177 lines
19 KiB
XML
177 lines
19 KiB
XML
<document id="E46BFB958EEBBF8C08FBC7BB15B6BAC4" ID="10.11646/zootaxa.4273.4.2SLASH11258" ID-CLB-Dataset="32892" ID-DOI="10.11646/zootaxa.4273.4.2" ID-GBIF-Dataset="316e1968-2a32-462e-a554-99165078f1fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="803688" ID-ZooBank="77091F8E-2A3C-4A9B-81C2-656B5BF72C22" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1496820183743" checkinUser="plazi" docAuthor="Majnon-Jahromi, Bahareh, Gheibi, Mehdi, Fallahzadeh, Majid, Kehlmaier, Christian & Hesami, Shahram" docDate="2017" docId="03F487A3B078FF9FB8D3FF39B77063D6" docLanguage="en" docName="zootaxa.4273.4.2SLASH11258.pdf" docOrigin="Zootaxa 4273 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tomosvaryella pistacia Majnon-Jahromi & Kehlmaier, sp. nov." docType="treatment" docVersion="7" lastPageNumber="498" masterDocId="FFCDFFDBB071FF95B844FC24B20A632E" masterDocTitle="Pipunculidae (Diptera) from southern Iran, including two new species of the genus Tomosvaryella Aczél" masterLastPageNumber="500" masterPageNumber="488" pageNumber="497" updateTime="1698440239418" updateUser="ExternalLinkService">
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<mods:mods id="7127AC21048507B9E45F8F346F21C469" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="314DC095A5A2DC726ED78CD71F870E64">
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<mods:title id="2D34D3CA3DE4145FF504C0F16105CF34">Pipunculidae (Diptera) from southern Iran, including two new species of the genus Tomosvaryella Aczél</mods:title>
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</mods:titleInfo>
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<mods:name id="824750BCE67161B97399EE094BB2A779" type="personal">
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<mods:role id="04412EA4C0D1BB5664FCF86B827E61B7">
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<mods:roleTerm id="5B4BD28AECA572B73D54DD6FE420C03D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4D16CFD4AF879B5FA13D50F13CA439AD">Majnon-Jahromi, Bahareh</mods:namePart>
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</mods:name>
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<mods:name id="4EE17C328974081CF0727CD14E7A0D16" type="personal">
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<mods:role id="AF71A6B6D2D5EF0FE0571918DB9C7663">
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<mods:roleTerm id="0EBA1E102D3770D813302386F9589101">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C1704D4A9D312449BB82687C2B57859D">Gheibi, Mehdi</mods:namePart>
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</mods:name>
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<mods:name id="7414929C3734BDF80D3FB016C91D6175" type="personal">
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<mods:roleTerm id="BD7D0661BA7373D1D7E486E8F26133BE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8DABD6BC248811751AE1381CE9F8DA00">Fallahzadeh, Majid</mods:namePart>
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</mods:name>
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<mods:name id="2CF493752B287395F0CEA329F958D6A7" type="personal">
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<mods:roleTerm id="B1EBB81B9ABC340C8850E20E2E417ED9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="696D6C8BC73B8652658AAD5F37CF6AC6">Kehlmaier, Christian</mods:namePart>
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</mods:name>
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<mods:name id="432BAC67F09F261921BD8C17AA0C6E54" type="personal">
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<mods:role id="BA53C78E3E547FD2F2651B2D431B52B8">
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<mods:roleTerm id="598CA10015D226DA2AACDA87E9F1A037">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FB00C90C249E7DF6F3D7A518C84D34E4">Hesami, Shahram</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="2286C56B6572983118486D6B07152C4A">text</mods:typeOfResource>
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<mods:title id="DA6812CD6383EB76550B9DFF50F2538E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E6D67749830F340D55484891E1F2747F">
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<mods:date id="27EFE2A13AFF168E9FB5EFEB3A42BCE9">2017</mods:date>
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<mods:detail id="9AB834DF2C90074ECB752B04D3F8E586" type="volume">
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<mods:number id="D535942B1F438E7D2263D5302A2290CE">4273</mods:number>
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</mods:detail>
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<mods:detail id="506C3712246A0DB95855A4A2ACBD7DFB" type="issue">
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<mods:number id="421094AC3C20CD76FB40D3397530A8FD">4</mods:number>
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<mods:start id="93D48D15DDD0E9932E4E65C7AF9FFC78">488</mods:start>
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<mods:end id="BE030FE3651ECF3F31B55E5989C67462">500</mods:end>
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<mods:classification id="574C05AA1285C4CE0097FC3389E2DE70">journal article</mods:classification>
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<mods:identifier id="92AE9BB3C5B0CA27BBE3C7DE900A932F" type="CLB-Dataset">32892</mods:identifier>
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<mods:identifier id="AF2E785BE19DF770EB1AD042AEB8E3D8" type="DOI">10.11646/zootaxa.4273.4.2</mods:identifier>
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<mods:identifier id="D1ADF12350C9C305A6AD7DB33511F384" type="GBIF-Dataset">316e1968-2a32-462e-a554-99165078f1fc</mods:identifier>
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<mods:identifier id="DBC9D60D2CF3BADF21397BD30363E0ED" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="ED9FA32061F174415EC5B2D9CAB23D10" type="Zenodo-Dep">803688</mods:identifier>
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<mods:identifier id="F9F549C74F25DC258B9C05E3F7571973" type="ZooBank">77091F8E-2A3C-4A9B-81C2-656B5BF72C22</mods:identifier>
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<treatment id="03F487A3B078FF9FB8D3FF39B77063D6" ID-DOI="http://doi.org/10.5281/zenodo.6020335" ID-GBIF-Taxon="131812472" ID-Zenodo-Dep="6020335" LSID="urn:lsid:plazi:treatment:03F487A3B078FF9FB8D3FF39B77063D6" httpUri="http://treatment.plazi.org/id/03F487A3B078FF9FB8D3FF39B77063D6" lastPageId="10" lastPageNumber="498" pageId="9" pageNumber="497">
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<subSubSection id="C347653EB078FF9CB8D3FF39B31E6074" pageId="9" pageNumber="497" type="nomenclature">
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<paragraph id="8BE236B5B078FF9CB8D3FF39B17C6018" blockId="9.[151,886,797,858]" box="[151,886,797,822]" pageId="9" pageNumber="497">
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<heading id="D0AA81D9B078FF9CB8D3FF39B17C6018" bold="true" box="[151,886,797,822]" fontSize="10" level="2" pageId="9" pageNumber="497" reason="2">
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||
<emphasis id="B929EAA7B078FF9CB8D3FF39B17C6018" bold="true" box="[151,886,797,822]" pageId="9" pageNumber="497">
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<taxonomicName id="4C5D4D36B078FF9CB8D3FF39B11C6018" ID-CoL="9HKNB" authority="Majnon-Jahromi & Kehlmaier" authorityName="Majnon-Jahromi & Kehlmaier" box="[151,790,797,822]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="497" phylum="Arthropoda" rank="species" species="pistacia" status="sp. nov.">
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||
<emphasis id="B929EAA7B078FF9CB8D3FF39B3976018" bold="true" box="[151,413,797,822]" italics="true" pageId="9" pageNumber="497">Tomosvaryella pistacia</emphasis>
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Majnon-Jahromi & Kehlmaier
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</taxonomicName>
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<taxonomicNameLabel id="A21A57DCB078FF9CBB58FF39B17C6018" box="[796,886,797,822]" pageId="9" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8BE236B5B078FF9CB8D3FF65B31E6074" blockId="9.[151,886,797,858]" box="[151,276,833,858]" pageId="9" pageNumber="497">
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<figureCitation id="13662A30B078FF9CB8D3FF65B31E6074" box="[151,276,833,858]" captionStart="FIGURES 15 – 18" captionStartId="10.[151,264,964,987]" captionTargetBox="[151,1429,303,934]" captionTargetId="figure@10.[151,1435,303,943]" captionTargetPageId="10" captionText="FIGURES 15 – 18. Holotype of Tomosvaryella pistacia Majnon-Jahromi & Kehlmaier sp. nov. (DNA CK 828). 15 — right surstylus and phallus, lateral view; 16 — syntergosternite 8, epandrium and surstyli, dorsal view; 17 — left surstylus, lateral view; 18 — phallus, ventral view. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/803698/files/figure.png" pageId="9" pageNumber="497">Figs 15–18</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C347653EB078FF9CB8D3FFADB6C5608C" box="[151,1231,905,930]" pageId="9" pageNumber="497" type="materials_examined">
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<paragraph id="8BE236B5B078FF9CB8D3FFADB6C5608C" blockId="9.[151,1437,905,2010]" box="[151,1231,905,930]" pageId="9" pageNumber="497">
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<emphasis id="B929EAA7B078FF9CB8D3FFADB34F608C" bold="true" box="[151,325,905,930]" pageId="9" pageNumber="497">Type material.</emphasis>
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HOLOTYPE: 1♂ (DNA CK828,
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<accessionNumber id="940EAB56B078FF9CBA80FFADB136608C" box="[708,828,905,930]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LT671788" pageId="9" pageNumber="497">LT671788</accessionNumber>
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), loc. 6,
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<date id="FFE31075B078FF9CBBDFFFADB66A608C" box="[923,1120,905,930]" pageId="9" pageNumber="497" value="2015-06-29" valueMax="2015-07-11" valueMin="2015-06-29">29.vi–11.vii.2015</date>
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(SMTD).
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</paragraph>
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</subSubSection>
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<subSubSection id="C347653EB078FF9CB883FF89B3076594" pageId="9" pageNumber="497" type="description">
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<paragraph id="8BE236B5B078FF9CB883FF89B3076594" blockId="9.[151,1437,905,2010]" pageId="9" pageNumber="497">
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<emphasis id="B929EAA7B078FF9CB883FF89B3BF60E8" bold="true" box="[199,437,941,966]" pageId="9" pageNumber="497">Description. MALE</emphasis>
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. Body length
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<quantity id="4CA59B50B078FF9CBA1DFF8AB0B960EB" box="[601,691,942,966]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="9" pageNumber="497" unit="mm" value="3.1">3.1 mm</quantity>
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; wing length
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<quantity id="4CA59B50B078FF9CBB14FF89B1A160EB" box="[848,939,941,966]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="497" unit="mm" value="3.5">3.5 mm</quantity>
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; eyes meeting for a distance slightly longer than the length of ocellar triangle, i.e., for approximately six ommatidia; pedicellus with two minute bristles dorsally and one ventrally; flagellum pointed, dark brown; thorax dark brown except for the yellow postpronotal lobe; dorsal bristles of thorax hardly discernible, black; wing membrane with a very faint brownish tinge; wing membrane covered in microtrichia except for the base, including the anal lobe basally; third costal segment 1/3 the length of fourth costal segment; crossvein r-m reaching cell dm very shortly after the middle; crossvein bm-cu interrupted right before reaching cell br on both wings; halter with dark base, whitish stem and light brown knob; leg dark brown except for the narrowly yellow apex of femora, basal 1/3 and apex of tibia and tarsal segments except distitarsus; hind trochanter ventrally smooth, without spines or projection; fore femora with posteroventral row of minute black spines in apical half; mid femora with antero- and posteroventral row of minute black spines for more than apical half; hind femora with anteroventral row of minute black spines in apical half; pulvilli as long as distitarsus; abdomen dark brown; tergite 1 with a lateral patch of about ten short black setae on each side, slightly shorter than the width of tarsal segments; dorsal setae of abdomen black, hardly discernible; sclerites dark brown, apparently evenly sclerotised; genitalia, in dorsal view, with short epandrium, broader than long, with a narrow longitudinal stripe of distinctly weaker sclerotisation centrally and, therefore, apparently divided at first sight; surstyli almost symmetrical, gradually tapering towards apices, which are slightly knobbed; left surstylus slightly protruding the right one; membrane of syntergosternite 8 large, almost reaching epandrium, caudally occupying half of syntergosternite 8; in lateral view, apical half of surstyli bent towards venter by almost 90° (see inner margin); phallus trifid, largely retracted and thus hard to discern even under at a very high magnification; all ejaculatory ducts of different length (although this difference in length might be due to the fact that some ducts were partly torn off), the longest one upright, wavy and with a small membranous nose towards apex.
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<emphasis id="B929EAA7B078FF9CBD59FA59B79F65B8" bold="true" box="[1309,1429,1661,1686]" pageId="9" pageNumber="497">FEMALE</emphasis>
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. Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C347653EB078FF9FB883FAE1B1A663FB" lastPageId="10" lastPageNumber="498" pageId="9" pageNumber="497" type="diagnosis">
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<paragraph id="8BE236B5B078FF9FB883FAE1B1A663FB" blockId="9.[151,1437,905,2010]" lastBlockId="10.[151,1436,151,248]" lastPageId="10" lastPageNumber="498" pageId="9" pageNumber="497">
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<emphasis id="B929EAA7B078FF9CB883FAE1B3C765F0" bold="true" box="[199,461,1733,1758]" pageId="9" pageNumber="497">Differential diagnosis.</emphasis>
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<taxonomicName id="4C5D4D36B078FF9CB997FAE1B0DC65F3" box="[467,726,1733,1758]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="497" phylum="Arthropoda" rank="species" species="pistacia" status="sp. nov.">
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<emphasis id="B929EAA7B078FF9CB997FAE1B0DC65F3" box="[467,726,1733,1758]" italics="true" pageId="9" pageNumber="497">Tomosvaryella pistacia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21A57DCB078FF9CBA98FAE1B13F65F3" box="[732,821,1733,1758]" pageId="9" pageNumber="497" rank="species">
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<emphasis id="B929EAA7B078FF9CBA98FAE1B0FC65F0" bold="true" box="[732,758,1733,1758]" pageId="9" pageNumber="497">sp</emphasis>
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.
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<emphasis id="B929EAA7B078FF9CBB40FAE1B12465F0" bold="true" box="[772,814,1733,1758]" pageId="9" pageNumber="497">nov</emphasis>
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.
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</taxonomicNameLabel>
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stands morphologically similar to
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<taxonomicName id="4C5D4D36B078FF9CBCF3FAE2B73C65F0" box="[1207,1334,1733,1758]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="497" phylum="Arthropoda" rank="species" species="angulata" status="sp. nov.">
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<emphasis id="B929EAA7B078FF9CBCF3FAE2B73C65F0" box="[1207,1334,1733,1758]" italics="true" pageId="9" pageNumber="497">T. angulata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21A57DCB078FF9CBD78FAE1B79E65F3" box="[1340,1428,1733,1758]" pageId="9" pageNumber="497" rank="species">
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<emphasis id="B929EAA7B078FF9CBD78FAE1B75C65F0" bold="true" box="[1340,1366,1733,1758]" pageId="9" pageNumber="497">sp</emphasis>
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.
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<emphasis id="B929EAA7B078FF9CBD20FAE1B78465F0" bold="true" box="[1380,1422,1733,1758]" pageId="9" pageNumber="497">nov</emphasis>
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.
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</taxonomicNameLabel>
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, but the male of the first one does not have such a strongly-curved and distally-knobbed surstyli. Although the membranous area of the syntergosternite 8 is clearly larger in
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<taxonomicName id="4C5D4D36B078FF9CBB0EFB2AB1B4640B" box="[842,958,1806,1829]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="497" phylum="Arthropoda" rank="species" species="pistacia" status="sp. nov.">
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<emphasis id="B929EAA7B078FF9CBB0EFB2AB1B4640B" box="[842,958,1806,1829]" italics="true" pageId="9" pageNumber="497">T. pistacia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21A57DCB078FF9CBB81FB29B6146408" box="[965,1054,1805,1830]" pageId="9" pageNumber="497" rank="species">
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<emphasis id="B929EAA7B078FF9CBB81FB29B6146408" bold="true" box="[965,1054,1805,1830]" pageId="9" pageNumber="497">sp. nov.</emphasis>
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</taxonomicNameLabel>
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, to separate the syntergosternite 8 from the abdomen is crucial in order to differentiate the two species with certainty. Both species share a small epandrium that seems to be less sclerotised dorsally, at its narrowest point (before the anal opening); this feature is apparently more developed in
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<taxonomicName id="4C5D4D36B078FF9CB9BDFB5EB07B64BF" box="[505,625,1914,1937]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="497" phylum="Arthropoda" rank="species" species="pistacia" status="sp. nov.">
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<emphasis id="B929EAA7B078FF9CB9BDFB5EB07B64BF" box="[505,625,1914,1937]" italics="true" pageId="9" pageNumber="497">T. pistacia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21A57DCB078FF9CBA3FFB5DB0D264BC" box="[635,728,1913,1938]" pageId="9" pageNumber="497" rank="species">
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<emphasis id="B929EAA7B078FF9CBA3FFB5DB0D264BC" bold="true" box="[635,728,1913,1938]" pageId="9" pageNumber="497">sp. nov.</emphasis>
|
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</taxonomicNameLabel>
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, judging from the
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<typeStatus id="54E68817B078FF9CBBF3FB5DB61764BC" box="[951,1053,1913,1938]" pageId="9" pageNumber="497" type="holotype">holotype</typeStatus>
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. The small-sized epandrium puts these two species in a close relation to the Afrotropical
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<taxonomicName id="4C5D4D36B078FF9CBB47FBBAB1D16498" authority="Hardy" authorityName="Hardy" box="[771,987,1949,1974]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="497" phylum="Arthropoda" rank="species" species="congoana">
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<emphasis id="B929EAA7B078FF9CBB47FBBAB187649B" box="[771,909,1950,1973]" italics="true" pageId="9" pageNumber="497">T. congoana</emphasis>
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Hardy
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</taxonomicName>
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, known from the
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<collectingCountry id="F34A7625B078FF9CBCDBFBBAB70E6498" box="[1183,1284,1950,1974]" name="Comoros" pageId="9" pageNumber="497">Comores</collectingCountry>
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Archipelago,
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<collectingCountry id="F34A7625B078FF9CB8D3FBE5B32664F4" box="[151,300,1985,2010]" name="South Africa" pageId="9" pageNumber="497">South Africa</collectingCountry>
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,
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<collectingCountry id="F34A7625B078FF9CB97FFBE6B37C64F7" box="[315,374,1986,2009]" name="Democratic Republic of the Congo" pageId="9" pageNumber="497">Zaïre</collectingCountry>
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and
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<collectingCountry id="F34A7625B078FF9CB9F6FBE5B00D64F4" box="[434,519,1985,2010]" name="Zambia" pageId="9" pageNumber="497">Zambia</collectingCountry>
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(redescribed and illustrated by De Meyer 1993).
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<taxonomicName id="4C5D4D36B078FF9CBC0CFBE5B76E64F7" box="[1096,1380,1985,2010]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="497" phylum="Arthropoda" rank="species" species="congoana">
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<emphasis id="B929EAA7B078FF9CBC0CFBE5B76E64F7" box="[1096,1380,1985,2010]" italics="true" pageId="9" pageNumber="497">Tomosvaryella congoana</emphasis>
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</taxonomicName>
|
||
also shares a similarly-shaped phallus and female ovipositor; the female ovipositor, however, has two protuberances at its ventral base, instead of one protuberance, as in
|
||
<taxonomicName id="4C5D4D36B07BFF9FBA88FC99B14163FB" box="[716,843,188,213]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="498" phylum="Arthropoda" rank="species" species="angulata" status="sp. nov.">
|
||
<emphasis id="B929EAA7B07BFF9FBA88FC99B14163FB" box="[716,843,188,213]" italics="true" pageId="10" pageNumber="498">T. angulata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57DCB07BFF9FBB16FC98B1A663FB" box="[850,940,188,213]" pageId="10" pageNumber="498" rank="species">
|
||
<emphasis id="B929EAA7B07BFF9FBB16FC98B1A663FB" bold="true" box="[850,940,188,213]" pageId="10" pageNumber="498">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347653EB07BFF9FB883FCFBB77063D6" box="[199,1402,223,248]" pageId="10" pageNumber="498" type="etymology">
|
||
<paragraph id="8BE236B5B07BFF9FB883FCFBB77063D6" blockId="10.[151,1436,151,248]" box="[199,1402,223,248]" pageId="10" pageNumber="498">
|
||
<emphasis id="B929EAA7B07BFF9FB883FCFBB34463D6" bold="true" box="[199,334,223,248]" pageId="10" pageNumber="498">Etymology.</emphasis>
|
||
The species is named
|
||
<taxonomicName id="4C5D4D36B07BFF9FBA0EFCC5B0A963D6" box="[586,675,225,248]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="498" phylum="Arthropoda" rank="species" species="pistacia">
|
||
<emphasis id="B929EAA7B07BFF9FBA0EFCC5B0A963D6" box="[586,675,225,248]" italics="true" pageId="10" pageNumber="498">pistacia</emphasis>
|
||
</taxonomicName>
|
||
(Latin for pistachio) as it was collected in a pistachio plantation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |