D4D44B638FFF4FEECF8880253F888" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[311,577,1842,1867]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="dongnaiensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B638FFF4FEECF88801F8F889" box="[311,490,1843,1867]" italics="true" pageId="3" pageNumber="404">A. dongnaiensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B638FFF4FDD4F8880230F888" box="[527,546,1843,1866]" italics="true" pageId="3" pageNumber="404">et</emphasis>
|
||
Li
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB638FFF4FD9CF88902B3F889" box="[583,673,1842,1867]" pageId="3" pageNumber="404" rank="species">
|
||
<emphasis id="B929EAD5B638FFF4FD9CF88902B3F889" bold="true" box="[583,673,1842,1867]" pageId="3" pageNumber="404">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
was labeled as “sp. 282” in the analysis of Li and Li (2018).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63FFFF3FF4CFF2C0219FF73" blockId="4.[151,523,151,211]" box="[151,523,151,178]" pageId="4" pageNumber="405">
|
||
<taxonomicName id="4C5D4D44B63FFFF3FF4CFF2C01B8FF70" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[151,426,151,178]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="guan" status="sp. nov.">
|
||
<emphasis id="B929EAD5B63FFFF3FF4CFF2C01B8FF70" bold="true" box="[151,426,151,178]" pageId="4" pageNumber="405">
|
||
<emphasis id="B929EAD5B63FFFF3FF4CFF2C015AFF73" bold="true" box="[151,328,151,177]" italics="true" pageId="4" pageNumber="405">Althepus guan</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B63FFFF3FEA9FF2C0195FF73" bold="true" box="[370,391,151,177]" italics="true" pageId="4" pageNumber="405">et</emphasis>
|
||
Li
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB63FFFF3FE6AFF230219FF73" box="[433,523,152,177]" pageId="4" pageNumber="405" rank="species">
|
||
<emphasis id="B929EAD5B63FFFF3FE6AFF230219FF73" bold="true" box="[433,523,152,177]" pageId="4" pageNumber="405">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63FFFF3FF4CFF01015EFF11" blockId="4.[151,523,151,211]" box="[151,332,186,211]" pageId="4" pageNumber="405">
|
||
<figureCitation id="13662A42B63FFFF3FF4CFF0100E8FF11" box="[151,250,186,211]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="19.[151,250,1917,1939]" captionStartId-1="20.[151,250,1933,1955]" captionTargetBox-0="[230,1372,193,1896]" captionTargetBox-1="[209,1378,193,1911]" captionTargetId-0="figure@19.[214,1372,193,1896]" captionTargetId-1="figure@20.[209,1378,193,1911]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURE 7. Althepus guan Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hookshaped spine; SB = serrated bristles. The scale bar in figures 7A and 7C as in figure 7D." captionText-1="FIGURE 8. Althepus guan Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 8D as in figure 8E." httpUri-0="https://zenodo.org/record/1439804/files/figure.png" httpUri-1="https://zenodo.org/record/1439806/files/figure.png" pageId="4" pageNumber="405">Figs 7–8</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B63FFFF3FEDDFF000136FF11" box="[262,292,187,211]" captionStart="FIGURE 30" captionStartId="42.[151,250,1917,1939]" captionTargetBox="[177,1410,193,1896]" captionTargetId="figure@42.[177,1410,193,1896]" captionTargetPageId="42" captionText="FIGURE 30. Left cheliceral retromargin of male holotype. A, Althepus bamensis Li et Li sp. nov.; B, A. changmao Li et Li sp. nov.; C, A. dongnaiensis Li et Li sp. nov.; D, A. guan Li et Li sp. nov.; E, A. maechamensis Li et Li sp. nov.; F, A. menglaensis Li et Li sp. nov.; G, A. naphongensis Li et Li sp. nov.; H, A. natmataungensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439850/files/figure.png" pageId="4" pageNumber="405">30</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B63FFFF3FEEBFF00015EFF11" box="[304,332,187,211]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="4" pageNumber="405">32</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63FFFF3FF4CFEB903B3FEA1" pageId="4" pageNumber="405" type="materials_examined">
|
||
<paragraph id="8BE236C7B63FFFF3FF4CFEB903B3FEA1" blockId="4.[151,1436,258,1219]" pageId="4" pageNumber="405">
|
||
<materialsCitation id="3B353C9AB63FFFF3FF4CFEB904E8FEFD" ID-GBIF-Occurrence="1913888991" collectingDate="2014-05-17" country="Indonesia" county="Payakumbuh" elevation="949" latitude="-0.06373333" location="Imam Bonjol Cave" longLatPrecision="1" longitude="100.34492" municipality="Koto Tinggi Village" pageId="4" pageNumber="405" specimenCount="2" stateProvince="Sumatra" typeStatus="holotype">
|
||
<emphasis id="B929EAD5B63FFFF3FF4CFEB901D5FED9" bold="true" box="[151,455,258,283]" pageId="4" pageNumber="405">
|
||
<typeStatus id="54E68865B63FFFF3FF4CFEB900C0FED9" box="[151,210,258,283]" pageId="4" pageNumber="405">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68865B63FFFF3FE8AFEB901D0FED9" box="[337,450,258,283]" pageId="4" pageNumber="405" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male (IZCAS-Ar 33996),
|
||
<collectingCountry id="F34A7657B63FFFF3FD20FEB90379FED9" box="[763,875,258,283]" name="Indonesia" pageId="4" pageNumber="405">Indonesia</collectingCountry>
|
||
,
|
||
<collectingRegion id="4999F825B63FFFF3FCA2FEB803CBFED9" box="[889,985,259,283]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="405">Sumatra</collectingRegion>
|
||
,
|
||
<collectingRegion id="4999F825B63FFFF3FC3CFEB804E6FED9" box="[999,1268,258,283]" country="Indonesia" name="Sumatera Barat" pageId="4" pageNumber="405">
|
||
West
|
||
<collectingRegion id="4999F825B63FFFF3FBF3FEB804E6FED9" box="[1064,1268,258,283]" inferredBySuffix="true" pageId="4" pageNumber="405">Sumatra Province</collectingRegion>
|
||
</collectingRegion>
|
||
,
|
||
<collectingCounty id="62834E4BB63FFFF3FAD8FEB9058AFED9" box="[1283,1432,258,283]" pageId="4" pageNumber="405">Payakumbuh</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6B86ACBDB63FFFF3FF4CFE9C016EFEFD" box="[151,380,294,319]" pageId="4" pageNumber="405">Koto Tinggi Village</collectingMunicipality>
|
||
,
|
||
<location id="8E82601CB63FFFF3FE53FE9C0273FEFD" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB63FFFF3FE53FE9C0273FEFD" box="[392,609,294,319]" country="Indonesia" county="Payakumbuh" latitude="-0.06373333" longLatPrecision="1" longitude="100.34492" municipality="Koto Tinggi Village" name="Imam Bonjol Cave" pageId="4" pageNumber="405" stateProvince="Sumatra">Imam Bonjol Cave</location>
|
||
,
|
||
<geoCoordinate id="EE695000B63FFFF3FDB6FE9C02EBFEFD" box="[621,761,295,319]" direction="south" orientation="latitude" pageId="4" pageNumber="405" precision="1" value="-0.06373333">00°03.824'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695000B63FFFF3FCDEFE9D03B3FEFD" box="[773,929,294,319]" direction="east" orientation="longitude" pageId="4" pageNumber="405" precision="1" value="100.34492">100°20.695′E</geoCoordinate>
|
||
,
|
||
<elevation id="0070D1F4B63FFFF3FC76FE9C03E7FEFC" box="[941,1013,295,319]" metricMagnitude="2" metricUnit="m" metricValue="9.49" pageId="4" pageNumber="405" unit="m" value="949.0">
|
||
<quantity id="4CA59B22B63FFFF3FC76FE9C03E7FEFC" box="[941,1013,295,319]" metricMagnitude="2" metricUnit="m" metricValue="9.49" pageId="4" pageNumber="405" unit="m" value="949.0">949 m</quantity>
|
||
</elevation>
|
||
altitude,
|
||
<collectingDate id="EFA7E9EFB63FFFF3FBBBFE9C04E8FEFD" box="[1120,1274,295,319]" pageId="4" pageNumber="405" value="2014-05-17">
|
||
<date id="FFE31007B63FFFF3FBBBFE9C04E8FEFD" box="[1120,1274,295,319]" pageId="4" pageNumber="405" value="2014-05-17">17 May 2014</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B353C9AB63FFFF3FADEFE9D03B3FEA1" ID-GBIF-Occurrence="1913888992" collectingDate="2014-05-17" collectionCode="IZCAS" country="Indonesia" county="Payakumbuh" elevation="949" latitude="-0.06373333" location="Ar" longLatPrecision="1" longitude="100.34492" municipality="Koto Tinggi Village" pageId="4" pageNumber="405" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sumatra" typeStatus="paratype">
|
||
<emphasis id="B929EAD5B63FFFF3FADEFE9D0597FEFD" bold="true" box="[1285,1413,294,319]" pageId="4" pageNumber="405">
|
||
<typeStatus id="54E68865B63FFFF3FADEFE9D0593FEFD" box="[1285,1409,294,319]" pageId="4" pageNumber="405" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D5BFD4EB63FFFF3FA55FE9C00DFFEA1" pageId="4" pageNumber="405" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="9D5BFD4EB63FFFF3FEDFFEF0017AFEA1" box="[260,360,330,355]" pageId="4" pageNumber="405" type="female">1 female</specimenCount>
|
||
, same data as holotype (
|
||
<collectionCode id="ED4CAE02B63FFFF3FDA3FEF002D9FEA1" box="[632,715,331,355]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="4" pageNumber="405">IZCAS</collectionCode>
|
||
<location id="8E82601CB63FFFF3FD09FEF002FDFEA1" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB63FFFF3FD09FEF002FDFEA1" box="[722,751,331,355]" name="Ar" pageId="4" pageNumber="405">Ar</location>
|
||
33997–33998).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63FFFF3FF1CFED502A5FE69" pageId="4" pageNumber="405" type="etymology">
|
||
<paragraph id="8BE236C7B63FFFF3FF1CFED502A5FE69" blockId="4.[151,1436,258,1219]" pageId="4" pageNumber="405">
|
||
<emphasis id="B929EAD5B63FFFF3FF1CFED5015CFE45" bold="true" box="[199,334,366,391]" pageId="4" pageNumber="405">Etymology.</emphasis>
|
||
The specific name is derived from the Chinese pinyin 'guǎn', which means 'tube', referring to the tube-like embolus (
|
||
<figureCitation id="13662A42B63FFFF3FEB4FE2901D8FE69" box="[367,458,402,427]" captionStart="FIGURE 7" captionStartId="19.[151,250,1917,1939]" captionTargetBox="[230,1372,193,1896]" captionTargetId="figure@19.[214,1372,193,1896]" captionTargetPageId="19" captionText="FIGURE 7. Althepus guan Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hookshaped spine; SB = serrated bristles. The scale bar in figures 7A and 7C as in figure 7D." httpUri="https://zenodo.org/record/1439804/files/figure.png" pageId="4" pageNumber="405">Fig. 7D</figureCitation>
|
||
); noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63FFFF3FF1CFE0D0480FE31" pageId="4" pageNumber="405" type="diagnosis">
|
||
<paragraph id="8BE236C7B63FFFF3FF1CFE0D0480FE31" blockId="4.[151,1436,258,1219]" pageId="4" pageNumber="405">
|
||
<emphasis id="B929EAD5B63FFFF3FF1CFE0D0153FE0D" bold="true" box="[199,321,438,463]" pageId="4" pageNumber="405">Diagnosis.</emphasis>
|
||
The species can be easily distinguished from all the other species by the tube-like (
|
||
<figureCitation id="13662A42B63FFFF3FAD2FE0D0571FE0D" box="[1289,1379,438,463]" captionStart="FIGURE 7" captionStartId="19.[151,250,1917,1939]" captionTargetBox="[230,1372,193,1896]" captionTargetId="figure@19.[214,1372,193,1896]" captionTargetPageId="19" captionText="FIGURE 7. Althepus guan Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hookshaped spine; SB = serrated bristles. The scale bar in figures 7A and 7C as in figure 7D." httpUri="https://zenodo.org/record/1439804/files/figure.png" pageId="4" pageNumber="405">Fig. 7D</figureCitation>
|
||
) and bifurcate (
|
||
<figureCitation id="13662A42B63FFFF3FED1FE61014DFE31" box="[266,351,474,499]" captionStart="FIGURE 7" captionStartId="19.[151,250,1917,1939]" captionTargetBox="[230,1372,193,1896]" captionTargetId="figure@19.[214,1372,193,1896]" captionTargetPageId="19" captionText="FIGURE 7. Althepus guan Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hookshaped spine; SB = serrated bristles. The scale bar in figures 7A and 7C as in figure 7D." httpUri="https://zenodo.org/record/1439804/files/figure.png" pageId="4" pageNumber="405">Fig. 7B</figureCitation>
|
||
) embolus in males and one single pair of small spermathecae in females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63FFFF3FF1CFE450508FB95" pageId="4" pageNumber="405" type="description">
|
||
<paragraph id="8BE236C7B63FFFF3FF1CFE45018AFC61" blockId="4.[151,1436,258,1219]" pageId="4" pageNumber="405">
|
||
<emphasis id="B929EAD5B63FFFF3FF1CFE450237FDD5" bold="true" box="[199,549,510,535]" pageId="4" pageNumber="405">
|
||
Description. Male (
|
||
<typeStatus id="54E68865B63FFFF3FE71FE450205FDD5" box="[426,535,510,535]" pageId="4" pageNumber="405" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 5.19; carapace 1.63 length, 1.64 width; abdomen 3.00 length, 1.38 width. Carapace round (
|
||
<figureCitation id="13662A42B63FFFF3FE7EFD9901EEFDF9" box="[421,508,546,571]" captionStart="FIGURE 8" captionStartId="20.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@20.[209,1378,193,1911]" captionTargetPageId="20" captionText="FIGURE 8. Althepus guan Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 8D as in figure 8E." httpUri="https://zenodo.org/record/1439806/files/figure.png" pageId="4" pageNumber="405">Fig. 8C</figureCitation>
|
||
), yellow, with three longitudinal brown bands of different widths, the middle band wider than the other bands. Anterior margin of thoracic region distinctly elevated. Clypeus brown. Cheliceral promargin with two teeth, followed by a lamina, retromargin with two small teeth, posterior surface of fang with 24 small denticles (
|
||
<figureCitation id="13662A42B63FFFF3FE82FD3501DAFD65" box="[345,456,654,679]" captionStart="FIGURE 30" captionStartId="42.[151,250,1917,1939]" captionTargetBox="[177,1410,193,1896]" captionTargetId="figure@42.[177,1410,193,1896]" captionTargetPageId="42" captionText="FIGURE 30. Left cheliceral retromargin of male holotype. A, Althepus bamensis Li et Li sp. nov.; B, A. changmao Li et Li sp. nov.; C, A. dongnaiensis Li et Li sp. nov.; D, A. guan Li et Li sp. nov.; E, A. maechamensis Li et Li sp. nov.; F, A. menglaensis Li et Li sp. nov.; G, A. naphongensis Li et Li sp. nov.; H, A. natmataungensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439850/files/figure.png" pageId="4" pageNumber="405">Fig. 30D</figureCitation>
|
||
). Labium brown. Sternum yellow, margin with a round of brown band. Abdomen elongate, with complex patterns of white and brown dorsally and ventrally. Legs brown, tibia with white annulations. Leg measurements: I missing, II 30.85 (7.69, 0.67, 7.69, 12.50, 2.30), III 18.70 (5.45, 0.65, 4.75, 6.60, 1.25), IV 27.35 (7.69, 0.67, 7.37, 9.62, 2.00). Male palp (
|
||
<figureCitation id="13662A42B63FFFF3FCF0FD4103A2FCD1" box="[811,944,762,787]" captionStart="FIGURE 7" captionStartId="19.[151,250,1917,1939]" captionTargetBox="[230,1372,193,1896]" captionTargetId="figure@19.[214,1372,193,1896]" captionTargetPageId="19" captionText="FIGURE 7. Althepus guan Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hookshaped spine; SB = serrated bristles. The scale bar in figures 7A and 7C as in figure 7D." httpUri="https://zenodo.org/record/1439804/files/figure.png" pageId="4" pageNumber="405">Figs 7A–D</figureCitation>
|
||
): tarsus with three slightly curved, serrated bristles at the top of the cymbial protrusion (
|
||
<figureCitation id="13662A42B63FFFF3FD40FCA502EAFCF5" box="[667,760,798,823]" captionStart="FIGURE 7" captionStartId="19.[151,250,1917,1939]" captionTargetBox="[230,1372,193,1896]" captionTargetId="figure@19.[214,1372,193,1896]" captionTargetPageId="19" captionText="FIGURE 7. Althepus guan Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hookshaped spine; SB = serrated bristles. The scale bar in figures 7A and 7C as in figure 7D." httpUri="https://zenodo.org/record/1439804/files/figure.png" pageId="4" pageNumber="405">Fig. 7A</figureCitation>
|
||
); one hooked spine with tip directed proximally (
|
||
<figureCitation id="13662A42B63FFFF3FAE9FCA5059CFCF5" box="[1330,1422,798,823]" captionStart="FIGURE 7" captionStartId="19.[151,250,1917,1939]" captionTargetBox="[230,1372,193,1896]" captionTargetId="figure@19.[214,1372,193,1896]" captionTargetPageId="19" captionText="FIGURE 7. Althepus guan Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hookshaped spine; SB = serrated bristles. The scale bar in figures 7A and 7C as in figure 7D." httpUri="https://zenodo.org/record/1439804/files/figure.png" pageId="4" pageNumber="405">Fig. 7A</figureCitation>
|
||
); bulb bright yellow, ovate; embolus arising distally from bulb, curved, distal part with a slender furcation; conductor arising prolatero-distally from bulb, shorter than embolus; embolus and conductor widely separated (distance less than diameter of bulb).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63FFFF3FF1CFC1502C2FBF1" blockId="4.[151,1436,258,1219]" pageId="4" pageNumber="405">
|
||
<emphasis id="B929EAD5B63FFFF3FF1CFC1501BFFC05" bold="true" box="[199,429,942,967]" pageId="4" pageNumber="405">
|
||
Female (
|
||
<typeStatus id="54E68865B63FFFF3FEF5FC1501B2FC05" box="[302,416,942,967]" pageId="4" pageNumber="405" type="paratype">paratype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Similar to male in color and general features (
|
||
<figureCitation id="13662A42B63FFFF3FC65FC15042FFC05" box="[958,1085,942,967]" captionStart="FIGURE 8" captionStartId="20.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@20.[209,1378,193,1911]" captionTargetPageId="20" captionText="FIGURE 8. Althepus guan Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 8D as in figure 8E." httpUri="https://zenodo.org/record/1439806/files/figure.png" pageId="4" pageNumber="405">Figs 8D–E</figureCitation>
|
||
), but smaller. Internal genitalia with a pair of spermathecae, distal part swollen (
|
||
<figureCitation id="13662A42B63FFFF3FD1BFC690309FC29" box="[704,795,978,1003]" captionStart="FIGURE 8" captionStartId="20.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@20.[209,1378,193,1911]" captionTargetPageId="20" captionText="FIGURE 8. Althepus guan Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 8D as in figure 8E." httpUri="https://zenodo.org/record/1439806/files/figure.png" pageId="4" pageNumber="405">Fig. 8A</figureCitation>
|
||
). Measurements: total length 4.05; carapace 1.25 length, 1.33 width; abdomen 2.72 length, 1.50 width. Leg measurements: I missing, II missing, III 13.83 (3.85, 0.48, 3.45, 4.65, 1.40), IV 19.57 (5.51, 0.50, 5.38, 6.54, 1.64).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63FFFF3FF1CFB850508FB95" blockId="4.[151,1436,258,1219]" box="[199,1306,1086,1111]" pageId="4" pageNumber="405">
|
||
<emphasis id="B929EAD5B63FFFF3FF1CFB85012EFB95" bold="true" box="[199,316,1086,1111]" pageId="4" pageNumber="405">Variation.</emphasis>
|
||
Males: carapace 1.24, 1.63 long, 1.28, 1.64 width; femur I: 8.97 (leg I lost in
|
||
<typeStatus id="54E68865B63FFFF3FB7CFB85051CFB95" box="[1191,1294,1086,1111]" pageId="4" pageNumber="405" type="holotype">holotype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63FFFF3FF1CFBD903C3FB5C" pageId="4" pageNumber="405" type="distribution">
|
||
<paragraph id="8BE236C7B63FFFF3FF1CFBD9034EFBB9" blockId="4.[151,1436,258,1219]" box="[199,860,1122,1147]" pageId="4" pageNumber="405">
|
||
<emphasis id="B929EAD5B63FFFF3FF1CFBD90172FBB9" bold="true" box="[199,352,1122,1147]" pageId="4" pageNumber="405">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54E68865B63FFFF3FDBBFBD80282FBB9" box="[608,656,1123,1147]" pageId="4" pageNumber="405">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13662A42B63FFFF3FD21FBD9035DFBB9" box="[762,847,1122,1147]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="4" pageNumber="405">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63FFFF3FF1CFB3D03C3FB5C" blockId="4.[151,1436,258,1219]" box="[199,977,1158,1183]" pageId="4" pageNumber="405">
|
||
<emphasis id="B929EAD5B63FFFF3FF1CFB3D0197FB5D" bold="true" box="[199,389,1158,1183]" pageId="4" pageNumber="405">Natural history.</emphasis>
|
||
Collected at a cave entrance at an altitude of
|
||
<specimenCount id="9D5BFD4EB63FFFF3FC58FB3C03C3FB5C" box="[899,977,1159,1183]" pageId="4" pageNumber="405" type="generic">
|
||
<quantity id="4CA59B22B63FFFF3FC58FB3C03DFFB5C" box="[899,973,1159,1183]" metricMagnitude="2" metricUnit="m" metricValue="9.49" pageId="4" pageNumber="405" unit="m" value="949.0">949 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63FFFF3FF1CFB110148FA90" pageId="4" pageNumber="405" type="discussion">
|
||
<paragraph id="8BE236C7B63FFFF3FF1CFB1104CDFB01" blockId="4.[151,1436,258,1219]" box="[199,1247,1194,1219]" pageId="4" pageNumber="405">
|
||
<emphasis id="B929EAD5B63FFFF3FF1CFB110122FB01" bold="true" box="[199,304,1194,1219]" pageId="4" pageNumber="405">Remark.</emphasis>
|
||
<taxonomicName id="4C5D4D44B63FFFF3FEECFB1001F7FB00" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[311,485,1194,1218]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="guan" status="sp. nov.">
|
||
<emphasis id="B929EAD5B63FFFF3FEECFB10019CFB00" box="[311,398,1195,1218]" italics="true" pageId="4" pageNumber="405">A. guan</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B63FFFF3FE68FB1001D4FB00" box="[435,454,1195,1218]" italics="true" pageId="4" pageNumber="405">et</emphasis>
|
||
Li
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB63FFFF3FE37FB110257FB01" box="[492,581,1194,1219]" pageId="4" pageNumber="405" rank="species">
|
||
<emphasis id="B929EAD5B63FFFF3FE37FB110257FB01" bold="true" box="[492,581,1194,1219]" pageId="4" pageNumber="405">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
was labeled as “sp. 88” in the analysis of Li and Li (2018).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63FFFF3FF4CFAAD026CFAF2" blockId="4.[151,638,1302,1362]" box="[151,638,1302,1328]" pageId="4" pageNumber="405">
|
||
<taxonomicName id="4C5D4D44B63FFFF3FF4CFAAD020CFAF2" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[151,542,1302,1328]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="maechamensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B63FFFF3FF4CFAAD020CFAF2" bold="true" box="[151,542,1302,1328]" pageId="4" pageNumber="405">
|
||
<emphasis id="B929EAD5B63FFFF3FF4CFAAD01AEFAF2" bold="true" box="[151,444,1302,1328]" italics="true" pageId="4" pageNumber="405">Althepus maechamensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B63FFFF3FE3EFAAD01E8FAF2" bold="true" box="[485,506,1302,1328]" italics="true" pageId="4" pageNumber="405">et</emphasis>
|
||
Li
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB63FFFF3FDFEFAAC026CFAF2" box="[549,638,1303,1328]" pageId="4" pageNumber="405" rank="species">
|
||
<emphasis id="B929EAD5B63FFFF3FDFEFAAC026CFAF2" bold="true" box="[549,638,1303,1328]" pageId="4" pageNumber="405">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63FFFF3FF4CFA820148FA90" blockId="4.[151,638,1302,1362]" box="[151,346,1337,1362]" pageId="4" pageNumber="405">
|
||
<figureCitation id="13662A42B63FFFF3FF4CFA82011AFA90" box="[151,264,1337,1362]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="21.[151,250,1917,1939]" captionStartId-1="22.[151,250,1933,1955]" captionTargetBox-0="[214,1372,209,1896]" captionTargetBox-1="[209,1378,207,1911]" captionTargetId-0="figure@21.[214,1372,193,1896]" captionTargetId-1="figure@22.[209,1378,193,1911]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURE 9. Althepus maechamensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulbs of A. maechamensis and A. stonei (modified from Deeleman-Reinhold, 1995), prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 9A and 9C as in figure 9D." captionText-1="FIGURE 10. Althepus maechamensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 10D as in figure 10E." httpUri-0="https://zenodo.org/record/1439808/files/figure.png" httpUri-1="https://zenodo.org/record/1439810/files/figure.png" pageId="4" pageNumber="405">Figs 9–10</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B63FFFF3FECFFA810120FA90" box="[276,306,1338,1362]" captionStart="FIGURE 30" captionStartId="42.[151,250,1917,1939]" captionTargetBox="[177,1410,193,1896]" captionTargetId="figure@42.[177,1410,193,1896]" captionTargetPageId="42" captionText="FIGURE 30. Left cheliceral retromargin of male holotype. A, Althepus bamensis Li et Li sp. nov.; B, A. changmao Li et Li sp. nov.; C, A. dongnaiensis Li et Li sp. nov.; D, A. guan Li et Li sp. nov.; E, A. maechamensis Li et Li sp. nov.; F, A. menglaensis Li et Li sp. nov.; G, A. naphongensis Li et Li sp. nov.; H, A. natmataungensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439850/files/figure.png" pageId="4" pageNumber="405">30</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B63FFFF3FEE5FA810148FA90" box="[318,346,1338,1362]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="4" pageNumber="405">32</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63FFFF3FF4CFA3A0221FA20" pageId="4" pageNumber="405" type="materials_examined">
|
||
<paragraph id="8BE236C7B63FFFF3FF4CFA3A0221FA20" blockId="4.[151,1436,1409,2010]" pageId="4" pageNumber="405">
|
||
<materialsCitation id="3B353C9AB63FFFF3FF4CFA3A0373FA7C" ID-GBIF-Occurrence="1913888989" collectingDate="2014-10-14" country="Thailand" elevation="1649" latitude="18.52795" location="Chiangmai Province" longLatPrecision="1" longitude="98.49938" municipality="Mae Cham District" pageId="4" pageNumber="405" specimenCount="2" typeStatus="holotype">
|
||
<emphasis id="B929EAD5B63FFFF3FF4CFA3A01C4FA58" bold="true" box="[151,470,1409,1434]" pageId="4" pageNumber="405">
|
||
<typeStatus id="54E68865B63FFFF3FF4CFA3A00C0FA58" box="[151,210,1409,1434]" pageId="4" pageNumber="405">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68865B63FFFF3FE84FA3A01C2FA58" box="[351,464,1409,1434]" pageId="4" pageNumber="405" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male (IZCAS-Ar 33999),
|
||
<collectingCountry id="F34A7657B63FFFF3FCFCFA3A039FFA58" box="[807,909,1409,1434]" name="Thailand" pageId="4" pageNumber="405">Thailand</collectingCountry>
|
||
,
|
||
<location id="8E82601CB63FFFF3FC78FA3A0484FA58" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB63FFFF3FC78FA3A0484FA58" box="[931,1174,1409,1434]" country="Thailand" latitude="18.52795" longLatPrecision="1" longitude="98.49938" municipality="Mae Cham District" name="Chiangmai Province" pageId="4" pageNumber="405">Chiangmai Province</location>
|
||
,
|
||
<collectingMunicipality id="6B86ACBDB63FFFF3FB71FA39058AFA58" box="[1194,1432,1409,1434]" pageId="4" pageNumber="405">Mae Cham District</collectingMunicipality>
|
||
,
|
||
<geoCoordinate id="EE695000B63FFFF3FF4CFA1E0135FA7C" box="[151,295,1445,1470]" direction="north" orientation="latitude" pageId="4" pageNumber="405" precision="1" value="18.52795">18°31.677'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695000B63FFFF3FEEDFA1E01D1FA7C" box="[310,451,1445,1470]" direction="east" orientation="longitude" pageId="4" pageNumber="405" precision="1" value="98.49938">98°29.963'E</geoCoordinate>
|
||
,
|
||
<elevation id="0070D1F4B63FFFF3FE09FA1E0238FA7F" box="[466,554,1445,1470]" metricMagnitude="3" metricUnit="m" metricValue="1.649" pageId="4" pageNumber="405" unit="m" value="1649.0">
|
||
<quantity id="4CA59B22B63FFFF3FE09FA1E0238FA7F" box="[466,554,1445,1470]" metricMagnitude="3" metricUnit="m" metricValue="1.649" pageId="4" pageNumber="405" unit="m" value="1649.0">1649 m</quantity>
|
||
</elevation>
|
||
altitude,
|
||
<collectingDate id="EFA7E9EFB63FFFF3FD41FA1D0373FA7C" box="[666,865,1445,1470]" pageId="4" pageNumber="405" value="2014-10-14">
|
||
<date id="FFE31007B63FFFF3FD41FA1D0373FA7C" box="[666,865,1445,1470]" pageId="4" pageNumber="405" value="2014-10-14">14 October 2014</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B353C9AB63FFFF3FCB5FA1E0221FA20" ID-GBIF-Occurrence="1913888990" collectingDate="2014-10-14" country="Thailand" elevation="1649" latitude="18.52795" location="Chiangmai Province" longLatPrecision="1" longitude="98.49938" municipality="Mae Cham District" pageId="4" pageNumber="405" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
|
||
<emphasis id="B929EAD5B63FFFF3FCB5FA1E03FDFA7C" bold="true" box="[878,1007,1445,1470]" pageId="4" pageNumber="405">
|
||
<typeStatus id="54E68865B63FFFF3FCB5FA1E03F8FA7C" box="[878,1002,1445,1470]" pageId="4" pageNumber="405" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D5BFD4EB63FFFF3FC21FA1D045BFA7C" box="[1018,1097,1445,1470]" pageId="4" pageNumber="405" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="9D5BFD4EB63FFFF3FB5DFA1D04EBFA7C" box="[1158,1273,1445,1470]" pageId="4" pageNumber="405" type="female">3 females</specimenCount>
|
||
, same data as holotype (IZCAS-Ar 34000–34003).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63FFFF3FF1CFA56040FF9C4" box="[199,1053,1517,1542]" pageId="4" pageNumber="405" type="etymology">
|
||
<paragraph id="8BE236C7B63FFFF3FF1CFA56040FF9C4" blockId="4.[151,1436,1409,2010]" box="[199,1053,1517,1542]" pageId="4" pageNumber="405">
|
||
<emphasis id="B929EAD5B63FFFF3FF1CFA56015CF9C4" bold="true" box="[199,334,1517,1542]" pageId="4" pageNumber="405">Etymology.</emphasis>
|
||
The specific name is an adjective and refers to the
|
||
<typeStatus id="54E68865B63FFFF3FC57FA5503AEF9C4" box="[908,956,1518,1542]" pageId="4" pageNumber="405">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63FFFF3FF1CF9AA0470F888" pageId="4" pageNumber="405" type="diagnosis">
|
||
<paragraph id="8BE236C7B63FFFF3FF1CF9AA0470F888" blockId="4.[151,1436,1409,2010]" pageId="4" pageNumber="405">
|
||
<emphasis id="B929EAD5B63FFFF3FF1CF9AA0153F9E8" bold="true" box="[199,321,1553,1578]" pageId="4" pageNumber="405">Diagnosis.</emphasis>
|
||
The species resembles
|
||
<taxonomicName id="4C5D4D44B63FFFF3FD85F9A903EDF9E8" authority="Deeleman-Reinhold, 1985" authorityName="Deeleman-Reinhold" authorityYear="1985" box="[606,1023,1553,1578]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="stonei">
|
||
<emphasis id="B929EAD5B63FFFF3FD85F9A902D4F9EB" box="[606,710,1554,1577]" italics="true" pageId="4" pageNumber="405">A. stonei</emphasis>
|
||
Deeleman-Reinhold, 1985
|
||
</taxonomicName>
|
||
(Deeleman-Reinhold 1985) and
|
||
<taxonomicName id="4C5D4D44B63FFFF3FA5FF9A90186F98F" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="phadaengensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B63FFFF3FA5FF9A9012FF98F" italics="true" pageId="4" pageNumber="405">A. phadaengensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B63FFFF3FEB9F98D0167F98F" box="[354,373,1590,1613]" italics="true" pageId="4" pageNumber="405">et</emphasis>
|
||
Li
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB63FFFF3FE41F98E01E6F98C" box="[410,500,1589,1614]" pageId="4" pageNumber="405" rank="species">
|
||
<emphasis id="B929EAD5B63FFFF3FE41F98E01E6F98C" bold="true" box="[410,500,1589,1614]" pageId="4" pageNumber="405">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13662A42B63FFFF3FDD9F98E0290F98C" box="[514,642,1589,1614]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStartId-0="28.[151,250,1917,1939]" captionStartId-1="29.[151,250,1933,1955]" captionTargetBox-0="[214,1372,206,1896]" captionTargetBox-1="[209,1378,193,1911]" captionTargetId-0="figure@28.[214,1372,193,1896]" captionTargetId-1="figure@29.[209,1378,193,1911]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionText-0="FIGURE 16. Althepus phadaengensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulbs of A. phadaengensis and A. stonei (modified from Deeleman-Reinhold, 1995), prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 16A and 16C as in figure 16D." captionText-1="FIGURE 17. Althepus phadaengensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 17D as in figure 17E." httpUri-0="https://zenodo.org/record/1439822/files/figure.png" httpUri-1="https://zenodo.org/record/1439824/files/figure.png" pageId="4" pageNumber="405">Figs 16–17</figureCitation>
|
||
) in having a similarly shaped conductor and embolus in males and two pairs of spermathecae in females. Males of the new species can be distinguished by the shorter embolus and conductor, the shorter curved part of the conductor (
|
||
<figureCitation id="13662A42B63FFFF3FD3CF9C60352F954" box="[743,832,1661,1686]" captionStart="FIGURE 9" captionStartId="21.[151,250,1917,1939]" captionTargetBox="[214,1372,209,1896]" captionTargetId="figure@21.[214,1372,193,1896]" captionTargetPageId="21" captionText="FIGURE 9. Althepus maechamensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulbs of A. maechamensis and A. stonei (modified from Deeleman-Reinhold, 1995), prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 9A and 9C as in figure 9D." httpUri="https://zenodo.org/record/1439808/files/figure.png" pageId="4" pageNumber="405">Fig. 9B</figureCitation>
|
||
), compared to the longer embolus and conductor, the longer curved part of the conductor in
|
||
<taxonomicName id="4C5D4D44B63FFFF3FD8FF9190309F97B" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[596,795,1697,1721]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="phadaengensis">
|
||
<emphasis id="B929EAD5B63FFFF3FD8FF9190309F97B" box="[596,795,1697,1721]" italics="true" pageId="4" pageNumber="405">A. phadaengensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13662A42B63FFFF3FCF6F91A0386F978" box="[813,916,1697,1722]" captionStart="FIGURE 16" captionStartId="28.[151,250,1917,1939]" captionTargetBox="[214,1372,206,1896]" captionTargetId="figure@28.[214,1372,193,1896]" captionTargetPageId="28" captionText="FIGURE 16. Althepus phadaengensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulbs of A. phadaengensis and A. stonei (modified from Deeleman-Reinhold, 1995), prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 16A and 16C as in figure 16D." httpUri="https://zenodo.org/record/1439822/files/figure.png" pageId="4" pageNumber="405">Fig. 16B</figureCitation>
|
||
), and by the sharp, upswept distal part of the conductor (
|
||
<figureCitation id="13662A42B63FFFF3FECCF97E0162F91C" box="[279,368,1733,1758]" captionStart="FIGURE 9" captionStartId="21.[151,250,1917,1939]" captionTargetBox="[214,1372,209,1896]" captionTargetId="figure@21.[214,1372,193,1896]" captionTargetPageId="21" captionText="FIGURE 9. Althepus maechamensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulbs of A. maechamensis and A. stonei (modified from Deeleman-Reinhold, 1995), prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 9A and 9C as in figure 9D." httpUri="https://zenodo.org/record/1439808/files/figure.png" pageId="4" pageNumber="405">Fig. 9B</figureCitation>
|
||
), compared with the blunt, pendulous distal part of the conductor in
|
||
<taxonomicName id="4C5D4D44B63FFFF3FBA0F97D04CCF91F" authorityName="Deeleman-Reinhold" authorityYear="1985" box="[1147,1246,1734,1757]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="stonei">
|
||
<emphasis id="B929EAD5B63FFFF3FBA0F97D04CCF91F" box="[1147,1246,1734,1757]" italics="true" pageId="4" pageNumber="405">A. stonei</emphasis>
|
||
</taxonomicName>
|
||
. Females can be distinguished by a more coiled spermathecae (
|
||
<figureCitation id="13662A42B63FFFF3FD76F9520305F8C0" box="[685,791,1769,1794]" captionStart="FIGURE 10" captionStartId="22.[151,250,1933,1955]" captionTargetBox="[209,1378,207,1911]" captionTargetId="figure@22.[209,1378,193,1911]" captionTargetPageId="22" captionText="FIGURE 10. Althepus maechamensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 10D as in figure 10E." httpUri="https://zenodo.org/record/1439810/files/figure.png" pageId="4" pageNumber="405">Fig. 10A</figureCitation>
|
||
), compared with the weakly twisted spermathecae in
|
||
<taxonomicName id="4C5D4D44B63FFFF3FA5FF95100C8F8E7" authorityName="Deeleman-Reinhold" authorityYear="1985" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="stonei">
|
||
<emphasis id="B929EAD5B63FFFF3FA5FF95100C8F8E7" italics="true" pageId="4" pageNumber="405">A. stonei</emphasis>
|
||
</taxonomicName>
|
||
, and by the lateral spermathecae having a more slender stalk than medial spermathecae, compared with the stalks of the spermathecae having the similar diameter in
|
||
<taxonomicName id="4C5D4D44B63FFFF3FCC0F88903CFF88B" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[795,989,1841,1865]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="maechamensis">
|
||
<emphasis id="B929EAD5B63FFFF3FCC0F88903CFF88B" box="[795,989,1841,1865]" italics="true" pageId="4" pageNumber="405">A. maechamensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13662A42B63FFFF3FC36F88A0444F888" box="[1005,1110,1841,1866]" captionStart="FIGURE 17" captionStartId="29.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@29.[209,1378,193,1911]" captionTargetPageId="29" captionText="FIGURE 17. Althepus phadaengensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 17D as in figure 17E." httpUri="https://zenodo.org/record/1439824/files/figure.png" pageId="4" pageNumber="405">Fig. 17A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63FFFF2FF1CF8EE02D0FD0F" lastPageId="5" lastPageNumber="406" pageId="4" pageNumber="405" type="description">
|
||
<paragraph id="8BE236C7B63FFFF2FF1CF8EE02E2FE12" blockId="4.[151,1436,1409,2010]" lastBlockId="5.[151,1436,151,824]" lastPageId="5" lastPageNumber="406" pageId="4" pageNumber="405">
|
||
<emphasis id="B929EAD5B63FFFF3FF1CF8EE0237F8AC" bold="true" box="[199,549,1877,1902]" pageId="4" pageNumber="405">
|
||
Description. Male (
|
||
<typeStatus id="54E68865B63FFFF3FE71F8EE0205F8AC" box="[426,535,1877,1902]" pageId="4" pageNumber="405" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 4.74; carapace 1.72 length, 1.70 width; abdomen 2.68 length, 1.51 width. Carapace round (
|
||
<figureCitation id="13662A42B63FFFF3FE73F8C2021CF850" box="[424,526,1913,1938]" captionStart="FIGURE 10" captionStartId="22.[151,250,1933,1955]" captionTargetBox="[209,1378,207,1911]" captionTargetId="figure@22.[209,1378,193,1911]" captionTargetPageId="22" captionText="FIGURE 10. Althepus maechamensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 10D as in figure 10E." httpUri="https://zenodo.org/record/1439810/files/figure.png" pageId="4" pageNumber="405">Fig. 10C</figureCitation>
|
||
), yellowish, with three longitudinal brown bands of different widths, the middle shortened to form a triangular band around eyes. Anterior margin of thoracic region distinctly elevated. Clypeus brown. Cheliceral promargin with two teeth, followed by a lamina, retromargin with two small teeth, posterior surface of fang with 23 small denticles (
|
||
<figureCitation id="13662A42B63EFFF2FD80FF2C02D3FF72" box="[603,705,151,176]" captionStart="FIGURE 30" captionStartId="42.[151,250,1917,1939]" captionTargetBox="[177,1410,193,1896]" captionTargetId="figure@42.[177,1410,193,1896]" captionTargetPageId="42" captionText="FIGURE 30. Left cheliceral retromargin of male holotype. A, Althepus bamensis Li et Li sp. nov.; B, A. changmao Li et Li sp. nov.; C, A. dongnaiensis Li et Li sp. nov.; D, A. guan Li et Li sp. nov.; E, A. maechamensis Li et Li sp. nov.; F, A. menglaensis Li et Li sp. nov.; G, A. naphongensis Li et Li sp. nov.; H, A. natmataungensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439850/files/figure.png" pageId="5" pageNumber="406">Fig. 30E</figureCitation>
|
||
). Labium brown. Sternum brown, with one irregular longitudinal yellow line in the middle. Abdomen elongate, with complex patterns of white and dark-brown dorsally and ventrally. Legs brown, femur and tibia with yellow annulations. Leg measurements: I 33.01 (8.46, 0.69, 8.33, 13.29, 2.24), II 22.00 (6.09, 0.69, 5.71, 8.14, 1.37), III 15.07 (4.25, 0.67, 3.84, 5.06, 1.25), IV 22.30 (6.41, 0.70, 6.15, 7.44, 1.60). Male palp (
|
||
<figureCitation id="13662A42B63EFFF2FE3FFE9C027AFE82" box="[484,616,295,320]" captionStart="FIGURE 9" captionStartId="21.[151,250,1917,1939]" captionTargetBox="[214,1372,209,1896]" captionTargetId="figure@21.[214,1372,193,1896]" captionTargetPageId="21" captionText="FIGURE 9. Althepus maechamensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulbs of A. maechamensis and A. stonei (modified from Deeleman-Reinhold, 1995), prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 9A and 9C as in figure 9D." httpUri="https://zenodo.org/record/1439808/files/figure.png" pageId="5" pageNumber="406">Figs 9A–D</figureCitation>
|
||
): tarsus distinct, extends distally and with three slightly curved, serrated bristles at the top of cymbial protrusion (
|
||
<figureCitation id="13662A42B63EFFF2FDB1FEF702D7FEA6" box="[618,709,332,357]" captionStart="FIGURE 9" captionStartId="21.[151,250,1917,1939]" captionTargetBox="[214,1372,209,1896]" captionTargetId="figure@21.[214,1372,193,1896]" captionTargetPageId="21" captionText="FIGURE 9. Althepus maechamensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulbs of A. maechamensis and A. stonei (modified from Deeleman-Reinhold, 1995), prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 9A and 9C as in figure 9D." httpUri="https://zenodo.org/record/1439808/files/figure.png" pageId="5" pageNumber="406">Fig. 9A</figureCitation>
|
||
), one hooked spine with tip directed proximally (
|
||
<figureCitation id="13662A42B63EFFF2FB22FEF70546FEA6" box="[1273,1364,332,357]" captionStart="FIGURE 9" captionStartId="21.[151,250,1917,1939]" captionTargetBox="[214,1372,209,1896]" captionTargetId="figure@21.[214,1372,193,1896]" captionTargetPageId="21" captionText="FIGURE 9. Althepus maechamensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulbs of A. maechamensis and A. stonei (modified from Deeleman-Reinhold, 1995), prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 9A and 9C as in figure 9D." httpUri="https://zenodo.org/record/1439808/files/figure.png" pageId="5" pageNumber="406">Fig. 9A</figureCitation>
|
||
); bulb yellow, ovate; embolus arising retrolatero-distally from bulb, proximal part flat and distinctly curved; conductor arising prolatero-distally from bulb, increasingly widening, curved at the distal part; embolus and conductor widely separated (distance almost equal to diameter of bulb).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63EFFF2FF1CFE670144FD46" blockId="5.[151,1436,151,824]" pageId="5" pageNumber="406">
|
||
<emphasis id="B929EAD5B63EFFF2FF1CFE6701BEFE37" bold="true" box="[199,428,476,501]" pageId="5" pageNumber="406">
|
||
Female (
|
||
<typeStatus id="54E68865B63EFFF2FEF6FE67018DFE37" box="[301,415,476,501]" pageId="5" pageNumber="406" type="paratype">paratype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Similar to male in color and general features (
|
||
<figureCitation id="13662A42B63EFFF2FC6FFE670453FE36" box="[948,1089,476,501]" captionStart="FIGURE 10" captionStartId="22.[151,250,1933,1955]" captionTargetBox="[209,1378,207,1911]" captionTargetId="figure@22.[209,1378,193,1911]" captionTargetPageId="22" captionText="FIGURE 10. Althepus maechamensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 10D as in figure 10E." httpUri="https://zenodo.org/record/1439810/files/figure.png" pageId="5" pageNumber="406">Figs 10D–E</figureCitation>
|
||
), but smaller. Internal genitalia with two spermathecae sitting on long, slender stalks on each side (
|
||
<figureCitation id="13662A42B63EFFF2FC78FE44041CFDDA" box="[931,1038,511,536]" captionStart="FIGURE 10" captionStartId="22.[151,250,1933,1955]" captionTargetBox="[209,1378,207,1911]" captionTargetId="figure@22.[209,1378,193,1911]" captionTargetPageId="22" captionText="FIGURE 10. Althepus maechamensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 10D as in figure 10E." httpUri="https://zenodo.org/record/1439810/files/figure.png" pageId="5" pageNumber="406">Fig. 10A</figureCitation>
|
||
). Measurements: total length 3.72; carapace 1.19 length, 1.36 width; abdomen 2.18 length, 1.31 width. Leg measurements: I 23.42 (5.77, 0.53, 5.90, 8.97, 2.25), II 15.07 (4.00, 0.52, 3.75, 5.45, 1.35), III 10.92 (3.13, 0.50, 2.66, 3.48, 1.15), IV 15.33 (4.35, 0.53, 4.25, 5.00, 1.20).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63EFFF2FF1CFD3402D0FD0F" blockId="5.[151,1436,151,824]" pageId="5" pageNumber="406">
|
||
<emphasis id="B929EAD5B63EFFF2FF1CFD34012EFD6A" bold="true" box="[199,316,655,680]" pageId="5" pageNumber="406">Variation.</emphasis>
|
||
Males: carapace 1.34, 1.72 length, 1.27, 1.70 width; femur I: 5.32, 8.46. Females: carapace 1.19– 1.30 length, 1.34–1.40 width; femur I: 5.32–5.90.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63EFFF2FF1CFD6C03F6FCD6" pageId="5" pageNumber="406" type="distribution">
|
||
<paragraph id="8BE236C7B63EFFF2FF1CFD6C034EFD32" blockId="5.[151,1436,151,824]" box="[199,860,727,752]" pageId="5" pageNumber="406">
|
||
<emphasis id="B929EAD5B63EFFF2FF1CFD6C0172FD32" bold="true" box="[199,352,727,752]" pageId="5" pageNumber="406">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54E68865B63EFFF2FDBBFD630282FD32" box="[608,656,728,752]" pageId="5" pageNumber="406">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13662A42B63EFFF2FD21FD6C035DFD32" box="[762,847,727,752]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="5" pageNumber="406">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63EFFF2FF1CFD4703F6FCD6" blockId="5.[151,1436,151,824]" box="[199,996,764,789]" pageId="5" pageNumber="406">
|
||
<emphasis id="B929EAD5B63EFFF2FF1CFD470197FCD7" bold="true" box="[199,389,764,789]" pageId="5" pageNumber="406">Natural history.</emphasis>
|
||
Collected in a tropical forest at an altitude of
|
||
<quantity id="4CA59B22B63EFFF2FC5CFD4703F2FCD6" box="[903,992,764,789]" metricMagnitude="3" metricUnit="m" metricValue="1.649" pageId="5" pageNumber="406" unit="m" value="1649.0">1649 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63EFFF2FF1CFCA40175FC05" pageId="5" pageNumber="406" type="discussion">
|
||
<paragraph id="8BE236C7B63EFFF2FF1CFCA4054AFCFA" blockId="5.[151,1436,151,824]" box="[199,1368,799,824]" pageId="5" pageNumber="406">
|
||
<emphasis id="B929EAD5B63EFFF2FF1CFCA40122FCFA" bold="true" box="[199,304,799,824]" pageId="5" pageNumber="406">Remark.</emphasis>
|
||
<taxonomicName id="4C5D4D44B63EFFF2FEECFC9B0242FCF5" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[311,592,799,824]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="maechamensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B63EFFF2FEECFC9B01E8FCFA" box="[311,506,800,824]" italics="true" pageId="5" pageNumber="406">A. maechamensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B63EFFF2FDC5FC9A0223FCFA" box="[542,561,801,824]" italics="true" pageId="5" pageNumber="406">et</emphasis>
|
||
Li
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB63EFFF2FD8CFCA402A3FCFA" box="[599,689,799,824]" pageId="5" pageNumber="406" rank="species">
|
||
<emphasis id="B929EAD5B63EFFF2FD8CFCA402A3FCFA" bold="true" box="[599,689,799,824]" pageId="5" pageNumber="406">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
was labeled as “sp. 170” in the analysis of Li and Li (2018).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63EFFF2FF4CFC370270FC67" blockId="5.[151,610,908,967]" box="[151,610,908,934]" pageId="5" pageNumber="406">
|
||
<taxonomicName id="4C5D4D44B63EFFF2FF4CFC370210FC64" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[151,514,908,934]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="menglaensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B63EFFF2FF4CFC370210FC64" bold="true" box="[151,514,908,934]" pageId="5" pageNumber="406">
|
||
<emphasis id="B929EAD5B63EFFF2FF4CFC3701B2FC64" bold="true" box="[151,416,908,934]" italics="true" pageId="5" pageNumber="406">Althepus menglaensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B63EFFF2FE11FC3701CDFC64" bold="true" box="[458,479,908,934]" italics="true" pageId="5" pageNumber="406">et</emphasis>
|
||
Li
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB63EFFF2FDD2FC370270FC67" box="[521,610,908,933]" pageId="5" pageNumber="406" rank="species">
|
||
<emphasis id="B929EAD5B63EFFF2FDD2FC370270FC67" bold="true" box="[521,610,908,933]" pageId="5" pageNumber="406">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63EFFF2FF4CFC150175FC05" blockId="5.[151,610,908,967]" box="[151,359,942,967]" pageId="5" pageNumber="406">
|
||
<figureCitation id="13662A42B63EFFF2FF4CFC150104FC05" box="[151,278,942,967]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStartId-0="23.[151,250,1933,1955]" captionStartId-1="24.[151,250,1933,1955]" captionTargetBox-0="[209,1378,205,1911]" captionTargetBox-1="[209,1378,193,1911]" captionTargetId-0="figure@23.[209,1378,193,1911]" captionTargetId-1="figure@24.[209,1378,193,1911]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionText-0="FIGURE 11. Althepus menglaensis Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 11A and 11C as in figure 11D." captionText-1="FIGURE 12. Althepus menglaensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 12D as in figure 12E." httpUri-0="https://zenodo.org/record/1439812/files/figure.png" httpUri-1="https://zenodo.org/record/1439814/files/figure.png" pageId="5" pageNumber="406">Figs 11–12</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B63EFFF2FEFAFC14012DFC05" box="[289,319,943,967]" captionStart="FIGURE 30" captionStartId="42.[151,250,1917,1939]" captionTargetBox="[177,1410,193,1896]" captionTargetId="figure@42.[177,1410,193,1896]" captionTargetPageId="42" captionText="FIGURE 30. Left cheliceral retromargin of male holotype. A, Althepus bamensis Li et Li sp. nov.; B, A. changmao Li et Li sp. nov.; C, A. dongnaiensis Li et Li sp. nov.; D, A. guan Li et Li sp. nov.; E, A. maechamensis Li et Li sp. nov.; F, A. menglaensis Li et Li sp. nov.; G, A. naphongensis Li et Li sp. nov.; H, A. natmataungensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439850/files/figure.png" pageId="5" pageNumber="406">30</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B63EFFF2FE90FC140175FC05" box="[331,359,943,967]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="5" pageNumber="406">32</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63EFFF2FF4CFC4D0164FBB9" pageId="5" pageNumber="406" type="materials_examined">
|
||
<paragraph id="8BE236C7B63EFFF2FF4CFC4D0164FBB9" blockId="5.[151,1437,1014,1903]" pageId="5" pageNumber="406">
|
||
<materialsCitation id="3B353C9AB63EFFF2FF4CFC4D0164FBB9" ID-GBIF-Occurrence="1913888968" collectingDate="2013-06-15" collectorName="S. Li" country="China" county="Mengla County" elevation="761" latitude="21.404217" location="Xishuangbanna Nature Reserve" longLatPrecision="1" longitude="101.6054" municipality="Xiaolongha Village" pageId="5" pageNumber="406" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan Province" typeStatus="holotype">
|
||
<emphasis id="B929EAD5B63EFFF2FF4CFC4D01C3FBCD" bold="true" box="[151,465,1014,1039]" pageId="5" pageNumber="406">
|
||
<typeStatus id="54E68865B63EFFF2FF4CFC4D00C0FBCD" box="[151,210,1014,1039]" pageId="5" pageNumber="406">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68865B63EFFF2FE80FC4D01DEFBCD" box="[347,460,1014,1039]" pageId="5" pageNumber="406" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male (IZCAS-Ar 34004),
|
||
<collectingCountry id="F34A7657B63EFFF2FCC2FC4D034DFBCD" box="[793,863,1014,1039]" name="China" pageId="5" pageNumber="406">China</collectingCountry>
|
||
,
|
||
<collectingRegion id="4999F825B63EFFF2FCA9FC4C042FFBCD" box="[882,1085,1014,1039]" country="China" name="Yunnan" pageId="5" pageNumber="406">Yunnan Province</collectingRegion>
|
||
,
|
||
<collectingCounty id="62834E4BB63EFFF2FB94FC4D0515FBCD" box="[1103,1287,1014,1039]" pageId="5" pageNumber="406">Mengla County</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6B86ACBDB63EFFF2FAC2FC4D00F9FBF1" pageId="5" pageNumber="406">Xiaolongha Village</collectingMunicipality>
|
||
,
|
||
<location id="8E82601CB63EFFF2FED8FBA10297FBF0" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB63EFFF2FED8FBA10297FBF0" box="[259,645,1050,1075]" country="China" county="Mengla County" latitude="21.404217" longLatPrecision="1" longitude="101.6054" municipality="Xiaolongha Village" name="Xishuangbanna Nature Reserve" pageId="5" pageNumber="406" stateProvince="Yunnan Province">Xishuangbanna Nature Reserve</location>
|
||
:
|
||
<location id="8E82601CB63EFFF2FD47FBA10411FBF1" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB63EFFF2FD47FBA10411FBF1" box="[668,1027,1050,1075]" country="China" county="Mengla County" latitude="21.404217" longLatPrecision="1" longitude="101.6054" municipality="Xiaolongha Village" name="Biological Diversity Corridor" pageId="5" pageNumber="406" stateProvince="Yunnan Province">Biological Diversity Corridor</location>
|
||
,
|
||
<location id="8E82601CB63EFFF2FBC7FBA1058AFBF0" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB63EFFF2FBC7FBA1058AFBF0" box="[1052,1432,1050,1075]" country="China" county="Mengla County" latitude="21.404217" longLatPrecision="1" longitude="101.6054" municipality="Xiaolongha Village" name="Tropical Seasonal Rain Forest" pageId="5" pageNumber="406" stateProvince="Yunnan Province">Tropical Seasonal Rain Forest</location>
|
||
,
|
||
<geoCoordinate id="EE695000B63EFFF2FF4CFB850135FB95" box="[151,295,1086,1111]" direction="north" orientation="latitude" pageId="5" pageNumber="406" precision="1" value="21.404217">21°24.253′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695000B63EFFF2FEEEFB8501C3FB95" box="[309,465,1086,1111]" direction="east" orientation="longitude" pageId="5" pageNumber="406" precision="1" value="101.6054">101°36.324′E</geoCoordinate>
|
||
,
|
||
<quantity id="4CA59B22B63EFFF2FE05FB850235FB94" box="[478,551,1086,1111]" metricMagnitude="2" metricUnit="m" metricValue="7.609999999999999" pageId="5" pageNumber="406" unit="m" value="761.0">
|
||
<elevation id="0070D1F4B63EFFF2FE05FB850235FB94" box="[478,551,1086,1111]" metricMagnitude="2" metricUnit="m" metricValue="7.609999999999999" pageId="5" pageNumber="406" unit="m" value="761.0">761 m</elevation>
|
||
</quantity>
|
||
altitude,
|
||
<collectingDate id="EFA7E9EFB63EFFF2FD48FB85033DFB95" box="[659,815,1086,1111]" pageId="5" pageNumber="406" value="2013-06-15">
|
||
<date id="FFE31007B63EFFF2FD48FB85033DFB95" box="[659,815,1086,1111]" pageId="5" pageNumber="406" value="2013-06-15">15 June 2013</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26A85311B63EFFF2FCE7FB840366FB95" box="[828,884,1086,1111]" pageId="5" pageNumber="406">S. Li</collectorName>
|
||
leg.
|
||
<emphasis id="B929EAD5B63EFFF2FC75FB850436FB95" bold="true" box="[942,1060,1086,1111]" pageId="5" pageNumber="406">
|
||
<typeStatus id="54E68865B63EFFF2FC75FB85040DFB95" box="[942,1055,1086,1111]" pageId="5" pageNumber="406" type="paratype">Paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D5BFD4EB63EFFF2FBF6FB840481FB95" box="[1069,1171,1086,1111]" pageId="5" pageNumber="406" type="female">1 female</specimenCount>
|
||
, same data as holotype (IZCAS-Ar 34005).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63EFFF2FF1CFB3D03FCFB5D" box="[199,1006,1158,1183]" pageId="5" pageNumber="406" type="etymology">
|
||
<paragraph id="8BE236C7B63EFFF2FF1CFB3D03FCFB5D" blockId="5.[151,1437,1014,1903]" box="[199,1006,1158,1183]" pageId="5" pageNumber="406">
|
||
<emphasis id="B929EAD5B63EFFF2FF1CFB3D015CFB5D" bold="true" box="[199,334,1158,1183]" pageId="5" pageNumber="406">Etymology.</emphasis>
|
||
The specific name is an adjective refers to the
|
||
<typeStatus id="54E68865B63EFFF2FC86FB3C039FFB5D" box="[861,909,1159,1183]" pageId="5" pageNumber="406">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63EFFF2FF1CFB1104CFFB25" pageId="5" pageNumber="406" type="diagnosis">
|
||
<paragraph id="8BE236C7B63EFFF2FF1CFB1104CFFB25" blockId="5.[151,1437,1014,1903]" pageId="5" pageNumber="406">
|
||
<emphasis id="B929EAD5B63EFFF2FF1CFB110153FB01" bold="true" box="[199,321,1194,1219]" pageId="5" pageNumber="406">Diagnosis.</emphasis>
|
||
This new species can be easily distinguished from all the other species by a ventral protrusion of the palpal tarsus (
|
||
<figureCitation id="13662A42B63EFFF2FE87FB7501F5FB25" box="[348,487,1230,1255]" captionStart="FIGURE 11" captionStartId="23.[151,250,1933,1955]" captionTargetBox="[209,1378,205,1911]" captionTargetId="figure@23.[209,1378,193,1911]" captionTargetPageId="23" captionText="FIGURE 11. Althepus menglaensis Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 11A and 11C as in figure 11D." httpUri="https://zenodo.org/record/1439812/files/figure.png" pageId="5" pageNumber="406">Figs 11A–D</figureCitation>
|
||
) and the wave-shaped margin of the conductor (
|
||
<figureCitation id="13662A42B63EFFF2FBDEFB75047AFB25" box="[1029,1128,1230,1255]" captionStart="FIGURE 11" captionStartId="23.[151,250,1933,1955]" captionTargetBox="[209,1378,205,1911]" captionTargetId="figure@23.[209,1378,193,1911]" captionTargetPageId="23" captionText="FIGURE 11. Althepus menglaensis Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 11A and 11C as in figure 11D." httpUri="https://zenodo.org/record/1439812/files/figure.png" pageId="5" pageNumber="406">Fig. 11B</figureCitation>
|
||
) in males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63EFFF2FF1CFB4902BCF8C1" pageId="5" pageNumber="406" type="description">
|
||
<paragraph id="8BE236C7B63EFFF2FF1CFB4902B4F9B1" blockId="5.[151,1437,1014,1903]" pageId="5" pageNumber="406">
|
||
<emphasis id="B929EAD5B63EFFF2FF1CFB490237FAC9" bold="true" box="[199,549,1266,1291]" pageId="5" pageNumber="406">
|
||
Description. Male (
|
||
<typeStatus id="54E68865B63EFFF2FE71FB490205FAC9" box="[426,535,1266,1291]" pageId="5" pageNumber="406" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 3.60; carapace 1.47 length, 1.48 width; abdomen 2.10 length, 0.98 width. Carapace round (
|
||
<figureCitation id="13662A42B63EFFF2FE7EFAAD0219FAED" box="[421,523,1302,1327]" captionStart="FIGURE 12" captionStartId="24.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@24.[209,1378,193,1911]" captionTargetPageId="24" captionText="FIGURE 12. Althepus menglaensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 12D as in figure 12E." httpUri="https://zenodo.org/record/1439814/files/figure.png" pageId="5" pageNumber="406">Fig. 12C</figureCitation>
|
||
), yellow, with three longitudinal brown bands of different widths, the middle one wider than the lateral ones. Clypeus brown. Cheliceral promargin with two teeth, followed by a lamina, retromargin with two small teeth, posterior surface of fang with 21 small denticles (
|
||
<figureCitation id="13662A42B63EFFF2FC1BFAE50437FAB5" box="[960,1061,1374,1399]" captionStart="FIGURE 30" captionStartId="42.[151,250,1917,1939]" captionTargetBox="[177,1410,193,1896]" captionTargetId="figure@42.[177,1410,193,1896]" captionTargetPageId="42" captionText="FIGURE 30. Left cheliceral retromargin of male holotype. A, Althepus bamensis Li et Li sp. nov.; B, A. changmao Li et Li sp. nov.; C, A. dongnaiensis Li et Li sp. nov.; D, A. guan Li et Li sp. nov.; E, A. maechamensis Li et Li sp. nov.; F, A. menglaensis Li et Li sp. nov.; G, A. naphongensis Li et Li sp. nov.; H, A. natmataungensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439850/files/figure.png" pageId="5" pageNumber="406">Fig. 30F</figureCitation>
|
||
). Labium brown. Sternum brown with triangular yellow spots. Abdomen elongate, with complex patterns of white and dark-brown dorsally and ventrally. Legs brown. Leg measurements: I 28.83 (7.50, 0.55, 7.43, 11.35, 2.00), II 19.67 (5.38, 0.51, 5.04, 6.86, 1.88), III missing, IV 19.63 (5.51, 0.53, 5.64, 6.47, 1.48). Male palp (
|
||
<figureCitation id="13662A42B63EFFF2FC3FFA71046BFA21" box="[996,1145,1482,1507]" captionStart="FIGURE 11" captionStartId="23.[151,250,1933,1955]" captionTargetBox="[209,1378,205,1911]" captionTargetId="figure@23.[209,1378,193,1911]" captionTargetPageId="23" captionText="FIGURE 11. Althepus menglaensis Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 11A and 11C as in figure 11D." httpUri="https://zenodo.org/record/1439812/files/figure.png" pageId="5" pageNumber="406">Figs 11A–D</figureCitation>
|
||
): tarsus distinct, extends distally,with three slightly curved, serrated bristles at the top of the cymbial protrusion (
|
||
<figureCitation id="13662A42B63EFFF2FB4DFA5504EDF9C5" box="[1174,1279,1518,1543]" captionStart="FIGURE 11" captionStartId="23.[151,250,1933,1955]" captionTargetBox="[209,1378,205,1911]" captionTargetId="figure@23.[209,1378,193,1911]" captionTargetPageId="23" captionText="FIGURE 11. Althepus menglaensis Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 11A and 11C as in figure 11D." httpUri="https://zenodo.org/record/1439812/files/figure.png" pageId="5" pageNumber="406">Fig. 11D</figureCitation>
|
||
), one hooked spine with tip directed proximally (
|
||
<figureCitation id="13662A42B63EFFF2FDF0F9A90286F9E9" box="[555,660,1554,1579]" captionStart="FIGURE 11" captionStartId="23.[151,250,1933,1955]" captionTargetBox="[209,1378,205,1911]" captionTargetId="figure@23.[209,1378,193,1911]" captionTargetPageId="23" captionText="FIGURE 11. Althepus menglaensis Li et Li sp. nov., right pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 11A and 11C as in figure 11D." httpUri="https://zenodo.org/record/1439812/files/figure.png" pageId="5" pageNumber="406">Fig. 11A</figureCitation>
|
||
); bulb yellow, ovate; embolus arising retrolaterally from bulb, short and straight; conductor arising distally from bulb, with the wave-shaped margin; embolus and conductor widely separated (distance less than diameter of bulb).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63EFFF2FF1CF9C502BCF8C1" blockId="5.[151,1437,1014,1903]" pageId="5" pageNumber="406">
|
||
<emphasis id="B929EAD5B63EFFF2FF1CF9C501BFF955" bold="true" box="[199,429,1662,1687]" pageId="5" pageNumber="406">
|
||
Female (
|
||
<typeStatus id="54E68865B63EFFF2FEF5F9C501B2F955" box="[302,416,1662,1687]" pageId="5" pageNumber="406" type="paratype">paratype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Similar to male in color and general features (
|
||
<figureCitation id="13662A42B63EFFF2FC18F9C5045EF955" box="[963,1100,1662,1687]" captionStart="FIGURE 12" captionStartId="24.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@24.[209,1378,193,1911]" captionTargetPageId="24" captionText="FIGURE 12. Althepus menglaensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 12D as in figure 12E." httpUri="https://zenodo.org/record/1439814/files/figure.png" pageId="5" pageNumber="406">Figs 12E–F</figureCitation>
|
||
), but larger. Internal genitalia with two pairs of spermathecae, long, curved, distal part spherical (
|
||
<figureCitation id="13662A42B63EFFF2FC79F919041FF979" box="[930,1037,1698,1723]" captionStart="FIGURE 12" captionStartId="24.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@24.[209,1378,193,1911]" captionTargetPageId="24" captionText="FIGURE 12. Althepus menglaensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 12D as in figure 12E." httpUri="https://zenodo.org/record/1439814/files/figure.png" pageId="5" pageNumber="406">Fig. 12A</figureCitation>
|
||
). Measurements: total length 3.80; carapace 1.05 length, 1.23 width; abdomen 2.50 length, 1.65 width. Leg measurements: I missing, II missing, III 10.03 (2.75, 0.31, 2.82, 3.13, 1.02), IV missing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63EFFF2FF1CF8B503EEF888" pageId="5" pageNumber="406" type="distribution">
|
||
<paragraph id="8BE236C7B63EFFF2FF1CF8B5034EF8E5" blockId="5.[151,1437,1014,1903]" box="[199,860,1806,1831]" pageId="5" pageNumber="406">
|
||
<emphasis id="B929EAD5B63EFFF2FF1CF8B50172F8E5" bold="true" box="[199,352,1806,1831]" pageId="5" pageNumber="406">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54E68865B63EFFF2FDBBF8B40282F8E5" box="[608,656,1807,1831]" pageId="5" pageNumber="406">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13662A42B63EFFF2FD21F8B5035DF8E5" box="[762,847,1806,1831]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="5" pageNumber="406">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63EFFF2FF1CF88903EEF888" blockId="5.[151,1437,1014,1903]" box="[199,1020,1842,1867]" pageId="5" pageNumber="406">
|
||
<emphasis id="B929EAD5B63EFFF2FF1CF8890197F889" bold="true" box="[199,389,1842,1867]" pageId="5" pageNumber="406">Natural history.</emphasis>
|
||
Collected among tree buttresses at an altitude of
|
||
<specimenCount id="9D5BFD4EB63EFFF2FC75F88903EEF888" box="[942,1020,1842,1867]" pageId="5" pageNumber="406" type="generic">
|
||
<quantity id="4CA59B22B63EFFF2FC75F88903EAF888" box="[942,1016,1842,1867]" metricMagnitude="2" metricUnit="m" metricValue="7.609999999999999" pageId="5" pageNumber="406" unit="m" value="761.0">761 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63EFFF1FF1CF8ED017AFF11" lastPageId="6" lastPageNumber="407" pageId="5" pageNumber="406" type="discussion">
|
||
<paragraph id="8BE236C7B63EFFF2FF1CF8ED0523F8AD" blockId="5.[151,1437,1014,1903]" box="[199,1329,1878,1903]" pageId="5" pageNumber="406">
|
||
<emphasis id="B929EAD5B63EFFF2FF1CF8ED0122F8AD" bold="true" box="[199,304,1878,1903]" pageId="5" pageNumber="406">Remark.</emphasis>
|
||
<taxonomicName id="4C5D4D44B63EFFF2FEECF8EC0224F8AC" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[311,566,1878,1902]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="menglaensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B63EFFF2FEECF8EC01F3F8AC" box="[311,481,1878,1902]" italics="true" pageId="5" pageNumber="406">A. menglaensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B63EFFF2FDDEF8EC020AF8AC" box="[517,536,1879,1902]" italics="true" pageId="5" pageNumber="406">et</emphasis>
|
||
Li
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB63EFFF2FDE5F8ED028AF8AD" box="[574,664,1878,1903]" pageId="5" pageNumber="406" rank="species">
|
||
<emphasis id="B929EAD5B63EFFF2FDE5F8ED028AF8AD" bold="true" box="[574,664,1878,1903]" pageId="5" pageNumber="406">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
was labeled as “sp. 47” in the analysis of Li and Li (2018).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63DFFF1FF4CFF2C0267FF73" blockId="6.[151,629,151,211]" box="[151,629,151,178]" pageId="6" pageNumber="407">
|
||
<taxonomicName id="4C5D4D44B63DFFF1FF4CFF2C0206FF70" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[151,532,151,178]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="naphongensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B63DFFF1FF4CFF2C0206FF70" bold="true" box="[151,532,151,178]" pageId="6" pageNumber="407">
|
||
<emphasis id="B929EAD5B63DFFF1FF4CFF2C01A3FF73" bold="true" box="[151,433,151,177]" italics="true" pageId="6" pageNumber="407">Althepus naphongensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B63DFFF1FE07FF2C01E3FF73" bold="true" box="[476,497,151,177]" italics="true" pageId="6" pageNumber="407">et</emphasis>
|
||
Li
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB63DFFF1FDC0FF230267FF73" box="[539,629,152,177]" pageId="6" pageNumber="407" rank="species">
|
||
<emphasis id="B929EAD5B63DFFF1FDC0FF230267FF73" bold="true" box="[539,629,152,177]" pageId="6" pageNumber="407">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63DFFF1FF4CFF01017AFF11" blockId="6.[151,629,151,211]" box="[151,360,186,211]" pageId="6" pageNumber="407">
|
||
<figureCitation id="13662A42B63DFFF1FF4CFF010105FF11" box="[151,279,186,211]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStartId-0="25.[151,250,1933,1955]" captionStartId-1="26.[151,250,1933,1955]" captionTargetBox-0="[209,1378,193,1911]" captionTargetBox-1="[209,1378,193,1911]" captionTargetId-0="figure@25.[209,1378,193,1911]" captionTargetId-1="figure@26.[209,1378,193,1911]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="FIGURE 13. Althepus naphongensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, retrolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 13A and 13C as in figure 13D." captionText-1="FIGURE 14. Althepus naphongensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus (abdomen missing), dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 14D as in figure 14E." httpUri-0="https://zenodo.org/record/1439816/files/figure.png" httpUri-1="https://zenodo.org/record/1439818/files/figure.png" pageId="6" pageNumber="407">Figs 13–14</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B63DFFF1FEF9FF000153FF11" box="[290,321,187,211]" captionStart="FIGURE 30" captionStartId="42.[151,250,1917,1939]" captionTargetBox="[177,1410,193,1896]" captionTargetId="figure@42.[177,1410,193,1896]" captionTargetPageId="42" captionText="FIGURE 30. Left cheliceral retromargin of male holotype. A, Althepus bamensis Li et Li sp. nov.; B, A. changmao Li et Li sp. nov.; C, A. dongnaiensis Li et Li sp. nov.; D, A. guan Li et Li sp. nov.; E, A. maechamensis Li et Li sp. nov.; F, A. menglaensis Li et Li sp. nov.; G, A. naphongensis Li et Li sp. nov.; H, A. natmataungensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439850/files/figure.png" pageId="6" pageNumber="407">30</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B63DFFF1FE97FF00017AFF11" box="[332,360,187,211]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="6" pageNumber="407">32</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63DFFF1FF4CFEB90164FEA1" pageId="6" pageNumber="407" type="materials_examined">
|
||
<paragraph id="8BE236C7B63DFFF1FF4CFEB90164FEA1" blockId="6.[151,1437,258,1255]" pageId="6" pageNumber="407">
|
||
<materialsCitation id="3B353C9AB63DFFF1FF4CFEB9039CFEFD" ID-GBIF-Occurrence="1913888972" collectingDate="2011-05-20" country="Vietnam" elevation="179" latitude="22.389866" location="Na Phong Cave" longLatPrecision="1" longitude="105.6137" municipality="Ba Be National Park" pageId="6" pageNumber="407" specimenCount="2" stateProvince="Bac Kan Province" typeStatus="holotype">
|
||
<emphasis id="B929EAD5B63DFFF1FF4CFEB901D1FED9" bold="true" box="[151,451,258,283]" pageId="6" pageNumber="407">
|
||
<typeStatus id="54E68865B63DFFF1FF4CFEB900C0FED9" box="[151,210,258,283]" pageId="6" pageNumber="407">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68865B63DFFF1FE97FEB901AFFED9" box="[332,445,258,283]" pageId="6" pageNumber="407" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male (IZCAS-Ar 34006),
|
||
<collectingCountry id="F34A7657B63DFFF1FD35FEB90342FED9" box="[750,848,258,283]" name="Vietnam" pageId="6" pageNumber="407">Vietnam</collectingCountry>
|
||
,
|
||
<collectingRegion id="4999F825B63DFFF1FC80FEB80439FED9" box="[859,1067,258,283]" country="Vietnam" name="Bac Kan" pageId="6" pageNumber="407">
|
||
Bac
|
||
<collectingRegion id="4999F825B63DFFF1FC55FEB80439FED9" box="[910,1067,258,283]" inferredBySuffix="true" pageId="6" pageNumber="407">Kan Province</collectingRegion>
|
||
</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B86ACBDB63DFFF1FBEDFEB80533FED9" box="[1078,1313,258,283]" pageId="6" pageNumber="407">Ba Be National Park</collectingMunicipality>
|
||
,
|
||
<location id="8E82601CB63DFFF1FAF7FEB800C1FEFD" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB63DFFF1FAF7FEB800C1FEFD" country="Vietnam" latitude="22.389866" longLatPrecision="1" longitude="105.6137" municipality="Ba Be National Park" name="Na Phong Cave" pageId="6" pageNumber="407" stateProvince="Bac Kan Province">Na Phong Cave</location>
|
||
,
|
||
<geoCoordinate id="EE695000B63DFFF1FF38FE9D0166FEFD" box="[227,372,294,319]" direction="north" orientation="latitude" pageId="6" pageNumber="407" precision="1" value="22.389866">22°23.392′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695000B63DFFF1FE5FFE9D0232FEFD" box="[388,544,294,319]" direction="east" orientation="longitude" pageId="6" pageNumber="407" precision="1" value="105.6137">105°36.822′E</geoCoordinate>
|
||
,
|
||
<elevation id="0070D1F4B63DFFF1FDEBFE9C026EFEFC" box="[560,636,295,319]" metricMagnitude="2" metricUnit="m" metricValue="1.7899999999999998" pageId="6" pageNumber="407" unit="m" value="179.0">
|
||
<quantity id="4CA59B22B63DFFF1FDEBFE9C026EFEFC" box="[560,636,295,319]" metricMagnitude="2" metricUnit="m" metricValue="1.7899999999999998" pageId="6" pageNumber="407" unit="m" value="179.0">179 m</quantity>
|
||
</elevation>
|
||
altitude,
|
||
<collectingDate id="EFA7E9EFB63DFFF1FD35FE9C039CFEFD" box="[750,910,295,319]" pageId="6" pageNumber="407" value="2011-05-20">
|
||
<date id="FFE31007B63DFFF1FD35FE9C039CFEFD" box="[750,910,295,319]" pageId="6" pageNumber="407" value="2011-05-20">20 May 2011</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B353C9AB63DFFF1FC46FE9D0164FEA1" ID-GBIF-Occurrence="1913888984" collectingDate="2011-05-20" country="Vietnam" elevation="179" latitude="22.389866" location="Na Phong Cave" longLatPrecision="1" longitude="105.6137" municipality="Ba Be National Park" pageId="6" pageNumber="407" specimenCount="1" specimenCount-female="1" stateProvince="Bac Kan Province" typeStatus="paratype">
|
||
<emphasis id="B929EAD5B63DFFF1FC46FE9D0401FEFD" bold="true" box="[925,1043,294,319]" pageId="6" pageNumber="407">
|
||
<typeStatus id="54E68865B63DFFF1FC46FE9D041CFEFD" box="[925,1038,294,319]" pageId="6" pageNumber="407" type="paratype">Paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D5BFD4EB63DFFF1FBC4FE9C049AFEFD" box="[1055,1160,294,319]" pageId="6" pageNumber="407" type="female">1 female</specimenCount>
|
||
, same data as holotype (IZCAS-Ar 34007).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63DFFF1FF1CFED5058FFE45" box="[199,1437,366,391]" pageId="6" pageNumber="407" type="etymology">
|
||
<paragraph id="8BE236C7B63DFFF1FF1CFED5058FFE45" blockId="6.[151,1437,258,1255]" box="[199,1437,366,391]" pageId="6" pageNumber="407">
|
||
<emphasis id="B929EAD5B63DFFF1FF1CFED5015CFE45" bold="true" box="[199,334,366,391]" pageId="6" pageNumber="407">Etymology.</emphasis>
|
||
The specific name is an adjective and refers to the name of the cave where this species was found.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63DFFF1FF1CFE29036FFE31" pageId="6" pageNumber="407" type="diagnosis">
|
||
<paragraph id="8BE236C7B63DFFF1FF1CFE29036FFE31" blockId="6.[151,1437,258,1255]" pageId="6" pageNumber="407">
|
||
<emphasis id="B929EAD5B63DFFF1FF1CFE290153FE69" bold="true" box="[199,321,402,427]" pageId="6" pageNumber="407">Diagnosis.</emphasis>
|
||
This new species can be easily distinguished from all other species by the slender male palp and the gradually widening conductor (
|
||
<figureCitation id="13662A42B63DFFF1FE2EFE0D026DFE0D" box="[501,639,438,463]" captionStart="FIGURE 13" captionStartId="25.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@25.[209,1378,193,1911]" captionTargetPageId="25" captionText="FIGURE 13. Althepus naphongensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, retrolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 13A and 13C as in figure 13D." httpUri="https://zenodo.org/record/1439816/files/figure.png" pageId="6" pageNumber="407">Figs 13C–D</figureCitation>
|
||
) in males and five rounded spermathecae sitting on long, slender stalks on right side (left side has six spermathecae,
|
||
<figureCitation id="13662A42B63DFFF1FD55FE6102E1FE31" box="[654,755,474,499]" captionStart="FIGURE 14" captionStartId="26.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@26.[209,1378,193,1911]" captionTargetPageId="26" captionText="FIGURE 14. Althepus naphongensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus (abdomen missing), dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 14D as in figure 14E." httpUri="https://zenodo.org/record/1439818/files/figure.png" pageId="6" pageNumber="407">Fig. 14A</figureCitation>
|
||
) in females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63DFFF1FF1CFE4503CEFBB9" pageId="6" pageNumber="407" type="description">
|
||
<paragraph id="8BE236C7B63DFFF1FF1CFE4502B4FC61" blockId="6.[151,1437,258,1255]" pageId="6" pageNumber="407">
|
||
<emphasis id="B929EAD5B63DFFF1FF1CFE45023AFDD5" bold="true" box="[199,552,510,535]" pageId="6" pageNumber="407">
|
||
Description. Male (
|
||
<typeStatus id="54E68865B63DFFF1FE75FE450209FDD5" box="[430,539,510,535]" pageId="6" pageNumber="407" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length -; carapace 1.33 length, 1.42 width; abdomen missing. Carapace round (
|
||
<figureCitation id="13662A42B63DFFF1FF37FD99014AFDF9" box="[236,344,546,571]" captionStart="FIGURE 14" captionStartId="26.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@26.[209,1378,193,1911]" captionTargetPageId="26" captionText="FIGURE 14. Althepus naphongensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus (abdomen missing), dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 14D as in figure 14E." httpUri="https://zenodo.org/record/1439818/files/figure.png" pageId="6" pageNumber="407">Fig. 14C</figureCitation>
|
||
), yellowish, with three longitudinal brown bands of different width, the middle shortened to triangular band around eyes. Anterior margin of thoracic region distinctly elevated. Clypeus brown. Cheliceral promargin with two teeth, followed by a lamina, retromargin with two small teeth, posterior surface of fang with 16 small denticles (
|
||
<figureCitation id="13662A42B63DFFF1FE8FFD3501D2FD65" box="[340,448,654,679]" captionStart="FIGURE 30" captionStartId="42.[151,250,1917,1939]" captionTargetBox="[177,1410,193,1896]" captionTargetId="figure@42.[177,1410,193,1896]" captionTargetPageId="42" captionText="FIGURE 30. Left cheliceral retromargin of male holotype. A, Althepus bamensis Li et Li sp. nov.; B, A. changmao Li et Li sp. nov.; C, A. dongnaiensis Li et Li sp. nov.; D, A. guan Li et Li sp. nov.; E, A. maechamensis Li et Li sp. nov.; F, A. menglaensis Li et Li sp. nov.; G, A. naphongensis Li et Li sp. nov.; H, A. natmataungensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439850/files/figure.png" pageId="6" pageNumber="407">Fig. 30G</figureCitation>
|
||
). Labium brown. Sternum yellow, with two irregular brown lines on each side. Legs brown, femur and tibia with white annulations. Leg measurements: I 36.45 (8.97, 0.54, 8.78, 14.81, 3.35), II 22.67 (6.03, 0.53, 5.38, 8.65, 2.08), III 15.36 (4.24, 0.50, 3.97, 5.13, 1.52), IV 23.00 (6.73, 0.55, 6.28, 7.44, 2.00). Male palp (
|
||
<figureCitation id="13662A42B63DFFF1FF03FD41017EFCD1" box="[216,364,762,787]" captionStart="FIGURE 13" captionStartId="25.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@25.[209,1378,193,1911]" captionTargetPageId="25" captionText="FIGURE 13. Althepus naphongensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, retrolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 13A and 13C as in figure 13D." httpUri="https://zenodo.org/record/1439816/files/figure.png" pageId="6" pageNumber="407">Figs 13A–D</figureCitation>
|
||
): tarsus with three slightly curved, serrated bristles at the top of the cymbial protrusion (
|
||
<figureCitation id="13662A42B63DFFF1FAB4FD4100D9FCF5" captionStart="FIGURE 13" captionStartId="25.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@25.[209,1378,193,1911]" captionTargetPageId="25" captionText="FIGURE 13. Althepus naphongensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, retrolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 13A and 13C as in figure 13D." httpUri="https://zenodo.org/record/1439816/files/figure.png" pageId="6" pageNumber="407">Fig. 13A</figureCitation>
|
||
); one hooked spine with tip directed distally (
|
||
<figureCitation id="13662A42B63DFFF1FD04FCA50358FCF5" box="[735,842,798,823]" captionStart="FIGURE 13" captionStartId="25.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@25.[209,1378,193,1911]" captionTargetPageId="25" captionText="FIGURE 13. Althepus naphongensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, retrolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 13A and 13C as in figure 13D." httpUri="https://zenodo.org/record/1439816/files/figure.png" pageId="6" pageNumber="407">Fig. 13A</figureCitation>
|
||
); tarsus and tibia with many bristles (
|
||
<figureCitation id="13662A42B63DFFF1FB24FCA5059DFCF5" box="[1279,1423,798,823]" captionStart="FIGURE 13" captionStartId="25.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@25.[209,1378,193,1911]" captionTargetPageId="25" captionText="FIGURE 13. Althepus naphongensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, retrolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 13A and 13C as in figure 13D." httpUri="https://zenodo.org/record/1439816/files/figure.png" pageId="6" pageNumber="407">Figs 13C–D</figureCitation>
|
||
); bulb bright yellow, ovate; embolus arising retrolaterally from bulb, short and slightly curved (
|
||
<figureCitation id="13662A42B63DFFF1FB6BFCF9050AFC99" box="[1200,1304,834,859]" captionStart="FIGURE 13" captionStartId="25.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@25.[209,1378,193,1911]" captionTargetPageId="25" captionText="FIGURE 13. Althepus naphongensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, retrolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 13A and 13C as in figure 13D." httpUri="https://zenodo.org/record/1439816/files/figure.png" pageId="6" pageNumber="407">Fig. 13A</figureCitation>
|
||
); conductor arising retrolatero-distally from bulb, distal part gradually widening (
|
||
<figureCitation id="13662A42B63DFFF1FC7BFCDD043CFCBD" box="[928,1070,870,895]" captionStart="FIGURE 13" captionStartId="25.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@25.[209,1378,193,1911]" captionTargetPageId="25" captionText="FIGURE 13. Althepus naphongensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, retrolateral view; C. Palp, retrolateral view; D. Palp, prolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 13A and 13C as in figure 13D." httpUri="https://zenodo.org/record/1439816/files/figure.png" pageId="6" pageNumber="407">Figs 13C–D</figureCitation>
|
||
); embolus and conductor widely separated (distance less than diameter of bulb).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63DFFF1FF1CFC1503CEFBB9" blockId="6.[151,1437,258,1255]" pageId="6" pageNumber="407">
|
||
<emphasis id="B929EAD5B63DFFF1FF1CFC1501BEFC05" bold="true" box="[199,428,942,967]" pageId="6" pageNumber="407">
|
||
Female (
|
||
<typeStatus id="54E68865B63DFFF1FEF6FC15018DFC05" box="[301,415,942,967]" pageId="6" pageNumber="407" type="paratype">paratype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Carapace similar to male in color and general features. Abdomen elongate (
|
||
<figureCitation id="13662A42B63DFFF1FADFFC150582FC05" box="[1284,1424,942,967]" captionStart="FIGURE 14" captionStartId="26.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@26.[209,1378,193,1911]" captionTargetPageId="26" captionText="FIGURE 14. Althepus naphongensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus (abdomen missing), dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 14D as in figure 14E." httpUri="https://zenodo.org/record/1439818/files/figure.png" pageId="6" pageNumber="407">Figs 14D–E</figureCitation>
|
||
), brown, with a yellow herringbone streak dorsally and some longitudinal yellow bands of different widths and lengths ventrally. Internal genitalia with five pairs of spermathecae, slender, distal part swollen (one side has six spermathecae,
|
||
<figureCitation id="13662A42B63DFFF1FEE6FBA101B6FBF1" box="[317,420,1050,1075]" captionStart="FIGURE 14" captionStartId="26.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@26.[209,1378,193,1911]" captionTargetPageId="26" captionText="FIGURE 14. Althepus naphongensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus (abdomen missing), dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 14D as in figure 14E." httpUri="https://zenodo.org/record/1439818/files/figure.png" pageId="6" pageNumber="407">Fig. 14A</figureCitation>
|
||
). Measurements: total length 3.80; carapace 1.35 length, 1.48 width; abdomen 2.22 length, 1.28 width. Leg measurements: I 32.45 (7.95, 0.54, 8.15, 12.69, 3.12), II 20.83 (5.38, 0.53, 5.38, 7.56, 1.98), III 13.87 (3.85, 0.49, 3.65, 4.55, 1.33), IV 20.61 (5.77, 0.54, 5.83, 6.67, 1.80).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63DFFF1FF1CFB3D02AEFB01" pageId="6" pageNumber="407" type="distribution">
|
||
<paragraph id="8BE236C7B63DFFF1FF1CFB3D034EFB5D" blockId="6.[151,1437,258,1255]" box="[199,860,1158,1183]" pageId="6" pageNumber="407">
|
||
<emphasis id="B929EAD5B63DFFF1FF1CFB3D0172FB5D" bold="true" box="[199,352,1158,1183]" pageId="6" pageNumber="407">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54E68865B63DFFF1FDBBFB3C0282FB5D" box="[608,656,1159,1183]" pageId="6" pageNumber="407">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13662A42B63DFFF1FD21FB3D035DFB5D" box="[762,847,1158,1183]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="6" pageNumber="407">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63DFFF1FF1CFB1102AEFB01" blockId="6.[151,1437,258,1255]" box="[199,700,1194,1219]" pageId="6" pageNumber="407">
|
||
<emphasis id="B929EAD5B63DFFF1FF1CFB110197FB01" bold="true" box="[199,389,1194,1219]" pageId="6" pageNumber="407">Natural history.</emphasis>
|
||
Collected on the cave wall.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63DFFF1FF1CFB75012CFAAA" pageId="6" pageNumber="407" type="discussion">
|
||
<paragraph id="8BE236C7B63DFFF1FF1CFB75055AFB25" blockId="6.[151,1437,258,1255]" box="[199,1352,1230,1255]" pageId="6" pageNumber="407">
|
||
<emphasis id="B929EAD5B63DFFF1FF1CFB750122FB25" bold="true" box="[199,304,1230,1255]" pageId="6" pageNumber="407">Remark.</emphasis>
|
||
<taxonomicName id="4C5D4D44B63DFFF1FEECFB740255FB24" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[311,583,1230,1254]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="naphongensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B63DFFF1FEECFB7401E2FB24" box="[311,496,1230,1254]" italics="true" pageId="6" pageNumber="407">A. naphongensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B63DFFF1FDCEFB74023AFB24" box="[533,552,1231,1254]" italics="true" pageId="6" pageNumber="407">et</emphasis>
|
||
Li
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB63DFFF1FD95FB7502B5FB25" box="[590,679,1230,1255]" pageId="6" pageNumber="407" rank="species">
|
||
<emphasis id="B929EAD5B63DFFF1FD95FB7502B5FB25" bold="true" box="[590,679,1230,1255]" pageId="6" pageNumber="407">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
was labeled as “sp. 50 ” in the analysis of Li and Li (2018).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63DFFF1FF4CFA960289FA85" blockId="6.[151,667,1325,1384]" box="[151,667,1325,1351]" pageId="6" pageNumber="407">
|
||
<taxonomicName id="4C5D4D44B63DFFF1FF4CFA960229FA85" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[151,571,1325,1351]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="natmataungensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B63DFFF1FF4CFA960229FA85" bold="true" box="[151,571,1325,1351]" pageId="6" pageNumber="407">
|
||
<emphasis id="B929EAD5B63DFFF1FF4CFA9601CAFA85" bold="true" box="[151,472,1325,1351]" italics="true" pageId="6" pageNumber="407">Althepus natmataungensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B63DFFF1FDD9FA960205FA85" bold="true" box="[514,535,1325,1351]" italics="true" pageId="6" pageNumber="407">et</emphasis>
|
||
Li
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB63DFFF1FD99FA950289FA85" box="[578,667,1326,1351]" pageId="6" pageNumber="407" rank="species">
|
||
<emphasis id="B929EAD5B63DFFF1FD99FA950289FA85" bold="true" box="[578,667,1326,1351]" pageId="6" pageNumber="407">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63DFFF1FF4CFAF4012CFAAA" blockId="6.[151,667,1325,1384]" box="[151,318,1359,1384]" pageId="6" pageNumber="407">
|
||
<figureCitation id="13662A42B63DFFF1FF4CFAF400FEFAAA" box="[151,236,1359,1384]" captionStart="FIGURE 15" captionStartId="27.[151,250,1933,1955]" captionTargetBox="[215,1378,193,1911]" captionTargetId="figure@27.[209,1378,193,1911]" captionTargetPageId="27" captionText="FIGURE 15. Althepus natmataungensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 15A and 15C as in figure 15D." httpUri="https://zenodo.org/record/1439820/files/figure.png" pageId="6" pageNumber="407">Figs 15</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B63DFFF1FF23FAEB0104FAAA" box="[248,278,1360,1384]" captionStart="FIGURE 30" captionStartId="42.[151,250,1917,1939]" captionTargetBox="[177,1410,193,1896]" captionTargetId="figure@42.[177,1410,193,1896]" captionTargetPageId="42" captionText="FIGURE 30. Left cheliceral retromargin of male holotype. A, Althepus bamensis Li et Li sp. nov.; B, A. changmao Li et Li sp. nov.; C, A. dongnaiensis Li et Li sp. nov.; D, A. guan Li et Li sp. nov.; E, A. maechamensis Li et Li sp. nov.; F, A. menglaensis Li et Li sp. nov.; G, A. naphongensis Li et Li sp. nov.; H, A. natmataungensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439850/files/figure.png" pageId="6" pageNumber="407">30</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B63DFFF1FEF9FAEB012CFAAA" box="[290,318,1360,1384]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="6" pageNumber="407">32</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63DFFF1FF4CFA2C0103FA3A" pageId="6" pageNumber="407" type="materials_examined">
|
||
<paragraph id="8BE236C7B63DFFF1FF4CFA2C0103FA3A" blockId="6.[151,1437,1431,2033]" pageId="6" pageNumber="407">
|
||
<materialsCitation id="3B353C9AB63DFFF1FF4CFA2C0103FA3A" ID-GBIF-Occurrence="1913888980" collectingDate="2014-05-16" collectorName="Road S & P. Jager" country="Myanmar" county="Nat Ma Taung National Park" elevation="2543" latitude="21.207983" location="Pristine Primary Forest" longLatPrecision="1" longitude="93.91486" municipality="Nat Ma Taung Summit" pageId="6" pageNumber="407" specimenCount="2" stateProvince="Chin" typeStatus="holotype">
|
||
<emphasis id="B929EAD5B63DFFF1FF4CFA2C01D6FA72" bold="true" box="[151,452,1431,1456]" pageId="6" pageNumber="407">
|
||
<typeStatus id="54E68865B63DFFF1FF4CFA2C00C0FA72" box="[151,210,1431,1456]" pageId="6" pageNumber="407">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68865B63DFFF1FE95FA2C01ADFA72" box="[334,447,1431,1456]" pageId="6" pageNumber="407" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male (IZCAS-Ar 34008),
|
||
<collectingCountry id="F34A7657B63DFFF1FD29FA230373FA72" box="[754,865,1432,1456]" name="Myanmar" pageId="6" pageNumber="407">Myanmar</collectingCountry>
|
||
,
|
||
<collectingRegion id="4999F825B63DFFF1FCB6FA2C03B1FA72" box="[877,931,1431,1456]" country="Myanmar" name="Chin" pageId="6" pageNumber="407">Chin</collectingRegion>
|
||
State,
|
||
<collectingCounty id="62834E4BB63DFFF1FC29FA23052FFA72" box="[1010,1341,1431,1456]" pageId="6" pageNumber="407">Nat Ma Taung National Park</collectingCounty>
|
||
,
|
||
<collectorName id="26A85311B63DFFF1FA92FA2C058EFA72" box="[1353,1436,1431,1456]" pageId="6" pageNumber="407">Road S</collectorName>
|
||
of
|
||
<collectingMunicipality id="6B86ACBDB63DFFF1FF63FA0601D6FA17" box="[184,452,1468,1493]" pageId="6" pageNumber="407">Nat Ma Taung Summit</collectingMunicipality>
|
||
,
|
||
<location id="8E82601CB63DFFF1FE0AFA0702CFFA16" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB63DFFF1FE0AFA0702CFFA16" box="[465,733,1468,1493]" country="Myanmar" county="Nat Ma Taung National Park" latitude="21.207983" longLatPrecision="1" longitude="93.91486" municipality="Nat Ma Taung Summit" name="Pristine Primary Forest" pageId="6" pageNumber="407" stateProvince="Chin">Pristine Primary Forest</location>
|
||
,
|
||
<geoCoordinate id="EE695000B63DFFF1FD30FA07036BFA16" box="[747,889,1468,1492]" direction="north" orientation="latitude" pageId="6" pageNumber="407" precision="1" value="21.207983">21°12.479'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695000B63DFFF1FC5CFA070403FA17" box="[903,1041,1468,1493]" direction="east" orientation="longitude" pageId="6" pageNumber="407" precision="1" value="93.91486">93°54.892'E</geoCoordinate>
|
||
,
|
||
<elevation id="0070D1F4B63DFFF1FBC4FA070464FA16" box="[1055,1142,1468,1493]" metricMagnitude="3" metricUnit="m" metricValue="2.543" pageId="6" pageNumber="407" unit="m" value="2543.0">
|
||
<quantity id="4CA59B22B63DFFF1FBC4FA070464FA16" box="[1055,1142,1468,1493]" metricMagnitude="3" metricUnit="m" metricValue="2.543" pageId="6" pageNumber="407" unit="m" value="2543.0">2543 m</quantity>
|
||
</elevation>
|
||
altitude,
|
||
<collectorName id="26A85311B63DFFF1FB3FFA06052FFA17" box="[1252,1341,1469,1493]" pageId="6" pageNumber="407">P. Jäger</collectorName>
|
||
leg.,
|
||
<collectingDate id="EFA7E9EFB63DFFF1FA5BFA07011FFA3A" pageId="6" pageNumber="407" value="2014-05-16">
|
||
<date id="FFE31007B63DFFF1FA5BFA07011FFA3A" pageId="6" pageNumber="407" value="2014-05-16">16 May 2014</date>
|
||
</collectingDate>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63DFFF1FF1CF9BF03FCF9DF" box="[199,1006,1540,1565]" pageId="6" pageNumber="407" type="etymology">
|
||
<paragraph id="8BE236C7B63DFFF1FF1CF9BF03FCF9DF" blockId="6.[151,1437,1431,2033]" box="[199,1006,1540,1565]" pageId="6" pageNumber="407">
|
||
<emphasis id="B929EAD5B63DFFF1FF1CF9BF015CF9DF" bold="true" box="[199,334,1540,1565]" pageId="6" pageNumber="407">Etymology.</emphasis>
|
||
The specific name is an adjective refers to the
|
||
<typeStatus id="54E68865B63DFFF1FC86F9BE039FF9DF" box="[861,909,1541,1565]" pageId="6" pageNumber="407">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63DFFF1FF1CF99C0317F9A7" pageId="6" pageNumber="407" type="diagnosis">
|
||
<paragraph id="8BE236C7B63DFFF1FF1CF99C0317F9A7" blockId="6.[151,1437,1431,2033]" pageId="6" pageNumber="407">
|
||
<emphasis id="B929EAD5B63DFFF1FF1CF99C0153F982" bold="true" box="[199,321,1575,1600]" pageId="6" pageNumber="407">Diagnosis.</emphasis>
|
||
This new species can be easily recognized from all the other species by the bifurcate distal part of the conductor and the wavelike embolus (
|
||
<figureCitation id="13662A42B63DFFF1FDB1F9F702E8F9A7" box="[618,762,1612,1637]" captionStart="FIGURE 15" captionStartId="27.[151,250,1933,1955]" captionTargetBox="[215,1378,193,1911]" captionTargetId="figure@27.[209,1378,193,1911]" captionTargetPageId="27" captionText="FIGURE 15. Althepus natmataungensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 15A and 15C as in figure 15D." httpUri="https://zenodo.org/record/1439820/files/figure.png" pageId="6" pageNumber="407">Figs 15A–D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63DFFF0FF1CF9D4018FFF72" lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407" type="description">
|
||
<paragraph id="8BE236C7B63DFFF1FF1CF9D40173F833" blockId="6.[151,1437,1431,2033]" pageId="6" pageNumber="407">
|
||
<emphasis id="B929EAD5B63DFFF1FF1CF9D40237F94A" bold="true" box="[199,549,1647,1672]" pageId="6" pageNumber="407">
|
||
Description. Male (
|
||
<typeStatus id="54E68865B63DFFF1FE71F9D40205F94A" box="[426,535,1647,1672]" pageId="6" pageNumber="407" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 3.85; carapace 1.30 length, 1.38 width; abdomen 2.13 length, 1.19 width. Carapace rounded, yellow, with three longitudinal brown bands of different width, the middle one wider than the lateral ones. Clypeus brown. Cheliceral promargin with two teeth, followed by a lamina, retromargin with two small teeth, posterior surface of fang with 19 small denticles (
|
||
<figureCitation id="13662A42B63DFFF1FC7FF9670402F936" box="[932,1040,1756,1781]" captionStart="FIGURE 30" captionStartId="42.[151,250,1917,1939]" captionTargetBox="[177,1410,193,1896]" captionTargetId="figure@42.[177,1410,193,1896]" captionTargetPageId="42" captionText="FIGURE 30. Left cheliceral retromargin of male holotype. A, Althepus bamensis Li et Li sp. nov.; B, A. changmao Li et Li sp. nov.; C, A. dongnaiensis Li et Li sp. nov.; D, A. guan Li et Li sp. nov.; E, A. maechamensis Li et Li sp. nov.; F, A. menglaensis Li et Li sp. nov.; G, A. naphongensis Li et Li sp. nov.; H, A. natmataungensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439850/files/figure.png" pageId="6" pageNumber="407">Fig. 30H</figureCitation>
|
||
). Labium brown. Sternum yellow, margin with brown band. Abdomen elongate, with complex pattern of white and brown dorsally and ventrally. Legs brown, femur and tibia with white annulations. Leg measurements: I 16.26 (4.50, 0.57, 4.10, 5.45, 1.64), II 15.88 (4.25, 0.56, 4.00, 5.45, 1.62), III 11.41 (3.20, 0.56, 3.00, 3.40, 1.25), IV missing. Male palp (
|
||
<figureCitation id="13662A42B63DFFF1FB24F8FC0582F8A2" box="[1279,1424,1863,1888]" captionStart="FIGURE 15" captionStartId="27.[151,250,1933,1955]" captionTargetBox="[215,1378,193,1911]" captionTargetId="figure@27.[209,1378,193,1911]" captionTargetPageId="27" captionText="FIGURE 15. Althepus natmataungensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 15A and 15C as in figure 15D." httpUri="https://zenodo.org/record/1439820/files/figure.png" pageId="6" pageNumber="407">Figs 15A–D</figureCitation>
|
||
): tarsus with three slightly curved, serrated bristles at the top (one of them missing,
|
||
<figureCitation id="13662A42B63DFFF1FBECF8D70488F847" box="[1079,1178,1900,1925]" captionStart="FIGURE 15" captionStartId="27.[151,250,1933,1955]" captionTargetBox="[215,1378,193,1911]" captionTargetId="figure@27.[209,1378,193,1911]" captionTargetPageId="27" captionText="FIGURE 15. Althepus natmataungensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 15A and 15C as in figure 15D." httpUri="https://zenodo.org/record/1439820/files/figure.png" pageId="6" pageNumber="407">Fig. 15C</figureCitation>
|
||
) and one large hooked spine with tip directed distally (
|
||
<figureCitation id="13662A42B63DFFF1FDDAF8340279F86A" box="[513,619,1935,1960]" captionStart="FIGURE 15" captionStartId="27.[151,250,1933,1955]" captionTargetBox="[215,1378,193,1911]" captionTargetId="figure@27.[209,1378,193,1911]" captionTargetPageId="27" captionText="FIGURE 15. Althepus natmataungensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 15A and 15C as in figure 15D." httpUri="https://zenodo.org/record/1439820/files/figure.png" pageId="6" pageNumber="407">Fig. 15A</figureCitation>
|
||
); bulb yellow, ovate; embolus arising distally from bulb, flat and wavy; conductor arising proximally from bulb, bifurcate; embolus and conductor widely separated (distance less than diameter of bulb).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63CFFF0FF1CFF2C018FFF72" blockId="7.[199,1394,151,285]" box="[199,413,151,176]" pageId="7" pageNumber="408">
|
||
<emphasis id="B929EAD5B63CFFF0FF1CFF2C0135FF72" bold="true" box="[199,295,151,176]" pageId="7" pageNumber="408">Female:</emphasis>
|
||
Unknown
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63CFFF0FF1CFF070388FF3A" pageId="7" pageNumber="408" type="distribution">
|
||
<paragraph id="8BE236C7B63CFFF0FF1CFF07034EFF16" blockId="7.[199,1394,151,285]" box="[199,860,188,213]" pageId="7" pageNumber="408">
|
||
<emphasis id="B929EAD5B63CFFF0FF1CFF070172FF17" bold="true" box="[199,352,188,213]" pageId="7" pageNumber="408">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54E68865B63CFFF0FDBBFF060282FF17" box="[608,656,189,213]" pageId="7" pageNumber="408">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13662A42B63CFFF0FD21FF07035DFF16" box="[762,847,188,213]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="7" pageNumber="408">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63CFFF0FF1CFF640388FF3A" blockId="7.[199,1394,151,285]" box="[199,922,223,248]" pageId="7" pageNumber="408">
|
||
<emphasis id="B929EAD5B63CFFF0FF1CFF640197FF3A" bold="true" box="[199,389,223,248]" pageId="7" pageNumber="408">Natural history.</emphasis>
|
||
Collected under leaf litter by hand and sieving.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63CFFF0FF1CFEBF017AFE69" pageId="7" pageNumber="408" type="discussion">
|
||
<paragraph id="8BE236C7B63CFFF0FF1CFEBF0560FEDE" blockId="7.[199,1394,151,285]" box="[199,1394,260,285]" pageId="7" pageNumber="408">
|
||
<emphasis id="B929EAD5B63CFFF0FF1CFEBF0122FEDF" bold="true" box="[199,304,260,285]" pageId="7" pageNumber="408">Remark.</emphasis>
|
||
<taxonomicName id="4C5D4D44B63CFFF0FEECFEBF0279FEDE" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[311,619,260,284]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="natmataungensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B63CFFF0FEECFEBF0206FEDE" box="[311,532,260,284]" italics="true" pageId="7" pageNumber="408">A. natmataungensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B63CFFF0FDE2FEBE025EFEDE" box="[569,588,261,284]" italics="true" pageId="7" pageNumber="408">et</emphasis>
|
||
Li
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB63CFFF0FDA9FEBF02D9FEDF" box="[626,715,260,285]" pageId="7" pageNumber="408" rank="species">
|
||
<emphasis id="B929EAD5B63CFFF0FDA9FEBF02D9FEDF" bold="true" box="[626,715,260,285]" pageId="7" pageNumber="408">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
was labeled as “sp. 210” in the analysis of Li and Li (2018).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63CFFF0FF4CFED40292FE4B" blockId="7.[151,640,367,427]" box="[151,640,367,394]" pageId="7" pageNumber="408">
|
||
<taxonomicName id="4C5D4D44B63CFFF0FF4CFED40232FE48" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[151,544,367,394]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="phadaengensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B63CFFF0FF4CFED40232FE48" bold="true" box="[151,544,367,394]" pageId="7" pageNumber="408">
|
||
<emphasis id="B929EAD5B63CFFF0FF4CFED401AFFE4B" bold="true" box="[151,445,367,393]" italics="true" pageId="7" pageNumber="408">Althepus phadaengensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B63CFFF0FE3CFED401EEFE4B" bold="true" box="[487,508,367,393]" italics="true" pageId="7" pageNumber="408">et</emphasis>
|
||
Li
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB63CFFF0FDFDFECB0292FE4B" box="[550,640,368,393]" pageId="7" pageNumber="408" rank="species">
|
||
<emphasis id="B929EAD5B63CFFF0FDFDFECB0292FE4B" bold="true" box="[550,640,368,393]" pageId="7" pageNumber="408">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63CFFF0FF4CFE29017AFE69" blockId="7.[151,640,367,427]" box="[151,360,402,427]" pageId="7" pageNumber="408">
|
||
<figureCitation id="13662A42B63CFFF0FF4CFE290105FE69" box="[151,279,402,427]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStartId-0="28.[151,250,1917,1939]" captionStartId-1="29.[151,250,1933,1955]" captionTargetBox-0="[214,1372,206,1896]" captionTargetBox-1="[209,1378,193,1911]" captionTargetId-0="figure@28.[214,1372,193,1896]" captionTargetId-1="figure@29.[209,1378,193,1911]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionText-0="FIGURE 16. Althepus phadaengensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulbs of A. phadaengensis and A. stonei (modified from Deeleman-Reinhold, 1995), prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 16A and 16C as in figure 16D." captionText-1="FIGURE 17. Althepus phadaengensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 17D as in figure 17E." httpUri-0="https://zenodo.org/record/1439822/files/figure.png" httpUri-1="https://zenodo.org/record/1439824/files/figure.png" pageId="7" pageNumber="408">Figs 16–17</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B63CFFF0FEF9FE28017AFE69" box="[290,360,403,427]" captionStart-0="FIGURE 31" captionStart-1="FIGURE 32" captionStartId-0="43.[151,250,1917,1939]" captionStartId-1="44.[151,250,1148,1170]" captionTargetBox-0="[177,1409,195,1896]" captionTargetBox-1="[393,1194,195,1124]" captionTargetId-0="figure@43.[177,1410,193,1896]" captionTargetId-1="figure@44.[393,1194,195,1124]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionText-0="FIGURE 31. Left cheliceral retromargin of male holotype (A–F), female holotype (G) and female paratype (H). A, Althepus phadaengensis Li et Li sp. nov.; B, A. shanhu Li et Li sp. nov.; C, A. suayaiensis Li et Li sp. nov.; D, A. tadetuensis Li et Li sp. nov.; E, A. tanhuang Li et Li sp. nov.; F, A. thanlaensis Li et Li sp. nov.; G, A. tharnlodensis Li et Li sp. nov.; H, A. viengkeoensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." captionText-1="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri-0="https://zenodo.org/record/1439852/files/figure.png" httpUri-1="https://zenodo.org/record/1439854/files/figure.png" pageId="7" pageNumber="408">31–32</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63CFFF0FF4CFE610444FDF9" pageId="7" pageNumber="408" type="materials_examined">
|
||
<paragraph id="8BE236C7B63CFFF0FF4CFE610444FDF9" blockId="7.[151,1437,474,1651]" pageId="7" pageNumber="408">
|
||
<materialsCitation id="3B353C9AB63CFFF0FF4CFE61058BFDD5" ID-GBIF-Occurrence="1913888982" collectingDate="2016-11-21" country="Thailand" county="Muang District" elevation="293" latitude="19.42325" location="Tham" longLatPrecision="1" longitude="97.98428" municipality="Jumpae Subdistrict" pageId="7" pageNumber="408" specimenCount="2" stateProvince="Mae Hong Son Province" typeStatus="holotype">
|
||
<emphasis id="B929EAD5B63CFFF0FF4CFE6101D6FE31" bold="true" box="[151,452,474,499]" pageId="7" pageNumber="408">
|
||
<typeStatus id="54E68865B63CFFF0FF4CFE6100C0FE31" box="[151,210,474,499]" pageId="7" pageNumber="408">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68865B63CFFF0FE95FE6101ADFE31" box="[334,447,474,499]" pageId="7" pageNumber="408" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male (IZCAS-Ar 34009),
|
||
<collectingCountry id="F34A7657B63CFFF0FD29FE61034AFE31" box="[754,856,474,499]" name="Thailand" pageId="7" pageNumber="408">Thailand</collectingCountry>
|
||
,
|
||
<collectingRegion id="4999F825B63CFFF0FCBEFE60046DFE31" box="[869,1151,474,499]" country="Thailand" name="Mae Hong Son" pageId="7" pageNumber="408">
|
||
Mae
|
||
<collectingRegion id="4999F825B63CFFF0FC44FE60046DFE31" box="[927,1151,474,499]" inferredBySuffix="true" pageId="7" pageNumber="408">Hong Son Province</collectingRegion>
|
||
</collectingRegion>
|
||
,
|
||
<collectingCounty id="62834E4BB63CFFF0FB57FE600528FE31" box="[1164,1338,474,499]" pageId="7" pageNumber="408">Muang District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6B86ACBDB63CFFF0FA9CFE600101FDD5" pageId="7" pageNumber="408">Jumpae Subdistrict</collectingMunicipality>
|
||
,
|
||
<location id="8E82601CB63CFFF0FEFEFE450174FDD5" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB63CFFF0FEFEFE450174FDD5" box="[293,358,510,535]" country="Thailand" county="Muang District" latitude="19.42325" longLatPrecision="1" longitude="97.98428" municipality="Jumpae Subdistrict" name="Tham" pageId="7" pageNumber="408" stateProvince="Mae Hong Son Province">Tham</location>
|
||
(cave)
|
||
<location id="8E82601CB63CFFF0FE12FE450286FDD5" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB63CFFF0FE12FE450286FDD5" box="[457,660,510,535]" country="Thailand" county="Muang District" latitude="19.42325" longLatPrecision="1" longitude="97.98428" municipality="Jumpae Subdistrict" name="Pha Daeng Cave" pageId="7" pageNumber="408" stateProvince="Mae Hong Son Province">Pha Daeng Cave</location>
|
||
,
|
||
<geoCoordinate id="EE695000B63CFFF0FD7CFE450325FDD5" box="[679,823,510,535]" direction="north" orientation="latitude" pageId="7" pageNumber="408" precision="1" value="19.42325">19°25.395'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695000B63CFFF0FC91FE4503C4FDD5" box="[842,982,510,535]" direction="east" orientation="longitude" pageId="7" pageNumber="408" precision="1" value="97.98428">97°59.057'E</geoCoordinate>
|
||
,
|
||
<elevation id="0070D1F4B63CFFF0FC32FE440425FDD4" box="[1001,1079,511,535]" metricMagnitude="2" metricUnit="m" metricValue="2.93" pageId="7" pageNumber="408" unit="m" value="293.0">
|
||
<quantity id="4CA59B22B63CFFF0FC32FE440425FDD4" box="[1001,1079,511,535]" metricMagnitude="2" metricUnit="m" metricValue="2.93" pageId="7" pageNumber="408" unit="m" value="293.0">293 m</quantity>
|
||
</elevation>
|
||
altitude,
|
||
<collectingDate id="EFA7E9EFB63CFFF0FB75FE44058BFDD5" box="[1198,1433,510,535]" pageId="7" pageNumber="408" value="2016-11-21">
|
||
<date id="FFE31007B63CFFF0FB75FE44058BFDD5" box="[1198,1433,510,535]" pageId="7" pageNumber="408" value="2016-11-21">21 November 2016</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B353C9AB63CFFF0FF4CFD990444FDF9" ID-GBIF-Occurrence="1913888981" box="[151,1110,546,571]" collectingDate="2016-11-21" country="Thailand" county="Muang District" elevation="293" latitude="19.42325" location="Tham" longLatPrecision="1" longitude="97.98428" municipality="Jumpae Subdistrict" pageId="7" pageNumber="408" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Mae Hong Son Province" typeStatus="paratype">
|
||
<emphasis id="B929EAD5B63CFFF0FF4CFD99010AFDF9" bold="true" box="[151,280,546,571]" pageId="7" pageNumber="408">
|
||
<typeStatus id="54E68865B63CFFF0FF4CFD990101FDF9" box="[151,275,546,571]" pageId="7" pageNumber="408" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D5BFD4EB63CFFF0FEC5FD980167FDF9" box="[286,373,546,571]" pageId="7" pageNumber="408" type="male">3 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D5BFD4EB63CFFF0FE70FD980208FDF9" box="[427,538,546,571]" pageId="7" pageNumber="408" type="female">2 females</specimenCount>
|
||
, same data as holotype (IZCAS-Ar 34010–34014).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63CFFF0FF1CFDFD0539FD9D" box="[199,1323,582,607]" pageId="7" pageNumber="408" type="etymology">
|
||
<paragraph id="8BE236C7B63CFFF0FF1CFDFD0539FD9D" blockId="7.[151,1437,474,1651]" box="[199,1323,582,607]" pageId="7" pageNumber="408">
|
||
<emphasis id="B929EAD5B63CFFF0FF1CFDFD015CFD9D" bold="true" box="[199,334,582,607]" pageId="7" pageNumber="408">Etymology.</emphasis>
|
||
The specific name is an adjective and refers to the cave in which the species was found.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63CFFF0FF1CFDD10438FCBD" pageId="7" pageNumber="408" type="diagnosis">
|
||
<paragraph id="8BE236C7B63CFFF0FF1CFDD10438FCBD" blockId="7.[151,1437,474,1651]" pageId="7" pageNumber="408">
|
||
<emphasis id="B929EAD5B63CFFF0FF1CFDD10153FD41" bold="true" box="[199,321,618,643]" pageId="7" pageNumber="408">Diagnosis.</emphasis>
|
||
The species resembles
|
||
<taxonomicName id="4C5D4D44B63CFFF0FD85FDD003EDFD41" authority="Deeleman-Reinhold, 1995" authorityName="Deeleman-Reinhold" authorityYear="1995" box="[606,1023,618,643]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="stonei">
|
||
<emphasis id="B929EAD5B63CFFF0FD85FDD002D4FD40" box="[606,710,619,642]" italics="true" pageId="7" pageNumber="408">A. stonei</emphasis>
|
||
Deeleman-Reinhold, 1995
|
||
</taxonomicName>
|
||
(Deeleman-Reinhold 1995) and
|
||
<taxonomicName id="4C5D4D44B63CFFF0FA5FFDD00181FD64" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="maechamensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B63CFFF0FA5FFDD00128FD64" italics="true" pageId="7" pageNumber="408">A. maechamensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B63CFFF0FEBAFD340167FD64" box="[353,373,655,678]" italics="true" pageId="7" pageNumber="408">et</emphasis>
|
||
Li
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB63CFFF0FE40FD3501E7FD65" box="[411,501,654,679]" pageId="7" pageNumber="408" rank="species">
|
||
<emphasis id="B929EAD5B63CFFF0FE40FD3501E7FD65" bold="true" box="[411,501,654,679]" pageId="7" pageNumber="408">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13662A42B63CFFF0FDDFFD350265FD65" box="[516,631,654,679]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="21.[151,250,1917,1939]" captionStartId-1="22.[151,250,1933,1955]" captionTargetBox-0="[214,1372,209,1896]" captionTargetBox-1="[209,1378,207,1911]" captionTargetId-0="figure@21.[214,1372,193,1896]" captionTargetId-1="figure@22.[209,1378,193,1911]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURE 9. Althepus maechamensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulbs of A. maechamensis and A. stonei (modified from Deeleman-Reinhold, 1995), prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 9A and 9C as in figure 9D." captionText-1="FIGURE 10. Althepus maechamensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 10D as in figure 10E." httpUri-0="https://zenodo.org/record/1439808/files/figure.png" httpUri-1="https://zenodo.org/record/1439810/files/figure.png" pageId="7" pageNumber="408">Figs 9–10</figureCitation>
|
||
) in having a similarly shaped conductor and embolus in males and two pairs of spermathecae in females, but can be distinguished by the longer embolus and conductor, the longer curved part of the conductor (
|
||
<figureCitation id="13662A42B63CFFF0FE4FFD6D01E8FD2D" box="[404,506,726,751]" captionStart="FIGURE 16" captionStartId="28.[151,250,1917,1939]" captionTargetBox="[214,1372,206,1896]" captionTargetId="figure@28.[214,1372,193,1896]" captionTargetPageId="28" captionText="FIGURE 16. Althepus phadaengensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulbs of A. phadaengensis and A. stonei (modified from Deeleman-Reinhold, 1995), prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 16A and 16C as in figure 16D." httpUri="https://zenodo.org/record/1439822/files/figure.png" pageId="7" pageNumber="408">Fig. 16B</figureCitation>
|
||
), compared with the shorter embolus and conductor, the shorter curved part of the conductor in
|
||
<taxonomicName id="4C5D4D44B63CFFF0FEF6FD4001E2FCD1" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[301,496,763,787]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="maechamensis">
|
||
<emphasis id="B929EAD5B63CFFF0FEF6FD4001E2FCD1" box="[301,496,763,787]" italics="true" pageId="7" pageNumber="408">A. maechamensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13662A42B63CFFF0FDDAFD41024BFCD1" box="[513,601,762,787]" captionStart="FIGURE 9" captionStartId="21.[151,250,1917,1939]" captionTargetBox="[214,1372,209,1896]" captionTargetId="figure@21.[214,1372,193,1896]" captionTargetPageId="21" captionText="FIGURE 9. Althepus maechamensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulbs of A. maechamensis and A. stonei (modified from Deeleman-Reinhold, 1995), prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 9A and 9C as in figure 9D." httpUri="https://zenodo.org/record/1439808/files/figure.png" pageId="7" pageNumber="408">Fig. 9B</figureCitation>
|
||
), by the sharp, upswept distal part of the conductor (
|
||
<figureCitation id="13662A42B63CFFF0FB6AFD41050AFCD1" box="[1201,1304,762,787]" captionStart="FIGURE 16" captionStartId="28.[151,250,1917,1939]" captionTargetBox="[214,1372,206,1896]" captionTargetId="figure@28.[214,1372,193,1896]" captionTargetPageId="28" captionText="FIGURE 16. Althepus phadaengensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulbs of A. phadaengensis and A. stonei (modified from Deeleman-Reinhold, 1995), prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 16A and 16C as in figure 16D." httpUri="https://zenodo.org/record/1439822/files/figure.png" pageId="7" pageNumber="408">Fig. 16B</figureCitation>
|
||
), compared with the blunt, pendulous distal part of the conductor in
|
||
<taxonomicName id="4C5D4D44B63CFFF0FC88FCA403AFFCF4" authorityName="Deeleman-Reinhold" authorityYear="1995" box="[851,957,799,822]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="stonei">
|
||
<emphasis id="B929EAD5B63CFFF0FC88FCA403AFFCF4" box="[851,957,799,822]" italics="true" pageId="7" pageNumber="408">A. stonei</emphasis>
|
||
</taxonomicName>
|
||
in the males, and by the stalks of the spermathecae having the similar diameter, compared with the lateral spermathecae having a more slender stalk than medial spermathecae in
|
||
<taxonomicName id="4C5D4D44B63CFFF0FE7DFCDC021AFCBC" authorityName="Deeleman-Reinhold" authorityYear="1995" box="[422,520,871,894]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="stonei">
|
||
<emphasis id="B929EAD5B63CFFF0FE7DFCDC021AFCBC" box="[422,520,871,894]" italics="true" pageId="7" pageNumber="408">A. stonei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5D4D44B63CFFF0FDE5FCDC0312FCBC" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[574,768,870,894]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="maechamensis">
|
||
<emphasis id="B929EAD5B63CFFF0FDE5FCDC0312FCBC" box="[574,768,870,894]" italics="true" pageId="7" pageNumber="408">A. maechamensis</emphasis>
|
||
</taxonomicName>
|
||
in the females (
|
||
<figureCitation id="13662A42B63CFFF0FC6FFCDD040FFCBD" box="[948,1053,870,895]" captionStart="FIGURE 17" captionStartId="29.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@29.[209,1378,193,1911]" captionTargetPageId="29" captionText="FIGURE 17. Althepus phadaengensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 17D as in figure 17E." httpUri="https://zenodo.org/record/1439824/files/figure.png" pageId="7" pageNumber="408">Fig. 17A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63CFFF0FF1CFC31042DF9E9" pageId="7" pageNumber="408" type="description">
|
||
<paragraph id="8BE236C7B63CFFF0FF1CFC3104DFFAED" blockId="7.[151,1437,474,1651]" pageId="7" pageNumber="408">
|
||
<emphasis id="B929EAD5B63CFFF0FF1CFC310237FC61" bold="true" box="[199,549,906,931]" pageId="7" pageNumber="408">
|
||
Description. Male (
|
||
<typeStatus id="54E68865B63CFFF0FE71FC310205FC61" box="[426,535,906,931]" pageId="7" pageNumber="408" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 4.25; carapace 1.44 length, 1.45 width; abdomen 2.34 length, 1.28 width. Carapace round (
|
||
<figureCitation id="13662A42B63CFFF0FE73FC15021DFC05" box="[424,527,942,967]" captionStart="FIGURE 17" captionStartId="29.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@29.[209,1378,193,1911]" captionTargetPageId="29" captionText="FIGURE 17. Althepus phadaengensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 17D as in figure 17E." httpUri="https://zenodo.org/record/1439824/files/figure.png" pageId="7" pageNumber="408">Fig. 17C</figureCitation>
|
||
), yellowish, with three longitudinal brown bands of different widths, the middle one shortened to form a triangular band around the eyes. Anterior margin of thoracic region distinctly elevated. Clypeus brown. Cheliceral promargin with two teeth, followed by a lamina, retromargin with one small tooth, posterior surface of fang with 24 small denticles (
|
||
<figureCitation id="13662A42B63CFFF0FD16FBA10324FBF1" box="[717,822,1050,1075]" captionStart="FIGURE 31" captionStartId="43.[151,250,1917,1939]" captionTargetBox="[177,1409,195,1896]" captionTargetId="figure@43.[177,1410,193,1896]" captionTargetPageId="43" captionText="FIGURE 31. Left cheliceral retromargin of male holotype (A–F), female holotype (G) and female paratype (H). A, Althepus phadaengensis Li et Li sp. nov.; B, A. shanhu Li et Li sp. nov.; C, A. suayaiensis Li et Li sp. nov.; D, A. tadetuensis Li et Li sp. nov.; E, A. tanhuang Li et Li sp. nov.; F, A. thanlaensis Li et Li sp. nov.; G, A. tharnlodensis Li et Li sp. nov.; H, A. viengkeoensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439852/files/figure.png" pageId="7" pageNumber="408">Fig. 31A</figureCitation>
|
||
). Labium brown. Sternum brown. Abdomen elongate, with complex pattern of white and dark-brown dorsally and ventrally. Legs brown. Leg measurements: I 45.23 (9.74, 0.62, 11.86, 19.42, 3.59), II 29.75 (7.37, 0.59, 7.37, 12.18, 2.24), III 18.81 (5.58, 0.59, 4.95, 6.41, 1.28), IV 27.65 (8.14, 0.59, 7.56, 9.42, 1.94). Male palp (
|
||
<figureCitation id="13662A42B63CFFF0FD65FB3D0340FB5D" box="[702,850,1158,1183]" captionStart="FIGURE 16" captionStartId="28.[151,250,1917,1939]" captionTargetBox="[214,1372,206,1896]" captionTargetId="figure@28.[214,1372,193,1896]" captionTargetPageId="28" captionText="FIGURE 16. Althepus phadaengensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulbs of A. phadaengensis and A. stonei (modified from Deeleman-Reinhold, 1995), prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 16A and 16C as in figure 16D." httpUri="https://zenodo.org/record/1439822/files/figure.png" pageId="7" pageNumber="408">Figs 16A–D</figureCitation>
|
||
): tarsus distinct, extends distally,with three slightly curved, serrated bristles at the top of the cymbial protrusion (
|
||
<figureCitation id="13662A42B63CFFF0FC85FB1103DBFB01" box="[862,969,1194,1219]" captionStart="FIGURE 16" captionStartId="28.[151,250,1917,1939]" captionTargetBox="[214,1372,206,1896]" captionTargetId="figure@28.[214,1372,193,1896]" captionTargetPageId="28" captionText="FIGURE 16. Althepus phadaengensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulbs of A. phadaengensis and A. stonei (modified from Deeleman-Reinhold, 1995), prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 16A and 16C as in figure 16D." httpUri="https://zenodo.org/record/1439822/files/figure.png" pageId="7" pageNumber="408">Fig. 16A</figureCitation>
|
||
), one hooked spine with the tip directed towards proximally (
|
||
<figureCitation id="13662A42B63CFFF0FE5EFB7501FDFB25" box="[389,495,1230,1255]" captionStart="FIGURE 16" captionStartId="28.[151,250,1917,1939]" captionTargetBox="[214,1372,206,1896]" captionTargetId="figure@28.[214,1372,193,1896]" captionTargetPageId="28" captionText="FIGURE 16. Althepus phadaengensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulbs of A. phadaengensis and A. stonei (modified from Deeleman-Reinhold, 1995), prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 16A and 16C as in figure 16D." httpUri="https://zenodo.org/record/1439822/files/figure.png" pageId="7" pageNumber="408">Fig. 16D</figureCitation>
|
||
); bulb bright yellow, ovate; embolus arising retrolatero-distally from bulb, slender and slightly curved; conductor arising prolatero-distally from bulb, long, increasingly widening and curved at the distal part; embolus and conductor widely separated (distance almost equal to diameter of bulb).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63CFFF0FF1CFA810144FA21" blockId="7.[151,1437,474,1651]" pageId="7" pageNumber="408">
|
||
<emphasis id="B929EAD5B63CFFF0FF1CFA8101BFFA91" bold="true" box="[199,429,1338,1363]" pageId="7" pageNumber="408">
|
||
Female (
|
||
<typeStatus id="54E68865B63CFFF0FEF5FA8101B2FA91" box="[302,416,1338,1363]" pageId="7" pageNumber="408" type="paratype">paratype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Similar to male in color and general features (
|
||
<figureCitation id="13662A42B63CFFF0FC1BFA81045FFA91" box="[960,1101,1338,1363]" captionStart="FIGURE 17" captionStartId="29.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@29.[209,1378,193,1911]" captionTargetPageId="29" captionText="FIGURE 17. Althepus phadaengensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 17D as in figure 17E." httpUri="https://zenodo.org/record/1439824/files/figure.png" pageId="7" pageNumber="408">Figs 17D–E</figureCitation>
|
||
), but larger. Internal genitalia with two rounded spermathecae sitting on long stalks on each side (
|
||
<figureCitation id="13662A42B63CFFF0FC7FFAE50402FAB5" box="[932,1040,1374,1399]" captionStart="FIGURE 17" captionStartId="29.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@29.[209,1378,193,1911]" captionTargetPageId="29" captionText="FIGURE 17. Althepus phadaengensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 17D as in figure 17E." httpUri="https://zenodo.org/record/1439824/files/figure.png" pageId="7" pageNumber="408">Fig. 17A</figureCitation>
|
||
). Measurements: total length 4.55; carapace 1.36 length, 1.42 width; abdomen 2.81 length, 1.48 width. Leg measurements: I 41.15 (9.62, 0.56, 10.45, 17.31, 3.21), II 25.81 (6.35, 0.56, 7.69, 9.23, 1.98), III 15.93 (4.70, 0.53, 4.10, 5.32, 1.28), IV 22.70 (6.41, 0.56, 6.41, 7.37, 1.95).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63CFFF0FF1CFA55042DF9E9" blockId="7.[151,1437,474,1651]" pageId="7" pageNumber="408">
|
||
<emphasis id="B929EAD5B63CFFF0FF1CFA55012EF9C5" bold="true" box="[199,316,1518,1543]" pageId="7" pageNumber="408">Variation.</emphasis>
|
||
Males: carapace 1.30–1.60 length, 1.42–1.50 width; femur I: 9.74–12.18. Females: carapace 1.36, 1.42 length, 1.33, 1.42 width; femur I: 9.62 (n = 2; leg I lost in the other specimen).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63CFFF0FF1CF98D03B3F9B0" pageId="7" pageNumber="408" type="distribution">
|
||
<paragraph id="8BE236C7B63CFFF0FF1CF98D034EF98D" blockId="7.[151,1437,474,1651]" box="[199,860,1590,1615]" pageId="7" pageNumber="408">
|
||
<emphasis id="B929EAD5B63CFFF0FF1CF98D0172F98D" bold="true" box="[199,352,1590,1615]" pageId="7" pageNumber="408">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54E68865B63CFFF0FDBBF98C0282F98D" box="[608,656,1591,1615]" pageId="7" pageNumber="408">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13662A42B63CFFF0FD21F98D035DF98D" box="[762,847,1590,1615]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="7" pageNumber="408">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63CFFF0FF1CF9E103B3F9B0" blockId="7.[151,1437,474,1651]" box="[199,929,1626,1651]" pageId="7" pageNumber="408">
|
||
<emphasis id="B929EAD5B63CFFF0FF1CF9E10197F9B1" bold="true" box="[199,389,1626,1651]" pageId="7" pageNumber="408">Natural history.</emphasis>
|
||
Collected on the wall near the entrance of cave.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63CFFF0FF4CF97D017AF8C0" pageId="7" pageNumber="408" type="description">
|
||
<paragraph id="8BE236C7B63CFFF0FF4CF97D023AF922" blockId="7.[151,552,1734,1794]" box="[151,552,1734,1760]" pageId="7" pageNumber="408">
|
||
<taxonomicName id="4C5D4D44B63CFFF0FF4CF97D01DAF922" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[151,456,1734,1760]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="shanhu" status="sp. nov.">
|
||
<emphasis id="B929EAD5B63CFFF0FF4CF97D01DAF922" bold="true" box="[151,456,1734,1760]" pageId="7" pageNumber="408">
|
||
<emphasis id="B929EAD5B63CFFF0FF4CF97D0177F922" bold="true" box="[151,357,1734,1760]" italics="true" pageId="7" pageNumber="408">Althepus shanhu</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B63CFFF0FE54F97D01B6F922" bold="true" box="[399,420,1734,1760]" italics="true" pageId="7" pageNumber="408">et</emphasis>
|
||
Li
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB63CFFF0FE15F97C023AF922" box="[462,552,1735,1760]" pageId="7" pageNumber="408" rank="species">
|
||
<emphasis id="B929EAD5B63CFFF0FE15F97C023AF922" bold="true" box="[462,552,1735,1760]" pageId="7" pageNumber="408">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B63CFFF0FF4CF952017AF8C0" blockId="7.[151,552,1734,1794]" box="[151,360,1769,1794]" pageId="7" pageNumber="408">
|
||
<figureCitation id="13662A42B63CFFF0FF4CF9520104F8C0" box="[151,278,1769,1794]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStartId-0="30.[151,250,1933,1955]" captionStartId-1="31.[151,250,1933,1955]" captionTargetBox-0="[228,1378,193,1911]" captionTargetBox-1="[209,1378,193,1911]" captionTargetId-0="figure@30.[209,1378,193,1911]" captionTargetId-1="figure@31.[209,1378,193,1911]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionText-0="FIGURE 18. Althepus shanhu Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 18A and 18C as in figure 18D." captionText-1="FIGURE 19. Althepus shanhu Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 19D as in figure 19E." httpUri-0="https://zenodo.org/record/1439826/files/figure.png" httpUri-1="https://zenodo.org/record/1439828/files/figure.png" pageId="7" pageNumber="408">Figs 18–19</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B63CFFF0FEF9F951017AF8C0" box="[290,360,1770,1794]" captionStart-0="FIGURE 31" captionStart-1="FIGURE 32" captionStartId-0="43.[151,250,1917,1939]" captionStartId-1="44.[151,250,1148,1170]" captionTargetBox-0="[177,1409,195,1896]" captionTargetBox-1="[393,1194,195,1124]" captionTargetId-0="figure@43.[177,1410,193,1896]" captionTargetId-1="figure@44.[393,1194,195,1124]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionText-0="FIGURE 31. Left cheliceral retromargin of male holotype (A–F), female holotype (G) and female paratype (H). A, Althepus phadaengensis Li et Li sp. nov.; B, A. shanhu Li et Li sp. nov.; C, A. suayaiensis Li et Li sp. nov.; D, A. tadetuensis Li et Li sp. nov.; E, A. tanhuang Li et Li sp. nov.; F, A. thanlaensis Li et Li sp. nov.; G, A. tharnlodensis Li et Li sp. nov.; H, A. viengkeoensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." captionText-1="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri-0="https://zenodo.org/record/1439852/files/figure.png" httpUri-1="https://zenodo.org/record/1439854/files/figure.png" pageId="7" pageNumber="408">31–32</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63CFFF0FF4CF88A0330F850" pageId="7" pageNumber="408" type="materials_examined">
|
||
<paragraph id="8BE236C7B63CFFF0FF4CF88A0330F850" blockId="7.[151,1436,1841,2010]" pageId="7" pageNumber="408">
|
||
<materialsCitation id="3B353C9AB63CFFF0FF4CF88A047AF8AC" ID-GBIF-Occurrence="1913888983" collectingDate="2016-12-17" country="Myanmar" county="Putao County" elevation="2252" latitude="27.609167" location="Hponkanrazi Wildlife Sanctuary Roadside" longLatPrecision="1" longitude="96.980835" pageId="7" pageNumber="408" specimenCount="2" stateProvince="Kachin" typeStatus="holotype">
|
||
<emphasis id="B929EAD5B63CFFF0FF4CF88A01D0F888" bold="true" box="[151,450,1841,1866]" pageId="7" pageNumber="408">
|
||
<typeStatus id="54E68865B63CFFF0FF4CF88A00C0F888" box="[151,210,1841,1866]" pageId="7" pageNumber="408">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68865B63CFFF0FE97F88A01AFF888" box="[332,445,1841,1866]" pageId="7" pageNumber="408" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male (IZCAS-Ar 34015),
|
||
<collectingCountry id="F34A7657B63CFFF0FD37F8890349F888" box="[748,859,1842,1866]" name="Myanmar" pageId="7" pageNumber="408">Myanmar</collectingCountry>
|
||
,
|
||
<collectingRegion id="4999F825B63CFFF0FCBEF88A03A4F888" box="[869,950,1841,1866]" country="Myanmar" name="Kachin" pageId="7" pageNumber="408">Kachin</collectingRegion>
|
||
State,
|
||
<collectingCounty id="62834E4BB63CFFF0FBD8F889048FF888" box="[1027,1181,1842,1866]" pageId="7" pageNumber="408">Putao County</collectingCounty>
|
||
,
|
||
<location id="8E82601CB63CFFF0FB73F88A016EF8AC" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB63CFFF0FB73F88A016EF8AC" country="Myanmar" county="Putao County" latitude="27.609167" longLatPrecision="1" longitude="96.980835" name="Hponkanrazi Wildlife Sanctuary Roadside" pageId="7" pageNumber="408" stateProvince="Kachin">Hponkanrazi Wildlife Sanctuary Roadside</location>
|
||
,
|
||
<geoCoordinate id="EE695000B63CFFF0FE50F8EE020BF8AC" box="[395,537,1877,1902]" direction="north" orientation="latitude" pageId="7" pageNumber="408" precision="1" value="27.609167">27°36.550'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695000B63CFFF0FDFCF8EE02A0F8AC" box="[551,690,1877,1902]" direction="east" orientation="longitude" pageId="7" pageNumber="408" precision="1" value="96.980835">96°58.850'E</geoCoordinate>
|
||
,
|
||
<elevation id="0070D1F4B63CFFF0FD1AF8EE030BF8AF" box="[705,793,1877,1902]" metricMagnitude="3" metricUnit="m" metricValue="2.252" pageId="7" pageNumber="408" unit="m" value="2252.0">
|
||
<quantity id="4CA59B22B63CFFF0FD1AF8EE030BF8AF" box="[705,793,1877,1902]" metricMagnitude="3" metricUnit="m" metricValue="2.252" pageId="7" pageNumber="408" unit="m" value="2252.0">2252 m</quantity>
|
||
</elevation>
|
||
altitude,
|
||
<collectingDate id="EFA7E9EFB63CFFF0FC52F8ED047AF8AC" box="[905,1128,1877,1902]" pageId="7" pageNumber="408" value="2016-12-17">
|
||
<date id="FFE31007B63CFFF0FC52F8ED047AF8AC" box="[905,1128,1877,1902]" pageId="7" pageNumber="408" value="2016-12-17">17 December 2016</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B353C9AB63CFFF0FBADF8EE0330F850" ID-GBIF-Occurrence="1913888988" collectingDate="2016-12-17" country="Myanmar" county="Putao County" elevation="2252" latitude="27.609167" location="Hponkanrazi Wildlife Sanctuary Roadside" longLatPrecision="1" longitude="96.980835" pageId="7" pageNumber="408" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Kachin" typeStatus="paratype">
|
||
<emphasis id="B929EAD5B63CFFF0FBADF8EE04E5F8AC" bold="true" box="[1142,1271,1877,1902]" pageId="7" pageNumber="408">
|
||
<typeStatus id="54E68865B63CFFF0FBADF8EE04E0F8AC" box="[1142,1266,1877,1902]" pageId="7" pageNumber="408" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D5BFD4EB63CFFF0FAD9F8ED0542F8AC" box="[1282,1360,1877,1902]" pageId="7" pageNumber="408" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="9D5BFD4EB63CFFF0FA55F8ED00F4F850" pageId="7" pageNumber="408" type="female">1 female</specimenCount>
|
||
, same data as holotype (IZCAS-Ar 34016–34017).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB63CFFF0FF1CF8260360F818" pageId="7" pageNumber="408" type="etymology">
|
||
<paragraph id="8BE236C7B63CFFF0FF1CF8260360F818" blockId="7.[151,1436,1841,2010]" pageId="7" pageNumber="408">
|
||
<emphasis id="B929EAD5B63CFFF0FF1CF826015CF874" bold="true" box="[199,334,1949,1974]" pageId="7" pageNumber="408">Etymology.</emphasis>
|
||
The specific name is derived from the Chinese pinyin 'shān hú', which means 'coral', referring to the female’s spermathecae looking like coral; noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB633FFFFFF1CFF2C037AFF3A" pageId="8" pageNumber="409" type="diagnosis">
|
||
<paragraph id="8BE236C7B633FFFFFF1CFF2C037AFF3A" blockId="8.[151,1437,151,933]" pageId="8" pageNumber="409">
|
||
<emphasis id="B929EAD5B633FFFFFF1CFF2C0153FF72" bold="true" box="[199,321,151,176]" pageId="8" pageNumber="409">Diagnosis.</emphasis>
|
||
This new species can be easily distinguished from all other species by many long bristles of the palpal tibia (
|
||
<figureCitation id="13662A42B633FFFFFEF9FF0701BDFF16" box="[290,431,188,213]" captionStart="FIGURE 18" captionStartId="30.[151,250,1933,1955]" captionTargetBox="[228,1378,193,1911]" captionTargetId="figure@30.[209,1378,193,1911]" captionTargetPageId="30" captionText="FIGURE 18. Althepus shanhu Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 18A and 18C as in figure 18D." httpUri="https://zenodo.org/record/1439826/files/figure.png" pageId="8" pageNumber="409">Figs 18C–D</figureCitation>
|
||
), the petaloid structures on the distal conductor as well as the short embolus in males (
|
||
<figureCitation id="13662A42B633FFFFFAB4FF0700DAFF3A" captionStart="FIGURE 18" captionStartId="30.[151,250,1933,1955]" captionTargetBox="[228,1378,193,1911]" captionTargetId="figure@30.[209,1378,193,1911]" captionTargetPageId="30" captionText="FIGURE 18. Althepus shanhu Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 18A and 18C as in figure 18D." httpUri="https://zenodo.org/record/1439826/files/figure.png" pageId="8" pageNumber="409">Fig. 18B</figureCitation>
|
||
), and by many pairs of spermathecae in females (
|
||
<figureCitation id="13662A42B633FFFFFD28FF640349FF3A" box="[755,859,223,248]" captionStart="FIGURE 19" captionStartId="31.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@31.[209,1378,193,1911]" captionTargetPageId="31" captionText="FIGURE 19. Althepus shanhu Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 19D as in figure 19E." httpUri="https://zenodo.org/record/1439828/files/figure.png" pageId="8" pageNumber="409">Fig. 19A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB633FFFFFF1CFEBF04D0FC9F" pageId="8" pageNumber="409" type="description">
|
||
<paragraph id="8BE236C7B633FFFFFF1CFEBF0173FD6A" blockId="8.[151,1437,151,933]" pageId="8" pageNumber="409">
|
||
<emphasis id="B929EAD5B633FFFFFF1CFEBF0237FEDF" bold="true" box="[199,549,260,285]" pageId="8" pageNumber="409">
|
||
Description. Male (
|
||
<typeStatus id="54E68865B633FFFFFE71FEBF0205FEDF" box="[426,535,260,285]" pageId="8" pageNumber="409" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 4.00; carapace 1.48 length, 1.48 width; abdomen 2.28 length, 1.42 width. Carapace round (
|
||
<figureCitation id="13662A42B633FFFFFE74FE9C020AFE82" box="[431,536,295,320]" captionStart="FIGURE 19" captionStartId="31.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@31.[209,1378,193,1911]" captionTargetPageId="31" captionText="FIGURE 19. Althepus shanhu Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 19D as in figure 19E." httpUri="https://zenodo.org/record/1439828/files/figure.png" pageId="8" pageNumber="409">Fig. 19C</figureCitation>
|
||
), yellow, with three longitudinal brown bands of different widths, the middle wider than the others. Anterior margin of thoracic region distinctly elevated. Clypeus brown. Cheliceral promargin with two teeth, followed by a lamina, retromargin with two small teeth, posterior surface of fang provided with 26 small denticles (
|
||
<figureCitation id="13662A42B633FFFFFE8EFE2F01ACFE6E" box="[341,446,404,429]" captionStart="FIGURE 31" captionStartId="43.[151,250,1917,1939]" captionTargetBox="[177,1409,195,1896]" captionTargetId="figure@43.[177,1410,193,1896]" captionTargetPageId="43" captionText="FIGURE 31. Left cheliceral retromargin of male holotype (A–F), female holotype (G) and female paratype (H). A, Althepus phadaengensis Li et Li sp. nov.; B, A. shanhu Li et Li sp. nov.; C, A. suayaiensis Li et Li sp. nov.; D, A. tadetuensis Li et Li sp. nov.; E, A. tanhuang Li et Li sp. nov.; F, A. thanlaensis Li et Li sp. nov.; G, A. tharnlodensis Li et Li sp. nov.; H, A. viengkeoensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439852/files/figure.png" pageId="8" pageNumber="409">Fig. 31B</figureCitation>
|
||
). Labium brown. Sternum yellow, with many brown bands. Abdomen elongate, with complex patterns of yellow and brown dorsally and ventrally. Legs brown. Leg measurements: I missing, II 17.45 (4.80, 0.53, 4.75, 6.28, 1.09), III 12.17 (3.60, 0.53, 3.13, 3.80, 1.11), IV 18.20 (5.19, 0.51, 5.19, 6.03, 1.28). Male palp (
|
||
<figureCitation id="13662A42B633FFFFFF0CFE44017AFDDA" box="[215,360,511,536]" captionStart="FIGURE 18" captionStartId="30.[151,250,1933,1955]" captionTargetBox="[228,1378,193,1911]" captionTargetId="figure@30.[209,1378,193,1911]" captionTargetPageId="30" captionText="FIGURE 18. Althepus shanhu Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 18A and 18C as in figure 18D." httpUri="https://zenodo.org/record/1439826/files/figure.png" pageId="8" pageNumber="409">Figs 18A–D</figureCitation>
|
||
): tarsus with three slightly curved, serrated bristles at the top of the cymbial protrusion (one of them missing,
|
||
<figureCitation id="13662A42B633FFFFFE99FD9F01BEFDFE" box="[322,428,548,573]" captionStart="FIGURE 18" captionStartId="30.[151,250,1933,1955]" captionTargetBox="[228,1378,193,1911]" captionTargetId="figure@30.[209,1378,193,1911]" captionTargetPageId="30" captionText="FIGURE 18. Althepus shanhu Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 18A and 18C as in figure 18D." httpUri="https://zenodo.org/record/1439826/files/figure.png" pageId="8" pageNumber="409">Fig. 18A</figureCitation>
|
||
), one hooked spine with tip directed proximally (
|
||
<figureCitation id="13662A42B633FFFFFC2BFD9F0448FDFE" box="[1008,1114,548,573]" captionStart="FIGURE 18" captionStartId="30.[151,250,1933,1955]" captionTargetBox="[228,1378,193,1911]" captionTargetId="figure@30.[209,1378,193,1911]" captionTargetPageId="30" captionText="FIGURE 18. Althepus shanhu Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 18A and 18C as in figure 18D." httpUri="https://zenodo.org/record/1439826/files/figure.png" pageId="8" pageNumber="409">Fig. 18C</figureCitation>
|
||
); bulb bright yellow, ovate; embolus arising retrolatero-distally from bulb, short and straight (
|
||
<figureCitation id="13662A42B633FFFFFC5BFDFC03F5FDA2" box="[896,999,583,608]" captionStart="FIGURE 18" captionStartId="30.[151,250,1933,1955]" captionTargetBox="[228,1378,193,1911]" captionTargetId="figure@30.[209,1378,193,1911]" captionTargetPageId="30" captionText="FIGURE 18. Althepus shanhu Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 18A and 18C as in figure 18D." httpUri="https://zenodo.org/record/1439826/files/figure.png" pageId="8" pageNumber="409">Fig. 18B</figureCitation>
|
||
); conductor arising distally from bulb, wide, and some petaloid laminas arising distally; embolus and conductor widely separated (distance less than diameter of bulb).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B633FFFFFF1CFD0F0494FCFA" blockId="8.[151,1437,151,933]" pageId="8" pageNumber="409">
|
||
<emphasis id="B929EAD5B633FFFFFF1CFD0F01BFFD0F" bold="true" box="[199,429,692,717]" pageId="8" pageNumber="409">
|
||
Female (
|
||
<typeStatus id="54E68865B633FFFFFEF5FD0F01B2FD0F" box="[302,416,692,717]" pageId="8" pageNumber="409" type="paratype">paratype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Similar to male in color and general features (
|
||
<figureCitation id="13662A42B633FFFFFC1EFD0F0442FD0E" box="[965,1104,692,717]" captionStart="FIGURE 19" captionStartId="31.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@31.[209,1378,193,1911]" captionTargetPageId="31" captionText="FIGURE 19. Althepus shanhu Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 19D as in figure 19E." httpUri="https://zenodo.org/record/1439828/files/figure.png" pageId="8" pageNumber="409">Figs 19D–E</figureCitation>
|
||
) but larger. Internal genitalia with many pairs of spermathecae, short, distal part swollen (
|
||
<figureCitation id="13662A42B633FFFFFC9AFD6C03B8FD32" box="[833,938,727,752]" captionStart="FIGURE 19" captionStartId="31.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@31.[209,1378,193,1911]" captionTargetPageId="31" captionText="FIGURE 19. Althepus shanhu Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 19D as in figure 19E." httpUri="https://zenodo.org/record/1439828/files/figure.png" pageId="8" pageNumber="409">Fig. 19A</figureCitation>
|
||
). Measurements: total length 4.10; carapace 1.24 length, 1.41 width; abdomen 2.81 length, 2.05 width. Leg measurements: I missing, II 13.50 (3.60, 0.56, 3.68, 4.45, 1.21), III 10.31 (2.94, 0.54, 2.66, 3.00, 1.17), IV 14.57 (4.30, 0.56, 3.96, 4.35, 1.40).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B633FFFFFF1CFCFF04D0FC9F" blockId="8.[151,1437,151,933]" box="[199,1218,836,861]" pageId="8" pageNumber="409">
|
||
<emphasis id="B929EAD5B633FFFFFF1CFCFF012EFC9F" bold="true" box="[199,316,836,861]" pageId="8" pageNumber="409">Variation.</emphasis>
|
||
Males: carapace 1.35, 1.48 length, 1.42, 1.48 width (leg I lost in all specimens).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB633FFFFFF1CFCDC0418FC66" pageId="8" pageNumber="409" type="distribution">
|
||
<paragraph id="8BE236C7B633FFFFFF1CFCDC034EFC42" blockId="8.[151,1437,151,933]" box="[199,860,871,896]" pageId="8" pageNumber="409">
|
||
<emphasis id="B929EAD5B633FFFFFF1CFCDC0172FC42" bold="true" box="[199,352,871,896]" pageId="8" pageNumber="409">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54E68865B633FFFFFDBBFCD30282FC42" box="[608,656,872,896]" pageId="8" pageNumber="409">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13662A42B633FFFFFD21FCDC035DFC42" box="[762,847,871,896]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="8" pageNumber="409">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B633FFFFFF1CFC370418FC66" blockId="8.[151,1437,151,933]" box="[199,1034,908,933]" pageId="8" pageNumber="409">
|
||
<bibRef id="52FDA231B633FFFFFF1CFC370418FC66" box="[199,1034,908,933]" pageId="8" pageNumber="409" refId="ref9478" type="journal volume">
|
||
<emphasis id="B929EAD5B633FFFFFF1CFC370197FC67" bold="true" box="[199,389,908,933]" pageId="8" pageNumber="409">
|
||
<author id="53E0DE22B633FFFFFF1CFC370136FC67" box="[199,292,908,933]" firstAuthorName="Natural" fns="*" inm="true" npo="*" pageId="8" pageNumber="409">Natural</author>
|
||
<title id="F959DDF1B633FFFFFEF0FC37016DFC67" box="[299,383,908,933]" pageId="8" pageNumber="409">history</title>
|
||
.
|
||
</emphasis>
|
||
<journalOrPublisher id="519C3C0BB633FFFFFE57FC3703B4FC67" box="[396,934,908,933]" pageId="8" pageNumber="409">Collected among tree buttresses at an altitude of</journalOrPublisher>
|
||
<quantity id="4CA59B22B633FFFFFC75FC370414FC66" box="[942,1030,908,933]" metricMagnitude="3" metricUnit="m" metricValue="2.252" pageId="8" pageNumber="409" unit="m" value="2252.0">
|
||
<part id="FFF99C5AB633FFFFFC75FC3703F4FC67" box="[942,998,908,933]" pageId="8" pageNumber="409" type="volume">2252</part>
|
||
m
|
||
</quantity>
|
||
.
|
||
</bibRef>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB633FFFFFF4CFC4C012DFBF1" pageId="8" pageNumber="409" type="description">
|
||
<paragraph id="8BE236C7B633FFFFFF4CFC4C0245FBD3" blockId="8.[151,599,1015,1075]" box="[151,599,1015,1042]" pageId="8" pageNumber="409">
|
||
<taxonomicName id="4C5D4D44B633FFFFFF4CFC4C01E5FBD0" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[151,503,1015,1042]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="suayaiensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B633FFFFFF4CFC4C01E5FBD0" bold="true" box="[151,503,1015,1042]" pageId="8" pageNumber="409">
|
||
<emphasis id="B929EAD5B633FFFFFF4CFC4C0186FBD3" bold="true" box="[151,404,1015,1041]" italics="true" pageId="8" pageNumber="409">Althepus suayaiensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B633FFFFFE65FC4C01C1FBD3" bold="true" box="[446,467,1015,1041]" italics="true" pageId="8" pageNumber="409">et</emphasis>
|
||
Li
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB633FFFFFE25FC430245FBD3" box="[510,599,1016,1041]" pageId="8" pageNumber="409" rank="species">
|
||
<emphasis id="B929EAD5B633FFFFFE25FC430245FBD3" bold="true" box="[510,599,1016,1041]" pageId="8" pageNumber="409">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B633FFFFFF4CFBA1012DFBF1" blockId="8.[151,599,1015,1075]" box="[151,319,1050,1075]" pageId="8" pageNumber="409">
|
||
<figureCitation id="13662A42B633FFFFFF4CFBA100FEFBF1" box="[151,236,1050,1075]" captionStart="FIGURE 20" captionStartId="32.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@32.[209,1378,193,1911]" captionTargetPageId="32" captionText="FIGURE 20. Althepus suayaiensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 20A and 20C as in figure 20D." httpUri="https://zenodo.org/record/1439830/files/figure.png" pageId="8" pageNumber="409">Figs 20</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B633FFFFFF23FBA0012DFBF1" box="[248,319,1051,1075]" captionStart-0="FIGURE 31" captionStart-1="FIGURE 32" captionStartId-0="43.[151,250,1917,1939]" captionStartId-1="44.[151,250,1148,1170]" captionTargetBox-0="[177,1409,195,1896]" captionTargetBox-1="[393,1194,195,1124]" captionTargetId-0="figure@43.[177,1410,193,1896]" captionTargetId-1="figure@44.[393,1194,195,1124]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionText-0="FIGURE 31. Left cheliceral retromargin of male holotype (A–F), female holotype (G) and female paratype (H). A, Althepus phadaengensis Li et Li sp. nov.; B, A. shanhu Li et Li sp. nov.; C, A. suayaiensis Li et Li sp. nov.; D, A. tadetuensis Li et Li sp. nov.; E, A. tanhuang Li et Li sp. nov.; F, A. thanlaensis Li et Li sp. nov.; G, A. tharnlodensis Li et Li sp. nov.; H, A. viengkeoensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." captionText-1="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri-0="https://zenodo.org/record/1439852/files/figure.png" httpUri-1="https://zenodo.org/record/1439854/files/figure.png" pageId="8" pageNumber="409">31–32</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB633FFFFFF4CFBD90305FB01" pageId="8" pageNumber="409" type="materials_examined">
|
||
<paragraph id="8BE236C7B633FFFFFF4CFBD90305FB01" blockId="8.[151,1437,1122,1939]" pageId="8" pageNumber="409">
|
||
<bibRef id="52FDA231B633FFFFFF4CFBD90305FB01" pageId="8" pageNumber="409" refId="ref9507" type="book chapter">
|
||
<materialsCitation id="3B353C9AB633FFFFFF4CFBD904E7FB5D" ID-GBIF-Occurrence="1913888975" collectingDate="2016-11-14" country="Thailand" county="Phop Phra District" elevation="466" latitude="16.672266" location="Tham Sua Yai Cave" longLatPrecision="1" longitude="98.66897" municipality="Mae Ku Subdistrict" pageId="8" pageNumber="409" specimenCount="2" stateProvince="Tak Province" typeStatus="holotype">
|
||
<emphasis id="B929EAD5B633FFFFFF4CFBD901C3FBB9" bold="true" box="[151,465,1122,1147]" pageId="8" pageNumber="409">
|
||
<typeStatus id="54E68865B633FFFFFF4CFBD900C0FBB9" box="[151,210,1122,1147]" pageId="8" pageNumber="409">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68865B633FFFFFE80FBD901DEFBB9" box="[347,460,1122,1147]" pageId="8" pageNumber="409" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male (IZCAS-Ar 34018),
|
||
<author id="53E0DE22B633FFFFFCC1FBD90420FBB9" box="[794,1074,1122,1147]" case="TC" fn="Tak Province" fnc="TC" fns="N" ln="Thailand" lnc="TC" npo="LnFn" pageId="8" pageNumber="409">
|
||
<collectingCountry id="F34A7657B633FFFFFCC1FBD90392FBB9" box="[794,896,1122,1147]" name="Thailand" pageId="8" pageNumber="409">Thailand</collectingCountry>
|
||
,
|
||
<collectingRegion id="4999F825B633FFFFFC48FBD90420FBB9" box="[915,1074,1122,1147]" country="Thailand" name="Tak" pageId="8" pageNumber="409">Tak Province</collectingRegion>
|
||
</author>
|
||
,
|
||
<author id="53E0DE22B633FFFFFB9EFBD90101FB5D" case="TC" fn="Mae Ku Subdistrict" fnc="TC" fns="N" ln="Phop Phra District" lnc="TC" npo="LnFn" pageId="8" pageNumber="409">
|
||
<collectingCounty id="62834E4BB633FFFFFB9EFBD90534FBB9" box="[1093,1318,1122,1147]" pageId="8" pageNumber="409">Phop Phra District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6B86ACBDB633FFFFFAE2FBD80101FB5D" pageId="8" pageNumber="409">Mae Ku Subdistrict</collectingMunicipality>
|
||
</author>
|
||
,
|
||
<location id="8E82601CB633FFFFFEF9FB3D021CFB5D" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB633FFFFFEF9FB3D021CFB5D" box="[290,526,1158,1183]" country="Thailand" county="Phop Phra District" latitude="16.672266" longLatPrecision="1" longitude="98.66897" municipality="Mae Ku Subdistrict" name="Tham Sua Yai Cave" pageId="8" pageNumber="409" stateProvince="Tak Province">
|
||
<author id="53E0DE22B633FFFFFEF9FB3D021CFB5D" box="[290,526,1158,1183]" case="TC" fn="Tham Sua Yai" fnc="TC" fns="N" ln="Cave" lnc="TC" npo="FnLn" pageId="8" pageNumber="409">Tham Sua Yai Cave</author>
|
||
</location>
|
||
,
|
||
<title id="F959DDF1B633FFFFFDC6FB3D0416FB5D" box="[541,1028,1158,1183]" jopScore="0.0017673048" pageId="8" pageNumber="409" type="_" typeScore="26">
|
||
<geoCoordinate id="EE695000B633FFFFFDC6FB3D02BFFB5D" box="[541,685,1158,1183]" direction="north" orientation="latitude" pageId="8" pageNumber="409" precision="1" value="16.672266">16°40.336'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695000B633FFFFFD67FB3C035BFB5D" box="[700,841,1159,1183]" direction="east" orientation="longitude" pageId="8" pageNumber="409" precision="1" value="98.66897">98°40.138'E</geoCoordinate>
|
||
,
|
||
<elevation id="0070D1F4B633FFFFFC83FB3D03B0FB5C" box="[856,930,1158,1183]" metricMagnitude="2" metricUnit="m" metricValue="4.66" pageId="8" pageNumber="409" unit="m" value="466.0">
|
||
<quantity id="4CA59B22B633FFFFFC83FB3D03B0FB5C" box="[856,930,1158,1183]" metricMagnitude="2" metricUnit="m" metricValue="4.66" pageId="8" pageNumber="409" unit="m" value="466.0">466 m</quantity>
|
||
</elevation>
|
||
altitude
|
||
</title>
|
||
,
|
||
<collectingDate id="EFA7E9EFB633FFFFFBC9FB3C04E7FB5D" box="[1042,1269,1158,1183]" pageId="8" pageNumber="409" value="2016-11-14">14 November 2016</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B353C9AB633FFFFFAD9FB3D0305FB01" ID-GBIF-Occurrence="1913888986" collectingDate="2016-11-14" country="Thailand" county="Phop Phra District" elevation="466" latitude="16.672266" location="Tham Sua Yai Cave" longLatPrecision="1" longitude="98.66897" municipality="Mae Ku Subdistrict" pageId="8" pageNumber="409" specimenCount="2" specimenCount-male="2" stateProvince="Tak Province" typeStatus="paratype">
|
||
<journalOrPublisher id="519C3C0BB633FFFFFAD9FB3D0277FB01" jopScore="0.008394699" pageId="8" pageNumber="409">
|
||
<emphasis id="B929EAD5B633FFFFFAD9FB3D0596FB5D" bold="true" box="[1282,1412,1158,1183]" pageId="8" pageNumber="409">
|
||
<typeStatus id="54E68865B633FFFFFAD9FB3D056DFB5D" box="[1282,1407,1158,1183]" pageId="8" pageNumber="409" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D5BFD4EB633FFFFFA56FB3C00C9FB01" pageId="8" pageNumber="409" type="male">2 males</specimenCount>
|
||
, same data as holotype (IZCAS-Ar
|
||
</journalOrPublisher>
|
||
<pagination id="B294C833B633FFFFFDB7FB10031EFB01" box="[620,780,1195,1219]" pageId="8" pageNumber="409" type="pagination">34019–34020</pagination>
|
||
).
|
||
</materialsCitation>
|
||
</bibRef>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB633FFFFFF1CFB75040FFB25" box="[199,1053,1230,1255]" pageId="8" pageNumber="409" type="etymology">
|
||
<paragraph id="8BE236C7B633FFFFFF1CFB75040FFB25" blockId="8.[151,1437,1122,1939]" box="[199,1053,1230,1255]" pageId="8" pageNumber="409">
|
||
<emphasis id="B929EAD5B633FFFFFF1CFB75015CFB25" bold="true" box="[199,334,1230,1255]" pageId="8" pageNumber="409">Etymology.</emphasis>
|
||
The specific name is an adjective and refers to the
|
||
<typeStatus id="54E68865B633FFFFFC57FB7403AEFB25" box="[908,956,1231,1255]" pageId="8" pageNumber="409">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB633FFFFFF1CFB49058FFAED" pageId="8" pageNumber="409" type="diagnosis">
|
||
<paragraph id="8BE236C7B633FFFFFF1CFB49058FFAED" blockId="8.[151,1437,1122,1939]" pageId="8" pageNumber="409">
|
||
<emphasis id="B929EAD5B633FFFFFF1CFB490153FAC9" bold="true" box="[199,321,1266,1291]" pageId="8" pageNumber="409">Diagnosis.</emphasis>
|
||
This new species can be easily distinguished from all other species by the irregularly serrate margin of the distal conductor and a small triangular protrusion on the middle part of the conductor in males (
|
||
<figureCitation id="13662A42B633FFFFFAD8FAAD0582FAED" box="[1283,1424,1302,1327]" captionStart="FIGURE 20" captionStartId="32.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@32.[209,1378,193,1911]" captionTargetPageId="32" captionText="FIGURE 20. Althepus suayaiensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 20A and 20C as in figure 20D." httpUri="https://zenodo.org/record/1439830/files/figure.png" pageId="8" pageNumber="409">Figs 20A–B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB633FFFFFF1CFA810471F8E5" pageId="8" pageNumber="409" type="description">
|
||
<paragraph id="8BE236C7B633FFFFFF1CFA810173F91D" blockId="8.[151,1437,1122,1939]" pageId="8" pageNumber="409">
|
||
<emphasis id="B929EAD5B633FFFFFF1CFA810237FA91" bold="true" box="[199,549,1338,1363]" pageId="8" pageNumber="409">
|
||
Description. Male (
|
||
<typeStatus id="54E68865B633FFFFFE71FA810205FA91" box="[426,535,1338,1363]" pageId="8" pageNumber="409" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 3.95; carapace 1.41 length, 1.36 width; abdomen 2.20 length, 1.13 width. Carapace round, light yellow, with three longitudinal brown bands of different widths, the middle one wider than the others. Clypeus brown. Cheliceral promargin with two teeth, followed by a lamina, retromargin with two small teeth, posterior surface of fang with 19 small denticles (
|
||
<figureCitation id="13662A42B633FFFFFC97FA1D03A1FA7D" box="[844,947,1446,1471]" captionStart="FIGURE 31" captionStartId="43.[151,250,1917,1939]" captionTargetBox="[177,1409,195,1896]" captionTargetId="figure@43.[177,1410,193,1896]" captionTargetPageId="43" captionText="FIGURE 31. Left cheliceral retromargin of male holotype (A–F), female holotype (G) and female paratype (H). A, Althepus phadaengensis Li et Li sp. nov.; B, A. shanhu Li et Li sp. nov.; C, A. suayaiensis Li et Li sp. nov.; D, A. tadetuensis Li et Li sp. nov.; E, A. tanhuang Li et Li sp. nov.; F, A. thanlaensis Li et Li sp. nov.; G, A. tharnlodensis Li et Li sp. nov.; H, A. viengkeoensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439852/files/figure.png" pageId="8" pageNumber="409">Fig. 31C</figureCitation>
|
||
). Labium brown. Sternum yellow, with two irregular brown lines on each side. Abdomen elongate, with complex patterns of white and dark-brown dorsally and ventrally. Legs brown. Leg measurements: I 33.12 (7.69, 0.56 8.34, 14.29, 2.24), II 21.86 (5.90, 0.55, 5.64, 8.14, 1.63), III 19.93 (4.00, 0.52, 5.64, 8.14, 1.63), IV missing. Male palp (
|
||
<figureCitation id="13662A42B633FFFFFC38F9A90466F9E9" box="[995,1140,1554,1579]" captionStart="FIGURE 20" captionStartId="32.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@32.[209,1378,193,1911]" captionTargetPageId="32" captionText="FIGURE 20. Althepus suayaiensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 20A and 20C as in figure 20D." httpUri="https://zenodo.org/record/1439830/files/figure.png" pageId="8" pageNumber="409">Figs 20A–D</figureCitation>
|
||
): tarsus with three slightly curved, serrated bristles at the top of the cymbial protrusion (two of them were missing,
|
||
<figureCitation id="13662A42B633FFFFFB42F98D0510F98D" box="[1177,1282,1590,1615]" captionStart="FIGURE 20" captionStartId="32.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@32.[209,1378,193,1911]" captionTargetPageId="32" captionText="FIGURE 20. Althepus suayaiensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 20A and 20C as in figure 20D." httpUri="https://zenodo.org/record/1439830/files/figure.png" pageId="8" pageNumber="409">Fig. 20C</figureCitation>
|
||
), one hooked spine with tip directed proximally (
|
||
<figureCitation id="13662A42B633FFFFFDFBF9E1029AF9B1" box="[544,648,1626,1651]" captionStart="FIGURE 20" captionStartId="32.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@32.[209,1378,193,1911]" captionTargetPageId="32" captionText="FIGURE 20. Althepus suayaiensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 20A and 20C as in figure 20D." httpUri="https://zenodo.org/record/1439830/files/figure.png" pageId="8" pageNumber="409">Fig. 20A</figureCitation>
|
||
); bulb yellow, ovate; embolus arising distally from bulb, proximal part distinctly wide and slightly curved (
|
||
<figureCitation id="13662A42B633FFFFFDEDF9C502B3F955" box="[566,673,1662,1687]" captionStart="FIGURE 20" captionStartId="32.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@32.[209,1378,193,1911]" captionTargetPageId="32" captionText="FIGURE 20. Althepus suayaiensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 20A and 20C as in figure 20D." httpUri="https://zenodo.org/record/1439830/files/figure.png" pageId="8" pageNumber="409">Fig. 20A</figureCitation>
|
||
); conductor arising distally from bulb, distinctly wide, middle and distal part with some small protrusions (
|
||
<figureCitation id="13662A42B633FFFFFD83F91902F7F979" box="[600,741,1698,1723]" captionStart="FIGURE 20" captionStartId="32.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@32.[209,1378,193,1911]" captionTargetPageId="32" captionText="FIGURE 20. Althepus suayaiensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 20A and 20C as in figure 20D." httpUri="https://zenodo.org/record/1439830/files/figure.png" pageId="8" pageNumber="409">Figs 20A–B</figureCitation>
|
||
); embolus and conductor widely separated (distance less than diameter of bulb).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B633FFFFFF1CF951018FF8C1" blockId="8.[151,1437,1122,1939]" box="[199,413,1770,1795]" pageId="8" pageNumber="409">
|
||
<emphasis id="B929EAD5B633FFFFFF1CF9510135F8C1" bold="true" box="[199,295,1770,1795]" pageId="8" pageNumber="409">Female:</emphasis>
|
||
Unknown
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B633FFFFFF1CF8B50471F8E5" blockId="8.[151,1437,1122,1939]" box="[199,1123,1806,1831]" pageId="8" pageNumber="409">
|
||
<emphasis id="B929EAD5B633FFFFFF1CF8B5012EF8E5" bold="true" box="[199,316,1806,1831]" pageId="8" pageNumber="409">Variation.</emphasis>
|
||
Males: carapace 1.13–1.41 length, 1.17–1.36 width, femur I 7.69–8.21.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB633FFFFFF1CF88900CCF851" pageId="8" pageNumber="409" type="distribution">
|
||
<paragraph id="8BE236C7B633FFFFFF1CF889034EF889" blockId="8.[151,1437,1122,1939]" box="[199,860,1842,1867]" pageId="8" pageNumber="409">
|
||
<emphasis id="B929EAD5B633FFFFFF1CF8890172F889" bold="true" box="[199,352,1842,1867]" pageId="8" pageNumber="409">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54E68865B633FFFFFDBBF8880282F889" box="[608,656,1843,1867]" pageId="8" pageNumber="409">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13662A42B633FFFFFD21F889035DF889" box="[762,847,1842,1867]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="8" pageNumber="409">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B633FFFFFF1CF8ED00CCF851" blockId="8.[151,1437,1122,1939]" pageId="8" pageNumber="409">
|
||
<emphasis id="B929EAD5B633FFFFFF1CF8ED0197F8AD" bold="true" box="[199,389,1878,1903]" pageId="8" pageNumber="409">Natural history.</emphasis>
|
||
Collected on the cave wall. The cave is located in the bamboo plant community of deciduous forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB632FFFEFF4CFF2C017AFF11" pageId="9" pageNumber="410" type="description">
|
||
<paragraph id="8BE236C7B632FFFEFF4CFF2C0241FF73" blockId="9.[151,595,151,211]" box="[151,595,151,178]" pageId="9" pageNumber="410">
|
||
<taxonomicName id="4C5D4D44B632FFFEFF4CFF2C01E1FF70" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[151,499,151,178]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="tadetuensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B632FFFEFF4CFF2C01E1FF70" bold="true" box="[151,499,151,178]" pageId="9" pageNumber="410">
|
||
<emphasis id="B929EAD5B632FFFEFF4CFF2C0182FF73" bold="true" box="[151,400,151,177]" italics="true" pageId="9" pageNumber="410">Althepus tadetuensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B632FFFEFE60FF2C01C2FF73" bold="true" box="[443,464,151,177]" italics="true" pageId="9" pageNumber="410">et</emphasis>
|
||
Li
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB632FFFEFE21FF230241FF73" box="[506,595,152,177]" pageId="9" pageNumber="410" rank="species">
|
||
<emphasis id="B929EAD5B632FFFEFE21FF230241FF73" bold="true" box="[506,595,152,177]" pageId="9" pageNumber="410">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B632FFFEFF4CFF01017AFF11" blockId="9.[151,595,151,211]" box="[151,360,186,211]" pageId="9" pageNumber="410">
|
||
<figureCitation id="13662A42B632FFFEFF4CFF010105FF11" box="[151,279,186,211]" captionStart-0="FIGURE 21" captionStart-1="FIGURE 22" captionStartId-0="33.[151,250,1933,1955]" captionStartId-1="34.[151,250,1933,1955]" captionTargetBox-0="[225,1378,193,1911]" captionTargetBox-1="[209,1378,193,1911]" captionTargetId-0="figure@33.[209,1378,193,1911]" captionTargetId-1="figure@34.[209,1378,193,1911]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionText-0="FIGURE 21. Althepus tadetuensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 21A and 21C as in figure 21D." captionText-1="FIGURE 22. Althepus tadetuensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 22D as in figure 22E." httpUri-0="https://zenodo.org/record/1439832/files/figure.png" httpUri-1="https://zenodo.org/record/1439834/files/figure.png" pageId="9" pageNumber="410">Figs 21–22</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B632FFFEFEF9FF00017AFF11" box="[290,360,187,211]" captionStart-0="FIGURE 31" captionStart-1="FIGURE 32" captionStartId-0="43.[151,250,1917,1939]" captionStartId-1="44.[151,250,1148,1170]" captionTargetBox-0="[177,1409,195,1896]" captionTargetBox-1="[393,1194,195,1124]" captionTargetId-0="figure@43.[177,1410,193,1896]" captionTargetId-1="figure@44.[393,1194,195,1124]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionText-0="FIGURE 31. Left cheliceral retromargin of male holotype (A–F), female holotype (G) and female paratype (H). A, Althepus phadaengensis Li et Li sp. nov.; B, A. shanhu Li et Li sp. nov.; C, A. suayaiensis Li et Li sp. nov.; D, A. tadetuensis Li et Li sp. nov.; E, A. tanhuang Li et Li sp. nov.; F, A. thanlaensis Li et Li sp. nov.; G, A. tharnlodensis Li et Li sp. nov.; H, A. viengkeoensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." captionText-1="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri-0="https://zenodo.org/record/1439852/files/figure.png" httpUri-1="https://zenodo.org/record/1439854/files/figure.png" pageId="9" pageNumber="410">31–32</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB632FFFEFF4CFEB90295FEA1" pageId="9" pageNumber="410" type="materials_examined">
|
||
<paragraph id="8BE236C7B632FFFEFF4CFEB90295FEA1" blockId="9.[151,1437,258,1291]" pageId="9" pageNumber="410">
|
||
<bibRef id="52FDA231B632FFFEFF4CFEB90295FEA1" pageId="9" pageNumber="410" refId="ref9986" type="book chapter">
|
||
<materialsCitation id="3B353C9AB632FFFEFF4CFEB903C2FEFD" ID-GBIF-Occurrence="1913888974" collectingDate="2012-11-17" country="Laos" county="Tad" elevation="889" latitude="15.192117" location="Houses" longLatPrecision="1" longitude="106.1034" municipality="Vacation Villages" pageId="9" pageNumber="410" specimenCount="2" stateProvince="Champasak Province" typeStatus="holotype">
|
||
<emphasis id="B929EAD5B632FFFEFF4CFEB901D6FED9" bold="true" box="[151,452,258,283]" pageId="9" pageNumber="410">
|
||
<typeStatus id="54E68865B632FFFEFF4CFEB900C0FED9" box="[151,210,258,283]" pageId="9" pageNumber="410">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68865B632FFFEFE95FEB901ADFED9" box="[334,447,258,283]" pageId="9" pageNumber="410" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male (IZCAS-Ar 34021),
|
||
<author id="53E0DE22B632FFFEFD29FEB8043BFED9" box="[754,1065,258,283]" case="TC" fn="Champasak Province" fnc="TC" fns="N" ln="Laos" lnc="TC" npo="LnFn" pageId="9" pageNumber="410">
|
||
<collectingCountry id="F34A7657B632FFFEFD29FEB80339FED8" box="[754,811,259,282]" name="Laos" pageId="9" pageNumber="410">Laos</collectingCountry>
|
||
,
|
||
<collectingRegion id="4999F825B632FFFEFCECFEB9043BFED9" box="[823,1065,258,283]" country="Laos" name="Champasak" pageId="9" pageNumber="410">Champasak Province</collectingRegion>
|
||
</author>
|
||
,
|
||
<title id="F959DDF1B632FFFEFBEEFEB902F1FEFD" jopScore="0.05457072" pageId="9" pageNumber="410" type="_" typeScore="26">
|
||
<collectingCounty id="62834E4BB632FFFEFBEEFEB9044DFED9" box="[1077,1119,258,283]" pageId="9" pageNumber="410">Tad</collectingCounty>
|
||
E-tu,
|
||
<collectingMunicipality id="6B86ACBDB632FFFEFB7DFEB90579FED9" box="[1190,1387,258,283]" pageId="9" pageNumber="410">Vacation Villages</collectingMunicipality>
|
||
and
|
||
<location id="8E82601CB632FFFEFF4CFE9C00FEFEFC" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB632FFFEFF4CFE9C00FEFEFC" box="[151,236,295,318]" country="Laos" county="Tad" latitude="15.192117" longLatPrecision="1" longitude="106.1034" municipality="Vacation Villages" name="Houses" pageId="9" pageNumber="410" stateProvince="Champasak Province">Houses</location>
|
||
,
|
||
<geoCoordinate id="EE695000B632FFFEFF22FE9D0194FEFD" box="[249,390,294,319]" direction="north" orientation="latitude" pageId="9" pageNumber="410" precision="1" value="15.192117">15°11.527'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695000B632FFFEFE4FFE9D023EFEFD" box="[404,556,294,319]" direction="east" orientation="longitude" pageId="9" pageNumber="410" precision="1" value="106.1034">106°06.204'E</geoCoordinate>
|
||
,
|
||
<elevation id="0070D1F4B632FFFEFDE2FE9C0291FEFC" box="[569,643,295,319]" metricMagnitude="2" metricUnit="m" metricValue="8.89" pageId="9" pageNumber="410" unit="m" value="889.0">
|
||
<quantity id="4CA59B22B632FFFEFDE2FE9C0291FEFC" box="[569,643,295,319]" metricMagnitude="2" metricUnit="m" metricValue="8.89" pageId="9" pageNumber="410" unit="m" value="889.0">889 m</quantity>
|
||
</elevation>
|
||
altitude
|
||
</title>
|
||
,
|
||
<collectingDate id="EFA7E9EFB632FFFEFD2BFE9C03C2FEFD" box="[752,976,294,319]" pageId="9" pageNumber="410" value="2012-11-17">17 November 2012</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B353C9AB632FFFEFC07FE9D0295FEA1" ID-GBIF-Occurrence="1913888985" collectingDate="2012-11-17" country="Laos" county="Tad" elevation="889" latitude="15.192117" location="Houses" longLatPrecision="1" longitude="106.1034" municipality="Vacation Villages" pageId="9" pageNumber="410" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Champasak Province" typeStatus="paratype">
|
||
<journalOrPublisher id="519C3C0BB632FFFEFC07FE9D01C7FEA1" jopScore="0.098552175" pageId="9" pageNumber="410">
|
||
<emphasis id="B929EAD5B632FFFEFC07FE9D044CFEFD" bold="true" box="[988,1118,294,319]" pageId="9" pageNumber="410">
|
||
<typeStatus id="54E68865B632FFFEFC07FE9D044BFEFD" box="[988,1113,294,319]" pageId="9" pageNumber="410" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D5BFD4EB632FFFEFBBDFE9C04A6FEFD" box="[1126,1204,294,319]" pageId="9" pageNumber="410" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="9D5BFD4EB632FFFEFB34FE9C0547FEFD" box="[1263,1365,294,319]" pageId="9" pageNumber="410" type="female">1 female</specimenCount>
|
||
, same data as holotype (IZCAS-Ar
|
||
</journalOrPublisher>
|
||
<pagination id="B294C833B632FFFEFE07FEF00269FEA1" box="[476,635,331,355]" pageId="9" pageNumber="410" type="pagination">34022–34023</pagination>
|
||
).
|
||
</materialsCitation>
|
||
</bibRef>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB632FFFEFF1CFED503FCFE45" box="[199,1006,366,391]" pageId="9" pageNumber="410" type="etymology">
|
||
<paragraph id="8BE236C7B632FFFEFF1CFED503FCFE45" blockId="9.[151,1437,258,1291]" box="[199,1006,366,391]" pageId="9" pageNumber="410">
|
||
<emphasis id="B929EAD5B632FFFEFF1CFED5015CFE45" bold="true" box="[199,334,366,391]" pageId="9" pageNumber="410">Etymology.</emphasis>
|
||
The specific name is an adjective refers to the
|
||
<typeStatus id="54E68865B632FFFEFC86FED4039FFE45" box="[861,909,367,391]" pageId="9" pageNumber="410">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB632FFFEFF1CFE29050EFE0D" pageId="9" pageNumber="410" type="diagnosis">
|
||
<paragraph id="8BE236C7B632FFFEFF1CFE29050EFE0D" blockId="9.[151,1437,258,1291]" pageId="9" pageNumber="410">
|
||
<emphasis id="B929EAD5B632FFFEFF1CFE290153FE69" bold="true" box="[199,321,402,427]" pageId="9" pageNumber="410">Diagnosis.</emphasis>
|
||
The species can be easily distinguished from all the other species by the four furcate distal part of the conductor in males (
|
||
<figureCitation id="13662A42B632FFFEFE7EFE0D0219FE0D" box="[421,523,438,463]" captionStart="FIGURE 21" captionStartId="33.[151,250,1933,1955]" captionTargetBox="[225,1378,193,1911]" captionTargetId="figure@33.[209,1378,193,1911]" captionTargetPageId="33" captionText="FIGURE 21. Althepus tadetuensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 21A and 21C as in figure 21D." httpUri="https://zenodo.org/record/1439832/files/figure.png" pageId="9" pageNumber="410">Fig. 21D</figureCitation>
|
||
) and by one pair of long, slender spermathecae in females (
|
||
<figureCitation id="13662A42B632FFFEFB7DFE0D051DFE0D" box="[1190,1295,438,463]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@34.[209,1378,193,1911]" captionTargetPageId="34" captionText="FIGURE 22. Althepus tadetuensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 22D as in figure 22E." httpUri="https://zenodo.org/record/1439834/files/figure.png" pageId="9" pageNumber="410">Fig. 22A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB632FFFEFF1CFE610100FB5D" pageId="9" pageNumber="410" type="description">
|
||
<paragraph id="8BE236C7B632FFFEFF1CFE61016CFC61" blockId="9.[151,1437,258,1291]" pageId="9" pageNumber="410">
|
||
<emphasis id="B929EAD5B632FFFEFF1CFE610237FE31" bold="true" box="[199,549,474,499]" pageId="9" pageNumber="410">
|
||
Description. Male (
|
||
<typeStatus id="54E68865B632FFFEFE71FE610205FE31" box="[426,535,474,499]" pageId="9" pageNumber="410" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 3.80; carapace 1.17 length, 1.30 width; abdomen 2.13 length, 1.06 width. Carapace round (
|
||
<figureCitation id="13662A42B632FFFEFE7CFE45021DFDD5" box="[423,527,510,535]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@34.[209,1378,193,1911]" captionTargetPageId="34" captionText="FIGURE 22. Althepus tadetuensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 22D as in figure 22E." httpUri="https://zenodo.org/record/1439834/files/figure.png" pageId="9" pageNumber="410">Fig. 22C</figureCitation>
|
||
), yellow, with four longitudinal brown bands of different widths, the middle one shortened to form a triangular band around the eyes. Anterior margin of thoracic region distinctly elevated. Clypeus brown. Cheliceral promargin with two teeth, followed by a lamina, retromargin with two small teeth, posterior surface of fang with 20 small denticles (
|
||
<figureCitation id="13662A42B632FFFEFD0FFDD1032CFD41" box="[724,830,618,643]" captionStart="FIGURE 31" captionStartId="43.[151,250,1917,1939]" captionTargetBox="[177,1409,195,1896]" captionTargetId="figure@43.[177,1410,193,1896]" captionTargetPageId="43" captionText="FIGURE 31. Left cheliceral retromargin of male holotype (A–F), female holotype (G) and female paratype (H). A, Althepus phadaengensis Li et Li sp. nov.; B, A. shanhu Li et Li sp. nov.; C, A. suayaiensis Li et Li sp. nov.; D, A. tadetuensis Li et Li sp. nov.; E, A. tanhuang Li et Li sp. nov.; F, A. thanlaensis Li et Li sp. nov.; G, A. tharnlodensis Li et Li sp. nov.; H, A. viengkeoensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439852/files/figure.png" pageId="9" pageNumber="410">Fig. 31D</figureCitation>
|
||
). Labium brown. Sternum yellow, with two irregular brown lines on each side. Abdomen elongate, with complex pattern of white and dark-brown dorsally and ventrally. Legs brown. Leg measurements: I 32.08 (8.01, 0.40, 7.69, 13.08, 2.95), II 30.26 (7.82, 0.40, 8.01, 11.15, 2.88), III 13.70 (3.95, 0.40, 3.50, 4.50, 1.35), IV 19.90 (5.71, 0.40, 5.45, 6.67, 1.67). Male palp (
|
||
<figureCitation id="13662A42B632FFFEFBA1FD6D051FFD2D" box="[1146,1293,726,751]" captionStart="FIGURE 21" captionStartId="33.[151,250,1933,1955]" captionTargetBox="[225,1378,193,1911]" captionTargetId="figure@33.[209,1378,193,1911]" captionTargetPageId="33" captionText="FIGURE 21. Althepus tadetuensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 21A and 21C as in figure 21D." httpUri="https://zenodo.org/record/1439832/files/figure.png" pageId="9" pageNumber="410">Figs 21A–D</figureCitation>
|
||
): tarsus with three slightly curved, serrated bristles at the top of the cymbial protrusion (one of them missing,
|
||
<figureCitation id="13662A42B632FFFEFB2EFD41054FFCD1" box="[1269,1373,762,787]" captionStart="FIGURE 21" captionStartId="33.[151,250,1933,1955]" captionTargetBox="[225,1378,193,1911]" captionTargetId="figure@33.[209,1378,193,1911]" captionTargetPageId="33" captionText="FIGURE 21. Althepus tadetuensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 21A and 21C as in figure 21D." httpUri="https://zenodo.org/record/1439832/files/figure.png" pageId="9" pageNumber="410">Fig. 21C</figureCitation>
|
||
); one hooked spine with tip directed proximally (
|
||
<figureCitation id="13662A42B632FFFEFD5DFCA502E2FCF5" box="[646,752,798,823]" captionStart="FIGURE 21" captionStartId="33.[151,250,1933,1955]" captionTargetBox="[225,1378,193,1911]" captionTargetId="figure@33.[209,1378,193,1911]" captionTargetPageId="33" captionText="FIGURE 21. Althepus tadetuensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 21A and 21C as in figure 21D." httpUri="https://zenodo.org/record/1439832/files/figure.png" pageId="9" pageNumber="410">Fig. 21A</figureCitation>
|
||
); bulb yellow, pyriform; embolus arising retrolaterally from bulb, slender and curved; conductor arising prolatero-distally from bulb, S-shaped, with a transparent lamina and three black divarications arising from distal part; embolus and conductor widely separated (distance almost equal to diameter of bulb).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B632FFFEFF1CFC150184FB95" blockId="9.[151,1437,258,1291]" pageId="9" pageNumber="410">
|
||
<emphasis id="B929EAD5B632FFFEFF1CFC1501BEFC05" bold="true" box="[199,428,942,967]" pageId="9" pageNumber="410">
|
||
Female (
|
||
<typeStatus id="54E68865B632FFFEFEF6FC15018DFC05" box="[301,415,942,967]" pageId="9" pageNumber="410" type="paratype">paratype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Similar to male in color and general features (
|
||
<figureCitation id="13662A42B632FFFEFC62FC150456FC05" box="[953,1092,942,967]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@34.[209,1378,193,1911]" captionTargetPageId="34" captionText="FIGURE 22. Althepus tadetuensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 22D as in figure 22E." httpUri="https://zenodo.org/record/1439834/files/figure.png" pageId="9" pageNumber="410">Figs 22D–E</figureCitation>
|
||
) but smaller. Internal genitalia with a pair of spermathecae, distinctively long, curved, distal part swollen (
|
||
<figureCitation id="13662A42B632FFFEFC37FC690444FC29" box="[1004,1110,978,1003]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@34.[209,1378,193,1911]" captionTargetPageId="34" captionText="FIGURE 22. Althepus tadetuensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 22D as in figure 22E." httpUri="https://zenodo.org/record/1439834/files/figure.png" pageId="9" pageNumber="410">Fig. 22A</figureCitation>
|
||
). Measurements: total length 3.60; carapace 1.02 length, 1.33 width; abdomen 2.13 length, 1.09 width. Leg measurements: I 24.00 (6.41, 0.46, 6.54, 10.90, 2.69), II 17.92 (4.55, 0.46, 4.40, 6.09, 2.42), III 12.44 (3.38, 0.46, 2.97, 3.91, 1.72), IV 16.75 (4.81, 0.46, 4.49, 5.45, 1.54).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B632FFFEFF1CFBD90100FB5D" blockId="9.[151,1437,258,1291]" pageId="9" pageNumber="410">
|
||
<emphasis id="B929EAD5B632FFFEFF1CFBD9012EFBB9" bold="true" box="[199,316,1122,1147]" pageId="9" pageNumber="410">Variation.</emphasis>
|
||
Males: carapace 1.13, 1.17 length, 1.22, 1.30 width; femur I: 8.01 (n = 2; leg I lost in one specimen).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB632FFFEFF1CFB1103D9FB24" pageId="9" pageNumber="410" type="distribution">
|
||
<paragraph id="8BE236C7B632FFFEFF1CFB11034EFB01" blockId="9.[151,1437,258,1291]" box="[199,860,1194,1219]" pageId="9" pageNumber="410">
|
||
<emphasis id="B929EAD5B632FFFEFF1CFB110172FB01" bold="true" box="[199,352,1194,1219]" pageId="9" pageNumber="410">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54E68865B632FFFEFDBBFB100282FB01" box="[608,656,1195,1219]" pageId="9" pageNumber="410">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13662A42B632FFFEFD21FB11035DFB01" box="[762,847,1194,1219]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="9" pageNumber="410">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B632FFFEFF1CFB7503D9FB24" blockId="9.[151,1437,258,1291]" box="[199,971,1230,1255]" pageId="9" pageNumber="410">
|
||
<emphasis id="B929EAD5B632FFFEFF1CFB750197FB25" bold="true" box="[199,389,1230,1255]" pageId="9" pageNumber="410">Natural history.</emphasis>
|
||
Collected in a pit of the hill at an altitude of
|
||
<specimenCount id="9D5BFD4EB632FFFEFCA6FB7403D9FB24" box="[893,971,1231,1255]" pageId="9" pageNumber="410" type="generic">
|
||
<quantity id="4CA59B22B632FFFEFCA6FB7403D5FB24" box="[893,967,1231,1255]" metricMagnitude="2" metricUnit="m" metricValue="8.89" pageId="9" pageNumber="410" unit="m" value="889.0">889 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB632FFFEFF1CFB49017AFA58" pageId="9" pageNumber="410" type="discussion">
|
||
<paragraph id="8BE236C7B632FFFEFF1CFB490536FAC9" blockId="9.[151,1437,258,1291]" box="[199,1316,1266,1291]" pageId="9" pageNumber="410">
|
||
<bibRef id="52FDA231B632FFFEFF1CFB490536FAC9" box="[199,1316,1266,1291]" pageId="9" pageNumber="410" refId="ref10593" type="book">
|
||
<emphasis id="B929EAD5B632FFFEFF1CFB490122FAC9" bold="true" box="[199,304,1266,1291]" pageId="9" pageNumber="410">
|
||
<author id="53E0DE22B632FFFEFF1CFB490139FAC9" box="[199,299,1266,1291]" firstAuthorName="Remark" fns="*" inm="true" npo="*" pageId="9" pageNumber="410">Remark</author>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B929EAD5B632FFFEFEECFB4801C6FAC9" box="[311,468,1267,1291]" italics="true" pageId="9" pageNumber="410">
|
||
<title id="F959DDF1B632FFFEFEECFB480159FAC8" box="[311,331,1267,1290]" jopScore="0.007833366" pageId="9" pageNumber="410">A</title>
|
||
.
|
||
<journalOrPublisher id="519C3C0BB632FFFEFE8DFB4801C6FAC9" box="[342,468,1267,1291]" jopScore="0.17233406" pageId="9" pageNumber="410">tadetuensis</journalOrPublisher>
|
||
</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B632FFFEFE22FB48021EFAC8" box="[505,524,1267,1290]" italics="true" pageId="9" pageNumber="410">et</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B632FFFEFDEAFB490299FAC9" bold="true" box="[561,651,1266,1291]" pageId="9" pageNumber="410">sp. nov.</emphasis>
|
||
was labeled as “sp. 25” in the analysis of Li and Li (
|
||
<year id="FFF57794B632FFFEFB00FB48050BFAC9" box="[1243,1305,1267,1291]" pageId="9" pageNumber="410">2018</year>
|
||
).
|
||
</bibRef>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B632FFFEFF4CFAE50250FABA" blockId="9.[151,578,1374,1434]" box="[151,578,1374,1400]" pageId="9" pageNumber="410">
|
||
<taxonomicName id="4C5D4D44B632FFFEFF4CFAE501F3FABA" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[151,481,1374,1400]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="tanhuang" status="sp. nov.">
|
||
<emphasis id="B929EAD5B632FFFEFF4CFAE501F3FABA" bold="true" box="[151,481,1374,1400]" pageId="9" pageNumber="410">
|
||
<emphasis id="B929EAD5B632FFFEFF4CFAE5016DFABA" bold="true" box="[151,383,1374,1400]" italics="true" pageId="9" pageNumber="410">Althepus tanhuang</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B632FFFEFE72FAE501ACFABA" bold="true" box="[425,446,1374,1400]" italics="true" pageId="9" pageNumber="410">et</emphasis>
|
||
Li
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB632FFFEFE33FAE40250FABA" box="[488,578,1375,1400]" pageId="9" pageNumber="410" rank="species">
|
||
<emphasis id="B929EAD5B632FFFEFE33FAE40250FABA" bold="true" box="[488,578,1375,1400]" pageId="9" pageNumber="410">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B632FFFEFF4CFA3A017AFA58" blockId="9.[151,578,1374,1434]" box="[151,360,1409,1434]" pageId="9" pageNumber="410">
|
||
<figureCitation id="13662A42B632FFFEFF4CFA3A0104FA58" box="[151,278,1409,1434]" captionStart-0="FIGURE 23" captionStart-1="FIGURE 24" captionStartId-0="35.[151,250,1933,1955]" captionStartId-1="36.[151,250,1933,1955]" captionTargetBox-0="[211,1378,193,1911]" captionTargetBox-1="[209,1378,193,1911]" captionTargetId-0="figure@35.[209,1378,193,1911]" captionTargetId-1="figure@36.[209,1378,193,1911]" captionTargetPageId-0="35" captionTargetPageId-1="36" captionText-0="FIGURE 23. Althepus tanhuang Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 23A and 23C as in figure 23D." captionText-1="FIGURE 24. Althepus tanhuang Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 24D as in figure 24E." httpUri-0="https://zenodo.org/record/1439836/files/figure.png" httpUri-1="https://zenodo.org/record/1439838/files/figure.png" pageId="9" pageNumber="410">Figs 23–24</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B632FFFEFEF9FA39017AFA58" box="[290,360,1410,1434]" captionStart-0="FIGURE 31" captionStart-1="FIGURE 32" captionStartId-0="43.[151,250,1917,1939]" captionStartId-1="44.[151,250,1148,1170]" captionTargetBox-0="[177,1409,195,1896]" captionTargetBox-1="[393,1194,195,1124]" captionTargetId-0="figure@43.[177,1410,193,1896]" captionTargetId-1="figure@44.[393,1194,195,1124]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionText-0="FIGURE 31. Left cheliceral retromargin of male holotype (A–F), female holotype (G) and female paratype (H). A, Althepus phadaengensis Li et Li sp. nov.; B, A. shanhu Li et Li sp. nov.; C, A. suayaiensis Li et Li sp. nov.; D, A. tadetuensis Li et Li sp. nov.; E, A. tanhuang Li et Li sp. nov.; F, A. thanlaensis Li et Li sp. nov.; G, A. tharnlodensis Li et Li sp. nov.; H, A. viengkeoensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." captionText-1="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri-0="https://zenodo.org/record/1439852/files/figure.png" httpUri-1="https://zenodo.org/record/1439854/files/figure.png" pageId="9" pageNumber="410">31–32</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB632FFFEFF4CFA720164F9E8" pageId="9" pageNumber="410" type="materials_examined">
|
||
<paragraph id="8BE236C7B632FFFEFF4CFA720164F9E8" blockId="9.[151,1437,1481,2010]" pageId="9" pageNumber="410">
|
||
<bibRef id="52FDA231B632FFFEFF4CFA720164F9E8" pageId="9" pageNumber="410" refId="ref10641" type="journal article">
|
||
<materialsCitation id="3B353C9AB632FFFEFF4CFA7203B3F9C4" ID-GBIF-Occurrence="1913888966" collectingDate="2016-11-09" country="Thailand" latitude="12.766666" location="Phrayanak Cave" longLatPrecision="1" longitude="101.79855" municipality="Kongdin Subdistrict" pageId="9" pageNumber="410" specimenCount="2" stateProvince="Rayong Province" typeStatus="holotype">
|
||
<title id="F959DDF1B632FFFEFF4CFA720213FA20" box="[151,513,1481,1506]" jopScore="2.3790642E-4" pageId="9" pageNumber="410">
|
||
<emphasis id="B929EAD5B632FFFEFF4CFA7201D1FA20" bold="true" box="[151,451,1481,1506]" pageId="9" pageNumber="410">
|
||
<typeStatus id="54E68865B632FFFEFF4CFA7200C0FA20" box="[151,210,1481,1506]" pageId="9" pageNumber="410">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68865B632FFFEFE96FA7201ACFA20" box="[333,446,1481,1506]" pageId="9" pageNumber="410" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male
|
||
</title>
|
||
(
|
||
<journalOrPublisher id="519C3C0BB632FFFEFDCBFA710298FA20" box="[528,650,1482,1506]" jopScore="4.7581285E-4" pageId="9" pageNumber="410">IZCAS-Ar</journalOrPublisher>
|
||
<part id="FFF99C5AB632FFFEFD49FA7102CEFA20" box="[658,732,1482,1506]" pageId="9" pageNumber="410" type="volume">34024</part>
|
||
),
|
||
<author id="53E0DE22B632FFFEFD2BFA72043AFA20" box="[752,1064,1481,1506]" case="TC" fn="Rayong Province" fnc="TC" fns="N" ln="Thailand" lnc="TC" npo="LnFn" pageId="9" pageNumber="410">
|
||
<collectingCountry id="F34A7657B632FFFEFD2BFA720344FA20" box="[752,854,1481,1506]" name="Thailand" pageId="9" pageNumber="410">Thailand</collectingCountry>
|
||
,
|
||
<collectingRegion id="4999F825B632FFFEFCB8FA71043AFA20" box="[867,1064,1481,1506]" country="Thailand" name="Rayong" pageId="9" pageNumber="410">Rayong Province</collectingRegion>
|
||
</author>
|
||
,
|
||
<author id="53E0DE22B632FFFEFBEFFA7200C1F9C4" case="TC" fn="Phrayanak Cave" fnc="TC" fns="N" ln="Kongdin Subdistrict" lnc="TC" npo="LnFn" pageId="9" pageNumber="410">
|
||
<collectingMunicipality id="6B86ACBDB632FFFEFBEFFA720508FA20" box="[1076,1306,1481,1506]" pageId="9" pageNumber="410">Kongdin Subdistrict</collectingMunicipality>
|
||
,
|
||
<location id="8E82601CB632FFFEFAFDFA7200C1F9C4" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB632FFFEFAFDFA7200C1F9C4" country="Thailand" latitude="12.766666" longLatPrecision="1" longitude="101.79855" municipality="Kongdin Subdistrict" name="Phrayanak Cave" pageId="9" pageNumber="410" stateProvince="Rayong Province">Phrayanak Cave</location>
|
||
</author>
|
||
,
|
||
<geoCoordinate id="EE695000B632FFFEFF3BFA560163F9C4" box="[224,369,1517,1542]" direction="north" orientation="latitude" pageId="9" pageNumber="410" precision="1" value="12.766666">12°46.000'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695000B632FFFEFEA5FA55020AF9C4" box="[382,536,1518,1542]" direction="east" orientation="longitude" pageId="9" pageNumber="410" precision="1" value="101.79855">101°47.913'E</geoCoordinate>
|
||
, 0 m altitude, 0
|
||
<collectingDate id="EFA7E9EFB632FFFEFD14FA5503B3F9C4" box="[719,929,1517,1542]" pageId="9" pageNumber="410" value="2016-11-09">9 November 2016</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B353C9AB632FFFEFC77FA560164F9E8" ID-GBIF-Occurrence="1913888965" collectingDate="2016-11-09" country="Thailand" latitude="12.766666" location="Phrayanak Cave" longLatPrecision="1" longitude="101.79855" municipality="Kongdin Subdistrict" pageId="9" pageNumber="410" specimenCount="1" specimenCount-female="1" stateProvince="Rayong Province" typeStatus="paratype">
|
||
<emphasis id="B929EAD5B632FFFEFC77FA560431F9C4" bold="true" box="[940,1059,1517,1542]" pageId="9" pageNumber="410">
|
||
<typeStatus id="54E68865B632FFFEFC77FA56040FF9C4" box="[940,1053,1517,1542]" pageId="9" pageNumber="410" type="paratype">Paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D5BFD4EB632FFFEFBF7FA550480F9C4" box="[1068,1170,1517,1542]" pageId="9" pageNumber="410" type="female">1 female</specimenCount>
|
||
, same data as holotype (IZCAS-Ar
|
||
<pagination id="B294C833B632FFFEFEC4F9AA0178F9E8" box="[287,362,1553,1578]" pageId="9" pageNumber="410" type="pagination">34025</pagination>
|
||
).
|
||
</materialsCitation>
|
||
</bibRef>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB632FFFEFF1CF98E03B6F9B0" pageId="9" pageNumber="410" type="etymology">
|
||
<paragraph id="8BE236C7B632FFFEFF1CF98E03B6F9B0" blockId="9.[151,1437,1481,2010]" pageId="9" pageNumber="410">
|
||
<emphasis id="B929EAD5B632FFFEFF1CF98E015CF98C" bold="true" box="[199,334,1589,1614]" pageId="9" pageNumber="410">Etymology.</emphasis>
|
||
The specific name is derived from the Chinese pinyin 'tán huáng', which means 'spring', referring to coiled, spring-shaped spermathecae (
|
||
<figureCitation id="13662A42B632FFFEFD89F9E202AAF9B0" box="[594,696,1625,1650]" captionStart="FIGURE 24" captionStartId="36.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@36.[209,1378,193,1911]" captionTargetPageId="36" captionText="FIGURE 24. Althepus tanhuang Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 24D as in figure 24E." httpUri="https://zenodo.org/record/1439838/files/figure.png" pageId="9" pageNumber="410">Fig. 24E</figureCitation>
|
||
); noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB632FFFEFF1CF9C6053AF978" pageId="9" pageNumber="410" type="diagnosis">
|
||
<paragraph id="8BE236C7B632FFFEFF1CF9C6053AF978" blockId="9.[151,1437,1481,2010]" pageId="9" pageNumber="410">
|
||
<emphasis id="B929EAD5B632FFFEFF1CF9C60153F954" bold="true" box="[199,321,1661,1686]" pageId="9" pageNumber="410">Diagnosis.</emphasis>
|
||
This new species can be easily distinguished from all other species by the distinctly wide distal part of the folded conductor in male (
|
||
<figureCitation id="13662A42B632FFFEFDDDF91A027FF978" box="[518,621,1697,1722]" captionStart="FIGURE 23" captionStartId="35.[151,250,1933,1955]" captionTargetBox="[211,1378,193,1911]" captionTargetId="figure@35.[209,1378,193,1911]" captionTargetPageId="35" captionText="FIGURE 23. Althepus tanhuang Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 23A and 23C as in figure 23D." httpUri="https://zenodo.org/record/1439836/files/figure.png" pageId="9" pageNumber="410">Fig. 23A</figureCitation>
|
||
) and by the spring-shaped spermathecae in female (
|
||
<figureCitation id="13662A42B632FFFEFB68F91A0509F978" box="[1203,1307,1697,1722]" captionStart="FIGURE 24" captionStartId="36.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@36.[209,1378,193,1911]" captionTargetPageId="36" captionText="FIGURE 24. Althepus tanhuang Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 24D as in figure 24E." httpUri="https://zenodo.org/record/1439838/files/figure.png" pageId="9" pageNumber="410">Fig. 24A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB632FFFDFF1CF97E02C3FE6F" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410" type="description">
|
||
<paragraph id="8BE236C7B632FFFDFF1CF97E0173FEDF" blockId="9.[151,1437,1481,2010]" lastBlockId="10.[151,1437,151,501]" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410">
|
||
<emphasis id="B929EAD5B632FFFEFF1CF97E0237F91C" bold="true" box="[199,549,1733,1758]" pageId="9" pageNumber="410">
|
||
Description. Male (
|
||
<typeStatus id="54E68865B632FFFEFE71F97E0205F91C" box="[426,535,1733,1758]" pageId="9" pageNumber="410" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 4.35; carapace 1.50 length, 1.70 width; abdomen 2.50 length, 1.70 width. Carapace round (
|
||
<figureCitation id="13662A42B632FFFEFE71F9520201F8C0" box="[426,531,1769,1794]" captionStart="FIGURE 24" captionStartId="36.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@36.[209,1378,193,1911]" captionTargetPageId="36" captionText="FIGURE 24. Althepus tanhuang Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 24D as in figure 24E." httpUri="https://zenodo.org/record/1439838/files/figure.png" pageId="9" pageNumber="410">Fig. 24C</figureCitation>
|
||
), yellowish, with three longitudinal brown bands of different width, the middle shortened to triangular band around eyes. Anterior margin of thoracic region distinctly elevated. Clypeus brown. Cheliceral promargin with two teeth, followed by a lamina, retromargin with two small teeth, posterior surface of fang provided with 15 small denticles (
|
||
<figureCitation id="13662A42B632FFFEFDABF8EE02CEF8AC" box="[624,732,1877,1902]" captionStart="FIGURE 31" captionStartId="43.[151,250,1917,1939]" captionTargetBox="[177,1409,195,1896]" captionTargetId="figure@43.[177,1410,193,1896]" captionTargetPageId="43" captionText="FIGURE 31. Left cheliceral retromargin of male holotype (A–F), female holotype (G) and female paratype (H). A, Althepus phadaengensis Li et Li sp. nov.; B, A. shanhu Li et Li sp. nov.; C, A. suayaiensis Li et Li sp. nov.; D, A. tadetuensis Li et Li sp. nov.; E, A. tanhuang Li et Li sp. nov.; F, A. thanlaensis Li et Li sp. nov.; G, A. tharnlodensis Li et Li sp. nov.; H, A. viengkeoensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439852/files/figure.png" pageId="9" pageNumber="410">Fig. 31E</figureCitation>
|
||
). Labium brown. Sternum brown. Abdomen elongate, with complex patterns of white and brown dorsally and ventrally. Legs brown. Leg measurements: I missing, II 32.32 (8.65, 0.71, 8.53, 12.18, 2.25), III 22.84 (6.54, 0.70, 6.09, 7.88, 1.63), IV 32.63 (8.97, 0.71, 9.29, 11.54, 2.12). Male palp (
|
||
<figureCitation id="13662A42B632FFFEFF0CF87A017AF818" box="[215,360,1985,2010]" captionStart="FIGURE 23" captionStartId="35.[151,250,1933,1955]" captionTargetBox="[211,1378,193,1911]" captionTargetId="figure@35.[209,1378,193,1911]" captionTargetPageId="35" captionText="FIGURE 23. Althepus tanhuang Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 23A and 23C as in figure 23D." httpUri="https://zenodo.org/record/1439836/files/figure.png" pageId="9" pageNumber="410">Figs 23A–D</figureCitation>
|
||
): tarsus with three slightly curved, serrated bristles at the top of cymbial protrusion (
|
||
<figureCitation id="13662A42B632FFFEFAF3F87A059DF818" box="[1320,1423,1985,2010]" captionStart="FIGURE 23" captionStartId="35.[151,250,1933,1955]" captionTargetBox="[211,1378,193,1911]" captionTargetId="figure@35.[209,1378,193,1911]" captionTargetPageId="35" captionText="FIGURE 23. Althepus tanhuang Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 23A and 23C as in figure 23D." httpUri="https://zenodo.org/record/1439836/files/figure.png" pageId="9" pageNumber="410">Fig. 23C</figureCitation>
|
||
), one hooked spine with tip directed proximally (
|
||
<figureCitation id="13662A42B631FFFDFD61FF2C0337FF72" box="[698,805,151,176]" captionStart="FIGURE 23" captionStartId="35.[151,250,1933,1955]" captionTargetBox="[211,1378,193,1911]" captionTargetId="figure@35.[209,1378,193,1911]" captionTargetPageId="35" captionText="FIGURE 23. Althepus tanhuang Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 23A and 23C as in figure 23D." httpUri="https://zenodo.org/record/1439836/files/figure.png" pageId="10" pageNumber="411">Fig. 23A</figureCitation>
|
||
); bulb yellow, ovate; embolus arising prolatero-distally from bulb, slender and curved (
|
||
<figureCitation id="13662A42B631FFFDFDDFFF070286FF16" box="[516,660,188,213]" captionStart="FIGURE 23" captionStartId="35.[151,250,1933,1955]" captionTargetBox="[211,1378,193,1911]" captionTargetId="figure@35.[209,1378,193,1911]" captionTargetPageId="35" captionText="FIGURE 23. Althepus tanhuang Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 23A and 23C as in figure 23D." httpUri="https://zenodo.org/record/1439836/files/figure.png" pageId="10" pageNumber="411">Figs 23C–D</figureCitation>
|
||
); conductor arising retrolatero-distally from bulb, distinct wide and distal part gradually narrow (
|
||
<figureCitation id="13662A42B631FFFDFE31FF640244FF3A" box="[490,598,223,248]" captionStart="FIGURE 23" captionStartId="35.[151,250,1933,1955]" captionTargetBox="[211,1378,193,1911]" captionTargetId="figure@35.[209,1378,193,1911]" captionTargetPageId="35" captionText="FIGURE 23. Althepus tanhuang Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 23A and 23C as in figure 23D." httpUri="https://zenodo.org/record/1439836/files/figure.png" pageId="10" pageNumber="411">Fig. 23A</figureCitation>
|
||
); embolus and conductor widely separated (the distance almost equal to diameter of bulb).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B631FFFDFF1CFE9C02C3FE6F" blockId="10.[151,1437,151,501]" pageId="10" pageNumber="411">
|
||
<emphasis id="B929EAD5B631FFFDFF1CFE9C01BEFE82" bold="true" box="[199,428,295,320]" pageId="10" pageNumber="411">
|
||
Female (
|
||
<typeStatus id="54E68865B631FFFDFEF6FE9C018DFE82" box="[301,415,295,320]" pageId="10" pageNumber="411" type="paratype">paratype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Similar to male in general features (
|
||
<figureCitation id="13662A42B631FFFDFC9DFE9C03C1FE82" box="[838,979,295,320]" captionStart="FIGURE 24" captionStartId="36.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@36.[209,1378,193,1911]" captionTargetPageId="36" captionText="FIGURE 24. Althepus tanhuang Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 24D as in figure 24E." httpUri="https://zenodo.org/record/1439838/files/figure.png" pageId="10" pageNumber="411">Figs 24D–E</figureCitation>
|
||
), but larger and darker. Internal genitalia with a pair of spermathecae, coiled, spring-shaped, the distal part swollen (
|
||
<figureCitation id="13662A42B631FFFDFC30FEF70447FEA6" box="[1003,1109,332,357]" captionStart="FIGURE 24" captionStartId="36.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@36.[209,1378,193,1911]" captionTargetPageId="36" captionText="FIGURE 24. Althepus tanhuang Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 24D as in figure 24E." httpUri="https://zenodo.org/record/1439838/files/figure.png" pageId="10" pageNumber="411">Fig. 24A</figureCitation>
|
||
). Measurements: total length 4.36; carapace 1.38 length, 1.48 width; abdomen 2.50 length, 1.20 width. Leg measurements: I missing, II missing, III missing, IV 16.09 (6.73, 0.56, 6.54, 8.01, 5.45).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB631FFFDFF1CFE0C03ECFE36" pageId="10" pageNumber="411" type="distribution">
|
||
<paragraph id="8BE236C7B631FFFDFF1CFE0C034EFE12" blockId="10.[151,1437,151,501]" box="[199,860,439,464]" pageId="10" pageNumber="411">
|
||
<emphasis id="B929EAD5B631FFFDFF1CFE0C0172FE12" bold="true" box="[199,352,439,464]" pageId="10" pageNumber="411">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54E68865B631FFFDFDBBFE030282FE12" box="[608,656,440,464]" pageId="10" pageNumber="411">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13662A42B631FFFDFD21FE0C035DFE12" box="[762,847,439,464]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="10" pageNumber="411">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B631FFFDFF1CFE6703ECFE36" blockId="10.[151,1437,151,501]" box="[199,1022,476,501]" pageId="10" pageNumber="411">
|
||
<emphasis id="B929EAD5B631FFFDFF1CFE670197FE37" bold="true" box="[199,389,476,501]" pageId="10" pageNumber="411">Natural history.</emphasis>
|
||
Collected around the entrance zone of a limestone cave.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB631FFFDFF4CFDFC017AFD41" pageId="10" pageNumber="411" type="description">
|
||
<paragraph id="8BE236C7B631FFFDFF4CFDFC0245FDA3" blockId="10.[151,599,583,643]" box="[151,599,583,610]" pageId="10" pageNumber="411">
|
||
<taxonomicName id="4C5D4D44B631FFFDFF4CFDFC01E5FDA0" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[151,503,583,610]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="thanlaensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B631FFFDFF4CFDFC01E5FDA0" bold="true" box="[151,503,583,610]" pageId="10" pageNumber="411">
|
||
<emphasis id="B929EAD5B631FFFDFF4CFDFC0186FDA3" bold="true" box="[151,404,583,609]" italics="true" pageId="10" pageNumber="411">Althepus thanlaensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B631FFFDFE65FDFC01C1FDA3" bold="true" box="[446,467,583,609]" italics="true" pageId="10" pageNumber="411">et</emphasis>
|
||
Li
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB631FFFDFE25FDF30245FDA3" box="[510,599,584,609]" pageId="10" pageNumber="411" rank="species">
|
||
<emphasis id="B929EAD5B631FFFDFE25FDF30245FDA3" bold="true" box="[510,599,584,609]" pageId="10" pageNumber="411">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B631FFFDFF4CFDD1017AFD41" blockId="10.[151,599,583,643]" box="[151,360,618,643]" pageId="10" pageNumber="411">
|
||
<figureCitation id="13662A42B631FFFDFF4CFDD10105FD41" box="[151,279,618,643]" captionStart-0="FIGURE 25" captionStart-1="FIGURE 26" captionStartId-0="37.[151,250,1933,1955]" captionStartId-1="38.[151,250,1933,1955]" captionTargetBox-0="[215,1378,193,1911]" captionTargetBox-1="[209,1378,193,1911]" captionTargetId-0="figure@37.[209,1378,193,1911]" captionTargetId-1="figure@38.[209,1378,193,1911]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionText-0="FIGURE 25. Althepus thanlaensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 25A and 25C as in figure 25D." captionText-1="FIGURE 26. Althepus thanlaensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 26D as in figure 26E." httpUri-0="https://zenodo.org/record/1439840/files/figure.png" httpUri-1="https://zenodo.org/record/1439842/files/figure.png" pageId="10" pageNumber="411">Figs 25–26</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B631FFFDFEF9FDD0017AFD41" box="[290,360,619,643]" captionStart-0="FIGURE 31" captionStart-1="FIGURE 32" captionStartId-0="43.[151,250,1917,1939]" captionStartId-1="44.[151,250,1148,1170]" captionTargetBox-0="[177,1409,195,1896]" captionTargetBox-1="[393,1194,195,1124]" captionTargetId-0="figure@43.[177,1410,193,1896]" captionTargetId-1="figure@44.[393,1194,195,1124]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionText-0="FIGURE 31. Left cheliceral retromargin of male holotype (A–F), female holotype (G) and female paratype (H). A, Althepus phadaengensis Li et Li sp. nov.; B, A. shanhu Li et Li sp. nov.; C, A. suayaiensis Li et Li sp. nov.; D, A. tadetuensis Li et Li sp. nov.; E, A. tanhuang Li et Li sp. nov.; F, A. thanlaensis Li et Li sp. nov.; G, A. tharnlodensis Li et Li sp. nov.; H, A. viengkeoensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." captionText-1="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri-0="https://zenodo.org/record/1439852/files/figure.png" httpUri-1="https://zenodo.org/record/1439854/files/figure.png" pageId="10" pageNumber="411">31–32</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB631FFFDFF4CFD090459FCD1" pageId="10" pageNumber="411" type="materials_examined">
|
||
<paragraph id="8BE236C7B631FFFDFF4CFD090459FCD1" blockId="10.[151,1437,690,1615]" pageId="10" pageNumber="411">
|
||
<bibRef id="52FDA231B631FFFDFF4CFD090459FCD1" pageId="10" pageNumber="411" refId="ref11189" type="book chapter">
|
||
<materialsCitation id="3B353C9AB631FFFDFF4CFD09058BFD2D" ID-GBIF-Occurrence="1913888976" collectingDate="2014-10-17" country="Thailand" county="Nakhothai District" elevation="510" latitude="16.9136" location="Tham" longLatPrecision="1" longitude="100.72045" municipality="Ban Kaset Sut Village" pageId="10" pageNumber="411" specimenCount="2" stateProvince="Phitsanulok Province" typeStatus="holotype">
|
||
<emphasis id="B929EAD5B631FFFDFF4CFD0901DEFD09" bold="true" box="[151,460,690,715]" pageId="10" pageNumber="411">
|
||
<typeStatus id="54E68865B631FFFDFF4CFD0900C0FD09" box="[151,210,690,715]" pageId="10" pageNumber="411">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68865B631FFFDFE8EFD0901D4FD09" box="[341,454,690,715]" pageId="10" pageNumber="411" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male (IZCAS-Ar 34026),
|
||
<author id="53E0DE22B631FFFDFCD1FD090465FD09" box="[778,1143,690,715]" case="TC" fn="Phitsanulok Province" fnc="TC" fns="N" ln="Thailand" lnc="TC" npo="LnFn" pageId="10" pageNumber="411">
|
||
<collectingCountry id="F34A7657B631FFFDFCD1FD090362FD09" box="[778,880,690,715]" name="Thailand" pageId="10" pageNumber="411">Thailand</collectingCountry>
|
||
,
|
||
<collectingRegion id="4999F825B631FFFDFC5BFD090465FD09" box="[896,1143,690,715]" country="Thailand" name="Phitsanulok" pageId="10" pageNumber="411">Phitsanulok Province</collectingRegion>
|
||
</author>
|
||
,
|
||
<author id="53E0DE22B631FFFDFB5CFD090174FD2D" case="TC" fn="Ban Kaset Sut Village" fnc="TC" fns="N" ln="Nakhothai District" lnc="TC" npo="LnFn" pageId="10" pageNumber="411">
|
||
<collectingCounty id="62834E4BB631FFFDFB5CFD09054DFD09" box="[1159,1375,690,715]" pageId="10" pageNumber="411">Nakhothai District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6B86ACBDB631FFFDFAB4FD080174FD2D" pageId="10" pageNumber="411">Ban Kaset Sut Village</collectingMunicipality>
|
||
</author>
|
||
,
|
||
<title id="F959DDF1B631FFFDFEAEFD6D04D0FD2D" box="[373,1218,726,751]" jopScore="0.15882303" pageId="10" pageNumber="411" type="_" typeScore="26">
|
||
<location id="8E82601CB631FFFDFEAEFD6D01A4FD2D" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB631FFFDFEAEFD6D01A4FD2D" box="[373,438,726,751]" country="Thailand" county="Nakhothai District" latitude="16.9136" longLatPrecision="1" longitude="100.72045" municipality="Ban Kaset Sut Village" name="Tham" pageId="10" pageNumber="411" stateProvince="Phitsanulok Province">Tham</location>
|
||
(cave)
|
||
<location id="8E82601CB631FFFDFDC9FD6D02AEFD2D" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB631FFFDFDC9FD6D02AEFD2D" box="[530,700,726,751]" country="Thailand" county="Nakhothai District" latitude="16.9136" longLatPrecision="1" longitude="100.72045" municipality="Ban Kaset Sut Village" name="Than La Cave" pageId="10" pageNumber="411" stateProvince="Phitsanulok Province">Than La Cave</location>
|
||
,
|
||
<geoCoordinate id="EE695000B631FFFDFD10FD6D034EFD2D" box="[715,860,726,751]" direction="north" orientation="latitude" pageId="10" pageNumber="411" precision="1" value="16.9136">16°54.816'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695000B631FFFDFCB1FD6C0417FD2D" box="[874,1029,727,751]" direction="east" orientation="longitude" pageId="10" pageNumber="411" precision="1" value="100.72045">100°43.227'E</geoCoordinate>
|
||
,
|
||
<elevation id="0070D1F4B631FFFDFBCEFD6D0472FD2C" box="[1045,1120,726,751]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="10" pageNumber="411" unit="m" value="510.0">
|
||
<quantity id="4CA59B22B631FFFDFBCEFD6D0472FD2C" box="[1045,1120,726,751]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="10" pageNumber="411" unit="m" value="510.0">510 m</quantity>
|
||
</elevation>
|
||
altitude
|
||
</title>
|
||
,
|
||
<collectingDate id="EFA7E9EFB631FFFDFB0AFD6C058BFD2D" box="[1233,1433,726,751]" pageId="10" pageNumber="411" value="2014-10-17">17 October 2014</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B353C9AB631FFFDFF4CFD410459FCD1" ID-GBIF-Occurrence="1913888964" box="[151,1099,762,787]" collectingDate="2014-10-17" country="Thailand" county="Nakhothai District" elevation="510" latitude="16.9136" location="Tham" longLatPrecision="1" longitude="100.72045" municipality="Ban Kaset Sut Village" pageId="10" pageNumber="411" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Phitsanulok Province" typeStatus="paratype">
|
||
<journalOrPublisher id="519C3C0BB631FFFDFF4CFD410388FCD1" box="[151,922,762,787]" jopScore="0.20507082" pageId="10" pageNumber="411">
|
||
<emphasis id="B929EAD5B631FFFDFF4CFD41010AFCD1" bold="true" box="[151,280,762,787]" pageId="10" pageNumber="411">
|
||
<typeStatus id="54E68865B631FFFDFF4CFD410101FCD1" box="[151,275,762,787]" pageId="10" pageNumber="411" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D5BFD4EB631FFFDFEC5FD400178FCD1" box="[286,362,762,787]" pageId="10" pageNumber="411" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="9D5BFD4EB631FFFDFE7BFD40021DFCD1" box="[416,527,762,787]" pageId="10" pageNumber="411" type="female">2 females</specimenCount>
|
||
, same data as holotype (IZCAS-Ar
|
||
</journalOrPublisher>
|
||
<pagination id="B294C833B631FFFDFC7BFD40042DFCD1" box="[928,1087,763,787]" pageId="10" pageNumber="411" type="pagination">34027–34029</pagination>
|
||
).
|
||
</materialsCitation>
|
||
</bibRef>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB631FFFDFF1CFCA5040FFCF5" box="[199,1053,798,823]" pageId="10" pageNumber="411" type="etymology">
|
||
<paragraph id="8BE236C7B631FFFDFF1CFCA5040FFCF5" blockId="10.[151,1437,690,1615]" box="[199,1053,798,823]" pageId="10" pageNumber="411">
|
||
<emphasis id="B929EAD5B631FFFDFF1CFCA5015CFCF5" bold="true" box="[199,334,798,823]" pageId="10" pageNumber="411">Etymology.</emphasis>
|
||
The specific name is an adjective and refers to the
|
||
<typeStatus id="54E68865B631FFFDFC57FCA403AEFCF5" box="[908,956,799,823]" pageId="10" pageNumber="411">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB631FFFDFF1CFCF904B7FCBD" pageId="10" pageNumber="411" type="diagnosis">
|
||
<paragraph id="8BE236C7B631FFFDFF1CFCF904B7FCBD" blockId="10.[151,1437,690,1615]" pageId="10" pageNumber="411">
|
||
<emphasis id="B929EAD5B631FFFDFF1CFCF90153FC99" bold="true" box="[199,321,834,859]" pageId="10" pageNumber="411">Diagnosis.</emphasis>
|
||
This new species can be easily distinguished from all other species by the blade-like conductor (
|
||
<figureCitation id="13662A42B631FFFDFF44FCDD0139FCBD" box="[159,299,870,895]" captionStart="FIGURE 25" captionStartId="37.[151,250,1933,1955]" captionTargetBox="[215,1378,193,1911]" captionTargetId="figure@37.[209,1378,193,1911]" captionTargetPageId="37" captionText="FIGURE 25. Althepus thanlaensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 25A and 25C as in figure 25D." httpUri="https://zenodo.org/record/1439840/files/figure.png" pageId="10" pageNumber="411">Figs 25A–D</figureCitation>
|
||
) in males, and two pairs of similarly sized spermathecae in females (
|
||
<figureCitation id="13662A42B631FFFDFBEBFCDD048BFCBD" box="[1072,1177,870,895]" captionStart="FIGURE 26" captionStartId="38.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@38.[209,1378,193,1911]" captionTargetPageId="38" captionText="FIGURE 26. Althepus thanlaensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 26D as in figure 26E." httpUri="https://zenodo.org/record/1439842/files/figure.png" pageId="10" pageNumber="411">Fig. 26A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB631FFFDFF1CFC310443FA21" pageId="10" pageNumber="411" type="description">
|
||
<paragraph id="8BE236C7B631FFFDFF1CFC3102B4FAC9" blockId="10.[151,1437,690,1615]" pageId="10" pageNumber="411">
|
||
<emphasis id="B929EAD5B631FFFDFF1CFC310237FC61" bold="true" box="[199,549,906,931]" pageId="10" pageNumber="411">
|
||
Description. Male (
|
||
<typeStatus id="54E68865B631FFFDFE71FC310205FC61" box="[426,535,906,931]" pageId="10" pageNumber="411" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 4.10; carapace 1.44 length, 1.53 width; abdomen 2.30 length, 1.22 width. Carapace rounded (
|
||
<figureCitation id="13662A42B631FFFDFE1FFC15023EFC05" box="[452,556,942,967]" captionStart="FIGURE 26" captionStartId="38.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@38.[209,1378,193,1911]" captionTargetPageId="38" captionText="FIGURE 26. Althepus thanlaensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 26D as in figure 26E." httpUri="https://zenodo.org/record/1439842/files/figure.png" pageId="10" pageNumber="411">Fig. 26C</figureCitation>
|
||
), yellow, with three longitudinal brown bands of different widths, the middle one wider than the others. Clypeus brown. Cheliceral promargin with two teeth, followed by a lamina, retromargin with two small teeth, posterior surface of fang with 20 small denticles (
|
||
<figureCitation id="13662A42B631FFFDFC60FC4D040DFBCD" box="[955,1055,1014,1039]" captionStart="FIGURE 31" captionStartId="43.[151,250,1917,1939]" captionTargetBox="[177,1409,195,1896]" captionTargetId="figure@43.[177,1410,193,1896]" captionTargetPageId="43" captionText="FIGURE 31. Left cheliceral retromargin of male holotype (A–F), female holotype (G) and female paratype (H). A, Althepus phadaengensis Li et Li sp. nov.; B, A. shanhu Li et Li sp. nov.; C, A. suayaiensis Li et Li sp. nov.; D, A. tadetuensis Li et Li sp. nov.; E, A. tanhuang Li et Li sp. nov.; F, A. thanlaensis Li et Li sp. nov.; G, A. tharnlodensis Li et Li sp. nov.; H, A. viengkeoensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439852/files/figure.png" pageId="10" pageNumber="411">Fig. 31F</figureCitation>
|
||
). Labium brown. Sternum brown. Abdomen elongate, with patterns of white and brown dorsally and ventrally. Legs brown, femur and tibia with white annulations. Leg measurements: I 47.75 (10.90, 0.63, 11.54, 21.15, 3.53), II 26.97 (7.05, 0.63, 6.92, 10.13, 2.24), III missing, IV 26.17 (7.37, 0.60, 7.31, 8.97, 1.92). Male palp (
|
||
<figureCitation id="13662A42B631FFFDFC0BFBD90477FBB9" box="[976,1125,1122,1147]" captionStart="FIGURE 25" captionStartId="37.[151,250,1933,1955]" captionTargetBox="[215,1378,193,1911]" captionTargetId="figure@37.[209,1378,193,1911]" captionTargetPageId="37" captionText="FIGURE 25. Althepus thanlaensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 25A and 25C as in figure 25D." httpUri="https://zenodo.org/record/1439840/files/figure.png" pageId="10" pageNumber="411">Figs 25A–D</figureCitation>
|
||
): tarsus with three slightly curved, serrated bristles at the top of cymbial protrusion (one of them missing,
|
||
<figureCitation id="13662A42B631FFFDFBDFFB3D0478FB5D" box="[1028,1130,1158,1183]" captionStart="FIGURE 25" captionStartId="37.[151,250,1933,1955]" captionTargetBox="[215,1378,193,1911]" captionTargetId="figure@37.[209,1378,193,1911]" captionTargetPageId="37" captionText="FIGURE 25. Althepus thanlaensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 25A and 25C as in figure 25D." httpUri="https://zenodo.org/record/1439840/files/figure.png" pageId="10" pageNumber="411">Fig. 25A</figureCitation>
|
||
), one hooked spine with tip directed proximally (
|
||
<figureCitation id="13662A42B631FFFDFE58FB1101FEFB01" box="[387,492,1194,1219]" captionStart="FIGURE 25" captionStartId="37.[151,250,1933,1955]" captionTargetBox="[215,1378,193,1911]" captionTargetId="figure@37.[209,1378,193,1911]" captionTargetPageId="37" captionText="FIGURE 25. Althepus thanlaensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, prolateral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 25A and 25C as in figure 25D." httpUri="https://zenodo.org/record/1439840/files/figure.png" pageId="10" pageNumber="411">Fig. 25A</figureCitation>
|
||
); bulb yellow, ovate; embolus arising distally from bulb, slender and slightly curved distally; conductor arising distally from bulb, blade-like and increasingly widening; embolus and conductor widely separated (distance less than diameter of bulb).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B631FFFDFF1CFAAD04A4FA59" blockId="10.[151,1437,690,1615]" pageId="10" pageNumber="411">
|
||
<emphasis id="B929EAD5B631FFFDFF1CFAAD01BFFAED" bold="true" box="[199,429,1302,1327]" pageId="10" pageNumber="411">
|
||
Female (
|
||
<typeStatus id="54E68865B631FFFDFEF5FAAD01B2FAED" box="[302,416,1302,1327]" pageId="10" pageNumber="411" type="paratype">paratype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Similar to male in color and general features (
|
||
<figureCitation id="13662A42B631FFFDFC1EFAAD0442FAED" box="[965,1104,1302,1327]" captionStart="FIGURE 26" captionStartId="38.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@38.[209,1378,193,1911]" captionTargetPageId="38" captionText="FIGURE 26. Althepus thanlaensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 26D as in figure 26E." httpUri="https://zenodo.org/record/1439842/files/figure.png" pageId="10" pageNumber="411">Figs 26D–E</figureCitation>
|
||
) but larger. Internal genitalia with two pairs of spermathecae, long, curved, distal part swollen (
|
||
<figureCitation id="13662A42B631FFFDFC45FA810418FA91" box="[926,1034,1338,1363]" captionStart="FIGURE 26" captionStartId="38.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@38.[209,1378,193,1911]" captionTargetPageId="38" captionText="FIGURE 26. Althepus thanlaensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 26D as in figure 26E." httpUri="https://zenodo.org/record/1439842/files/figure.png" pageId="10" pageNumber="411">Fig. 26A</figureCitation>
|
||
). Measurements: total length 4.50; carapace 1.36 length, 1.28 width; abdomen 2.78 length, 1.62 width. Leg measurements: I 38.06 (8.65, 0.57, 9.29, 16.34, 3.21), II 21.25 (5.77, 0.54, 5.64, 7.50, 1.80), III missing, IV - (6.03, 0.57, 5.96, 6.54, -).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B631FFFDFF1CFA1D0443FA21" blockId="10.[151,1437,690,1615]" pageId="10" pageNumber="411">
|
||
<emphasis id="B929EAD5B631FFFDFF1CFA1D012EFA7D" bold="true" box="[199,316,1446,1471]" pageId="10" pageNumber="411">Variation.</emphasis>
|
||
Males: carapace 1.42, 1.44 length, 1.50, 1.53 width; femur I: 10.90 (n = 2; leg I lost in one specimen). Prosoma: females 1.36, 1.48 length, 1.28, 1.44 width; femur I: 5.77, 8.65.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB631FFFDFF1CFA5502AEF9E9" pageId="10" pageNumber="411" type="distribution">
|
||
<paragraph id="8BE236C7B631FFFDFF1CFA55034EF9C5" blockId="10.[151,1437,690,1615]" box="[199,860,1518,1543]" pageId="10" pageNumber="411">
|
||
<emphasis id="B929EAD5B631FFFDFF1CFA550172F9C5" bold="true" box="[199,352,1518,1543]" pageId="10" pageNumber="411">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54E68865B631FFFDFDBBFA540282F9C5" box="[608,656,1519,1543]" pageId="10" pageNumber="411">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13662A42B631FFFDFD21FA55035DF9C5" box="[762,847,1518,1543]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="10" pageNumber="411">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B631FFFDFF1CF9A902AEF9E9" blockId="10.[151,1437,690,1615]" box="[199,700,1554,1579]" pageId="10" pageNumber="411">
|
||
<emphasis id="B929EAD5B631FFFDFF1CF9A90197F9E9" bold="true" box="[199,389,1554,1579]" pageId="10" pageNumber="411">Natural history.</emphasis>
|
||
Collected on the cave wall.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB631FFFDFF1CF98D012DF91C" pageId="10" pageNumber="411" type="discussion">
|
||
<paragraph id="8BE236C7B631FFFDFF1CF98D0526F98D" blockId="10.[151,1437,690,1615]" box="[199,1332,1590,1615]" pageId="10" pageNumber="411">
|
||
<bibRef id="52FDA231B631FFFDFF1CF98D0526F98D" box="[199,1332,1590,1615]" pageId="10" pageNumber="411" refId="ref11763" type="book">
|
||
<emphasis id="B929EAD5B631FFFDFF1CF98D0122F98D" bold="true" box="[199,304,1590,1615]" pageId="10" pageNumber="411">
|
||
<author id="53E0DE22B631FFFDFF1CF98D0139F98D" box="[199,299,1590,1615]" firstAuthorName="Remark" fns="*" inm="true" npo="*" pageId="10" pageNumber="411">Remark</author>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B929EAD5B631FFFDFEECF98C01C4F98C" box="[311,470,1590,1614]" italics="true" pageId="10" pageNumber="411">
|
||
<title id="F959DDF1B631FFFDFEECF98C0159F98C" box="[311,331,1591,1614]" jopScore="0.007833366" pageId="10" pageNumber="411">A</title>
|
||
.
|
||
<journalOrPublisher id="519C3C0BB631FFFDFE8DF98D01C4F98C" box="[342,470,1590,1614]" jopScore="0.17233406" pageId="10" pageNumber="411">thanlaensis</journalOrPublisher>
|
||
</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B631FFFDFE20F98C021CF98C" box="[507,526,1591,1614]" italics="true" pageId="10" pageNumber="411">et</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B631FFFDFDE8F98D029FF98D" bold="true" box="[563,653,1590,1615]" pageId="10" pageNumber="411">sp. nov.</emphasis>
|
||
was labeled as “sp. 173” in the analysis of Li and Li (
|
||
<year id="FFF57794B631FFFDFB31F98C053AF98D" box="[1258,1320,1591,1615]" pageId="10" pageNumber="411">2018</year>
|
||
).
|
||
</bibRef>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B631FFFDFF4CF9190263F97E" blockId="10.[151,625,1698,1758]" box="[151,625,1698,1725]" pageId="10" pageNumber="411">
|
||
<taxonomicName id="4C5D4D44B631FFFDFF4CF9190203F97F" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[151,529,1698,1725]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="tharnlodensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B631FFFDFF4CF9190203F97F" bold="true" box="[151,529,1698,1725]" pageId="10" pageNumber="411">
|
||
<emphasis id="B929EAD5B631FFFDFF4CF91901BCF97E" bold="true" box="[151,430,1698,1724]" italics="true" pageId="10" pageNumber="411">Althepus tharnlodensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B631FFFDFE03F91901FFF97E" bold="true" box="[472,493,1698,1724]" italics="true" pageId="10" pageNumber="411">et</emphasis>
|
||
Li
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB631FFFDFDC3F9180263F97E" box="[536,625,1699,1724]" pageId="10" pageNumber="411" rank="species">
|
||
<emphasis id="B929EAD5B631FFFDFDC3F9180263F97E" bold="true" box="[536,625,1699,1724]" pageId="10" pageNumber="411">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B631FFFDFF4CF97E012DF91C" blockId="10.[151,625,1698,1758]" box="[151,319,1733,1758]" pageId="10" pageNumber="411">
|
||
<figureCitation id="13662A42B631FFFDFF4CF97E00FEF91C" box="[151,236,1733,1758]" captionStart="FIGURE 27" captionStartId="39.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@39.[209,1378,193,1911]" captionTargetPageId="39" captionText="FIGURE 27. Althepus tharnlodensis Li et Li sp. nov., female holotype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Female habitus, retrolateral view; D. Female habitus, dorsal view; E. Female habitus, ventral view; SP = spermatheca; PP = pore plate. The scale bar in figures 27C and 27E as in figure 27D." httpUri="https://zenodo.org/record/1439844/files/figure.png" pageId="10" pageNumber="411">Figs 27</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B631FFFDFF23F97D012DF91C" box="[248,319,1734,1758]" captionStart-0="FIGURE 31" captionStart-1="FIGURE 32" captionStartId-0="43.[151,250,1917,1939]" captionStartId-1="44.[151,250,1148,1170]" captionTargetBox-0="[177,1409,195,1896]" captionTargetBox-1="[393,1194,195,1124]" captionTargetId-0="figure@43.[177,1410,193,1896]" captionTargetId-1="figure@44.[393,1194,195,1124]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionText-0="FIGURE 31. Left cheliceral retromargin of male holotype (A–F), female holotype (G) and female paratype (H). A, Althepus phadaengensis Li et Li sp. nov.; B, A. shanhu Li et Li sp. nov.; C, A. suayaiensis Li et Li sp. nov.; D, A. tadetuensis Li et Li sp. nov.; E, A. tanhuang Li et Li sp. nov.; F, A. thanlaensis Li et Li sp. nov.; G, A. tharnlodensis Li et Li sp. nov.; H, A. viengkeoensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." captionText-1="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri-0="https://zenodo.org/record/1439852/files/figure.png" httpUri-1="https://zenodo.org/record/1439854/files/figure.png" pageId="10" pageNumber="411">31–32</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB631FFFDFF4CF8B60509F888" pageId="10" pageNumber="411" type="materials_examined">
|
||
<paragraph id="8BE236C7B631FFFDFF4CF8B60509F888" blockId="10.[151,1436,1805,2010]" pageId="10" pageNumber="411">
|
||
<bibRef id="52FDA231B631FFFDFF4CF8B60509F888" pageId="10" pageNumber="411" refId="ref11809" type="journal article">
|
||
<materialsCitation id="3B353C9AB631FFFDFF4CF8B60509F888" ID-GBIF-Occurrence="1913888967" collectingDate="2014-10-27" country="Thailand" county="Tap Pud District" elevation="116" latitude="8.518967" location="Tharn Lod Cave" longLatPrecision="1" longitude="98.56525" municipality="Tham Thong Lang Subdistrict" pageId="10" pageNumber="411" specimenCount="2" stateProvince="Phang Nga Province" typeStatus="holotype">
|
||
<title id="F959DDF1B631FFFDFF4CF8B6031AF8E4" box="[151,776,1805,1830]" jopScore="0.0018726592" pageId="10" pageNumber="411">
|
||
<emphasis id="B929EAD5B631FFFDFF4CF8B601DBF8E4" bold="true" box="[151,457,1805,1830]" pageId="10" pageNumber="411">
|
||
<typeStatus id="54E68865B631FFFDFF4CF8B600C0F8E4" box="[151,210,1805,1830]" pageId="10" pageNumber="411">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68865B631FFFDFE88F8B601D6F8E4" box="[339,452,1805,1830]" pageId="10" pageNumber="411" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
female (IZCAS-Ar 34030)
|
||
</title>
|
||
,
|
||
<collectingCountry id="F34A7657B631FFFDFCCCF8B6036FF8E4" box="[791,893,1805,1830]" name="Thailand" pageId="10" pageNumber="411">Thailand</collectingCountry>
|
||
,
|
||
<author id="53E0DE22B631FFFDFC57F8B6055EF8E4" box="[908,1356,1805,1830]" case="TC" fn="Tap Pud District" fnc="TC" fns="N" ln="Phang Nga Province" lnc="TC" npo="LnFn" pageId="10" pageNumber="411">
|
||
<collectingRegion id="4999F825B631FFFDFC57F8B6046FF8E4" box="[908,1149,1805,1830]" country="Thailand" name="Phangnga" pageId="10" pageNumber="411">
|
||
<collectingRegion id="4999F825B631FFFDFC57F8B603C0F8E4" box="[908,978,1805,1830]" country="Malaysia" name="Pahang" pageId="10" pageNumber="411">Phang</collectingRegion>
|
||
<collectingRegion id="4999F825B631FFFDFC06F8B5046FF8E4" box="[989,1149,1805,1830]" inferredBySuffix="true" pageId="10" pageNumber="411">Nga Province</collectingRegion>
|
||
</collectingRegion>
|
||
,
|
||
<collectingCounty id="62834E4BB631FFFDFB50F8B5055EF8E4" box="[1163,1356,1805,1830]" pageId="10" pageNumber="411">Tap Pud District</collectingCounty>
|
||
</author>
|
||
,
|
||
<author id="53E0DE22B631FFFDFA80F8B6027BF888" case="TC" fn="Tharn Lod Cave" fnc="TC" fns="N" ln="Tham Thong Lang Subdistrict" lnc="TC" npo="LnFn" pageId="10" pageNumber="411">
|
||
<collectingMunicipality id="6B86ACBDB631FFFDFA80F8B601B1F888" pageId="10" pageNumber="411">Tham Thong Lang Subdistrict</collectingMunicipality>
|
||
,
|
||
<location id="8E82601CB631FFFDFE74F88A027BF888" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB631FFFDFE74F88A027BF888" box="[431,617,1841,1866]" country="Thailand" county="Tap Pud District" latitude="8.518967" longLatPrecision="1" longitude="98.56525" municipality="Tham Thong Lang Subdistrict" name="Tharn Lod Cave" pageId="10" pageNumber="411" stateProvince="Phang Nga Province">Tharn Lod Cave</location>
|
||
</author>
|
||
,
|
||
<journalOrPublisher id="519C3C0BB631FFFDFDAFF8890317F888" box="[628,773,1842,1866]" jopScore="0.0411985" pageId="10" pageNumber="411" type="_" typeScore="19">
|
||
<geoCoordinate id="EE695000B631FFFDFDAFF8890317F888" box="[628,773,1842,1866]" direction="north" orientation="latitude" pageId="10" pageNumber="411" precision="1" value="8.518967">08°31.138'N</geoCoordinate>
|
||
</journalOrPublisher>
|
||
,
|
||
<geoCoordinate id="EE695000B631FFFDFCCBF88A038EF888" box="[784,924,1841,1866]" direction="east" orientation="longitude" pageId="10" pageNumber="411" precision="1" value="98.56525">
|
||
<part id="FFF99C5AB631FFFDFCCBF88A033FF888" box="[784,813,1841,1866]" pageId="10" pageNumber="411" type="volume">98</part>
|
||
°33.915'E
|
||
</geoCoordinate>
|
||
,
|
||
<elevation id="0070D1F4B631FFFDFC73F88A03FCF88B" box="[936,1006,1841,1866]" metricMagnitude="2" metricUnit="m" metricValue="1.16" pageId="10" pageNumber="411" unit="m" value="116.0">
|
||
<quantity id="4CA59B22B631FFFDFC73F88A03FCF88B" box="[936,1006,1841,1866]" metricMagnitude="2" metricUnit="m" metricValue="1.16" pageId="10" pageNumber="411" unit="m" value="116.0">
|
||
<pagination id="B294C833B631FFFDFC73F88A03C3F888" box="[936,977,1841,1866]" pageId="10" pageNumber="411" type="pagination">116</pagination>
|
||
m
|
||
</quantity>
|
||
</elevation>
|
||
altitude,
|
||
<collectingDate id="EFA7E9EFB631FFFDFB8CF8890505F888" box="[1111,1303,1841,1866]" pageId="10" pageNumber="411" value="2014-10-27">27 October 2014</collectingDate>
|
||
.
|
||
</materialsCitation>
|
||
</bibRef>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB631FFFDFF4CF8EE03FFF8AC" box="[151,1005,1877,1902]" pageId="10" pageNumber="411" type="etymology">
|
||
<paragraph id="8BE236C7B631FFFDFF4CF8EE03FFF8AC" blockId="10.[151,1436,1805,2010]" box="[151,1005,1877,1902]" pageId="10" pageNumber="411">
|
||
<emphasis id="B929EAD5B631FFFDFF4CF8EE010CF8AC" bold="true" box="[151,286,1877,1902]" pageId="10" pageNumber="411">Etymology.</emphasis>
|
||
The specific name is an adjective and refers to the
|
||
<typeStatus id="54E68865B631FFFDFC87F8ED039EF8AC" box="[860,908,1878,1902]" pageId="10" pageNumber="411">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB631FFFCFF1CF8C201B6FF17" lastPageId="11" lastPageNumber="412" pageId="10" pageNumber="411" type="diagnosis">
|
||
<paragraph id="8BE236C7B631FFFCFF1CF8C201B6FF17" blockId="10.[151,1436,1805,2010]" lastBlockId="11.[151,1437,151,717]" lastPageId="11" lastPageNumber="412" pageId="10" pageNumber="411">
|
||
<bibRef id="52FDA231B631FFFCFF1CF8C201B6FF17" lastPageId="11" lastPageNumber="412" pageId="10" pageNumber="411" refId="ref11874" type="book">
|
||
<emphasis id="B929EAD5B631FFFDFF1CF8C20153F850" bold="true" box="[199,321,1913,1938]" pageId="10" pageNumber="411">
|
||
<author id="53E0DE22B631FFFDFF1CF8C2012EF850" box="[199,316,1913,1938]" firstAuthorName="Diagnosis" fns="*" inm="true" npo="*" pageId="10" pageNumber="411">Diagnosis</author>
|
||
.
|
||
</emphasis>
|
||
<title id="F959DDF1B631FFFDFE92F8C203D0F850" box="[329,962,1913,1938]" jopScore="0.106122784" pageId="10" pageNumber="411">
|
||
The species resembles
|
||
<emphasis id="B929EAD5B631FFFDFD97F8C1030DF853" box="[588,799,1913,1937]" italics="true" pageId="10" pageNumber="411">A. lakmueangensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B631FFFDFC9CF8C10349F853" box="[839,859,1914,1937]" italics="true" pageId="10" pageNumber="411">et</emphasis>
|
||
Li, 2017
|
||
</title>
|
||
(
|
||
<journalOrPublisher id="519C3C0BB631FFFDFC09F8C20425F853" box="[978,1079,1913,1938]" jopScore="0.16593583" pageId="10" pageNumber="411" type="_" typeScore="19">
|
||
Liu
|
||
<emphasis id="B929EAD5B631FFFDFBD9F8C10425F853" box="[1026,1079,1913,1937]" italics="true" pageId="10" pageNumber="411">et al</emphasis>
|
||
</journalOrPublisher>
|
||
.
|
||
<year id="FFF57794B631FFFDFB99F8C1046EF850" box="[1090,1148,1914,1938]" pageId="10" pageNumber="411">2017</year>
|
||
) and
|
||
<emphasis id="B929EAD5B631FFFDFB60F8C1054DF853" box="[1211,1375,1914,1937]" italics="true" pageId="10" pageNumber="411">A. muangensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B631FFFDFA53F8C1058EF853" box="[1416,1436,1914,1937]" italics="true" pageId="10" pageNumber="411">et</emphasis>
|
||
Li, 2017 (Liu
|
||
<emphasis id="B929EAD5B631FFFDFEE2F8250166F877" box="[313,372,1949,1973]" italics="true" pageId="10" pageNumber="411">et al.</emphasis>
|
||
2017) in having one pair of spermathecae in females, but can be distinguished by the longer stalks of spermathecae (
|
||
<figureCitation id="13662A42B631FFFDFE7EF87A021CF818" box="[421,526,1985,2010]" captionStart="FIGURE 27" captionStartId="39.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@39.[209,1378,193,1911]" captionTargetPageId="39" captionText="FIGURE 27. Althepus tharnlodensis Li et Li sp. nov., female holotype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Female habitus, retrolateral view; D. Female habitus, dorsal view; E. Female habitus, ventral view; SP = spermatheca; PP = pore plate. The scale bar in figures 27C and 27E as in figure 27D." httpUri="https://zenodo.org/record/1439844/files/figure.png" pageId="10" pageNumber="411">Fig. 27A</figureCitation>
|
||
), compared with the shorter stalks of spermathecae in
|
||
<emphasis id="B929EAD5B631FFFDFBAFF879050AF81B" box="[1140,1304,1986,2009]" italics="true" pageId="10" pageNumber="411">A. muangensis</emphasis>
|
||
, and by the larger, arch-shaped pore plates (
|
||
<figureCitation id="13662A42B630FFFCFDD2FF2C0267FF72" box="[521,629,151,176]" captionStart="FIGURE 27" captionStartId="39.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@39.[209,1378,193,1911]" captionTargetPageId="39" captionText="FIGURE 27. Althepus tharnlodensis Li et Li sp. nov., female holotype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Female habitus, retrolateral view; D. Female habitus, dorsal view; E. Female habitus, ventral view; SP = spermatheca; PP = pore plate. The scale bar in figures 27C and 27E as in figure 27D." httpUri="https://zenodo.org/record/1439844/files/figure.png" pageId="11" pageNumber="412">Fig. 27A</figureCitation>
|
||
), compared with the smaller pore plates in
|
||
<emphasis id="B929EAD5B630FFFCFBAAFF230555FF72" box="[1137,1351,152,176]" italics="true" pageId="11" pageNumber="412">A. lakmueangensis</emphasis>
|
||
and
|
||
<emphasis id="B929EAD5B630FFFCFA5FFF230109FF16" italics="true" pageId="11" pageNumber="412">A. muangensis</emphasis>
|
||
in females..
|
||
</bibRef>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB630FFFCFF1CFF640198FDA2" pageId="11" pageNumber="412" type="description">
|
||
<paragraph id="8BE236C7B630FFFCFF1CFF6400C2FDFF" blockId="11.[151,1437,151,717]" pageId="11" pageNumber="412">
|
||
<emphasis id="B929EAD5B630FFFCFF1CFF640256FF3A" bold="true" box="[199,580,223,248]" pageId="11" pageNumber="412">
|
||
Description. Female (
|
||
<typeStatus id="54E68865B630FFFCFE12FF640224FF3A" box="[457,566,223,248]" pageId="11" pageNumber="412" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 3.80; carapace 1.18 length, 1.28 width; abdomen 2.13 length, 0.90 width. Carapace round (
|
||
<figureCitation id="13662A42B630FFFCFE33FEBF0267FEDE" box="[488,629,260,285]" captionStart="FIGURE 27" captionStartId="39.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@39.[209,1378,193,1911]" captionTargetPageId="39" captionText="FIGURE 27. Althepus tharnlodensis Li et Li sp. nov., female holotype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Female habitus, retrolateral view; D. Female habitus, dorsal view; E. Female habitus, ventral view; SP = spermatheca; PP = pore plate. The scale bar in figures 27C and 27E as in figure 27D." httpUri="https://zenodo.org/record/1439844/files/figure.png" pageId="11" pageNumber="412">Figs 27C–E</figureCitation>
|
||
), yellow, with three longitudinal brown bands of different widths, the middle one wider than the others. Anterior margin of thoracic region distinctly elevated. Clypeus brown. Cheliceral promargin with one tooth, followed by a lamina, retromargin with two small teeth, posterior surface of fang with 24 small denticles (
|
||
<figureCitation id="13662A42B630FFFCFE8CFED401D6FE4A" box="[343,452,367,392]" captionStart="FIGURE 31" captionStartId="43.[151,250,1917,1939]" captionTargetBox="[177,1409,195,1896]" captionTargetId="figure@43.[177,1410,193,1896]" captionTargetPageId="43" captionText="FIGURE 31. Left cheliceral retromargin of male holotype (A–F), female holotype (G) and female paratype (H). A, Althepus phadaengensis Li et Li sp. nov.; B, A. shanhu Li et Li sp. nov.; C, A. suayaiensis Li et Li sp. nov.; D, A. tadetuensis Li et Li sp. nov.; E, A. tanhuang Li et Li sp. nov.; F, A. thanlaensis Li et Li sp. nov.; G, A. tharnlodensis Li et Li sp. nov.; H, A. viengkeoensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439852/files/figure.png" pageId="11" pageNumber="412">Fig. 31G</figureCitation>
|
||
). Labium brown. Sternum brown. Abdomen elongate, brown, with one longitudinal yellow band and two latitudinal yellow bands dorsally, and two narrow white bands ventrally. Legs brown, femur and tibia with white annulations. Leg measurements: I - (8.61, 0.53, 9.94, 22.44, -), II 22.72 (5.50, 0.50, 5.60, 9.62, 1.50), III 14.33 (4.00, 0.51, 3.75, 4.80, 1.27), IV 19.57 (5.77, 0.50, 5.51, 6.60, 1.19). Female epigyne as in
|
||
<figureCitation id="13662A42B630FFFCFAABFE6700D8FDDA" captionStart="FIGURE 27" captionStartId="39.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1911]" captionTargetId="figure@39.[209,1378,193,1911]" captionTargetPageId="39" captionText="FIGURE 27. Althepus tharnlodensis Li et Li sp. nov., female holotype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Female habitus, retrolateral view; D. Female habitus, dorsal view; E. Female habitus, ventral view; SP = spermatheca; PP = pore plate. The scale bar in figures 27C and 27E as in figure 27D." httpUri="https://zenodo.org/record/1439844/files/figure.png" pageId="11" pageNumber="412">Fig. 27A</figureCitation>
|
||
. Internal genitalia with a pair of spermathecae, curved, distal part swollen, each side with a pore plate at its base.
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B630FFFCFF1CFDFC0198FDA2" blockId="11.[151,1437,151,717]" box="[199,394,583,608]" pageId="11" pageNumber="412">
|
||
<emphasis id="B929EAD5B630FFFCFF1CFDFC011FFDA2" bold="true" box="[199,269,583,608]" pageId="11" pageNumber="412">Male:</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB630FFFCFF1CFDD702AEFD6A" pageId="11" pageNumber="412" type="distribution">
|
||
<paragraph id="8BE236C7B630FFFCFF1CFDD7034EFD46" blockId="11.[151,1437,151,717]" box="[199,860,620,645]" pageId="11" pageNumber="412">
|
||
<emphasis id="B929EAD5B630FFFCFF1CFDD70172FD47" bold="true" box="[199,352,620,645]" pageId="11" pageNumber="412">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54E68865B630FFFCFDBBFDD60282FD47" box="[608,656,621,645]" pageId="11" pageNumber="412">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13662A42B630FFFCFD21FDD7035DFD46" box="[762,847,620,645]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="11" pageNumber="412">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B630FFFCFF1CFD3402AEFD6A" blockId="11.[151,1437,151,717]" box="[199,700,655,680]" pageId="11" pageNumber="412">
|
||
<emphasis id="B929EAD5B630FFFCFF1CFD340197FD6A" bold="true" box="[199,389,655,680]" pageId="11" pageNumber="412">Natural history.</emphasis>
|
||
Collected on the cave wall.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB630FFFCFF1CFD0F017AFC99" pageId="11" pageNumber="412" type="discussion">
|
||
<paragraph id="8BE236C7B630FFFCFF1CFD0F055FFD0E" blockId="11.[151,1437,151,717]" box="[199,1357,692,717]" pageId="11" pageNumber="412">
|
||
<bibRef id="52FDA231B630FFFCFF1CFD0F055FFD0E" box="[199,1357,692,717]" pageId="11" pageNumber="412" refId="ref12236" type="book">
|
||
<emphasis id="B929EAD5B630FFFCFF1CFD0F0122FD0F" bold="true" box="[199,304,692,717]" pageId="11" pageNumber="412">
|
||
<author id="53E0DE22B630FFFCFF1CFD0F0139FD0F" box="[199,299,692,717]" firstAuthorName="Remark" fns="*" inm="true" npo="*" pageId="11" pageNumber="412">Remark</author>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B929EAD5B630FFFCFEECFD0F01FDFD0E" box="[311,495,692,716]" italics="true" pageId="11" pageNumber="412">
|
||
<title id="F959DDF1B630FFFCFEECFD0F0159FD09" box="[311,331,692,715]" jopScore="0.007833366" pageId="11" pageNumber="412">A</title>
|
||
.
|
||
<journalOrPublisher id="519C3C0BB630FFFCFE8DFD0F01FDFD0E" box="[342,495,692,716]" jopScore="0.17233406" pageId="11" pageNumber="412">tharnlodensis</journalOrPublisher>
|
||
</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B630FFFCFDCFFD0E023AFD0E" box="[532,552,693,716]" italics="true" pageId="11" pageNumber="412">et</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B630FFFCFD97FD0F02B4FD0F" bold="true" box="[588,678,692,717]" pageId="11" pageNumber="412">sp. nov.</emphasis>
|
||
was labeled as “sp. 360” in the analysis of Li and Li (
|
||
<year id="FFF57794B630FFFCFAD8FD0F0553FD0E" box="[1283,1345,692,716]" pageId="11" pageNumber="412">2018</year>
|
||
).
|
||
</bibRef>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B630FFFCFF4CFCA40266FCFB" blockId="11.[151,628,799,859]" box="[151,628,799,826]" pageId="11" pageNumber="412">
|
||
<taxonomicName id="4C5D4D44B630FFFCFF4CFCA40206FCF8" authority="Li et Li" authorityName="Li & Liu & Wongprom & Li" authorityYear="2018" box="[151,532,799,826]" class="Arachnida" family="Ochyroceratidae" genus="Althepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="viengkeoensis" status="sp. nov.">
|
||
<emphasis id="B929EAD5B630FFFCFF4CFCA40206FCF8" bold="true" box="[151,532,799,826]" pageId="11" pageNumber="412">
|
||
<emphasis id="B929EAD5B630FFFCFF4CFCA401A0FCFB" bold="true" box="[151,434,799,825]" italics="true" pageId="11" pageNumber="412">Althepus viengkeoensis</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B630FFFCFE07FCA401E3FCFB" bold="true" box="[476,497,799,825]" italics="true" pageId="11" pageNumber="412">et</emphasis>
|
||
Li
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57AEB630FFFCFDC0FC9B0266FCFB" box="[539,628,800,825]" pageId="11" pageNumber="412" rank="species">
|
||
<emphasis id="B929EAD5B630FFFCFDC0FC9B0266FCFB" bold="true" box="[539,628,800,825]" pageId="11" pageNumber="412">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B630FFFCFF4CFCF9017AFC99" blockId="11.[151,628,799,859]" box="[151,360,834,859]" pageId="11" pageNumber="412">
|
||
<figureCitation id="13662A42B630FFFCFF4CFCF90105FC99" box="[151,279,834,859]" captionStart-0="FIGURE 28" captionStart-1="FIGURE 29" captionStartId-0="40.[151,250,1933,1955]" captionStartId-1="41.[151,250,1933,1955]" captionTargetBox-0="[220,1340,193,1911]" captionTargetBox-1="[209,1378,193,1909]" captionTargetId-0="figure@40.[209,1378,193,1911]" captionTargetId-1="figure@41.[209,1378,193,1911]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionText-0="FIGURE 28. Althepus viengkeoensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 28A and 28C as in figure 28D." captionText-1="FIGURE 29. Althepus viengkeoensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 29D as in figure 29E." httpUri-0="https://zenodo.org/record/1439846/files/figure.png" httpUri-1="https://zenodo.org/record/1439848/files/figure.png" pageId="11" pageNumber="412">Figs 28–29</figureCitation>
|
||
,
|
||
<figureCitation id="13662A42B630FFFCFEF9FCF8017AFC99" box="[290,360,835,859]" captionStart-0="FIGURE 31" captionStart-1="FIGURE 32" captionStartId-0="43.[151,250,1917,1939]" captionStartId-1="44.[151,250,1148,1170]" captionTargetBox-0="[177,1409,195,1896]" captionTargetBox-1="[393,1194,195,1124]" captionTargetId-0="figure@43.[177,1410,193,1896]" captionTargetId-1="figure@44.[393,1194,195,1124]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionText-0="FIGURE 31. Left cheliceral retromargin of male holotype (A–F), female holotype (G) and female paratype (H). A, Althepus phadaengensis Li et Li sp. nov.; B, A. shanhu Li et Li sp. nov.; C, A. suayaiensis Li et Li sp. nov.; D, A. tadetuensis Li et Li sp. nov.; E, A. tanhuang Li et Li sp. nov.; F, A. thanlaensis Li et Li sp. nov.; G, A. tharnlodensis Li et Li sp. nov.; H, A. viengkeoensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." captionText-1="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri-0="https://zenodo.org/record/1439852/files/figure.png" httpUri-1="https://zenodo.org/record/1439854/files/figure.png" pageId="11" pageNumber="412">31–32</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB630FFFCFF4CFC3102DFFC29" pageId="11" pageNumber="412" type="materials_examined">
|
||
<paragraph id="8BE236C7B630FFFCFF4CFC3102DFFC29" blockId="11.[151,1437,906,1759]" pageId="11" pageNumber="412">
|
||
<bibRef id="52FDA231B630FFFCFF4CFC3102DFFC29" pageId="11" pageNumber="412" refId="ref12284" type="journal article">
|
||
<materialsCitation id="3B353C9AB630FFFCFF4CFC3104E1FC05" ID-GBIF-Occurrence="1913888987" collectingDate="2012-12-02" country="Laos" county="Lom Cave" elevation="314" latitude="18.957634" location="Vien Tiane Province" longLatPrecision="1" longitude="102.437416" municipality="North of Vieng Keo Village" pageId="11" pageNumber="412" specimenCount="2" typeStatus="holotype">
|
||
<title id="F959DDF1B630FFFCFF4CFC31021AFC61" box="[151,520,906,931]" jopScore="2.3790642E-4" pageId="11" pageNumber="412">
|
||
<emphasis id="B929EAD5B630FFFCFF4CFC3101DAFC61" bold="true" box="[151,456,906,931]" pageId="11" pageNumber="412">
|
||
<typeStatus id="54E68865B630FFFCFF4CFC3100C0FC61" box="[151,210,906,931]" pageId="11" pageNumber="412">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E68865B630FFFCFE89FC3101D1FC61" box="[338,451,906,931]" pageId="11" pageNumber="412" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male
|
||
</title>
|
||
(
|
||
<journalOrPublisher id="519C3C0BB630FFFCFDC1FC300286FC61" box="[538,660,907,931]" jopScore="4.7581285E-4" pageId="11" pageNumber="412">IZCAS-Ar</journalOrPublisher>
|
||
<part id="FFF99C5AB630FFFCFD45FC3002FAFC61" box="[670,744,907,931]" pageId="11" pageNumber="412" type="volume">34031</part>
|
||
),
|
||
<author id="53E0DE22B630FFFCFD24FC300427FC61" box="[767,1077,906,931]" case="TC" fn="Vien Tiane Province" fnc="TC" fns="N" ln="Laos" lnc="TC" npo="LnFn" pageId="11" pageNumber="412">
|
||
<collectingCountry id="F34A7657B630FFFCFD24FC30032AFC60" box="[767,824,907,930]" name="Laos" pageId="11" pageNumber="412">Laos</collectingCountry>
|
||
,
|
||
<location id="8E82601CB630FFFCFC9DFC310427FC61" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB630FFFCFC9DFC310427FC61" box="[838,1077,906,931]" country="Laos" county="Lom Cave" latitude="18.957634" longLatPrecision="1" longitude="102.437416" municipality="North of Vieng Keo Village" name="Vien Tiane Province" pageId="11" pageNumber="412">Vien Tiane Province</location>
|
||
</author>
|
||
,
|
||
<location id="8E82601CB630FFFCFB98FC300539FC61" LSID="urn:lsid:plazi:treatment:03F487D1B63AFFFCFF4CF9EF0550F91D:8E82601CB630FFFCFB98FC300539FC61" box="[1091,1323,906,931]" country="Laos" county="Lom Cave" latitude="18.957634" longLatPrecision="1" longitude="102.437416" municipality="North of Vieng Keo Village" name="Vang Vieng District" pageId="11" pageNumber="412">
|
||
<author id="53E0DE22B630FFFCFB98FC300539FC61" box="[1091,1323,906,931]" case="TC" fn="Vang Vieng" fnc="TC" fns="N" ln="District" lnc="TC" npo="FnLn" pageId="11" pageNumber="412">Vang Vieng District</author>
|
||
</location>
|
||
,
|
||
<author id="53E0DE22B630FFFCFAE2FC3101EDFC05" case="TC" fn="Lom Cave" fnc="TC" fns="N" ln="North of Vieng Keo Village" lnc="TC" npo="LnFn" pageId="11" pageNumber="412">
|
||
<collectingMunicipality id="6B86ACBDB630FFFCFAE2FC310161FC05" pageId="11" pageNumber="412">North of Vieng Keo Village</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62834E4BB630FFFCFE59FC1401EDFC05" box="[386,511,943,967]" pageId="11" pageNumber="412">Lom Cave</collectingCounty>
|
||
</author>
|
||
,
|
||
<geoCoordinate id="EE695000B630FFFCFDD4FC15028FFC05" box="[527,669,942,967]" direction="north" orientation="latitude" pageId="11" pageNumber="412" precision="1" value="18.957634">18°57.458'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695000B630FFFCFD77FC150357FC05" box="[684,837,942,967]" direction="east" orientation="longitude" pageId="11" pageNumber="412" precision="1" value="102.437416">102°26.245'E</geoCoordinate>
|
||
,
|
||
<elevation id="0070D1F4B630FFFCFC8FFC1403B2FC04" box="[852,928,943,967]" metricMagnitude="2" metricUnit="m" metricValue="3.1399999999999997" pageId="11" pageNumber="412" unit="m" value="314.0">
|
||
<quantity id="4CA59B22B630FFFCFC8FFC1403B2FC04" box="[852,928,943,967]" metricMagnitude="2" metricUnit="m" metricValue="3.1399999999999997" pageId="11" pageNumber="412" unit="m" value="314.0">314 m</quantity>
|
||
</elevation>
|
||
altitude, 0
|
||
<collectingDate id="EFA7E9EFB630FFFCFBC4FC1404E1FC05" box="[1055,1267,942,967]" pageId="11" pageNumber="412" value="2012-12-02">2 December 2012</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B353C9AB630FFFCFADAFC1502DFFC29" ID-GBIF-Occurrence="1913888969" collectingDate="2012-12-02" country="Laos" county="Lom Cave" elevation="314" latitude="18.957634" location="Vien Tiane Province" longLatPrecision="1" longitude="102.437416" municipality="North of Vieng Keo Village" pageId="11" pageNumber="412" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<emphasis id="B929EAD5B630FFFCFADAFC150591FC05" bold="true" box="[1281,1411,942,967]" pageId="11" pageNumber="412">
|
||
<typeStatus id="54E68865B630FFFCFADAFC15056CFC05" box="[1281,1406,942,967]" pageId="11" pageNumber="412" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D5BFD4EB630FFFCFA56FC1400F4FC29" pageId="11" pageNumber="412" type="female">1 female</specimenCount>
|
||
, same data as holotype (IZCAS-Ar
|
||
<pagination id="B294C833B630FFFCFDADFC6802D3FC29" box="[630,705,979,1003]" pageId="11" pageNumber="412" type="pagination">34032</pagination>
|
||
).
|
||
</materialsCitation>
|
||
</bibRef>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB630FFFCFF1CFC4D055AFBCC" box="[199,1352,1014,1039]" pageId="11" pageNumber="412" type="etymology">
|
||
<paragraph id="8BE236C7B630FFFCFF1CFC4D055AFBCC" blockId="11.[151,1437,906,1759]" box="[199,1352,1014,1039]" pageId="11" pageNumber="412">
|
||
<emphasis id="B929EAD5B630FFFCFF1CFC4D015CFBCD" bold="true" box="[199,334,1014,1039]" pageId="11" pageNumber="412">Etymology.</emphasis>
|
||
The specific name is an adjective and refers to the
|
||
<typeStatus id="54E68865B630FFFCFC57FC4C03AEFBCD" box="[908,956,1015,1039]" pageId="11" pageNumber="412">type</typeStatus>
|
||
locality; adjective in genitive case.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB630FFFCFF1CFBA104ADFB95" pageId="11" pageNumber="412" type="diagnosis">
|
||
<paragraph id="8BE236C7B630FFFCFF1CFBA104ADFB95" blockId="11.[151,1437,906,1759]" pageId="11" pageNumber="412">
|
||
<emphasis id="B929EAD5B630FFFCFF1CFBA10153FBF1" bold="true" box="[199,321,1050,1075]" pageId="11" pageNumber="412">Diagnosis.</emphasis>
|
||
This new species can be easily distinguished from all other species by the depigmented carapace and a sigmoid conductor (
|
||
<figureCitation id="13662A42B630FFFCFE61FB850232FB95" box="[442,544,1086,1111]" captionStart="FIGURE 28" captionStartId="40.[151,250,1933,1955]" captionTargetBox="[220,1340,193,1911]" captionTargetId="figure@40.[209,1378,193,1911]" captionTargetPageId="40" captionText="FIGURE 28. Althepus viengkeoensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 28A and 28C as in figure 28D." httpUri="https://zenodo.org/record/1439846/files/figure.png" pageId="11" pageNumber="412">Fig. 28A</figureCitation>
|
||
) in males and one pair of duct-like spermathecae in female.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB630FFFCFF1CFBD902C2F9B1" pageId="11" pageNumber="412" type="description">
|
||
<paragraph id="8BE236C7B630FFFCFF1CFBD90555FA21" blockId="11.[151,1437,906,1759]" pageId="11" pageNumber="412">
|
||
<emphasis id="B929EAD5B630FFFCFF1CFBD90237FBB9" bold="true" box="[199,549,1122,1147]" pageId="11" pageNumber="412">
|
||
Description. Male (
|
||
<typeStatus id="54E68865B630FFFCFE71FBD90205FBB9" box="[426,535,1122,1147]" pageId="11" pageNumber="412" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 4.60; carapace 1.62 length, 1.80 width; abdomen 2.50 length, 1.63 width. Carapace rounded (
|
||
<figureCitation id="13662A42B630FFFCFE1CFB3D0239FB5D" box="[455,555,1158,1183]" captionStart="FIGURE 29" captionStartId="41.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1909]" captionTargetId="figure@41.[209,1378,193,1911]" captionTargetPageId="41" captionText="FIGURE 29. Althepus viengkeoensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 29D as in figure 29E." httpUri="https://zenodo.org/record/1439848/files/figure.png" pageId="11" pageNumber="412">Fig 29A</figureCitation>
|
||
), light yellow, with three longitudinal brown bands, the middle shortened to form a triangular band around the eyes. Anterior margin of thoracic region distinctly elevated. Clypeus brown. Cheliceral promargin with two teeth, followed by a lamina, retromargin with two small teeth, posterior surface of fang with 22 small denticles (
|
||
<figureCitation id="13662A42B630FFFCFE23FB490277FAC9" box="[504,613,1266,1291]" captionStart="FIGURE 31" captionStartId="43.[151,250,1917,1939]" captionTargetBox="[177,1409,195,1896]" captionTargetId="figure@43.[177,1410,193,1896]" captionTargetPageId="43" captionText="FIGURE 31. Left cheliceral retromargin of male holotype (A–F), female holotype (G) and female paratype (H). A, Althepus phadaengensis Li et Li sp. nov.; B, A. shanhu Li et Li sp. nov.; C, A. suayaiensis Li et Li sp. nov.; D, A. tadetuensis Li et Li sp. nov.; E, A. tanhuang Li et Li sp. nov.; F, A. thanlaensis Li et Li sp. nov.; G, A. tharnlodensis Li et Li sp. nov.; H, A. viengkeoensis Li et Li sp. nov. PT = promarginal teeth; RT = retromarginal teeth; CL = cheliceral lamina." httpUri="https://zenodo.org/record/1439852/files/figure.png" pageId="11" pageNumber="412">Fig. 31H</figureCitation>
|
||
). Labium brown, with two light, round areas basally. Sternum brown. Abdomen elongate, brown, with a yellow humanoid streak dorsally and two yellow bands ventrally. Legs brown. Leg measurements: I missing, II missing, III - (6.54, -, -, -, -), IV - (8.59, 0.68, 9.68, -, -). Male palp (
|
||
<figureCitation id="13662A42B630FFFCFADBFA810582FA91" box="[1280,1424,1338,1363]" captionStart="FIGURE 28" captionStartId="40.[151,250,1933,1955]" captionTargetBox="[220,1340,193,1911]" captionTargetId="figure@40.[209,1378,193,1911]" captionTargetPageId="40" captionText="FIGURE 28. Althepus viengkeoensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 28A and 28C as in figure 28D." httpUri="https://zenodo.org/record/1439846/files/figure.png" pageId="11" pageNumber="412">Figs 28A–D</figureCitation>
|
||
): tarsus with three slightly curved, serrated bristles at the top (
|
||
<figureCitation id="13662A42B630FFFCFC9BFAE503B4FAB5" box="[832,934,1374,1399]" captionStart="FIGURE 28" captionStartId="40.[151,250,1933,1955]" captionTargetBox="[220,1340,193,1911]" captionTargetId="figure@40.[209,1378,193,1911]" captionTargetPageId="40" captionText="FIGURE 28. Althepus viengkeoensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 28A and 28C as in figure 28D." httpUri="https://zenodo.org/record/1439846/files/figure.png" pageId="11" pageNumber="412">Fig. 28A</figureCitation>
|
||
) and one hooked spine slightly bent with tip directed proximally (
|
||
<figureCitation id="13662A42B630FFFCFE53FA3901E6FA59" box="[392,500,1410,1435]" captionStart="FIGURE 28" captionStartId="40.[151,250,1933,1955]" captionTargetBox="[220,1340,193,1911]" captionTargetId="figure@40.[209,1378,193,1911]" captionTargetPageId="40" captionText="FIGURE 28. Althepus viengkeoensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 28A and 28C as in figure 28D." httpUri="https://zenodo.org/record/1439846/files/figure.png" pageId="11" pageNumber="412">Fig. 28A</figureCitation>
|
||
); bulb light yellow, ovate; embolus arising retrolatero-distally from bulb, slightly curved distally; conductor arising prolatero-distally from bulb, slightly curved, with distal part curling until it nearly closes (
|
||
<figureCitation id="13662A42B630FFFCFEECFA7101D7FA21" box="[311,453,1482,1507]" captionStart="FIGURE 28" captionStartId="40.[151,250,1933,1955]" captionTargetBox="[220,1340,193,1911]" captionTargetId="figure@40.[209,1378,193,1911]" captionTargetPageId="40" captionText="FIGURE 28. Althepus viengkeoensis Li et Li sp. nov., left pedipalp of male holotype. A. Palp, ventral view; B. Palpal bulb, ventral view; C. Palp, prolateral view; D. Palp, retrolateral view. CO = conductor; EM = embolus; CP = cymbial protrusion; HS = hook-shaped spine; SB = serrated bristles. The scale bar in figures 28A and 28C as in figure 28D." httpUri="https://zenodo.org/record/1439846/files/figure.png" pageId="11" pageNumber="412">Figs 28A–B</figureCitation>
|
||
); embolus and conductor widely separated (distance less than diameter of bulb).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B630FFFCFF1CFA5502C2F9B1" blockId="11.[151,1437,906,1759]" pageId="11" pageNumber="412">
|
||
<emphasis id="B929EAD5B630FFFCFF1CFA5501BFF9C5" bold="true" box="[199,429,1518,1543]" pageId="11" pageNumber="412">
|
||
Female (
|
||
<typeStatus id="54E68865B630FFFCFEF5FA5501B2F9C5" box="[302,416,1518,1543]" pageId="11" pageNumber="412" type="paratype">paratype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Similar to male in color and general features (
|
||
<figureCitation id="13662A42B630FFFCFC1EFA550442F9C5" box="[965,1104,1518,1543]" captionStart="FIGURE 29" captionStartId="41.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1909]" captionTargetId="figure@41.[209,1378,193,1911]" captionTargetPageId="41" captionText="FIGURE 29. Althepus viengkeoensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 29D as in figure 29E." httpUri="https://zenodo.org/record/1439848/files/figure.png" pageId="11" pageNumber="412">Figs 29D–E</figureCitation>
|
||
) but larger. Internal genitalia with a pair of spermathecae, slender, transparent (
|
||
<figureCitation id="13662A42B630FFFCFD1BF9A90335F9E9" box="[704,807,1554,1579]" captionStart="FIGURE 29" captionStartId="41.[151,250,1933,1955]" captionTargetBox="[209,1378,193,1909]" captionTargetId="figure@41.[209,1378,193,1911]" captionTargetPageId="41" captionText="FIGURE 29. Althepus viengkeoensis Li et Li sp. nov., male holotype and female paratype. A. Internal genitalia, dorsal view; B. Female epigastric furrow, ventral view; C. Male habitus, dorsal view; D. Female habitus, dorsal view; E. Female habitus, ventral view. SP = spermatheca. The scale bar in figure 29D as in figure 29E." httpUri="https://zenodo.org/record/1439848/files/figure.png" pageId="11" pageNumber="412">Fig. 29A</figureCitation>
|
||
). Measurements: total length 4.84; carapace 1.50 length, 1.48 width; abdomen 2.60 length, 2.03 width. Leg measurements: I missing, II missing, III 18.22 (5.45, 0.65, 4.50, 6.09, 1.53), IV 24.47 (7.18, 0.67, 7.05, 7.69, 1.88).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB630FFFCFF1CF9C5032CF978" pageId="11" pageNumber="412" type="distribution">
|
||
<paragraph id="8BE236C7B630FFFCFF1CF9C5034EF955" blockId="11.[151,1437,906,1759]" box="[199,860,1662,1687]" pageId="11" pageNumber="412">
|
||
<emphasis id="B929EAD5B630FFFCFF1CF9C50172F955" bold="true" box="[199,352,1662,1687]" pageId="11" pageNumber="412">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54E68865B630FFFCFDBBF9C40282F955" box="[608,656,1663,1687]" pageId="11" pageNumber="412">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13662A42B630FFFCFD21F9C5035DF955" box="[762,847,1662,1687]" captionStart="FIGURE 32" captionStartId="44.[151,250,1148,1170]" captionTargetBox="[393,1194,195,1124]" captionTargetId="figure@44.[393,1194,195,1124]" captionTargetPageId="44" captionText="FIGURE 32. Known distribution of new Althepus species from Southeast Asia. 1, A. shanhu Li et Li sp. nov.; 2, A. natmataungensis Li et Li sp. nov.; 3, A. menglaensis Li et Li sp. nov.; 4, A. naphongensis Li et Li sp. nov.; 5, A. phadaengensis Li et Li sp. nov.; 6, A. changmao Li et Li sp. nov.; 7, A. viengkeoensis Li et Li sp. nov.; 8, A. maechamensis Li et Li sp. nov.; 9, A. bamensis Li et Li sp. nov.; 10, A. suayaiensis Li et Li sp. nov.; 11, A. thanlaensis Li et Li sp. nov.; 12, A. tadetuensis Li et Li sp. nov.; 13, A. tanhuang Li et Li sp. nov.; 14, A. dongnaiensis Li et Li sp. nov.; 15, A. tharnlodensis Li et Li sp. nov.; 16, A. guan Li et Li sp. nov." httpUri="https://zenodo.org/record/1439854/files/figure.png" pageId="11" pageNumber="412">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236C7B630FFFCFF1CF919032CF978" blockId="11.[151,1437,906,1759]" box="[199,830,1698,1723]" pageId="11" pageNumber="412">
|
||
<emphasis id="B929EAD5B630FFFCFF1CF9190197F979" bold="true" box="[199,389,1698,1723]" pageId="11" pageNumber="412">Natural history.</emphasis>
|
||
Collected on the wall deep in the cave.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347654CB630FFFCFF1CF97D0550F91D" box="[199,1346,1734,1759]" pageId="11" pageNumber="412" type="discussion">
|
||
<paragraph id="8BE236C7B630FFFCFF1CF97D0550F91D" blockId="11.[151,1437,906,1759]" box="[199,1346,1734,1759]" pageId="11" pageNumber="412">
|
||
<bibRef id="52FDA231B630FFFCFF1CF97D0550F91D" box="[199,1346,1734,1759]" pageId="11" pageNumber="412" refId="ref12798" type="book">
|
||
<emphasis id="B929EAD5B630FFFCFF1CF97D0122F91D" bold="true" box="[199,304,1734,1759]" pageId="11" pageNumber="412">
|
||
<author id="53E0DE22B630FFFCFF1CF97D0139F91D" box="[199,299,1734,1759]" firstAuthorName="Remark" fns="*" inm="true" npo="*" pageId="11" pageNumber="412">Remark</author>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B929EAD5B630FFFCFEECF97C01E0F91C" box="[311,498,1734,1758]" italics="true" pageId="11" pageNumber="412">
|
||
<title id="F959DDF1B630FFFCFEECF97C0159F91C" box="[311,331,1735,1758]" jopScore="0.007833366" pageId="11" pageNumber="412">A</title>
|
||
.
|
||
<journalOrPublisher id="519C3C0BB630FFFCFE8DF97D01E0F91C" box="[342,498,1734,1758]" jopScore="0.17233406" pageId="11" pageNumber="412">viengkeoensis</journalOrPublisher>
|
||
</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B630FFFCFDCCF97C0238F91C" box="[535,554,1735,1758]" italics="true" pageId="11" pageNumber="412">et</emphasis>
|
||
Li
|
||
<emphasis id="B929EAD5B630FFFCFD94F97D02BBF91D" bold="true" box="[591,681,1734,1759]" pageId="11" pageNumber="412">sp. nov.</emphasis>
|
||
was labeled as “sp. 22” in the analysis of Li and Li (
|
||
<year id="FFF57794B630FFFCFB23F97C0524F91D" box="[1272,1334,1735,1759]" pageId="11" pageNumber="412">2018</year>
|
||
).
|
||
</bibRef>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |