172 lines
17 KiB
XML
172 lines
17 KiB
XML
<document id="4D00327E54B21EA31F6F5464A0F3F553" ID-CLB-Dataset="45658" ID-DOI="10.11646/zootaxa.3802.4.1" ID-GBIF-Dataset="43524c34-a5b7-4f7e-908a-3e3b61ffc6db" ID-ISSN="1175-5326" ID-Zenodo-Dep="252490" ID-ZooBank="CF479CC3-C014-460D-9C71-3A6C2AB2778B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460427181093" checkinUser="plazi" docAuthor="Jørgensen, Aslak, Boesgaard, Tom M., Møbjerg, Nadja & Kristensen, Reinhardt M." docDate="2014" docId="03F487B7FFB0FFEA68CE1A41D2A9A3C7" docLanguage="en" docName="zt03802p443.pdf" docOrigin="Zootaxa 3802 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Raiarctus Renaud-Mornant 1981" docType="treatment" docVersion="10" lastPageNumber="424" masterDocId="FFCDFFCFFFA7FFFD68591B0DD61FA000" masterDocTitle="The tardigrade fauna of Australian marine caves: With descriptions of nine new species of Arthrotardigrada" masterLastPageNumber="443" masterPageNumber="401" pageNumber="424" updateTime="1698330854638" updateUser="ExternalLinkService">
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<mods:mods id="10BC4430E4089BD8D0D63E0A8CEBC2F0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="7DEF4C20AD527B410597DED27E90A6A9">
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<mods:title id="73C063BFB01012AFEB075F68EB4882D9">The tardigrade fauna of Australian marine caves: With descriptions of nine new species of Arthrotardigrada</mods:title>
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<mods:namePart id="78010023EF4B7B4A643176FF08E6354E">Jørgensen, Aslak</mods:namePart>
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<mods:namePart id="C6ECE7E92D3977B715F5F07F6431D39E">Boesgaard, Tom M.</mods:namePart>
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<mods:namePart id="A19ED113AE057ECD7D552DED4AA5298B">Møbjerg, Nadja</mods:namePart>
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<mods:name id="2BC8FC78C0E31C9AC91018C898A6806D" type="personal">
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<mods:namePart id="BF553F5703D4CF29348C9A8CB3EDB1A0">Kristensen, Reinhardt M.</mods:namePart>
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<mods:title id="164DE4ECF427594D2F8806D0D3E9BF72">Zootaxa</mods:title>
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<mods:date id="2B2FE047D0C1AE608EF30B716DB81847">2014</mods:date>
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<mods:number id="1997D45E5A4E62BFBD61E54C79BEF2E3">3802</mods:number>
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<treatment id="03F487B7FFB0FFEA68CE1A41D2A9A3C7" ID-DOI="http://doi.org/10.5281/zenodo.5691093" ID-GBIF-Taxon="119428905" ID-Zenodo-Dep="5691093" LSID="urn:lsid:plazi:treatment:03F487B7FFB0FFEA68CE1A41D2A9A3C7" httpUri="http://treatment.plazi.org/id/03F487B7FFB0FFEA68CE1A41D2A9A3C7" lastPageNumber="424" pageId="23" pageNumber="424">
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<subSubSection id="C347652AFFB0FFEA68CE1A41D48FA166" box="[151,656,332,358]" pageId="23" pageNumber="424" type="nomenclature">
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<paragraph id="8BE236A1FFB0FFEA68CE1A41D48FA166" blockId="23.[151,656,332,358]" box="[151,656,332,358]" pageId="23" pageNumber="424">
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<heading id="D0AA81CDFFB0FFEA68CE1A41D48FA166" bold="true" box="[151,656,332,358]" fontSize="11" level="1" pageId="23" pageNumber="424" reason="1">
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<emphasis id="B929EAB3FFB0FFEA68CE1A41D48FA166" bold="true" box="[151,656,332,358]" pageId="23" pageNumber="424">
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Genus
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<taxonomicName id="4C5D4D22FFB0FFEA68B41A41D48FA166" ID-CoL="642GW" authority="Renaud-Mornant, 1981" authorityName="Renaud-Mornant" authorityYear="1981" box="[237,656,332,358]" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="genus">
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<emphasis id="B929EAB3FFB0FFEA68B41A41D77DA166" bold="true" box="[237,354,332,358]" italics="true" pageId="23" pageNumber="424">Raiarctus</emphasis>
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Renaud-Mornant, 1981
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C347652AFFB0FFEA68CE1A9FD2E2A23B" pageId="23" pageNumber="424" type="diagnosis">
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<paragraph id="8BE236A1FFB0FFEA68CE1A9FD2E2A23B" blockId="23.[151,1437,402,967]" pageId="23" pageNumber="424">
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<emphasis id="B929EAB3FFB0FFEA68CE1A9FD78EA1AB" bold="true" box="[151,401,402,427]" pageId="23" pageNumber="424">Diagnosis (emended)</emphasis>
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:
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<taxonomicName id="4C5D4D22FFB0FFEA69F81A9FD450A1AB" box="[417,591,402,427]" class="Heterotardigrada" family="Halechiniscidae" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="subFamily" subFamily="Styraconyxinae">Styraconyxinae</taxonomicName>
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with the dorso-lateral epicuticular coat supported by long thin pillars 5–6 µm regularly arranged in close parallel lines. With 3-pointed claws, with or without heart-shaped pads on the internal digits. With or without peduncles on every tarsus. Minor primary clava with a cone or tube-like shape; the short, secondary clavae either reduced or minor. Body dorso-ventrally flattened. With lateral tube-like openings to the seminal receptacles. Halechiniscid male gonopore (oval papilla with a crescent-shaped opening).
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</paragraph>
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</subSubSection>
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<subSubSection id="C347652AFFB0FFEA689E194BD463A2A7" pageId="23" pageNumber="424" type="materials_examined">
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<paragraph id="8BE236A1FFB0FFEA689E194BD521A25F" blockId="23.[151,1437,402,967]" box="[199,830,582,607]" pageId="23" pageNumber="424">
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<emphasis id="B929EAB3FFB0FFEA689E194BD77BA25F" bold="true" box="[199,356,582,607]" pageId="23" pageNumber="424">
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<typeStatus id="54E68803FFB0FFEA689E194BD71DA25F" box="[199,258,582,607]" pageId="23" pageNumber="424">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C5D4D22FFB0FFEA6932194AD521A25F" authority="Renaud-Mornant, 1981" authorityName="Renaud-Mornant" authorityYear="1981" box="[363,830,582,607]" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="species" species="colurus">
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<emphasis id="B929EAB3FFB0FFEA6932194AD42DA25F" box="[363,562,582,607]" italics="true" pageId="23" pageNumber="424">Raiarctus colurus</emphasis>
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Renaud-Mornant, 1981
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</taxonomicName>
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</paragraph>
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<paragraph id="8BE236A1FFB0FFEA689E1967D463A2A7" blockId="23.[151,1437,402,967]" pageId="23" pageNumber="424">
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<emphasis id="B929EAB3FFB0FFEA689E1967D7B0A283" bold="true" box="[199,431,618,643]" pageId="23" pageNumber="424">Additional species:</emphasis>
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<taxonomicName id="4C5D4D22FFB0FFEA69E21961D5DAA283" authority="Renaud-Mornant, 1981" authorityName="Renaud-Mornant" authorityYear="1981" box="[443,965,618,643]" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="species" species="aureolatus">
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<emphasis id="B929EAB3FFB0FFEA69E21961D4B4A283" box="[443,683,618,643]" italics="true" pageId="23" pageNumber="424">Raiarctus aureolatus</emphasis>
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Renaud-Mornant, 1981
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</taxonomicName>
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;
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<taxonomicName id="4C5D4D22FFB0FFEA6B8F1961D6E3A2A7" authority="D'Addabbo Gallo, Grimaldi" authorityName="D'Addabbo Gallo, Grimaldi" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="species" species="variabilis">
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<emphasis id="B929EAB3FFB0FFEA6B8F1961D2A5A283" box="[982,1210,618,643]" italics="true" pageId="23" pageNumber="424">Raiarctus variabilis</emphasis>
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D’Addabbo Gallo, Grimaldi
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</taxonomicName>
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de Zio & Morone De
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<bibRefCitation id="EFCC4B50FFB0FFEA69AF1983D463A2A7" author="Lucia" box="[502,636,654,679]" pageId="23" pageNumber="424" refString="D'Addabbo Gallo, M., Grimaldi de Zio, S. & Morone de Lucia, M. R. (1986) Description of two species of Halechiniscidae belonging to the sub-families Halechiniscinae and Styraconyxinae (Tardigrada: Heterotardigrada). Oebalia, 13, 69 - 75." type="journal article" year="1986">Lucia, 1986</bibRefCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C347652AFFB0FFEA689E19BFD2A9A3C7" pageId="23" pageNumber="424" type="discussion">
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||
<paragraph id="8BE236A1FFB0FFEA689E19BFD2A9A3C7" blockId="23.[151,1437,402,967]" pageId="23" pageNumber="424">
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<emphasis id="B929EAB3FFB0FFEA689E19BFD724A2CB" bold="true" box="[199,315,690,715]" pageId="23" pageNumber="424">Remarks.</emphasis>
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The genus
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<taxonomicName id="4C5D4D22FFB0FFEA699A19B9D430A2CB" box="[451,559,692,715]" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="genus">
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<emphasis id="B929EAB3FFB0FFEA699A19B9D430A2CB" box="[451,559,692,715]" italics="true" pageId="23" pageNumber="424">Raiarctus</emphasis>
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</taxonomicName>
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currently consists of
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<taxonomicName id="4C5D4D22FFB0FFEA6B7119B9D5EEA2CB" box="[808,1009,690,715]" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="species" species="colurus">
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<emphasis id="B929EAB3FFB0FFEA6B7119B9D5EEA2CB" box="[808,1009,690,715]" italics="true" pageId="23" pageNumber="424">Raiarctus colurus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C5D4D22FFB0FFEA6C5819B9D286A2CB" box="[1025,1177,690,715]" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="species" species="aureolatus">
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<emphasis id="B929EAB3FFB0FFEA6C5819B9D286A2CB" box="[1025,1177,690,715]" italics="true" pageId="23" pageNumber="424">R. aureolatus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C5D4D22FFB0FFEA6C8D19B9D37FA2CB" box="[1236,1376,690,715]" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="species" species="variabilis">
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<emphasis id="B929EAB3FFB0FFEA6C8D19B9D37FA2CB" box="[1236,1376,690,715]" italics="true" pageId="23" pageNumber="424">R. variabilis</emphasis>
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</taxonomicName>
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. The large variation in size and shape of several characters such as primary clava, cuticular pillars and cirrus E, meant that
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<taxonomicName id="4C5D4D22FFB0FFEA689119F1D723A313" box="[200,316,762,787]" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="species" species="colurus">
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<emphasis id="B929EAB3FFB0FFEA689119F1D723A313" box="[200,316,762,787]" italics="true" pageId="23" pageNumber="424">R. colurus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C5D4D22FFB0FFEA692D19F1D415A313" box="[372,522,762,787]" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="species" species="aureolatus">
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<emphasis id="B929EAB3FFB0FFEA692D19F1D415A313" box="[372,522,762,787]" italics="true" pageId="23" pageNumber="424">R. aureolatus</emphasis>
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</taxonomicName>
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were considered as variable forms of
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<taxonomicName id="4C5D4D22FFB0FFEA6BE319F1D25BA313" box="[954,1092,762,787]" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="species" species="variabilis">
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<emphasis id="B929EAB3FFB0FFEA6BE319F1D25BA313" box="[954,1092,762,787]" italics="true" pageId="23" pageNumber="424">R. variabilis</emphasis>
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</taxonomicName>
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(Grimaldi de Zio
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<emphasis id="B929EAB3FFB0FFEA6D4A19F1D353A313" box="[1299,1356,762,787]" italics="true" pageId="23" pageNumber="424">et al.</emphasis>
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1983). After a more detailed study of a larger number of
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<taxonomicName id="4C5D4D22FFB0FFEA6A831812D559A336" box="[730,838,799,822]" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="genus">
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<emphasis id="B929EAB3FFB0FFEA6A831812D559A336" box="[730,838,799,822]" italics="true" pageId="23" pageNumber="424">Raiarctus</emphasis>
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</taxonomicName>
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samples the taxonomic status was re-established by D’Addabbo
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<bibRefCitation id="EFCC4B50FFB0FFEA697F184FD41AA35B" box="[294,517,834,859]" pageId="23" pageNumber="424" refString="D'Addabbo Gallo, M., Grimaldi de Zio, S. & Morone de Lucia, M. R. (1986) Description of two species of Halechiniscidae belonging to the sub-families Halechiniscinae and Styraconyxinae (Tardigrada: Heterotardigrada). Oebalia, 13, 69 - 75." type="journal article">
|
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Gallo
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||
<emphasis id="B929EAB3FFB0FFEA69291849D7B2A35B" box="[368,429,834,859]" italics="true" pageId="23" pageNumber="424">et al.</emphasis>
|
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(1986)
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||
</bibRefCitation>
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||
. The description of
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<taxonomicName id="4C5D4D22FFB0FFEA6AAF1849D599A35B" box="[758,902,834,859]" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="species" species="variabilis">
|
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<emphasis id="B929EAB3FFB0FFEA6AAF1849D599A35B" box="[758,902,834,859]" italics="true" pageId="23" pageNumber="424">R. variabilis</emphasis>
|
||
</taxonomicName>
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fits on several characters to both of the new Australian
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<taxonomicName id="4C5D4D22FFB0FFEA694D186AD79FA37E" box="[276,384,871,894]" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="genus">
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<emphasis id="B929EAB3FFB0FFEA694D186AD79FA37E" box="[276,384,871,894]" italics="true" pageId="23" pageNumber="424">Raiarctus</emphasis>
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||
</taxonomicName>
|
||
species, e.g. the size and arrangement of the pillars in parallel lines. Genital papillae, which are openings for the seminal receptacles, as described in
|
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<taxonomicName id="4C5D4D22FFB0FFEA6B391881D5EBA3A3" box="[864,1012,906,931]" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="species" species="variabilis">
|
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<emphasis id="B929EAB3FFB0FFEA6B391881D5EBA3A3" box="[864,1012,906,931]" italics="true" pageId="23" pageNumber="424">R. variabilis</emphasis>
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</taxonomicName>
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and seen in several specimens of Mediterranean
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<taxonomicName id="4C5D4D22FFB0FFEA691818A2D7AAA3C7" box="[321,437,942,967]" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="species" species="colurus">
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<emphasis id="B929EAB3FFB0FFEA691818A2D7AAA3C7" box="[321,437,942,967]" italics="true" pageId="23" pageNumber="424">R. colurus</emphasis>
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</taxonomicName>
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and
|
||
<taxonomicName id="4C5D4D22FFB0FFEA69B218A2D49EA3C7" box="[491,641,942,967]" class="Heterotardigrada" family="Halechiniscidae" genus="Raiarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="23" pageNumber="424" phylum="Tardigrada" rank="species" species="aureolatus">
|
||
<emphasis id="B929EAB3FFB0FFEA69B218A2D49EA3C7" box="[491,641,942,967]" italics="true" pageId="23" pageNumber="424">R. aureolatus</emphasis>
|
||
</taxonomicName>
|
||
, were not clearly visible in the Australian species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |