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<document id="FE2A52AA1864E87F35F5FDDDDE6A8482" ID-CLB-Dataset="57380" ID-DOI="10.11646/phytotaxa.585.4.1" ID-GBIF-Dataset="aa4b6a5e-18fa-4915-a746-95d1cb74d648" ID-ISSN="1179-3163" ID-Zenodo-Dep="7703837" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1678177070991" checkinUser="plazi" docAuthor="Leong-Škorničková, Jana, Bình, Nguyén Qu ốc, Đăng, Tr ần H ữu, Trư ờng, Lưu H ồng &amp; Nuraliev, Maxim S." docDate="2023" docId="03F487B8FFECFFDB9EB9FB24FC4245DF" docLanguage="en" docName="phytotaxa.585.4.1.pdf" docOrigin="Phytotaxa 585 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.585.4.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Geostachys pierreana Gagnepain 1906" docType="treatment" docVersion="7" lastPageNumber="257" masterDocId="FFCDFFC0FFE5FFD79E31FFAAFFF3431B" masterDocTitle="A revision of Geostachys (Zingiberaceae: Alpinioideae) in Cambodia, Laos and Vietnam, including a new species, G. aristata" masterLastPageNumber="260" masterPageNumber="245" pageNumber="254" updateTime="1699476210154" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="AEE5C2ACD8120FD706D5E4A48690AEF8">A revision of Geostachys (Zingiberaceae: Alpinioideae) in Cambodia, Laos and Vietnam, including a new species, G. aristata</mods:title>
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<mods:namePart id="7F7C659241C0B221EF617BA8F4049C8A">Leong-Škorničková, Jana</mods:namePart>
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<mods:namePart id="995F21F35BAD41EF2F7117E658356FC4">Bình, Nguyén Qu ốc</mods:namePart>
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<mods:affiliation id="E8E137D315FF5ABFFFF5D382124919E3">Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18, Hoàng Qu ṍc Vi ệt Street, Cǻu Giǻy, 122300 Hà N ội, Vietnam &amp; binhzing @ gmail. com; https: // orcid. org / 0000 - 0002 - 6267 - 839 X</mods:affiliation>
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<mods:namePart id="46103CB4DB0ABB3398642FE003A14B10">Đăng, Tr ần H ữu</mods:namePart>
<mods:affiliation id="3EFB54CB5324887812C1644B485CC956">Becamex Institute of Research and Development, Becamex IDC Corp., 08, Hùng Vuõng Street, Hòa Phú Ward, Th ủ Dǻu M ột City, Bình Duõng Province, Vietnam &amp; Southern Institute of Ecology, Institute of Applied Materials Science, Vietnam Academy of Science and Technology, 1 D, TL 29 Street, District 12, H ồ Chí Minh City, Vietnam</mods:affiliation>
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<mods:namePart id="072AA3AA199E1707FF3FD88E611793AD">Trư ờng, Lưu H ồng</mods:namePart>
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<mods:affiliation id="7FEFBFA30A404CB888BE853B089A68FA">Southern Institute of Ecology, Institute of Applied Materials Science, Vietnam Academy of Science and Technology, 1 D, TL 29 Street, District 12, H ồ Chí Minh City, Vietnam &amp; hongtruongluu @ gmail. com; https: // orcid. org / 0000 - 0002 - 7036 - 7081</mods:affiliation>
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<mods:namePart id="94FD514F429DC09D484FA829EE8F3902">Nuraliev, Maxim S.</mods:namePart>
<mods:affiliation id="9A383B3F698DB46302D7DD79F3BA7247">Joint Russian-Vietnamese Tropical Scientific and Technological Center, Nguy ễn Vãn Huyên street, Cǻu Giǻy, Hà N ội, Vietnam Department of Higher Plants, Biological Faculty, M. V. Lomonosov Moscow State University, 1, 12, Leninskie Gory, 119234 Moscow, Russia</mods:affiliation>
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<mods:title id="DD036BC1416CAF3EB0F6733DCBA180BF">Phytotaxa</mods:title>
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<mods:date id="E0267E78C5D13CC1EE0BC21B78186B13">2023</mods:date>
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<mods:number id="E152A2EC9E8614B0673121AAD8247BA4">585</mods:number>
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<treatment id="03F487B8FFECFFDB9EB9FB24FC4245DF" ID-DOI="http://doi.org/10.5281/zenodo.7703852" ID-GBIF-Taxon="209416337" ID-Zenodo-Dep="7703852" LSID="urn:lsid:plazi:treatment:03F487B8FFECFFDB9EB9FB24FC4245DF" httpUri="http://treatment.plazi.org/id/03F487B8FFECFFDB9EB9FB24FC4245DF" lastPageId="12" lastPageNumber="257" pageId="9" pageNumber="254">
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<treatmentCitationGroup id="AB4D1180FFECFFDE9EB9FB24FCDF47B3" box="[136,812,1166,1193]" pageId="9" pageNumber="254">
<treatmentCitation id="0AFC10BFFFECFFDE9EB9FB24FD6A47B3" author="Gagnepain, F." box="[136,665,1166,1193]" page=")" pageId="9" pageNumber="254" year="1906">
<taxonomicName id="4C5D4D2DFFECFFDE9EB9FB24FD6A47B3" ID-CoL="3FSCQ" authority="Gagnepain (1906 a: 147)" authorityName="Gagnepain" authorityPageNumber="147" authorityYear="1906" box="[136,665,1166,1193]" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="9" pageNumber="254" phylum="Tracheophyta" rank="species" species="pierreana">
<emphasis id="B929EABCFFECFFDE9EB9FB24FE7147B3" bold="true" box="[136,386,1166,1193]" italics="true" pageId="9" pageNumber="254">Geostachys pierreana</emphasis>
<bibRefCitation id="EFCC4B5FFFECFFDE9FB8FB24FD6A47B3" author="Gagnepain, F." box="[393,665,1166,1192]" pageId="9" pageNumber="254" pagination="132 - 150" refId="ref10054" refString="Gagnepain, F. (1906 a) Zingiberacees nouvelles de l'herbier du Museum (15 ° note). Bulletin de la Societe Botanique de France 53 (2): 132 - 150. https: // doi. org / 10.1080 / 00378941.1906.10831155" type="journal article" year="1906">Gagnepain (1906a: 147)</bibRefCitation>
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(
<figureCitation id="13662A2BFFECFFDE9C99FB24FD1847B3" box="[680,747,1166,1192]" captionStart="FIGURE 1" captionStartId="3.[136,229,2003,2025]" captionTargetBox="[175,1412,190,1979]" captionTargetId="figure-18@3.[175,1412,190,1979]" captionTargetPageId="3" captionText="FIGURE 1. Distribution map of Geostachys species in Cambodia, Laos and Vietnam." figureDoi="http://doi.org/10.5281/zenodo.7703843" httpUri="https://zenodo.org/record/7703843/files/figure.png" pageId="9" pageNumber="254">Fig. 1</figureCitation>
,
<figureCitation id="13662A2BFFECFFDE9CC8FB24FCFA47B2" box="[761,777,1166,1193]" captionStart="FIGURE 5" captionStartId="10.[136,229,1816,1838]" captionTargetBox="[151,1435,425,1792]" captionTargetId="figure-137@10.[151,1435,425,1792]" captionTargetPageId="10" captionText="FIGURE 5. Geostachys pierreana. A. Leafy shoot (from above). B. Basal part of plant with inflorescences. C. Detail of ligule. D. Inflorescences. E. Flower with labellum removed (staminodes remain attached to floral tube). F. Cincinnus with one open flower (side view). G. Flower (front view).All from Netopilová &amp; Faltová s.n. Photo by M. Netopilová." figureDoi="http://doi.org/10.5281/zenodo.7703856" httpUri="https://zenodo.org/record/7703856/files/figure.png" pageId="9" pageNumber="254">5</figureCitation>
,
<figureCitation id="13662A2BFFECFFDE9D24FB24FCD747B3" box="[789,804,1166,1192]" captionStart="FIGURE 6" captionStartId="11.[136,229,1282,1304]" captionTargetBox="[151,1435,302,1272]" captionTargetId="figure-79@11.[151,1435,302,1272]" captionTargetPageId="11" captionText="FIGURE 6. Geostachys pierreana. A. Leafy shoot (view from below). B. Pseudostem with ligules and leaf bases. C. Basal part of plant with inflorescences. D. Cross section of cincinnus showing total of five flowers/fruits. E. Stamen (adaxial and abaxial view). F. Inflorescence, top view. All from Luu s.n. Photo by H.T. Luu." figureDoi="http://doi.org/10.5281/zenodo.7703858" httpUri="https://zenodo.org/record/7703858/files/figure.png" pageId="9" pageNumber="254">6</figureCitation>
)
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<emphasis id="B929EABCFFECFFDE9EB9FB1CFF3147D0" bold="true" box="[136,194,1206,1227]" pageId="9" pageNumber="254">Type:</emphasis>
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Country unknown [S. Vietnam, Cambodia or Thailand], 18651877,
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5851
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(
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, designated here,
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:
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!;
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<typeStatus id="54E6880CFFECFFDE9E8CFB70FEC947EB" box="[189,314,1242,1264]" pageId="9" pageNumber="254" type="isolectotype">isolectotypes</typeStatus>
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:
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! and
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!; flowering)
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.
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<paragraph id="8BE236AEFFECFFDD9EB9FAB4FB0C428B" blockId="9.[136,1452,1310,2093]" lastBlockId="10.[136,1452,158,401]" lastPageId="10" lastPageNumber="255" pageId="9" pageNumber="254">
Clump-forming herb up to
<quantity id="4CA59B4BFFECFFDE9F84FAB4FE2C4622" box="[437,479,1310,1337]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="254" unit="m" value="1.0">1 m</quantity>
tall.
<emphasis id="B929EABCFFECFFDE9C27FAB4FD884622" bold="true" box="[534,635,1310,1337]" italics="true" pageId="9" pageNumber="254">Rhizome</emphasis>
supported by stilt roots, sheathed by dry brown scales.
<emphasis id="B929EABCFFECFFDE9AD2FAB4FA804622" bold="true" box="[1251,1395,1310,1337]" italics="true" pageId="9" pageNumber="254">Leafy shoots</emphasis>
with 613 leaves per pseudostem, bladeless at basal 1/3;
<emphasis id="B929EABCFFECFFDE9CFFFAE8FC614647" box="[718,914,1346,1372]" italics="true" pageId="9" pageNumber="254">bladeless sheaths</emphasis>
26; most basal (outermost) sheaths very short, soon becoming dry and brown, inner sheaths distally gradually longer, green with reddish tinge, abaxially glabrous, striate (more prominent in dry material);
<emphasis id="B929EABCFFECFFDE9C7EFA20FD2046BF" box="[591,723,1418,1444]" italics="true" pageId="9" pageNumber="254">leaf sheaths</emphasis>
similar to bladeless sheaths;
<emphasis id="B929EABCFFECFFDE9A27FA20FBA546BF" box="[1046,1110,1418,1444]" italics="true" pageId="9" pageNumber="254">ligule</emphasis>
ca.
<quantity id="4CA59B4BFFECFFDE9AB3FA20FB3146BE" box="[1154,1218,1418,1445]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="254" unit="mm" value="3.0">3 mm</quantity>
long, triangular with obtuse apex, green with reddish to brown tinge (darker at base), glabrous;
<emphasis id="B929EABCFFECFFDE9D81FA04FC0F46D3" box="[944,1020,1454,1480]" italics="true" pageId="9" pageNumber="254">petiole</emphasis>
<quantity id="4CA59B4BFFECFFDE9A30FA04FB9946D2" box="[1025,1130,1454,1481]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="9" pageNumber="254" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">510 mm</quantity>
long, shallowly canaliculate, mostly green, adaxially puberulent and abaxially glabrous to sparsely puberulent;
<emphasis id="B929EABCFFECFFDE9A14FA78FB6746F7" box="[1061,1172,1490,1516]" italics="true" pageId="9" pageNumber="254">leaf blade</emphasis>
narrowly elliptic,
<quantity id="4CA59B4BFFECFFDE9B54FA78FF59450A" metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="9" pageNumber="254" unit="cm" value="30.0" valueMax="35.0" valueMin="25.0">2535 cm</quantity>
long,
<quantity id="4CA59B4BFFECFFDE9EC1FA5CFEB1450A" box="[240,322,1526,1553]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="9" pageNumber="254" unit="cm" value="5.0" valueMax="6.0" valueMin="4.0">46 cm</quantity>
wide, weakly plicate, adaxially green to dark green, somewhat shiny, glabrous, abaxially paler green, glabrous, base narrowly obtuse, apex acuminate, midrib adaxially sunken, pale green to ochraceous, glabrous, abaxially protruding, pale green, glabrous.
<emphasis id="B929EABCFFECFFDE9C34F994FD514542" bold="true" box="[517,674,1598,1625]" italics="true" pageId="9" pageNumber="254">Inflorescence</emphasis>
radical;
<emphasis id="B929EABCFFECFFDE9D39F994FC9E4543" box="[776,877,1598,1624]" italics="true" pageId="9" pageNumber="254">peduncle</emphasis>
erect, covered by 57 sheathing bracts;
<emphasis id="B929EABCFFECFFDE9B0FF994FF3D4567" italics="true" pageId="9" pageNumber="254">sheathing bracts</emphasis>
distichous, ovate to narrowly ovate, overlapping, basal ones shorter, distal ones significantly longer (sometimes developing blade vestiges at the apex, see
<figureCitation id="13662A2BFFECFFDE9C5DF92CFD3045BA" box="[620,707,1670,1697]" captionStart="FIGURE 5" captionStartId="10.[136,229,1816,1838]" captionTargetBox="[151,1435,425,1792]" captionTargetId="figure-137@10.[151,1435,425,1792]" captionTargetPageId="10" captionText="FIGURE 5. Geostachys pierreana. A. Leafy shoot (from above). B. Basal part of plant with inflorescences. C. Detail of ligule. D. Inflorescences. E. Flower with labellum removed (staminodes remain attached to floral tube). F. Cincinnus with one open flower (side view). G. Flower (front view).All from Netopilová &amp; Faltová s.n. Photo by M. Netopilová." figureDoi="http://doi.org/10.5281/zenodo.7703856" httpUri="https://zenodo.org/record/7703856/files/figure.png" pageId="9" pageNumber="254">Fig. 5D</figureCitation>
on the right), soon becoming brown and papery, striate, glabrous on both sides;
<emphasis id="B929EABCFFECFFDE9F1DF900FE8345DF" box="[300,368,1706,1732]" italics="true" pageId="9" pageNumber="254">thyrse</emphasis>
lax, many-flowered;
<emphasis id="B929EABCFFECFFDE9C52F900FD5A45DF" box="[611,681,1706,1732]" italics="true" pageId="9" pageNumber="254">rachis</emphasis>
<quantity id="4CA59B4BFFECFFDE9C80F900FCF745DE" box="[689,772,1706,1733]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="9" pageNumber="254" unit="cm" value="6.0" valueMax="8.0" valueMin="4.0">48 cm</quantity>
long, decurved, dull green with dark red tinge (turning dark red to maroon in fruiting stage), hirsute, shortly branched;
<emphasis id="B929EABCFFECFFDE9D2BF964FC9345F3" box="[794,864,1742,1768]" italics="true" pageId="9" pageNumber="254">bracts</emphasis>
ovate, broadly ovate to semi-circular,
<quantity id="4CA59B4BFFECFFDE9B3EF964FA9F45F2" box="[1295,1388,1742,1769]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="9" pageNumber="254" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">13 mm</quantity>
long,
<quantity id="4CA59B4BFFECFFDE9EB9F958FF154416" box="[136,230,1778,1805]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="9" pageNumber="254" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
wide, brownish-maroon at base, greenish to whitish distally, glabrous, caducous, each subtending cincinnus of 15 flowers on
<quantity id="4CA59B4BFFECFFDE9F6BF8BCFE35442A" box="[346,454,1814,1841]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="15.0" metricValueMin="4.0" pageId="9" pageNumber="254" unit="mm" value="9.5" valueMax="15.0" valueMin="4.0">415 mm</quantity>
long branch;
<emphasis id="B929EABCFFECFFDE9C55F8BCFD38442B" box="[612,715,1814,1840]" italics="true" pageId="9" pageNumber="254">branches</emphasis>
secund, of the same colour as rachis, hirsute;
<emphasis id="B929EABCFFECFFDE9AE8F8BCFABD442B" box="[1241,1358,1814,1840]" italics="true" pageId="9" pageNumber="254">bracteoles</emphasis>
broadly ovate to broadly elliptic, spathaceous, enveloping the flower with the rest of the cincinnus,
<quantity id="4CA59B4BFFECFFDE9AB3F890FB08444E" box="[1154,1275,1850,1877]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="3.0" metricValueMin="1.3" pageId="9" pageNumber="254" unit="mm" value="21.5" valueMax="30.0" valueMin="13.0">1330 mm</quantity>
long,
<quantity id="4CA59B4BFFECFFDE9B70F890FA5F444E" box="[1345,1452,1850,1877]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.7" metricValueMin="0.4" pageId="9" pageNumber="254" unit="mm" value="10.5" valueMax="17.0" valueMin="4.0">417 mm</quantity>
wide, pale pink-orange with greenish apex when young, soon scarious and pale brown to ochraceous, somewhat striate (visible in dry material), mostly glabrous, apex mucronate and puberulent.
<emphasis id="B929EABCFFECFFDE9DC1F828FBB14486" bold="true" box="[1008,1090,1922,1949]" italics="true" pageId="9" pageNumber="254">Flower</emphasis>
gullet-type;
<emphasis id="B929EABCFFECFFDE9AE9F828FADA4487" box="[1240,1321,1922,1948]" italics="true" pageId="9" pageNumber="254">pedicel</emphasis>
negligible;
<emphasis id="B929EABCFFECFFDE9EB9F80CFF3044DB" box="[136,195,1958,1984]" italics="true" pageId="9" pageNumber="254">calyx</emphasis>
tubular,
<quantity id="4CA59B4BFFECFFDE9F1FF80CFE5844DA" box="[302,427,1958,1985]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="9" pageNumber="254" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">1215 mm</quantity>
long, apex with single lobe ending with prominent ciliate mucro, semi-translucent pale yellow with red tinge, mostly glabrous, apex puberulent;
<emphasis id="B929EABCFFECFFDE9D23F860FC7B44FF" box="[786,904,1994,2020]" italics="true" pageId="9" pageNumber="254">floral tube</emphasis>
yellow;
<emphasis id="B929EABCFFECFFDE9DDCF860FB3744FF" box="[1005,1220,1994,2020]" italics="true" pageId="9" pageNumber="254">dorsal corolla lobe</emphasis>
elliptic-oblong with cucullate apex, yellow with white semi-translucent veins, glabrous;
<emphasis id="B929EABCFFECFFDE9DBDF844FB814B13" box="[908,1138,2030,2056]" italics="true" pageId="9" pageNumber="254">lateral corolla lobes</emphasis>
elliptic-oblong with obtuse apex, yellow with white semi-translucent veins, glabrous;
<emphasis id="B929EABCFFECFFDE9D26F7B8FC8A4B37" box="[791,889,2066,2092]" italics="true" pageId="9" pageNumber="254">labellum</emphasis>
obovate, apex slightly bilobed, yellow with semitranslucent veins, adaxially with short glandular hair, abaxially glabrous;
<emphasis id="B929EABCFFEFFFDD9DDEFF34FB3743A3" box="[1007,1220,158,184]" italics="true" pageId="10" pageNumber="255">lateral staminodes</emphasis>
irregularly obovate with round apex, nearly entirely connate to labellum, yellow with semi-translucent veins;
<emphasis id="B929EABCFFEFFFDD9AA8FF69FB1B43C7" box="[1177,1256,195,220]" italics="true" pageId="10" pageNumber="255">stamen</emphasis>
yellow;
<emphasis id="B929EABCFFEFFFDD9B61FF68FA5F43C7" box="[1360,1452,194,220]" italics="true" pageId="10" pageNumber="255">filament</emphasis>
dorso-ventrally compressed, pale yellow with glandular hair;
<emphasis id="B929EABCFFEFFFDD9D08FF4CFBF1421B" box="[825,1026,230,256]" italics="true" pageId="10" pageNumber="255">anther connective</emphasis>
yellow, with glandular hair;
<emphasis id="B929EABCFFEFFFDD9B74FF4DFA89421B" box="[1349,1402,231,256]" italics="true" pageId="10" pageNumber="255">crest</emphasis>
<quantity id="4CA59B4BFFEFFFDD9BB0FF4CFF47423E" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="10" pageNumber="255" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long, bluntly obscurely three-lobed, yellow;
<emphasis id="B929EABCFFEFFFDD9C88FEA0FCA7423F" box="[697,852,266,292]" italics="true" pageId="10" pageNumber="255">anther thecae</emphasis>
dehiscing through the entire length;
<emphasis id="B929EABCFFEFFFDD9ACDFEA1FACF423F" box="[1276,1340,267,292]" italics="true" pageId="10" pageNumber="255">ovary</emphasis>
trilocular with central placentation, cylindrical, greenish or with reddish tinge, glabrous;
<emphasis id="B929EABCFFEFFFDD9A3BFE85FB3C4253" box="[1034,1231,302,328]" italics="true" pageId="10" pageNumber="255">epigynous glands</emphasis>
two;
<emphasis id="B929EABCFFEFFFDD9B23FE84FAB64253" box="[1298,1349,302,328]" italics="true" pageId="10" pageNumber="255">style</emphasis>
filiform, cream white;
<emphasis id="B929EABCFFEFFFDD9F17FEF9FE824277" box="[294,369,339,364]" italics="true" pageId="10" pageNumber="255">stigma</emphasis>
capitulate, cream white; ostiole transverse, forward-facing, ciliate.
<emphasis id="B929EABCFFEFFFDD9A4DFEF8FB4A4277" bold="true" box="[1148,1209,338,364]" italics="true" pageId="10" pageNumber="255">Fruit</emphasis>
(immature) ellipsoid hesperidium, ca.
<quantity id="4CA59B4BFFEFFFDD9F76FEDCFE65428A" box="[327,406,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="255" unit="mm" value="10.0">10 mm</quantity>
long, ca.
<quantity id="4CA59B4BFFEFFFDD9C35FEDCFDB6428A" box="[516,581,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="255" unit="mm" value="6.0">6 mm</quantity>
in diam., red, glabrous, with persistent calyx;
<emphasis id="B929EABCFFEFFFDD9A7DFEDCFB7A428B" box="[1100,1161,374,400]" italics="true" pageId="10" pageNumber="255">seeds</emphasis>
unknown.
</paragraph>
</subSubSection>
<caption id="DF226626FFEFFFDD9EB9F8B2FC09446D" ID-DOI="http://doi.org/10.5281/zenodo.7703856" ID-Zenodo-Dep="7703856" httpUri="https://zenodo.org/record/7703856/files/figure.png" pageId="10" pageNumber="255" startId="10.[136,229,1816,1838]" targetBox="[151,1435,425,1792]" targetPageId="10">
<paragraph id="8BE236AEFFEFFFDD9EB9F8B2FC09446D" blockId="10.[136,1452,1816,1910]" pageId="10" pageNumber="255">
<emphasis id="B929EABCFFEFFFDD9EB9F8B2FEF24435" bold="true" box="[136,257,1816,1838]" pageId="10" pageNumber="255">FIGURE 5.</emphasis>
<taxonomicName id="4C5D4D2DFFEFFFDD9F3BF8B2FE134435" authorityName="Gagnepain" authorityYear="1906" box="[266,480,1816,1838]" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="10" pageNumber="255" phylum="Tracheophyta" rank="species" species="pierreana">
<emphasis id="B929EABCFFEFFFDD9F3BF8B2FE134435" box="[266,480,1816,1838]" italics="true" pageId="10" pageNumber="255">Geostachys pierreana</emphasis>
</taxonomicName>
.
<emphasis id="B929EABCFFEFFFDD9FDEF8B2FDF54436" bold="true" box="[495,518,1816,1837]" pageId="10" pageNumber="255">A.</emphasis>
Leafy shoot (from above).
<emphasis id="B929EABCFFEFFFDD9D2FF8B3FCC74435" bold="true" box="[798,820,1817,1838]" pageId="10" pageNumber="255">B.</emphasis>
Basal part of plant with inflorescences.
<emphasis id="B929EABCFFEFFFDD9AFCF8B2FB174435" bold="true" box="[1229,1252,1816,1838]" pageId="10" pageNumber="255">C.</emphasis>
Detail of ligule.
<emphasis id="B929EABCFFEFFFDD9BA5F8B3FA5F4435" bold="true" box="[1428,1452,1817,1838]" pageId="10" pageNumber="255">D.</emphasis>
Inflorescences.
<emphasis id="B929EABCFFEFFFDD9F11F897FEC54449" bold="true" box="[288,310,1853,1874]" pageId="10" pageNumber="255">E.</emphasis>
Flower with labellum removed (staminodes remain attached to floral tube).
<emphasis id="B929EABCFFEFFFDD9A13F897FBC74449" bold="true" box="[1058,1076,1853,1874]" pageId="10" pageNumber="255">F.</emphasis>
Cincinnus with one open flower (side view).
<emphasis id="B929EABCFFEFFFDD9EFAF8CAFF17446D" bold="true" box="[203,228,1888,1910]" pageId="10" pageNumber="255">G.</emphasis>
Flower (front view). All from
<emphasis id="B929EABCFFEFFFDD9C3AF8CAFD0C446D" box="[523,767,1888,1910]" italics="true" pageId="10" pageNumber="255">Netopilová &amp; Faltová s.n</emphasis>
. Photo by M. Netopilová.
</paragraph>
</caption>
<subSubSection id="C3476525FFEFFFDD9E8CF80FFD7044F8" pageId="10" pageNumber="255" type="etymology">
<paragraph id="8BE236AEFFEFFFDD9E8CF80FFD7044F8" blockId="10.[136,1452,1957,2091]" pageId="10" pageNumber="255">
<emphasis id="B929EABCFFEFFFDD9E8CF80FFEC944A4" bold="true" box="[189,314,1957,1983]" pageId="10" pageNumber="255">Eponymy:</emphasis>
—This species is named after a French botanist Jean Baptiste Louis Pierre (
<date id="FFE3106EFFEFFFDD9A8BF80FFA7344A4" box="[1210,1408,1957,1983]" pageId="10" pageNumber="255" value="1833-10-23">23 October 1833</date>
<date id="FFE3106EFFEFFFDD9BBEF80FFED144FF" pageId="10" pageNumber="255" value="1905-10-30">30 October 1905</date>
) who made the
<typeStatus id="54E6880CFFEFFFDD9FE7F863FDF544F8" box="[470,518,1993,2019]" pageId="10" pageNumber="255">type</typeStatus>
collection.
</paragraph>
</subSubSection>
<subSubSection id="C3476525FFEFFFDC9E8CF847FE67423F" lastPageId="11" lastPageNumber="256" pageId="10" pageNumber="255" type="distribution">
<paragraph id="8BE236AEFFEFFFDC9E8CF847FE67423F" blockId="10.[136,1452,1957,2091]" lastBlockId="11.[136,1451,158,293]" lastPageId="11" lastPageNumber="256" pageId="10" pageNumber="255">
<emphasis id="B929EABCFFEFFFDD9E8CF847FD164B1C" bold="true" box="[189,741,2029,2055]" pageId="10" pageNumber="255">Distribution &amp; IUCN preliminary assessment:</emphasis>
—Currently known only from Bokor National Park,
<collectingCountry id="F34A763EFFEFFFDD9B00F847FA5B4B1C" box="[1329,1448,2029,2055]" name="Cambodia" pageId="10" pageNumber="255">Cambodia</collectingCountry>
.
<taxonomicName id="4C5D4D2DFFEFFFDD9EB9F7BBFE8D4B30" authorityName="Gagnepain" authorityYear="1906" box="[136,382,2065,2091]" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="10" pageNumber="255" phylum="Tracheophyta" rank="species" species="pierreana">
<emphasis id="B929EABCFFEFFFDD9EB9F7BBFE8D4B30" box="[136,382,2065,2091]" italics="true" pageId="10" pageNumber="255">Geostachys pierreana</emphasis>
</taxonomicName>
is currently assessed as Data Deficient on IUCN Red List (
<bibRefCitation id="EFCC4B5FFFEFFFDD9A11F7BBFB3E4B30" author="Olander, S. B." box="[1056,1229,2065,2091]" pageId="10" pageNumber="255" pagination="117327601" refId="ref11324" refString="Olander, S. B. (2019 b) Geostachys pierreana. The IUCN Red List of Threatened Species 2019: e. T 117327601 A 124282562. https: // dx. doi. org / 10.2305 / IUCN. UK. 2019 - 2. RLTS. T 117327601 A 124282562. en." type="journal article" year="2019">Olander 2019b</bibRefCitation>
). Although the only known location in Bokor National Park is under legal protection, there is no information about the number or size of populations. The Popokvil Waterfall is accessible to public, and the effect of tourism on the vegetation surrounding this area needs to be assessed first for an evidence-based assessment. We therefore suggest continuing to treat this species as Data Deficient (DD).
</paragraph>
</subSubSection>
<caption id="DF226626FFEEFFDC9EB9FAA8FD254644" ID-DOI="http://doi.org/10.5281/zenodo.7703858" ID-Zenodo-Dep="7703858" httpUri="https://zenodo.org/record/7703858/files/figure.png" pageId="11" pageNumber="256" startId="11.[136,229,1282,1304]" targetBox="[151,1435,302,1272]" targetPageId="11">
<paragraph id="8BE236AEFFEEFFDC9EB9FAA8FD254644" blockId="11.[136,1451,1282,1376]" pageId="11" pageNumber="256">
<emphasis id="B929EABCFFEEFFDC9EB9FAA8FEF2460C" bold="true" box="[136,257,1282,1304]" pageId="11" pageNumber="256">FIGURE 6.</emphasis>
<taxonomicName id="4C5D4D2DFFEEFFDC9F3BFAA8FE2C4603" authorityName="Gagnepain" authorityYear="1906" box="[266,479,1282,1304]" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="11" pageNumber="256" phylum="Tracheophyta" rank="species" species="pierreana">
<emphasis id="B929EABCFFEEFFDC9F3BFAA8FE2C4603" box="[266,479,1282,1304]" italics="true" pageId="11" pageNumber="256">Geostachys pierreana</emphasis>
</taxonomicName>
.
<emphasis id="B929EABCFFEEFFDC9FDFFAA8FDF5460C" bold="true" box="[494,518,1282,1303]" pageId="11" pageNumber="256">A.</emphasis>
Leafy shoot (view from below).
<emphasis id="B929EABCFFEEFFDC9D67FAA8FC9F460C" bold="true" box="[854,876,1282,1303]" pageId="11" pageNumber="256">B.</emphasis>
Pseudostem with ligules and leaf bases.
<emphasis id="B929EABCFFEEFFDC9B3BFAA8FAD24603" bold="true" box="[1290,1313,1282,1304]" pageId="11" pageNumber="256">C.</emphasis>
Basal part of plant with inflorescences.
<emphasis id="B929EABCFFEEFFDC9FBBFA8CFE514620" bold="true" box="[394,418,1318,1339]" pageId="11" pageNumber="256">D.</emphasis>
Cross section of cincinnus showing total of five flowers/fruits.
<emphasis id="B929EABCFFEEFFDC9A28FA8CFBDC4620" bold="true" box="[1049,1071,1318,1339]" pageId="11" pageNumber="256">E.</emphasis>
Stamen (adaxial and abaxial view).
<emphasis id="B929EABCFFEEFFDC9BA8FA8CFA584620" bold="true" box="[1433,1451,1318,1339]" pageId="11" pageNumber="256">F.</emphasis>
Inflorescence, top view. All from
<emphasis id="B929EABCFFEEFFDC9FFAFAE1FDE0467B" box="[459,531,1355,1376]" italics="true" pageId="11" pageNumber="256">Luu s.n</emphasis>
. Photo by
<emphasis id="B929EABCFFEEFFDC9C4AFAE1FD23467B" box="[635,720,1355,1376]" italics="true" pageId="11" pageNumber="256">H.T. Luu</emphasis>
.
</paragraph>
</caption>
<subSubSection id="C3476525FFEEFFDC9E8CFA22FBA646DD" pageId="11" pageNumber="256" type="biology_ecology">
<paragraph id="8BE236AEFFEEFFDC9E8CFA22FBA646DD" blockId="11.[136,1452,1415,2090]" pageId="11" pageNumber="256">
<emphasis id="B929EABCFFEEFFDC9E8CFA22FE3F46B9" bold="true" box="[189,460,1415,1442]" pageId="11" pageNumber="256">Ecology &amp; phenology:</emphasis>
—Growing in broadleaved evergreen forest, including its edges and slightly disturbed areas, at elevations of ca.
<quantity id="4CA59B4BFFEEFFDC9F98FA06FE0D46DD" box="[425,510,1452,1478]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="256" unit="m" value="1000.0">1000 m</quantity>
a.s.l. Flowering has been observed in April and June.
</paragraph>
</subSubSection>
<subSubSection id="C3476525FFEEFFDC9E8CFA65FC9345FD" pageId="11" pageNumber="256" type="materials_examined">
<paragraph id="8BE236AEFFEEFFDC9E8CFA65FE8A4585" blockId="11.[136,1452,1415,2090]" pageId="11" pageNumber="256">
<emphasis id="B929EABCFFEEFFDC9E8CFA65FDAC46F2" bold="true" box="[189,607,1487,1513]" pageId="11" pageNumber="256">Additional specimens examined:—</emphasis>
<materialsCitation id="3B353CF3FFEEFFDC9C6EFA65FBD84515" ID-GBIF-Occurrence="4048240323" collectingDate="2017-06-01" collectionCode="P" collectorName="P. Dy Phon" country="Cambodia" county="Popokvil" elevation="970" location="Bokor National Park" municipality="April" pageId="11" pageNumber="256" specimenCode="P02203163" specimenCount="1" stateProvince="Kampot province">
<emphasis id="B929EABCFFEEFFDC9C6EFA65FC1A46F2" bold="true" box="[607,1001,1487,1514]" pageId="11" pageNumber="256">
<collectingCountry id="F34A763EFFEEFFDC9C6EFA65FCF346F2" box="[607,768,1487,1513]" name="Cambodia" pageId="11" pageNumber="256">CAMBODIA</collectingCountry>
.
<collectingRegion id="4999F84CFFEEFFDC9D3CFA7AFC1746F2" box="[781,996,1487,1514]" country="Cambodia" name="Kampot" pageId="11" pageNumber="256">Kampot province</collectingRegion>
:
</emphasis>
[
<location id="8E826075FFEEFFDC9DC6FA7AFB1946F1" LSID="urn:lsid:plazi:treatment:03F487B8FFECFFDB9EB9FB24FC4245DF:8E826075FFEEFFDC9DC6FA7AFB1946F1" box="[1015,1258,1488,1514]" country="Cambodia" county="Popokvil" municipality="April" name="Bokor National Park" pageId="11" pageNumber="256" stateProvince="Kampot province">Bokor National Park</location>
],
<location id="8E826075FFEEFFDC9ACCFA7AFA9646F1" LSID="urn:lsid:plazi:treatment:03F487B8FFECFFDB9EB9FB24FC4245DF:8E826075FFEEFFDC9ACCFA7AFA9646F1" box="[1277,1381,1488,1514]" country="Cambodia" county="Popokvil" municipality="April" name="Popokvil" pageId="11" pageNumber="256" stateProvince="Kampot province">Popokvil</location>
, April [year not given, inferred as 1967?],
<emphasis id="B929EABCFFEEFFDC9C19FA5FFD194515" box="[552,746,1524,1550]" italics="true" pageId="11" pageNumber="256">
<collectorName id="26A85378FFEEFFDC9C19FA5FFD584515" box="[552,683,1524,1550]" pageId="11" pageNumber="256">P. Dy Phon</collectorName>
1157
</emphasis>
(
<collectionCode id="ED4CAE6BFFEEFFDC9CCCFA5EFCE34515" box="[765,784,1524,1550]" pageId="11" pageNumber="256">P</collectionCode>
:
<specimenCode id="DBFB9ED5FFEEFFDC9D2FFA5EFC534515" box="[798,928,1524,1550]" pageId="11" pageNumber="256">P02203163</specimenCode>
, flowering);
</materialsCitation>
<materialsCitation id="3B353CF3FFEEFFDC9A04FA5EFE864585" ID-GBIF-Occurrence="4048240325" collectingDate="2017-06-01" collectorName="M. Netopilova &amp; I. Faltova" country="Cambodia" county="Popokvil" elevation="970" location="Bokor National Park" municipality="April" pageId="11" pageNumber="256" specimenCount="1" stateProvince="Kampot">
<location id="8E826075FFEEFFDC9A04FA5EFAD74515" LSID="urn:lsid:plazi:treatment:03F487B8FFECFFDB9EB9FB24FC4245DF:8E826075FFEEFFDC9A04FA5EFAD74515" box="[1077,1316,1524,1550]" country="Cambodia" county="Popokvil" municipality="April" name="Bokor National Park" pageId="11" pageNumber="256" stateProvince="Kampot">Bokor National Park</location>
, ca.
<locationDeviation id="9A502F8CFFEEFFDC9B6AFA5EFE404529" pageId="11" pageNumber="256">
<quantity id="4CA59B4BFFEEFFDC9B6AFA5EFA5F4515" box="[1371,1452,1524,1551]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="256" unit="km" value="1.5">1.5 km</quantity>
east of Popokvil waterfall
</locationDeviation>
, ca.
<elevation id="0070D19DFFEEFFDC9FDAF9B2FD874529" box="[491,628,1560,1586]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="11" pageNumber="256" unit="m" value="970.0">
<quantity id="4CA59B4BFFEEFFDC9FDAF9B2FDC64529" box="[491,565,1560,1586]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="11" pageNumber="256" unit="m" value="970.0">970 m</quantity>
a.s.l.
</elevation>
,
<date id="FFE3106EFFEEFFDC9CB2F9B2FCE04529" box="[643,787,1560,1586]" pageId="11" pageNumber="256" value="2017-06-01">
<collectingDate id="EFA7E986FFEEFFDC9CB2F9B2FCE04529" box="[643,787,1560,1586]" pageId="11" pageNumber="256" value="2017-06-01">1 June 2017</collectingDate>
</date>
,
<emphasis id="B929EABCFFEEFFDC9D11F9B3FB7D4529" box="[800,1166,1560,1586]" italics="true" pageId="11" pageNumber="256">
<collectorName id="26A85378FFEEFFDC9D11F9B3FC304529" box="[800,963,1560,1586]" pageId="11" pageNumber="256">M. Netopilová</collectorName>
&amp;
<collectorName id="26A85378FFEEFFDC9DDDF9B3FBAE4529" box="[1004,1117,1560,1586]" pageId="11" pageNumber="256">I. Faltová</collectorName>
s.n.
</emphasis>
(mounted over 2 sheets, Accession Nos.
<accessionNumber id="940EAB4DFFEEFFDC9F73F996FE2C454D" box="[322,479,1596,1622]" pageId="11" pageNumber="256">02602KBFR1</accessionNumber>
and
<accessionNumber id="940EAB4DFFEEFFDC9C2AF996FD49454D" box="[539,698,1596,1623]" pageId="11" pageNumber="256">02526KBFR6</accessionNumber>
, herbarium of the Department of Botany and Plant Physiology of the Faculty of Agrobiology, Food and Natural Resources of the Czech University of Life Sciences Prague, Czech Republic, flowering)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE236AEFFEEFFDC9E8CF90DFC9345FD" blockId="11.[136,1452,1415,2090]" pageId="11" pageNumber="256">
<emphasis id="B929EABCFFEEFFDC9E8CF90DFE3E45DA" bold="true" box="[189,461,1703,1729]" pageId="11" pageNumber="256">Other field records:—</emphasis>
<materialsCitation id="3B353CF3FFEEFFDC9FFCF90DFCAF45FD" ID-GBIF-Occurrence="4048240308" collectingDate="2014-06-05" collectionCode="LE" collectorName="H. T. Luu" country="Cambodia" location="Bokor National Park" pageId="11" pageNumber="256" specimenCode="LE01093259" specimenCount="1" stateProvince="Kampot province">
<emphasis id="B929EABCFFEEFFDC9FFCF90DFCAC45DA" bold="true" box="[461,863,1703,1730]" pageId="11" pageNumber="256">
<collectingCountry id="F34A763EFFEEFFDC9FFCF90DFD8345DA" box="[461,624,1703,1729]" name="Cambodia" pageId="11" pageNumber="256">CAMBODIA</collectingCountry>
.
<collectingRegion id="4999F84CFFEEFFDC9CB0F902FCA945DA" box="[641,858,1703,1730]" country="Cambodia" name="Kampot" pageId="11" pageNumber="256">Kampot province</collectingRegion>
:
</emphasis>
<location id="8E826075FFEEFFDC9D5AF902FBAD45D9" LSID="urn:lsid:plazi:treatment:03F487B8FFECFFDB9EB9FB24FC4245DF:8E826075FFEEFFDC9D5AF902FBAD45D9" box="[875,1118,1704,1730]" country="Cambodia" name="Bokor National Park" pageId="11" pageNumber="256" stateProvince="Kampot province">Bokor National Park</location>
,
<date id="FFE3106EFFEEFFDC9A5FF902FAF245D9" box="[1134,1281,1704,1731]" pageId="11" pageNumber="256" value="2014-06-05">
<collectingDate id="EFA7E986FFEEFFDC9A5FF902FAF245D9" box="[1134,1281,1704,1731]" pageId="11" pageNumber="256" value="2014-06-05">5 June 2014</collectingDate>
</date>
,
<emphasis id="B929EABCFFEEFFDC9B20F903FA5845D9" box="[1297,1451,1705,1730]" italics="true" pageId="11" pageNumber="256">
<collectorName id="26A85378FFEEFFDC9B20F903FA8A45D9" box="[1297,1401,1705,1730]" pageId="11" pageNumber="256">H.T. Luu</collectorName>
s.n.
</emphasis>
(flowering, photo uploaded to
<collectionCode id="ED4CAE6BFFEEFFDC9FECF966FDF045FD" box="[477,515,1740,1766]" pageId="11" pageNumber="256">LE</collectionCode>
:
<specimenCode id="DBFB9ED5FFEEFFDC9C3FF966FD5745FC" box="[526,676,1740,1767]" pageId="11" pageNumber="256">LE01093259</specimenCode>
; see also
<figureCitation id="13662A2BFFEEFFDC9D3FF966FCA145FD" box="[782,850,1740,1766]" captionStart="FIGURE 6" captionStartId="11.[136,229,1282,1304]" captionTargetBox="[151,1435,302,1272]" captionTargetId="figure-79@11.[151,1435,302,1272]" captionTargetPageId="11" captionText="FIGURE 6. Geostachys pierreana. A. Leafy shoot (view from below). B. Pseudostem with ligules and leaf bases. C. Basal part of plant with inflorescences. D. Cross section of cincinnus showing total of five flowers/fruits. E. Stamen (adaxial and abaxial view). F. Inflorescence, top view. All from Luu s.n. Photo by H.T. Luu." figureDoi="http://doi.org/10.5281/zenodo.7703858" httpUri="https://zenodo.org/record/7703858/files/figure.png" pageId="11" pageNumber="256">Fig. 6</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3476525FFEEFFDB9E8CF95AFC4245DF" lastPageId="12" lastPageNumber="257" pageId="11" pageNumber="256" type="discussion">
<paragraph id="8BE236AEFFEEFFDC9E8CF95AFD854B31" blockId="11.[136,1452,1415,2090]" pageId="11" pageNumber="256">
<emphasis id="B929EABCFFEEFFDC9E8CF95AFE324411" bold="true" box="[189,449,1776,1802]" pageId="11" pageNumber="256">Nomenclatural notes:</emphasis>
<bibRefCitation id="EFCC4B5FFFEEFFDC9FEFF95AFD444411" author="Gagnepain, F." box="[478,695,1776,1802]" pageId="11" pageNumber="256" pagination="132 - 150" refId="ref10054" refString="Gagnepain, F. (1906 a) Zingiberacees nouvelles de l'herbier du Museum (15 ° note). Bulletin de la Societe Botanique de France 53 (2): 132 - 150. https: // doi. org / 10.1080 / 00378941.1906.10831155" type="journal article" year="1906">Gagnepain (1906a)</bibRefCitation>
included a single gathering,
<emphasis id="B929EABCFFEEFFDC9A39F95BFB634411" box="[1032,1168,1776,1802]" italics="true" pageId="11" pageNumber="256">Pierre 5851</emphasis>
, in the protologue of
<taxonomicName id="4C5D4D2DFFEEFFDC9BA1F95AFF064435" authorityName="Gagnepain" authorityYear="1906" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="11" pageNumber="256" phylum="Tracheophyta" rank="species" species="pierreana">
<emphasis id="B929EABCFFEEFFDC9BA1F95AFF064435" italics="true" pageId="11" pageNumber="256">G. pierreana</emphasis>
</taxonomicName>
. From the title of Gagnepains paper it is evident that the specimens examined by him were located at P. Although the three available sheets at P are currently labelled as type (sensu
<typeStatus id="54E6880CFFEEFFDC9A33F892FB954449" box="[1026,1126,1848,1874]" pageId="11" pageNumber="256" type="holotype">holotype</typeStatus>
) and two isotypes, with this perception repeated by
<bibRefCitation id="EFCC4B5FFFEEFFDC9FA0F8F6FD11446D" author="Saensouk, P. &amp; Saensouk, S." box="[401,738,1884,1910]" pageId="11" pageNumber="256" pagination="3057 - 3067" refId="ref11672" refString="Saensouk, P. &amp; Saensouk, S. (2021) Taxonomic studies and three new records on genus Geostachys (Baker) Ridl. in Thailand. Biodiversitas 22 (7): 3057 - 3067. https: // doi. org / 10.13057 / biodiv / d 220760" type="journal article" year="2021">Saensouk &amp; Saensouk (2021)</bibRefCitation>
, they have to be considered
<typeStatus id="54E6880CFFEEFFDC9A14F8F6FB78446D" box="[1061,1163,1884,1910]" pageId="11" pageNumber="256" type="syntype">syntypes</typeStatus>
. All
<specimenCount id="9D5BFD27FFEEFFDC9AF1F8F6FA91446D" box="[1216,1378,1884,1910]" pageId="11" pageNumber="256" type="generic" typeStatus="syntypes">three syntypes</specimenCount>
are of good quality, each including leafy shoot and one or two inflorescences. One of the sheets (P00686635) also has several flower buds and one neatly pressed flower in an enclosed envelope, which was most likely used in the preparation of pencil drawing attached to the sheet P032698, and subsequently reproduced in Fig.
<quantity id="4CA59B4BFFEEFFDC9ABAF862FB3A44F9" box="[1163,1225,1992,2018]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="256" unit="in" value="10.0">10 in</quantity>
<bibRefCitation id="EFCC4B5FFFEEFFDC9AE4F862FA5B44F9" author="Gagnepain, F." box="[1237,1448,1992,2018]" pageId="11" pageNumber="256" pagination="25 - 121" refId="ref10146" refString="Gagnepain, F. (1908) Zingiberacees. In: Lecomte, H. (Ed.) Flore Generale de l'Indo-Chine, vol. 6. Masson and Co, Paris, pp. 25 - 121." type="book chapter" year="1908">Gagnepain (1908)</bibRefCitation>
. We designate this particular specimen (barcode P00686635) as the
<typeStatus id="54E6880CFFEEFFDC9DA8F846FBF04B1D" box="[921,1027,2028,2054]" pageId="11" pageNumber="256" type="lectotype">lectotype</typeStatus>
; the remaining two sheets (barcodes P00686634 and P032698) are isolectotypes.
</paragraph>
<paragraph id="8BE236AEFFE9FFDB9E8CFF34FAF4428B" blockId="12.[136,1452,158,1733]" pageId="12" pageNumber="257">
<emphasis id="B929EABCFFE9FFDB9E8CFF34FE6343A3" bold="true" box="[189,400,158,184]" pageId="12" pageNumber="257">Taxonomic notes:</emphasis>
<bibRefCitation id="EFCC4B5FFFE9FFDB9F9CFF35FDB843A3" author="Larsen, K." box="[429,587,158,185]" pageId="12" pageNumber="257" pagination="31 - 33" refId="ref10331" refString="Larsen, K. (1986) Geostachys angustifolia sp. nov. (Zingiberaceae) from Thailand. Nordic Journal of Botany 6 (1): 31 - 33. https: // doi. org / 10.1111 / j. 1756 - 1051.1986. tb 00857. x" type="journal article" year="1986">Larsen (1986)</bibRefCitation>
stated that
<taxonomicName id="4C5D4D2DFFE9FFDB9CFEFF35FCAC43A3" authorityName="Gagnepain" authorityYear="1906" box="[719,863,159,185]" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="257" phylum="Tracheophyta" rank="species" species="pierreana">
<emphasis id="B929EABCFFE9FFDB9CFEFF35FCAC43A3" box="[719,863,159,185]" italics="true" pageId="12" pageNumber="257">G. pierreana</emphasis>
</taxonomicName>
is described from
<collectingCountry id="F34A763EFFE9FFDB9A05FF34FB6543A3" box="[1076,1174,158,184]" name="Vietnam" pageId="12" pageNumber="257">Vietnam</collectingCountry>
, whereas
<bibRefCitation id="EFCC4B5FFFE9FFDB9B34FF35FF2043C7" author="Newman, M. &amp; Ketphanh, S. &amp; Svengsuksa, B. &amp; Thomas, P. &amp; Sengdala, K. &amp; Lamxay, V. &amp; Armstong, K." pageId="12" pageNumber="257" refId="ref11113" refString="Newman, M., Ketphanh, S., Svengsuksa, B., Thomas, P., Sengdala, K., Lamxay, V. &amp; Armstong, K. (2007) A Checklist of the Vascular Plants of Lao PDR. Royal Botanic Garden Edinburgh, Edinburgh, 394 pp." type="book" year="2007">
Newman
<emphasis id="B929EABCFFE9FFDB9B43FF35FA5F43A3" box="[1394,1452,158,184]" italics="true" pageId="12" pageNumber="257">et al.</emphasis>
(2007)
</bibRefCitation>
assumed it to be collected in
<collectingCountry id="F34A763EFFE9FFDB9C11FF69FDA943C6" box="[544,602,195,221]" name="Laos" pageId="12" pageNumber="257">Laos</collectingCountry>
.
<bibRefCitation id="EFCC4B5FFFE9FFDB9C55FF68FC4343C7" author="Saensouk, P. &amp; Saensouk, S." box="[612,944,194,220]" pageId="12" pageNumber="257" pagination="3057 - 3067" refId="ref11672" refString="Saensouk, P. &amp; Saensouk, S. (2021) Taxonomic studies and three new records on genus Geostachys (Baker) Ridl. in Thailand. Biodiversitas 22 (7): 3057 - 3067. https: // doi. org / 10.13057 / biodiv / d 220760" type="journal article" year="2021">Saensouk &amp; Saensouk (2021)</bibRefCitation>
produced three conflicting statements within a single paper that the
<typeStatus id="54E6880CFFE9FFDB9FB9FF4DFE4B421A" box="[392,440,231,257]" pageId="12" pageNumber="257">type</typeStatus>
locality is in
<collectingCountry id="F34A763EFFE9FFDB9C62FF4DFD7A421A" box="[595,649,231,257]" name="Laos" pageId="12" pageNumber="257">Laos</collectingCountry>
(as stated in the figure legend on p. 3059), in
<collectingCountry id="F34A763EFFE9FFDB9AA5FF4CFB09421B" box="[1172,1274,230,256]" name="Thailand" pageId="12" pageNumber="257">Thailand</collectingCountry>
,
<collectingRegion id="4999F84CFFE9FFDB9B36FF4CFA70421B" box="[1287,1411,230,256]" country="Thailand" name="Chanthaburi" pageId="12" pageNumber="257">Chantaburi</collectingRegion>
(as stated in species citation on p. 3064) or in
<collectingCountry id="F34A763EFFE9FFDB9C6DFEA0FD3C423F" box="[604,719,266,292]" name="Cambodia" pageId="12" pageNumber="257">Cambodia</collectingCountry>
(as implied in the notes p. 3065). In spite of the previous guesses, the original collection locality of
<taxonomicName id="4C5D4D2DFFE9FFDB9FCDFE85FD794253" authorityName="Gagnepain" authorityYear="1906" box="[508,650,303,329]" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="257" phylum="Tracheophyta" rank="species" species="pierreana">
<emphasis id="B929EABCFFE9FFDB9FCDFE85FD794253" box="[508,650,303,329]" italics="true" pageId="12" pageNumber="257">G. pierreana</emphasis>
</taxonomicName>
remains unknown as already clearly noted by
<bibRefCitation id="EFCC4B5FFFE9FFDB9AA1FE84FA954253" author="Gagnepain, F." box="[1168,1382,302,328]" pageId="12" pageNumber="257" pagination="132 - 150" refId="ref10054" refString="Gagnepain, F. (1906 a) Zingiberacees nouvelles de l'herbier du Museum (15 ° note). Bulletin de la Societe Botanique de France 53 (2): 132 - 150. https: // doi. org / 10.1080 / 00378941.1906.10831155" type="journal article" year="1906">Gagnepain (1906a)</bibRefCitation>
in the protologue: INDO-CHINE.—
<superScript id="7C289BE6FFE9FFDB9FD2FEF9FDF3427B" attach="left" box="[483,512,338,365]" fontSize="6" pageId="12" pageNumber="257">No</superScript>
5851 [
<emphasis id="B929EABCFFE9FFDB9C62FEF9FD694277" box="[595,666,339,364]" italics="true" pageId="12" pageNumber="257">Pierre</emphasis>
] sans désignation de localité, ni date. In
<emphasis id="B929EABCFFE9FFDB9A4DFEF8FF38428B" italics="true" pageId="12" pageNumber="257">Flore Generale de LIndoChine</emphasis>
, the distribution is given, with no further explanation, as “Cochinchine” (
<bibRefCitation id="EFCC4B5FFFE9FFDB9A34FEDCFB09428B" author="Gagnepain, F." box="[1029,1274,374,400]" pageId="12" pageNumber="257" pagination="25 - 121" refId="ref10146" refString="Gagnepain, F. (1908) Zingiberacees. In: Lecomte, H. (Ed.) Flore Generale de l'Indo-Chine, vol. 6. Masson and Co, Paris, pp. 25 - 121." type="book chapter" year="1908">Gagnepain 1908: 102</bibRefCitation>
).
</paragraph>
<paragraph id="8BE236AEFFE9FFDB9E8CFE30FAC8479F" blockId="12.[136,1452,158,1733]" pageId="12" pageNumber="257">
There are three sheets of
<emphasis id="B929EABCFFE9FFDB9FD6FE31FD9C42AF" box="[487,623,410,436]" italics="true" pageId="12" pageNumber="257">
Pierre
<quantity id="4CA59B4BFFE9FFDB9C06FE30FD9C42AF" box="[567,623,410,436]" metricMagnitude="2" metricUnit="m" metricValue="1.486154" pageId="12" pageNumber="257" unit="in" value="5851.0">5851</quantity>
</emphasis>
in P. They all have standard label common to herbarium collections of L. Pierre, with pre-printed locality given as Indochina; however, there is remark on all of them etiquette perdue indicating that the original label was lost. The date of the collection is accordingly given as 18651877, i.e., the entire 12-year long period when Pierre was based in
<collectingRegion id="4999F84CFFE9FFDB9CBDFDACFD29413B" box="[652,730,518,544]" country="Vietnam" name="Ho Chi Minh City" pageId="12" pageNumber="257">Saigon</collectingRegion>
(now
<collectingRegion id="4999F84CFFE9FFDB9D10FDADFC1D413B" box="[801,1006,518,545]" country="Vietnam" name="Ho Chi Minh City" pageId="12" pageNumber="257">Ho Chi Minh City</collectingRegion>
) as the Director of the Jardin Botanique et Zoologique. It is known that during this period Pierre made many expeditions not only in southern
<collectingCountry id="F34A763EFFE9FFDB9AD8FD80FAB8415F" box="[1257,1355,554,580]" name="Vietnam" pageId="12" pageNumber="257">Vietnam</collectingCountry>
, but also in neighbouring
<collectingCountry id="F34A763EFFE9FFDB9F77FDE4FE4A4173" box="[326,441,590,616]" name="Cambodia" pageId="12" pageNumber="257">Cambodia</collectingCountry>
and even
<collectingCountry id="F34A763EFFE9FFDB9C00FDE4FD664173" box="[561,661,590,616]" name="Thailand" pageId="12" pageNumber="257">Thailand</collectingCountry>
(former
<collectingCountry id="F34A763EFFE9FFDB9CCCFDE4FCCB4173" box="[765,824,590,616]" name="Thailand" pageId="12" pageNumber="257">Siam</collectingCountry>
) (
<bibRefCitation id="EFCC4B5FFFE9FFDB9D61FDE4FBEC4173" author="Gagnepain, F." box="[848,1055,590,616]" pageId="12" pageNumber="257" pagination="54 - 59" refId="ref10100" refString="Gagnepain, F. (1906 b) Notice biographique sur J. - B. - Louis Pierre. Bulletin de la Societe Botanique de France 53 (1): 54 - 59. https: // doi. org / 10.1080 / 00378941.1906.10831140" type="journal article" year="1906">Gagnepain 1906b</bibRefCitation>
). The numbering of the specimens is of no help to deduce the location or date as it appears that final numbers on Pierres standard labels were applied as the collections were sorted/worked through, and these differ from his provisional field numbers. We assume this based on examination of several sheets of collection numbers close to 5851 (i.e. 5848, 5852, 5854), which all belong to
<taxonomicName id="4C5D4D2DFFE9FFDB9EB9FD74FED941E3" authorityName="Martinov" authorityYear="1820" box="[136,298,734,760]" class="Liliopsida" family="Zingiberaceae" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="257" phylum="Tracheophyta" rank="family">Zingiberaceae</taxonomicName>
, but all were collected at different locations and in different years. There is no sheet of
<taxonomicName id="4C5D4D2DFFE9FFDB9ADFFD74FA9C41E3" authorityName="Ridley" authorityYear="1899" baseAuthorityName="Baker" baseAuthorityYear="1892" box="[1262,1391,734,760]" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="257" phylum="Tracheophyta" rank="genus">
<emphasis id="B929EABCFFE9FFDB9ADFFD74FA9C41E3" box="[1262,1391,734,760]" italics="true" pageId="12" pageNumber="257">Geostachys</emphasis>
</taxonomicName>
from
<collectingCountry id="F34A763EFFE9FFDB9EB9FCA8FF144007" box="[136,231,770,796]" name="Vietnam" pageId="12" pageNumber="257">Vietnam</collectingCountry>
present in any herbaria we have examined, which would be in good match with the
<typeStatus id="54E6880CFFE9FFDB9ABEFCA9FB4C4006" box="[1167,1215,771,797]" pageId="12" pageNumber="257">type</typeStatus>
of
<taxonomicName id="4C5D4D2DFFE9FFDB9AD5FCA9FA814007" authorityName="Gagnepain" authorityYear="1906" box="[1252,1394,771,797]" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="257" phylum="Tracheophyta" rank="species" species="pierreana">
<emphasis id="B929EABCFFE9FFDB9AD5FCA9FA814007" box="[1252,1394,771,797]" italics="true" pageId="12" pageNumber="257">G. pierreana</emphasis>
</taxonomicName>
. The only specimens which we consider to be in good match with the
<typeStatus id="54E6880CFFE9FFDB9D65FC8DFC77405A" box="[852,900,807,833]" pageId="12" pageNumber="257">type</typeStatus>
is collection by
<emphasis id="B929EABCFFE9FFDB9A0AFC8DFB2A405B" box="[1083,1241,806,832]" italics="true" pageId="12" pageNumber="257">Dy Phon 1157</emphasis>
and collection and photographs made by
<emphasis id="B929EABCFFE9FFDB9FB4FCE0FD54407F" box="[389,679,842,869]" italics="true" pageId="12" pageNumber="257">Netopilová &amp; Faltová s.n</emphasis>
., both from Bokor National Park,
<collectingCountry id="F34A763EFFE9FFDB9A03FCE0FB5A407F" box="[1074,1193,842,868]" name="Cambodia" pageId="12" pageNumber="257">Cambodia</collectingCountry>
. These two collections match the
<typeStatus id="54E6880CFFE9FFDB9ECFFCC5FEDD4092" box="[254,302,879,905]" pageId="12" pageNumber="257">type</typeStatus>
and the protologue in all major aspects, such as the shape and indumentum of leaf blades and ligules, short petioles, length and decurvation of the rachis and secund arrangement of the flower-bearing branches, number of flowers in cincinnus, as well as more specific details such as hirsute rachis and flower-bearing branches, shape and size of bracts, spathaceous glabrous bracteoles with mucronate puberulent apices and calyx with a single lobe ending in a sharply mucronate apex. From other labelled collections, it is known that Pierre visited and made collections in the same area (
<collectingRegion id="4999F84CFFE9FFDB9F03FB88FE7D4727" box="[306,398,1058,1084]" country="Cambodia" name="Kampot" pageId="12" pageNumber="257">Kampot</collectingRegion>
city and surrounding hills). Thus with all the evidence listed above, the most likely origin of the
<typeStatus id="54E6880CFFE9FFDB9E80FBEDFF12477A" box="[177,225,1095,1121]" pageId="12" pageNumber="257">type</typeStatus>
of
<taxonomicName id="4C5D4D2DFFE9FFDB9F36FBEDFE65477B" authorityName="Gagnepain" authorityYear="1906" box="[263,406,1095,1121]" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="257" phylum="Tracheophyta" rank="species" species="pierreana">
<emphasis id="B929EABCFFE9FFDB9F36FBEDFE65477B" box="[263,406,1095,1121]" italics="true" pageId="12" pageNumber="257">G. pierreana</emphasis>
</taxonomicName>
is
<collectingCountry id="F34A763EFFE9FFDB9F89FBECFDDC477B" box="[440,559,1094,1120]" name="Cambodia" pageId="12" pageNumber="257">Cambodia</collectingCountry>
. The above description, although based on the protologue and the three original sheets, is therefore also supplemented with observations from
<emphasis id="B929EABCFFE9FFDB9D73FBC1FC12479F" box="[834,993,1130,1156]" italics="true" pageId="12" pageNumber="257">Dy Phon 1157</emphasis>
and
<emphasis id="B929EABCFFE9FFDB9A29FBC0FAC8479F" box="[1048,1339,1130,1157]" italics="true" pageId="12" pageNumber="257">Netopilová &amp; Faltová s.n.</emphasis>
</paragraph>
<paragraph id="8BE236AEFFE9FFDB9E8CFB24FBD046BF" blockId="12.[136,1452,158,1733]" pageId="12" pageNumber="257">
The indication of
<taxonomicName id="4C5D4D2DFFE9FFDB9FB9FB25FDEB47B3" authorityName="Gagnepain" authorityYear="1906" box="[392,536,1167,1193]" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="257" phylum="Tracheophyta" rank="species" species="pierreana">
<emphasis id="B929EABCFFE9FFDB9FB9FB25FDEB47B3" box="[392,536,1167,1193]" italics="true" pageId="12" pageNumber="257">G. pierreana</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFCC4B5FFFE9FFDB9C75FB24FD2547B3" author="Ph am, H. H. &amp; Vietnamese" box="[580,726,1166,1192]" pageId="12" pageNumber="257" refId="ref11381" refString="Ph am, H. H. (2003) Cay c o Vi et Nam, an illustrated Flora of Vietnam. Vol. 3. Youth Publication, Ho Chi Minh City, 1020 pp. [in Vietnamese]" type="book" year="2003">Phạm (2003)</bibRefCitation>
as present in
<collectingCountry id="F34A763EFFE9FFDB9D43FB24FC2247B3" box="[882,977,1166,1192]" name="Vietnam" pageId="12" pageNumber="257">Vietnam</collectingCountry>
originated from the uncertainty regarding the
<typeStatus id="54E6880CFFE9FFDB9E80FB19FF1247D6" box="[177,225,1203,1229]" pageId="12" pageNumber="257">type</typeStatus>
collection, as follows from his reference to Gagnepain. The same is probably true for Nguyễn (2005). Nguyễn (2017) listed several additional Vietnamese collections under
<taxonomicName id="4C5D4D2DFFE9FFDB9D02FB7DFC3247EB" authorityName="Gagnepain" authorityYear="1906" box="[819,961,1239,1265]" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="257" phylum="Tracheophyta" rank="species" species="pierreana">
<emphasis id="B929EABCFFE9FFDB9D02FB7DFC3247EB" box="[819,961,1239,1265]" italics="true" pageId="12" pageNumber="257">G. pierreana</emphasis>
</taxonomicName>
whose identifications remained preliminary as they were based solely on the herbarium material that does not show important features of floral structure. For two collections from
<collectingRegion id="4999F84CFFE9FFDB9F77FAB4FDDB4623" box="[326,552,1310,1339]" country="Vietnam" name="Ninh Binh" pageId="12" pageNumber="257">Ninh BÌnh province</collectingRegion>
,
<emphasis id="B929EABCFFE9FFDB9C02FAB5FC0F4623" box="[563,1020,1310,1337]" italics="true" pageId="12" pageNumber="257">Averyanov et al. HAL 1638 &amp; HAL 1639</emphasis>
, photographic material appeared to be available, which allowed to identify them as
<taxonomicName id="4C5D4D2DFFE9FFDB9CBCFAE8FC1D4647" box="[653,1006,1346,1373]" class="Liliopsida" family="Zingiberaceae" genus="Wurfbainia" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="257" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B929EABCFFE9FFDB9CBCFAE8FCF84647" box="[653,779,1346,1372]" italics="true" pageId="12" pageNumber="257">Wurfbainia</emphasis>
sp. (Zingiberaceae)
</taxonomicName>
and
<taxonomicName id="4C5D4D2DFFE9FFDB9A18FAE8FEDA469B" authority="(Lindl.) Blume" authorityName="Blume" baseAuthorityName="Lindl." class="Liliopsida" family="Orchidaceae" genus="Tropidia" kingdom="Plantae" order="Asparagales" pageId="12" pageNumber="257" phylum="Tracheophyta" rank="species" species="angulosa">
<emphasis id="B929EABCFFE9FFDB9A18FAE8FB0A4647" box="[1065,1273,1346,1372]" italics="true" pageId="12" pageNumber="257">Tropidia angulosa</emphasis>
(Lindl.) Blume (Orchidaceae)
</taxonomicName>
, respectively. The other collections listed by Nguyễn (2017), all from Lâm Đồng province (
<emphasis id="B929EABCFFE9FFDB9B12FACDFA57469B" box="[1315,1444,1382,1411]" italics="true" pageId="12" pageNumber="257">Hà Tuḗ 458</emphasis>
,
<emphasis id="B929EABCFFE9FFDB9EB9FA20FF0546BF" box="[136,246,1418,1444]" italics="true" pageId="12" pageNumber="257">Nhan 785</emphasis>
,
<emphasis id="B929EABCFFE9FFDB9F35FA21FEB546BF" box="[260,326,1418,1444]" italics="true" pageId="12" pageNumber="257">Tṵ 82</emphasis>
,
<emphasis id="B929EABCFFE9FFDB9F65FA20FE3546BF" box="[340,454,1418,1444]" italics="true" pageId="12" pageNumber="257">Biên 1228</emphasis>
), are re-identified here as
<taxonomicName id="4C5D4D2DFFE9FFDB9CDCFA21FC6046BF" box="[749,915,1419,1445]" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="257" phylum="Tracheophyta" rank="species" species="annamensis">
<emphasis id="B929EABCFFE9FFDB9CDCFA21FC6046BF" box="[749,915,1419,1445]" italics="true" pageId="12" pageNumber="257">G. annamensis</emphasis>
</taxonomicName>
(see above).
</paragraph>
<paragraph id="8BE236AEFFE9FFDB9E8CFA04FC4245DF" blockId="12.[136,1452,158,1733]" pageId="12" pageNumber="257">
<bibRefCitation id="EFCC4B5FFFE9FFDB9E8CFA04FDF746D3" author="Saensouk, P. &amp; Saensouk, S." box="[189,516,1454,1480]" pageId="12" pageNumber="257" pagination="3057 - 3067" refId="ref11672" refString="Saensouk, P. &amp; Saensouk, S. (2021) Taxonomic studies and three new records on genus Geostachys (Baker) Ridl. in Thailand. Biodiversitas 22 (7): 3057 - 3067. https: // doi. org / 10.13057 / biodiv / d 220760" type="journal article" year="2021">Saensouk &amp; Saensouk (2021)</bibRefCitation>
have reported
<taxonomicName id="4C5D4D2DFFE9FFDB9C97FA05FCC046D3" authorityName="Gagnepain" authorityYear="1906" box="[678,819,1455,1481]" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="257" phylum="Tracheophyta" rank="species" species="pierreana">
<emphasis id="B929EABCFFE9FFDB9C97FA05FCC046D3" box="[678,819,1455,1481]" italics="true" pageId="12" pageNumber="257">G. pierreana</emphasis>
</taxonomicName>
as a new record for
<collectingCountry id="F34A763EFFE9FFDB9A3CFA04FB8046D3" box="[1037,1139,1454,1480]" name="Thailand" pageId="12" pageNumber="257">Thailand</collectingCountry>
, based on a single collection made by
<emphasis id="B929EABCFFE9FFDB9EC2FA79FE8546F6" box="[243,374,1491,1517]" italics="true" pageId="12" pageNumber="257">Nai Noe 77</emphasis>
(BK, K, P: P00599363) in
<collectingRegion id="4999F84CFFE9FFDB9C84FA78FC5D46F7" box="[693,942,1490,1516]" country="Thailand" name="Chanthaburi" pageId="12" pageNumber="257">Chanthaburi province</collectingRegion>
. Although the specimen has been previously annotated as
<taxonomicName id="4C5D4D2DFFE9FFDB9F2CFA5CFDCB450B" authorityName="Gagnepain" authorityYear="1906" box="[285,568,1526,1552]" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" isUncertain="true" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="257" phylum="Tracheophyta" rank="species" species="pierreana">
<emphasis id="B929EABCFFE9FFDB9F2CFA5CFE6D450B" box="[285,414,1526,1552]" italics="true" pageId="12" pageNumber="257">Geostachys</emphasis>
cf.
<emphasis id="B929EABCFFE9FFDB9FFAFA5DFDCB450B" box="[459,568,1527,1552]" italics="true" pageId="12" pageNumber="257">pierreana</emphasis>
</taxonomicName>
on the sheet deposited in P, it is clearly visible that the inflorescence is erect, with cincinni/flower-bearing branches evenly distributed around the rachis and thus fails to fit
<taxonomicName id="4C5D4D2DFFE9FFDB9A91F9B1FADD452F" authorityName="Gagnepain" authorityYear="1906" box="[1184,1326,1563,1589]" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="257" phylum="Tracheophyta" rank="species" species="pierreana">
<emphasis id="B929EABCFFE9FFDB9A91F9B1FADD452F" box="[1184,1326,1563,1589]" italics="true" pageId="12" pageNumber="257">G. pierreana</emphasis>
</taxonomicName>
, which has decurved inflorescences with secund arrangement of flower-bearing branches. Although determination to the species level with absolute certainty is challenging on the herbarium material, the collection
<emphasis id="B929EABCFFE9FFDB9A50F9C9FB154566" box="[1121,1254,1635,1661]" italics="true" pageId="12" pageNumber="257">Nai Noe 77</emphasis>
matches well
<taxonomicName id="4C5D4D2DFFE9FFDB9BA1F9C9FF1B45BB" authorityName="Picheansoonthon &amp; Mayoe" authorityYear="2011" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="257" phylum="Tracheophyta" rank="species" species="tratensis">
<emphasis id="B929EABCFFE9FFDB9BA1F9C9FF1B45BB" italics="true" pageId="12" pageNumber="257">G. tratensis</emphasis>
</taxonomicName>
, a species described from a location distanced only about
<quantity id="4CA59B4BFFE9FFDB9D41F92CFC4445BB" box="[880,951,1670,1696]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="257" unit="km" value="80.0">80 km</quantity>
(air distance) from it. We therefore conclude that there is currently no evidence, that
<taxonomicName id="4C5D4D2DFFE9FFDB9C72F901FD2145DF" authorityName="Gagnepain" authorityYear="1906" box="[579,722,1707,1733]" class="Liliopsida" family="Zingiberaceae" genus="Geostachys" kingdom="Plantae" order="Zingiberales" pageId="12" pageNumber="257" phylum="Tracheophyta" rank="species" species="pierreana">
<emphasis id="B929EABCFFE9FFDB9C72F901FD2145DF" box="[579,722,1707,1733]" italics="true" pageId="12" pageNumber="257">G. pierreana</emphasis>
</taxonomicName>
occurs in
<collectingCountry id="F34A763EFFE9FFDB9D77F900FC5E45DF" box="[838,941,1706,1732]" name="Thailand" pageId="12" pageNumber="257">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>