treatments-xml/data/03/F4/87/03F487D0A37A4D5EA2DBFF43FD75EB71.xml

<document ID-DOI="10.11646/zootaxa.5277.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7889695" ID-ZooBank="73ED69F3-3887-460D-86DE-C9F1302C8EC3" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1683099613832" checkinUser="plazi" docAuthor="Kluge, Nikita, Srinivasan, Pandiarajan, Sivaruban, T., Barathy, S. &amp; Isack, Rajasekaran" docDate="2023" docId="03F487D0A37A4D5EA2DBFF43FD75EB71" docLanguage="en" docName="zootaxa.5277.2.1.pdf" docOrigin="Zootaxa 5277 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5277.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Baetis (Tenuibaetis) hissaricus Novikova 1991" docType="treatment" docVersion="3" lastPageNumber="217" masterDocId="FFCDFFA8A3764D4EA24CFFD4FFABE864" masterDocTitle="Contribution to the knowledge of the subgenus Tenuibaetis Kang &amp; Yang 1994 (Ephemeroptera, Baetidae, Baetis s. l.)" masterLastPageNumber="258" masterPageNumber="201" pageNumber="213" updateTime="1683138298321" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title>Contribution to the knowledge of the subgenus Tenuibaetis Kang &amp; Yang 1994 (Ephemeroptera, Baetidae, Baetis s. l.)</mods:title>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kluge, Nikita</mods:namePart>
<mods:affiliation>Department of Entomology, Saint Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia. Website: http: // insecta. bio. spbu. ru</mods:affiliation>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Srinivasan, Pandiarajan</mods:namePart>
<mods:affiliation>PG and Research department of Zoology, the American College, Madurai- 625002, India</mods:affiliation>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sivaruban, T.</mods:namePart>
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<mods:affiliation>PG and Research department of Zoology, the American College, Madurai- 625002, India &amp; sivaruban 270 @ gmail. com; https: // orcid. org / 0000 - 0001 - 8997 - 9355</mods:affiliation>
<mods:nameIdentifier type="email">sivaruban270@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Barathy, S.</mods:namePart>
<mods:affiliation>Department of Zoology, Fatima College, Madurai- 625018, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Isack, Rajasekaran</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-9952-4335</mods:nameIdentifier>
<mods:affiliation>PG and Research department of Zoology, the American College, Madurai- 625002, India &amp; iceisack 143 @ gmail. com; https: // orcid. org / 0000 - 0002 - 9952 - 4335</mods:affiliation>
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<mods:date>2023</mods:date>
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<mods:number>2023-05-02</mods:number>
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<mods:number>2</mods:number>
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<subSubSection box="[151,744,151,178]" pageId="12" pageNumber="213" type="nomenclature">
<paragraph blockId="12.[151,744,151,214]" box="[151,744,151,178]" pageId="12" pageNumber="213">
<heading bold="true" box="[151,744,151,178]" fontSize="11" level="1" pageId="12" pageNumber="213" reason="1">
<emphasis bold="true" box="[151,744,151,178]" pageId="12" pageNumber="213">
2. 
<taxonomicName authority="Novikova 1991" authorityName="Novikova" authorityYear="1991" box="[181,744,151,178]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="hissaricus" subGenus="Tenuibaetis">
<emphasis bold="true" box="[181,256,151,177]" italics="true" pageId="12" pageNumber="213">Baetis</emphasis>
(
<emphasis bold="true" box="[273,412,151,178]" italics="true" pageId="12" pageNumber="213">Tenuibaetis</emphasis>
) 
<emphasis bold="true" box="[429,551,151,178]" italics="true" pageId="12" pageNumber="213">hissaricus</emphasis>
<bibRefCitation author="Novikova, E. A." box="[558,744,151,177]" pageId="12" pageNumber="213" pagination="793 - 795" refId="ref19156" refString="Novikova, E. A. (1991) [A new subspecies of mayflies Baetis ursinus hissaricus subsp. n. (Ephemeroptera, Baetidae) from Tadjikistan]. Entomologicheskoe Obozrenie, 70 (4), 793 - 795." type="journal article" year="1991">Novikova 1991</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,295,187,214]" pageId="12" pageNumber="213" type="description">
<paragraph blockId="12.[151,744,151,214]" box="[151,295,187,214]" pageId="12" pageNumber="213">
(
<figureCitation box="[159,288,187,214]" captionStart-0="FIGURES 44–46" captionStart-1="FIGURES 47–49" captionStart-2="FIGURES 50–51" captionStart-3="FIGURES 52–59" captionStartId-0="12.[151,264,1987,2011]" captionStartId-1="13.[151,264,1964,1988]" captionStartId-2="14.[151,264,1999,2023]" captionStartId-3="15.[151,264,1834,1858]" captionTargetBox-0="[169,1418,365,1963]" captionTargetBox-1="[151,1436,309,1941]" captionTargetBox-2="[224,1363,524,1975]" captionTargetBox-3="[151,1435,181,1810]" captionTargetId-0="figure-86@12.[169,1418,365,1963]" captionTargetId-1="figure-97@13.[151,1436,309,1941]" captionTargetId-2="figure-186@14.[224,1363,524,1975]" captionTargetId-3="figure-19@15.[151,1435,181,1810]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionText-0="FIGURES 44–46. Baetis (Tenuibaetis) hissaricus, larval exuviae. 44, head; 45, labrum; 46, labium (44, 46, holotype)." captionText-1="FIGURES 47–49. Baetis (Tenuibaetis) hissaricus, exuviae of larval fore, middle and hind femora. Abbreviation: 2-can, twochannel setae." captionText-2="FIGURES 50–51. Baetis (Tenuibaetis) hissaricus, exuviae of larval hind tibia. Abbreviations: 2-can, two-channel setae." captionText-3="FIGURES 52–59. Baetis (Tenuibaetis) hissaricus. 52–53, fore and middle legs of male imago; 54–55, genitalia of males imago; 56, hind wing of male; 57, hind wing of female; 58, subimaginal exuviae of female fore leg; 59, subimaginal exuviae of half of mesonotum (57, 58, holotype)." figureDoi-0="http://doi.org/10.5281/zenodo.7889721" figureDoi-1="http://doi.org/10.5281/zenodo.7889725" figureDoi-2="http://doi.org/10.5281/zenodo.7889729" figureDoi-3="http://doi.org/10.5281/zenodo.7889731" httpUri-0="https://zenodo.org/record/7889721/files/figure.png" httpUri-1="https://zenodo.org/record/7889725/files/figure.png" httpUri-2="https://zenodo.org/record/7889729/files/figure.png" httpUri-3="https://zenodo.org/record/7889731/files/figure.png" pageId="12" pageNumber="213">Figs 44–59</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="213" type="reference_group">
<paragraph blockId="12.[151,1364,255,341]" box="[151,1364,255,282]" pageId="12" pageNumber="213">
<treatmentCitationGroup box="[151,1364,255,282]" pageId="12" pageNumber="213">
<treatmentCitation author="Novikova, E. A." box="[151,619,256,280]" page="793" pageId="12" pageNumber="213" year="1991">
<taxonomicName authority="Novikova 1991: 793" authorityName="Novikova" authorityPageNumber="793" authorityYear="1991" box="[151,619,256,280]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="subSpecies" species="ursinus" subSpecies="hissaricus">
<emphasis box="[151,405,256,280]" italics="true" pageId="12" pageNumber="213">Baetis ursinus hissaricus</emphasis>
<bibRefCitation author="Novikova, E. A." box="[411,619,256,280]" pageId="12" pageNumber="213" pagination="793 - 795" refId="ref19156" refString="Novikova, E. A. (1991) [A new subspecies of mayflies Baetis ursinus hissaricus subsp. n. (Ephemeroptera, Baetidae) from Tadjikistan]. Entomologicheskoe Obozrenie, 70 (4), 793 - 795." type="journal article" year="1991">Novikova 1991: 793</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(larva, ♁ imago, 
<specimenCount box="[799,818,256,279]" pageId="12" pageNumber="213" type="female">♀</specimenCount>
imago, subimago); 
<treatmentCitation author="Kluge, N. J." box="[1020,1178,255,280]" page="12" pageId="12" pageNumber="213" year="1995">
<bibRefCitation author="Kluge, N. J." box="[1020,1178,255,280]" pageId="12" pageNumber="213" refId="ref18061" refString="Kluge, N. J. (1995) A catalogue of the type-specimens in the collection of the Zoological Institute, Russian Academy of Sciences. Insecta, Ephemeroptera. Zoological Institute of Russian Academy of Sciences, St. Petersburg, 52 pp. [in Russian]" type="book" year="1995">Kluge 1995: 12</bibRefCitation>
</treatmentCitation>
(
<typeStatus box="[1191,1236,256,280]" pageId="12" pageNumber="213">type</typeStatus>
specimens).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="12.[151,1364,255,341]" box="[151,768,286,310]" pageId="12" pageNumber="213">
<treatmentCitationGroup box="[151,768,286,310]" pageId="12" pageNumber="213">
<treatmentCitation author="Kluge, N. J." box="[151,687,286,310]" page="194" pageId="12" pageNumber="213" year="1997">
<taxonomicName authority=": Kluge 1997 b: 194" authorityName="Kluge" authorityPageNumber="194" authorityYear="1997" box="[151,687,286,310]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="subSpecies" species="ursinus" subGenus="Baetis" subSpecies="hissaricus">
<emphasis box="[151,214,287,310]" italics="true" pageId="12" pageNumber="213">Baetis</emphasis>
(
<emphasis box="[229,292,287,310]" italics="true" pageId="12" pageNumber="213">Baetis</emphasis>
) 
<emphasis box="[307,491,286,310]" italics="true" pageId="12" pageNumber="213">ursinus hissaricus</emphasis>
: 
<bibRefCitation author="Kluge, N. J." box="[504,687,286,310]" pageId="12" pageNumber="213" refId="ref18180" refString="Kluge, N. J. (1997 b) Order Ephemeroptera. In: Tsalolikhin, S. J. (Ed.), Key to freshwater invertebrates of Russia and adjacent lands. Vol. 3. Nauka, S-Petersburg, pp. 176 - 220." type="book" year="1997">Kluge 1997b: 194</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(larva).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="12.[151,1364,255,341]" box="[151,775,317,341]" pageId="12" pageNumber="213">
<taxonomicName authority="Novikova 1991" authorityName="Novikova" authorityYear="1991" box="[151,646,317,341]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="hissaricus" subGenus="Tenuibaetis">
<emphasis box="[151,214,318,341]" italics="true" pageId="12" pageNumber="213">Baetis</emphasis>
(
<emphasis box="[229,345,317,341]" italics="true" pageId="12" pageNumber="213">Tenuibaetis</emphasis>
) 
<emphasis box="[360,463,317,341]" italics="true" pageId="12" pageNumber="213">hissaricus</emphasis>
: Kluge 2022: 166
</taxonomicName>
(subimago).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889721" ID-Zenodo-Dep="7889721" box="[151,1352,1987,2011]" httpUri="https://zenodo.org/record/7889721/files/figure.png" pageId="12" pageNumber="213" startId="12.[151,264,1987,2011]" targetBox="[169,1418,365,1963]" targetPageId="12" targetType="figure">
<paragraph blockId="12.[151,1352,1987,2011]" box="[151,1352,1987,2011]" pageId="12" pageNumber="213">
<emphasis bold="true" box="[151,340,1987,2011]" pageId="12" pageNumber="213">FIGURES 44–46.</emphasis>
<taxonomicName authorityName=": Kluge" authorityYear="2022" box="[346,658,1987,2011]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="hissaricus" subGenus="Tenuibaetis">
<emphasis box="[346,409,1988,2011]" italics="true" pageId="12" pageNumber="213">Baetis</emphasis>
(
<emphasis box="[424,540,1987,2011]" italics="true" pageId="12" pageNumber="213">Tenuibaetis</emphasis>
) 
<emphasis box="[555,658,1987,2011]" italics="true" pageId="12" pageNumber="213">hissaricus</emphasis>
</taxonomicName>
, larval exuviae. 44, head; 45, labrum; 46, labium (44, 46, holotype).
</paragraph>
</caption>
<subSubSection lastPageId="16" lastPageNumber="217" pageId="13" pageNumber="214" type="materials_examined">
<paragraph blockId="13.[151,1436,150,287]" pageId="13" pageNumber="214">
<emphasis bold="true" box="[151,381,150,176]" pageId="13" pageNumber="214">Material examined</emphasis>
(
<collectionCode box="[399,450,151,177]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="13" pageNumber="214" type="Museum">ZIN</collectionCode>
, 
<typeStatus box="[465,563,151,177]" pageId="13" pageNumber="214">holotype</typeStatus>
and 
<typeStatus box="[624,737,151,177]" pageId="13" pageNumber="214">paratypes</typeStatus>
). 
<materialsCitation collectingDate="1981-08-15" collectionCode="VIII" collectorName="N. Kluge" country="Tajikistan" location="Garm" pageId="13" pageNumber="214" specimenCount="18" specimenCount-larva="18">
<collectingCountry box="[758,922,151,177]" name="Tajikistan" pageId="13" pageNumber="214">TAJIKISTAN</collectingCountry>
: 
<location LSID="urn:lsid:plazi:treatment:03F487D0A37A4D5EA2DBFF43FD75EB71:8E82601DA37B4D43A1E1FF43FC5BE8D5" box="[941,1008,151,177]" country="Tajikistan" name="Garm" pageId="13" pageNumber="214">Garm</location>
, small stream, 
<date box="[1186,1334,150,177]" pageId="13" pageNumber="214" value="1981-08-15">
<collectingDate box="[1186,1334,150,177]" pageId="13" pageNumber="214" value="1981-08-15">
15. 
<collectionCode box="[1221,1274,151,177]" pageId="13" pageNumber="214">VIII</collectionCode>
.1981
</collectingDate>
</date>
, coll. 
<collectorName pageId="13" pageNumber="214">N. Kluge</collectorName>
: 
<specimenCount box="[239,351,187,213]" pageId="13" pageNumber="214" type="larva">18 larvae</specimenCount>
</materialsCitation>
; 
<materialsCitation collectingDate="1981-08-15" collectingDateMax="1981-08-22" collectingDateMin="1981-08-15" collectionCode="VIII" collectorName="N. Kluge &amp; E. Novikova" country="Tajikistan" location="river Sardai-Miona" pageId="13" pageNumber="214" specimenCount="16" specimenCount-female="1" specimenCount-larva="15">
basin of 
<location LSID="urn:lsid:plazi:treatment:03F487D0A37A4D5EA2DBFF43FD75EB71:8E82601DA37B4D43A395FF6FFD18E8B1" box="[473,691,187,213]" country="Tajikistan" name="river Sardai-Miona" pageId="13" pageNumber="214">river Sardai-Miona</location>
<locationDeviation box="[703,976,186,213]" location="Ramit" pageId="13" pageNumber="214">
<quantity box="[703,779,186,213]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="214" unit="km" value="15.0">15 km</quantity>
upstream Ramit
</locationDeviation>
, 
<date box="[992,1182,186,213]" pageId="13" pageNumber="214" value="1981-08-15" valueMax="1981-08-22" valueMin="1981-08-15">
<collectingDate box="[992,1182,186,213]" pageId="13" pageNumber="214" value="1981-08-15" valueMax="1981-08-22" valueMin="1981-08-15">
15–22. 
<collectionCode box="[1069,1122,187,213]" pageId="13" pageNumber="214">VIII</collectionCode>
.1981
</collectingDate>
</date>
, coll. 
<collectorName box="[1259,1366,187,213]" pageId="13" pageNumber="214">N. Kluge</collectorName>
&amp; 
<collectorName pageId="13" pageNumber="214">E. Novikova</collectorName>
: 1 L-S♁, 3 L-S 
<specimenCount box="[456,471,223,249]" pageId="13" pageNumber="214" type="female">♀</specimenCount>
, 
<specimenCount box="[486,596,222,249]" pageId="13" pageNumber="214" type="larva">15 larvae</specimenCount>
</materialsCitation>
; 
<materialsCitation collectingDate="1981-08-18" collectingDateMax="1981-08-21" collectingDateMin="1981-08-18" collectionCode="VIII" collectorName="N. Kluge &amp; E. Novikova" country="Tajikistan" county="river Sardai-Miona" location="valley Divdara" municipality="Zangob" pageId="13" pageNumber="214" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:03F487D0A37A4D5EA2DBFF43FD75EB71:8E82601DA37B4D43A02EFF0BFCA6E89D" box="[610,781,223,249]" country="Tajikistan" county="river Sardai-Miona" municipality="Zangob" name="valley Divdara" pageId="13" pageNumber="214">valley Divdara</location>
, stream 
<collectingMunicipality box="[879,967,223,249]" pageId="13" pageNumber="214">Zangob</collectingMunicipality>
, 
<locationDeviation box="[981,1245,223,249]" location="Ramit" pageId="13" pageNumber="214">
<quantity box="[981,1055,223,249]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="214" unit="km" value="12.0">12 km</quantity>
upstream Ramit
</locationDeviation>
by 
<collectingCounty pageId="13" pageNumber="214">river Sardai-Miona</collectingCounty>
, 
<date box="[236,422,259,285]" pageId="13" pageNumber="214" value="1981-08-18" valueMax="1981-08-21" valueMin="1981-08-18">
<collectingDate box="[236,422,259,285]" pageId="13" pageNumber="214" value="1981-08-18" valueMax="1981-08-21" valueMin="1981-08-18">
18–21. 
<collectionCode box="[310,363,259,285]" pageId="13" pageNumber="214">VIII</collectionCode>
.1981
</collectingDate>
</date>
, coll. 
<collectorName box="[489,592,259,285]" pageId="13" pageNumber="214">N. Kluge</collectorName>
&amp; 
<collectorName box="[628,774,259,285]" pageId="13" pageNumber="214">E. Novikova</collectorName>
: 2 L-S-I♁, 5 L-S-I 
<specimenCount box="[992,1006,259,285]" pageId="13" pageNumber="214" type="female">♀</specimenCount>
</materialsCitation>
, 3 L-S♁, 10 L-S 
<specimenCount box="[1201,1216,259,285]" pageId="13" pageNumber="214" type="female">♀</specimenCount>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889725" ID-Zenodo-Dep="7889725" httpUri="https://zenodo.org/record/7889725/files/figure.png" pageId="13" pageNumber="214" startId="13.[151,264,1964,1988]" targetBox="[151,1436,309,1941]" targetPageId="13" targetType="figure">
<paragraph blockId="13.[151,1436,1964,2024]" pageId="13" pageNumber="214">
<emphasis bold="true" box="[151,342,1964,1988]" pageId="13" pageNumber="214">FIGURES 47–49.</emphasis>
<taxonomicName authorityName=": Kluge" authorityYear="2022" box="[349,663,1964,1988]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="hissaricus" subGenus="Tenuibaetis">
<emphasis box="[349,412,1965,1988]" italics="true" pageId="13" pageNumber="214">Baetis</emphasis>
(
<emphasis box="[428,544,1964,1988]" italics="true" pageId="13" pageNumber="214">Tenuibaetis</emphasis>
) 
<emphasis box="[560,663,1964,1988]" italics="true" pageId="13" pageNumber="214">hissaricus</emphasis>
</taxonomicName>
, exuviae of larval fore, middle and hind femora. Abbreviation: 2-can, twochannel setae.
</paragraph>
</caption>
<paragraph blockId="14.[151,1437,150,501]" box="[199,479,150,176]" pageId="14" pageNumber="215">
<emphasis bold="true" box="[199,479,150,176]" pageId="14" pageNumber="215">Additional descriptions</emphasis>
</paragraph>
<paragraph blockId="14.[151,1437,150,501]" pageId="14" pageNumber="215">
<emphasis bold="true" box="[199,275,186,213]" pageId="14" pageNumber="215">
<emphasis bold="true" box="[199,267,187,213]" italics="true" pageId="14" pageNumber="215">Larva</emphasis>
.
</emphasis>
CUTICULAR COLORATION (
<figureCitation box="[657,743,186,212]" captionStart="FIGURES 44–46" captionStartId="12.[151,264,1987,2011]" captionTargetBox="[169,1418,365,1963]" captionTargetId="figure-86@12.[169,1418,365,1963]" captionTargetPageId="12" captionText="FIGURES 44–46. Baetis (Tenuibaetis) hissaricus, larval exuviae. 44, head; 45, labrum; 46, labium (44, 46, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889721" httpUri="https://zenodo.org/record/7889721/files/figure.png" pageId="14" pageNumber="215">Figs 44</figureCitation>
, 
<figureCitation box="[753,827,186,212]" captionStart="FIGURES 47–49" captionStartId="13.[151,264,1964,1988]" captionTargetBox="[151,1436,309,1941]" captionTargetId="figure-97@13.[151,1436,309,1941]" captionTargetPageId="13" captionText="FIGURES 47–49. Baetis (Tenuibaetis) hissaricus, exuviae of larval fore, middle and hind femora. Abbreviation: 2-can, twochannel setae." figureDoi="http://doi.org/10.5281/zenodo.7889725" httpUri="https://zenodo.org/record/7889725/files/figure.png" pageId="14" pageNumber="215">47–49</figureCitation>
: Described and figured by 
<bibRefCitation author="Novikova, E. A." box="[1121,1307,186,212]" pageId="14" pageNumber="215" pagination="793 - 795" refId="ref19156" refString="Novikova, E. A. (1991) [A new subspecies of mayflies Baetis ursinus hissaricus subsp. n. (Ephemeroptera, Baetidae) from Tadjikistan]. Entomologicheskoe Obozrenie, 70 (4), 793 - 795." type="journal article" year="1991">Novikova (1991</bibRefCitation>
: figs 1–16); pronotum with darker sigilla on lighter background (
<bibRefCitation author="Novikova, E. A." box="[740,919,222,248]" pageId="14" pageNumber="215" pagination="793 - 795" refId="ref19156" refString="Novikova, E. A. (1991) [A new subspecies of mayflies Baetis ursinus hissaricus subsp. n. (Ephemeroptera, Baetidae) from Tadjikistan]. Entomologicheskoe Obozrenie, 70 (4), 793 - 795." type="journal article" year="1991">Novikova 1991</bibRefCitation>
: figs 1–2).
</paragraph>
<paragraph blockId="14.[151,1437,150,501]" box="[200,780,258,285]" pageId="14" pageNumber="215">HYPODERMAL COLORATION: Not expressed.</paragraph>
<paragraph blockId="14.[151,1437,150,501]" box="[200,1242,294,321]" pageId="14" pageNumber="215">
<collectionCode box="[200,507,294,320]" pageId="14" pageNumber="215">SHAPE AND SETATION</collectionCode>
: Mouthparts— 
<figureCitation box="[682,809,294,320]" captionStart="FIGURES 44–46" captionStartId="12.[151,264,1987,2011]" captionTargetBox="[169,1418,365,1963]" captionTargetId="figure-86@12.[169,1418,365,1963]" captionTargetPageId="12" captionText="FIGURES 44–46. Baetis (Tenuibaetis) hissaricus, larval exuviae. 44, head; 45, labrum; 46, labium (44, 46, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889721" httpUri="https://zenodo.org/record/7889721/files/figure.png" pageId="14" pageNumber="215">Figs 44–46</figureCitation>
. Hind protoptera present (as in 
<figureCitation box="[1161,1229,294,320]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1916,1940]" captionTargetBox="[151,1436,885,1892]" captionTargetId="figure-386@2.[151,1436,885,1892]" captionTargetPageId="2" captionText="FIGURES 1–4. Setae on larval legs of Baetis (Tenuibaetis) bialatus sp. n.. 1, two-channel and hirudiform setae on outer side of middle tibia under scanning electron microscope; 2–3, hirudiform seta; 4, two-channel seta on anterior side of femur in transmitted light. Abbreviations: 2-can, two-channel seta; hirud, hirudiform seta." figureDoi="http://doi.org/10.5281/zenodo.7889697" httpUri="https://zenodo.org/record/7889697/files/figure.png" pageId="14" pageNumber="215">Fig. 2</figureCitation>
).
</paragraph>
<paragraph blockId="14.[151,1437,150,501]" pageId="14" pageNumber="215">
Two-channel setae forming row along outer side of femur (including two subapical setae) elongate, blunt, distally widened and flattened; stout setae on anterior side of femur two-channel, from moderately long to short (
<figureCitation box="[1295,1425,366,392]" captionStart="FIGURES 47–49" captionStartId="13.[151,264,1964,1988]" captionTargetBox="[151,1436,309,1941]" captionTargetId="figure-97@13.[151,1436,309,1941]" captionTargetPageId="13" captionText="FIGURES 47–49. Baetis (Tenuibaetis) hissaricus, exuviae of larval fore, middle and hind femora. Abbreviation: 2-can, twochannel setae." figureDoi="http://doi.org/10.5281/zenodo.7889725" httpUri="https://zenodo.org/record/7889725/files/figure.png" pageId="14" pageNumber="215">Figs 47–49</figureCitation>
).
</paragraph>
<paragraph blockId="14.[151,1437,150,501]" pageId="14" pageNumber="215">
Tibia of each leg with small, stout two-channel setae on inner side and along patella-tibial suture (
<figureCitation box="[1276,1358,402,429]" captionStart="FIGURES 50–51" captionStartId="14.[151,264,1999,2023]" captionTargetBox="[224,1363,524,1975]" captionTargetId="figure-186@14.[224,1363,524,1975]" captionTargetPageId="14" captionText="FIGURES 50–51. Baetis (Tenuibaetis) hissaricus, exuviae of larval hind tibia. Abbreviations: 2-can, two-channel setae." figureDoi="http://doi.org/10.5281/zenodo.7889729" httpUri="https://zenodo.org/record/7889729/files/figure.png" pageId="14" pageNumber="215">Fig. 51</figureCitation>
). Stout setae on outer side of tibia pseudo-bifurcate and very dense, forming not one row, but stripe of irregularly situated setae 2–4 setae width (
<figureCitation box="[405,490,474,501]" captionStart="FIGURES 50–51" captionStartId="14.[151,264,1999,2023]" captionTargetBox="[224,1363,524,1975]" captionTargetId="figure-186@14.[224,1363,524,1975]" captionTargetPageId="14" captionText="FIGURES 50–51. Baetis (Tenuibaetis) hissaricus, exuviae of larval hind tibia. Abbreviations: 2-can, two-channel setae." figureDoi="http://doi.org/10.5281/zenodo.7889729" httpUri="https://zenodo.org/record/7889729/files/figure.png" pageId="14" pageNumber="215">Fig. 50</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889729" ID-Zenodo-Dep="7889729" box="[151,1370,1999,2023]" httpUri="https://zenodo.org/record/7889729/files/figure.png" pageId="14" pageNumber="215" startId="14.[151,264,1999,2023]" targetBox="[224,1363,524,1975]" targetPageId="14" targetType="figure">
<paragraph blockId="14.[151,1370,1999,2023]" box="[151,1370,1999,2023]" pageId="14" pageNumber="215">
<emphasis bold="true" box="[151,340,1999,2023]" pageId="14" pageNumber="215">FIGURES 50–51.</emphasis>
<taxonomicName authorityName=": Kluge" authorityYear="2022" box="[346,658,1999,2023]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="hissaricus" subGenus="Tenuibaetis">
<emphasis box="[346,409,2000,2023]" italics="true" pageId="14" pageNumber="215">Baetis</emphasis>
(
<emphasis box="[424,540,1999,2023]" italics="true" pageId="14" pageNumber="215">Tenuibaetis</emphasis>
) 
<emphasis box="[555,658,1999,2023]" italics="true" pageId="14" pageNumber="215">hissaricus</emphasis>
</taxonomicName>
, exuviae of larval hind tibia. Abbreviations: 2-can, two-channel setae.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889731" ID-Zenodo-Dep="7889731" httpUri="https://zenodo.org/record/7889731/files/figure.png" pageId="15" pageNumber="216" startId="15.[151,264,1834,1858]" targetBox="[151,1435,181,1810]" targetPageId="15" targetType="figure">
<paragraph blockId="15.[151,1436,1834,1930]" pageId="15" pageNumber="216">
<emphasis bold="true" box="[151,339,1834,1858]" pageId="15" pageNumber="216">FIGURES 52–59.</emphasis>
<taxonomicName authorityName=": Kluge" authorityYear="2022" box="[344,654,1834,1858]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="hissaricus" subGenus="Tenuibaetis">
<emphasis box="[344,407,1835,1858]" italics="true" pageId="15" pageNumber="216">Baetis</emphasis>
(
<emphasis box="[421,537,1834,1858]" italics="true" pageId="15" pageNumber="216">Tenuibaetis</emphasis>
) 
<emphasis box="[551,654,1834,1858]" italics="true" pageId="15" pageNumber="216">hissaricus</emphasis>
</taxonomicName>
. 52–53, fore and middle legs of male imago; 54–55, genitalia of males imago; 56, hind wing of male; 57, hind wing of female; 58, subimaginal exuviae of female fore leg; 59, subimaginal exuviae of half of mesonotum (57, 58, holotype).
</paragraph>
</caption>
<paragraph blockId="16.[151,1437,151,789]" pageId="16" pageNumber="217">
Denticles on posterior margins of terga I–X bluntly triangular (
<bibRefCitation author="Novikova, E. A." box="[903,1082,151,177]" pageId="16" pageNumber="217" pagination="793 - 795" refId="ref19156" refString="Novikova, E. A. (1991) [A new subspecies of mayflies Baetis ursinus hissaricus subsp. n. (Ephemeroptera, Baetidae) from Tadjikistan]. Entomologicheskoe Obozrenie, 70 (4), 793 - 795." type="journal article" year="1991">Novikova 1991</bibRefCitation>
: fig. 30). Denticles on posterior margin of sternum small and irregular, present only on sternum 
<collectionCode box="[866,898,187,213]" pageId="16" pageNumber="217">IX</collectionCode>
, sometimes also on sterna 
<collectionCode box="[1197,1245,187,213]" pageId="16" pageNumber="217">VIII</collectionCode>
and 
<collectionCode box="[1299,1341,187,213]" pageId="16" pageNumber="217">VII</collectionCode>
; in male row of denticles not interrupted by protogonostyli. Paraproct with denticles on most part of inner margin (as in 
<figureCitation captionStart="FIGURES 22–30" captionStartId="8.[151,264,1852,1876]" captionTargetBox="[151,1436,181,1827]" captionTargetId="figure-36@8.[151,1436,181,1828]" captionTargetPageId="8" captionText="FIGURES 22–30. Baetis (Tenuibaetis) ursinus, larval exuviae (24–29, one and the same magnification). 22, labrum; 23, maxilla and labium; 24, abdomen, 25–27, middle and hind legs; 28, half of pronotum and mesonotum; 29, head; 30, paraproct." figureDoi="http://doi.org/10.5281/zenodo.7889707" httpUri="https://zenodo.org/record/7889707/files/figure.png" pageId="16" pageNumber="217">Fig. 30</figureCitation>
). Paracercus significantly smaller than cerci.
</paragraph>
<paragraph blockId="16.[151,1437,151,789]" pageId="16" pageNumber="217">
<emphasis bold="true" box="[199,323,294,321]" pageId="16" pageNumber="217">
<emphasis bold="true" box="[199,316,294,321]" italics="true" pageId="16" pageNumber="217">Subimago</emphasis>
.
</emphasis>
CUTICULAR COLORATION: Head colorless, antennae brown. Pronotum light brown. Mesonotum with medioscutum and submedioscutum mostly ochre, lateral areas mostly light brownish (
<figureCitation box="[1337,1424,330,357]" captionStart="FIGURES 52–59" captionStartId="15.[151,264,1834,1858]" captionTargetBox="[151,1435,181,1810]" captionTargetId="figure-19@15.[151,1435,181,1810]" captionTargetPageId="15" captionText="FIGURES 52–59. Baetis (Tenuibaetis) hissaricus. 52–53, fore and middle legs of male imago; 54–55, genitalia of males imago; 56, hind wing of male; 57, hind wing of female; 58, subimaginal exuviae of female fore leg; 59, subimaginal exuviae of half of mesonotum (57, 58, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889731" httpUri="https://zenodo.org/record/7889731/files/figure.png" pageId="16" pageNumber="217">Fig. 59</figureCitation>
). Thoracic pleura and sterna with light brownish and dark brown areas (as in 
<figureCitation box="[1041,1142,366,393]" captionStart="FIGURES 215–224" captionStartId="43.[151,264,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-19@43.[151,1435,181,1746]" captionTargetPageId="43" captionText="FIGURES 215–224. Baetis (Tenuibaetis) bialatus sp. n. 215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217–218, exuviae of fore and hind legs of male subimago; 219–220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIII–IX; 224, egg (215–220, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889793" httpUri="https://zenodo.org/record/7889793/files/figure.png" pageId="16" pageNumber="217">Fig. 215</figureCitation>
). Legs of all pairs nearly colorless, with small, contrasting, brown marking near base of tibia (
<figureCitation box="[931,1017,402,429]" captionStart="FIGURES 52–59" captionStartId="15.[151,264,1834,1858]" captionTargetBox="[151,1435,181,1810]" captionTargetId="figure-19@15.[151,1435,181,1810]" captionTargetPageId="15" captionText="FIGURES 52–59. Baetis (Tenuibaetis) hissaricus. 52–53, fore and middle legs of male imago; 54–55, genitalia of males imago; 56, hind wing of male; 57, hind wing of female; 58, subimaginal exuviae of female fore leg; 59, subimaginal exuviae of half of mesonotum (57, 58, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889731" httpUri="https://zenodo.org/record/7889731/files/figure.png" pageId="16" pageNumber="217">Fig. 58</figureCitation>
). All abdominal terga and sterna I–X light brownish.
</paragraph>
<paragraph blockId="16.[151,1437,151,789]" box="[200,756,475,501]" pageId="16" pageNumber="217">HYPODERMAL COLORATION: As in imago.</paragraph>
<paragraph blockId="16.[151,1437,151,789]" pageId="16" pageNumber="217">
<emphasis bold="true" box="[199,351,510,537]" italics="true" pageId="16" pageNumber="217">Imago, male.</emphasis>
Described by 
<bibRefCitation author="Novikova, E. A." box="[517,713,511,537]" pageId="16" pageNumber="217" pagination="793 - 795" refId="ref19156" refString="Novikova, E. A. (1991) [A new subspecies of mayflies Baetis ursinus hissaricus subsp. n. (Ephemeroptera, Baetidae) from Tadjikistan]. Entomologicheskoe Obozrenie, 70 (4), 793 - 795." type="journal article" year="1991">Novikova (1991)</bibRefCitation>
. Middle and hind tarsi with 2 apical spines (on 1st+2nd and 3rd segments). Gonovectes gently bent, brown; sterno-styligeral muscle vestigial (
<figureCitation box="[1029,1113,546,573]" captionStart="FIGURES 52–59" captionStartId="15.[151,264,1834,1858]" captionTargetBox="[151,1435,181,1810]" captionTargetId="figure-19@15.[151,1435,181,1810]" captionTargetPageId="15" captionText="FIGURES 52–59. Baetis (Tenuibaetis) hissaricus. 52–53, fore and middle legs of male imago; 54–55, genitalia of males imago; 56, hind wing of male; 57, hind wing of female; 58, subimaginal exuviae of female fore leg; 59, subimaginal exuviae of half of mesonotum (57, 58, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889731" httpUri="https://zenodo.org/record/7889731/files/figure.png" pageId="16" pageNumber="217">Fig. 54</figureCitation>
).
</paragraph>
<paragraph blockId="16.[151,1437,151,789]" pageId="16" pageNumber="217">
<emphasis bold="true" box="[199,377,582,609]" italics="true" pageId="16" pageNumber="217">Imago, female.</emphasis>
Described by 
<bibRefCitation author="Novikova, E. A." box="[551,751,583,609]" pageId="16" pageNumber="217" pagination="793 - 795" refId="ref19156" refString="Novikova, E. A. (1991) [A new subspecies of mayflies Baetis ursinus hissaricus subsp. n. (Ephemeroptera, Baetidae) from Tadjikistan]. Entomologicheskoe Obozrenie, 70 (4), 793 - 795." type="journal article" year="1991">Novikova (1991)</bibRefCitation>
. All tarsi with 2 apical spines (fore tarsus on 2nd and 3rd segments, middle and hind tarsi on 1st+2nd and 3rd segments).
</paragraph>
<paragraph blockId="16.[151,1437,151,789]" box="[199,639,654,681]" pageId="16" pageNumber="217">
<emphasis bold="true" box="[199,334,654,680]" pageId="16" pageNumber="217">Dimension.</emphasis>
Fore wing length 
<quantity box="[539,635,654,681]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="16" pageNumber="217" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="217" type="distribution">
<paragraph blockId="16.[151,1437,151,789]" box="[199,663,690,717]" pageId="16" pageNumber="217">
<emphasis bold="true" box="[199,353,690,716]" pageId="16" pageNumber="217">Distribution.</emphasis>
Mountains of Central Asia.
</paragraph>
<paragraph blockId="16.[151,1437,151,789]" pageId="16" pageNumber="217">
<emphasis bold="true" box="[199,354,726,752]" pageId="16" pageNumber="217">Comparison.</emphasis>
Larva of 
<taxonomicName authorityName=": Kluge" authorityYear="2022" box="[466,659,727,753]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="hissaricus" subGenus="Tenuibaetis">
<emphasis box="[466,483,727,752]" italics="true" pageId="16" pageNumber="217">B</emphasis>
. (
<emphasis box="[508,529,727,752]" italics="true" pageId="16" pageNumber="217">T.</emphasis>
) 
<emphasis box="[545,659,727,753]" italics="true" pageId="16" pageNumber="217">hissaricus</emphasis>
</taxonomicName>
differs from other examined species by very dense stripe of pseudo-bifurcate setae on outer sides of tibiae (
<figureCitation box="[591,722,762,789]" captionStart="FIGURES 50–51" captionStartId="14.[151,264,1999,2023]" captionTargetBox="[224,1363,524,1975]" captionTargetId="figure-186@14.[224,1363,524,1975]" captionTargetPageId="14" captionText="FIGURES 50–51. Baetis (Tenuibaetis) hissaricus, exuviae of larval hind tibia. Abbreviations: 2-can, two-channel setae." figureDoi="http://doi.org/10.5281/zenodo.7889729" httpUri="https://zenodo.org/record/7889729/files/figure.png" pageId="16" pageNumber="217">Figs 50–51</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>