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<document id="CF24E5A91254B250B4EACDA24BBE58F0" ID-DOI="10.1080/00222930802354803" ID-ISSN="1464-5262" ID-Zenodo-Dep="4590517" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615293098108" checkinUser="carolina" docAuthor="Park, Sun-Jae &amp; Ahn, Kee-Jeong" docDate="2008" docId="03F4879AFFA745121BE2F9FCFF6F13D4" docLanguage="en" docName="JNATHIST.42.39-40.2501-2514.pdf" docOrigin="Journal of Natural History 42 (39 - 40)" docSource="http://dx.doi.org/10.1080/00222930802354803" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Neochthebius granulosus" docType="treatment" docVersion="3" lastPageNumber="2510" masterDocId="FFCDFFE2FFA045191A22FFD7FF8B157E" masterDocTitle="Redescription of the littoral genus and species of Neochthebius granulosus (Satô), and the first larval description for the genus (Coleoptera: Hydraenidae: Ochthebiinae)" masterLastPageNumber="2514" masterPageNumber="2501" pageNumber="2506" updateTime="1698881818549" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F773CD24E787FEEACC25D3028AEE77BA">Redescription of the littoral genus and species of Neochthebius granulosus (Satô), and the first larval description for the genus (Coleoptera: Hydraenidae: Ochthebiinae)</mods:title>
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<mods:namePart id="0723A51E48413EF85AF0E073DBE51FB5">Park, Sun-Jae</mods:namePart>
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<mods:namePart id="4922F9DC38A31E8FC9B18FD7C13BC509">Ahn, Kee-Jeong</mods:namePart>
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<mods:title id="A211C635312D58DA716175A8AB18B94F">Journal of Natural History</mods:title>
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<mods:date id="DC274CEB9111BCEF3DEECC93479B746E">2008</mods:date>
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<mods:number id="E13D1D5F0866D7A21241FBC0E7231929">2008-10-31</mods:number>
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<mods:number id="E65C072385528C6EDE8A8A305C7B5C5A">42</mods:number>
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<mods:number id="B6644DEFA6B923211C4D727E77FA2738">39 - 40</mods:number>
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<mods:identifier id="419B8F889DDBAF4C37DD69BA9B3B5F19" type="ISSN">1464-5262</mods:identifier>
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<subSubSection id="C3476507FFA7451E1BE2F9FCFCAA133D" box="[448,801,1579,1603]" pageId="7" pageNumber="2506" type="nomenclature">
<paragraph id="8BE2368CFFA7451E1BE2F9FCFCAA133D" blockId="7.[448,801,1579,1603]" box="[448,801,1579,1603]" pageId="7" pageNumber="2506">
<heading id="D0AA81E0FFA7451E1BE2F9FCFCAA133D" box="[448,801,1579,1603]" centered="true" fontSize="36" level="2" pageId="7" pageNumber="2506" reason="7">
<taxonomicName id="4C5D4D0FFFA7451E1BE2F9FCFCAA133D" authority="(Sato)" baseAuthorityName="Sato" baseAuthorityYear="1963" box="[448,801,1579,1603]" class="Insecta" family="Hydraenidae" genus="Neochthebius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2506" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis id="B929EA9EFFA7451E1BE2F9FCFD58133D" bold="true" box="[448,723,1579,1603]" italics="true" pageId="7" pageNumber="2506">Neochthebius granulosus</emphasis>
(Satô)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3476507FFA7451E1AD5F986FD4713F7" pageId="7" pageNumber="2506" type="description">
<paragraph id="8BE2368CFFA7451E1AD5F986FD4713F7" blockId="7.[149,1003,1617,1706]" pageId="7" pageNumber="2506">
(
<figureCitation id="13662A09FFA7451E1ADDF986FEEC1317" box="[255,359,1617,1641]" captionStart="Figures 12" captionStartId="3.[146,222,837,859]" captionTargetBox="[146,1103,165,800]" captionTargetId="figure-164@3.[146,1103,165,800]" captionTargetPageId="3" captionText="Figures 12. SEM habitus photos. (1) Neochthebius granulosus; (2) N. vandykei." figureDoi="http://doi.org/10.5281/zenodo.4590519" httpUri="https://zenodo.org/record/4590519/files/figure.png" pageId="7" pageNumber="2506">Figures 1</figureCitation>
,
<figureCitation id="13662A09FFA7451E1B55F986FE2A1317" box="[375,417,1617,1641]" captionStart="Figures 38" captionStartId="4.[213,289,1532,1554]" captionTargetBox="[292,1092,165,1495]" captionTargetId="figure-0@4.[292,1092,165,1495]" captionTargetPageId="4" captionText="Figures 38. Neochthebius granulosus. (3) antenna, dorsal aspect; (4) labrum, dorsal aspect; (5) mandible, dorsal aspect; (6) maxilla, ventral aspect; (7) aedeagus, dorsal aspect; (8) aedeagus, lateral aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590523" httpUri="https://zenodo.org/record/4590523/files/figure.png" pageId="7" pageNumber="2506">38</figureCitation>
, 910, 1213, 15, 1718, 1920, 22, 24, and 2636)
<taxonomicName id="4C5D4D0FFFA7451E1AB7F9A5FD4C13F7" authority="Sato, 1963: 129" authorityName="Sato" authorityPageNumber="129" authorityYear="1963" box="[149,711,1649,1674]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2506" phylum="Arthropoda" rank="species" species="granulosus" subGenus="Cobalius">
<emphasis id="B929EA9EFFA7451E1AB7F9A5FE9B13F4" box="[149,272,1650,1674]" italics="true" pageId="7" pageNumber="2506">Ochthebius</emphasis>
(
<emphasis id="B929EA9EFFA7451E1B00F9A5FE0813F4" box="[290,387,1650,1674]" italics="true" pageId="7" pageNumber="2506">Cobalius</emphasis>
)
<emphasis id="B929EA9EFFA7451E1BB7F9A5FD8013F4" box="[405,523,1650,1674]" italics="true" pageId="7" pageNumber="2506">granulosus</emphasis>
<bibRefCitation id="EFCC4B7DFFA7451E1831F9A6FD4C13F7" author="Sato M" box="[531,711,1649,1673]" pageId="7" pageNumber="2506" pagination="129 - 132" refId="ref3928" refString="Sato M. 1963. A new inter-tidal species of the genus Ochthebius Leach from Japan (Coleoptera: Hydeaenidae). Trans Shikoku Entomol Soc. 7 (4): 129 - 132." type="journal article" year="1963">Satô, 1963: 129</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3476507FFA7451E1AB7F945FCD913D4" box="[149,850,1682,1706]" pageId="7" pageNumber="2506" type="reference_group">
<paragraph id="8BE2368CFFA7451E1AB7F945FCD913D4" blockId="7.[149,1003,1617,1706]" box="[149,850,1682,1706]" pageId="7" pageNumber="2506">
<treatmentCitationGroup id="AB4D11A2FFA7451E1AB7F945FCD913D4" box="[149,850,1682,1706]" pageId="7" pageNumber="2506">
<treatmentCitation id="0AFC109DFFA7451E1AB7F945FD0613D4" author="Perkins PD" box="[149,653,1682,1706]" page="409" pageId="7" pageNumber="2506" year="1980">
<taxonomicName id="4C5D4D0FFFA7451E1AB7F945FD0613D4" authority=": Perkins, 1980: 409" authorityName=": Perkins" authorityPageNumber="409" authorityYear="1980" box="[149,653,1682,1706]" class="Insecta" family="Hydraenidae" genus="Neochthebius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2506" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis id="B929EA9EFFA7451E1AB7F945FE2313D4" box="[149,424,1682,1706]" italics="true" pageId="7" pageNumber="2506">Neochthebius granulosus</emphasis>
:
<bibRefCitation id="EFCC4B7DFFA7451E1B9AF945FD0613D4" author="Perkins PD" box="[440,653,1682,1706]" pageId="7" pageNumber="2506" pagination="3 - 554" refId="ref3841" refString="Perkins PD. 1980. Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaest Entomol. 16: 3 - 554." type="journal article" year="1980">Perkins, 1980: 409</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AFC109DFFA7451E18B8F945FCC513D4" author="Jach MA" box="[666,846,1682,1706]" page="113" pageId="7" pageNumber="2506" year="2004">
<bibRefCitation id="EFCC4B7DFFA7451E18B8F945FCC513D4" author="Jach MA" box="[666,846,1682,1706]" pageId="7" pageNumber="2506" pagination="102 - 122" refId="ref3799" refString="Jach MA. 2004. Hydraenidae. In: Lobl I, Smetana A, editors. Catalogue of palaearctic Coleoptera. Vol. 2. Hydrophiloidea-Histeroidea-Staphylinoidea. Stenstrup (Denmark): Apollo Books. p. 102 - 122." type="book chapter" year="2004">Jäch, 2004: 113</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3476507FFA845121AF7FF7EFF6F13D4" lastPageId="11" lastPageNumber="2510" pageId="8" pageNumber="2507" type="description">
<paragraph id="8BE2368CFFA845111AF7FF7EFEDF15BF" blockId="8.[213,340,169,193]" box="[213,340,169,193]" pageId="8" pageNumber="2507">
<emphasis id="B929EA9EFFA845111AF7FF7EFEDF15BF" box="[213,340,169,193]" italics="true" pageId="8" pageNumber="2507">Description</emphasis>
</paragraph>
<paragraph id="8BE2368CFFA845111AF7FF06FEF11492" blockId="8.[213,1169,209,492]" pageId="8" pageNumber="2507">
<emphasis id="B929EA9EFFA845111AF7FF06FE961597" box="[213,285,209,233]" italics="true" pageId="8" pageNumber="2507">Adults</emphasis>
. Length
<quantity id="4CA59B69FFA845111BA4FF06FD841597" box="[390,527,209,233]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="8" pageNumber="2507" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.41.6 mm</quantity>
. Body (
<figureCitation id="13662A09FFA84511184AFF06FD431597" box="[616,712,209,233]" captionStart="Figures 12" captionStartId="3.[146,222,837,859]" captionTargetBox="[146,1103,165,800]" captionTargetId="figure-164@3.[146,1103,165,800]" captionTargetPageId="3" captionText="Figures 12. SEM habitus photos. (1) Neochthebius granulosus; (2) N. vandykei." figureDoi="http://doi.org/10.5281/zenodo.4590519" httpUri="https://zenodo.org/record/4590519/files/figure.png" pageId="8" pageNumber="2507">Figure 1</figureCitation>
) almost unicolour, dark brown to dull black, antenna slightly lighter; about 2.8 times longer than wide; dorsum covered with some distinct white setae. Pronotum (
<figureCitation id="13662A09FFA8451118F4FEC5FCCA1454" box="[726,833,274,298]" captionStart="Figures 12" captionStartId="3.[146,222,837,859]" captionTargetBox="[146,1103,165,800]" captionTargetId="figure-164@3.[146,1103,165,800]" captionTargetPageId="3" captionText="Figures 12. SEM habitus photos. (1) Neochthebius granulosus; (2) N. vandykei." figureDoi="http://doi.org/10.5281/zenodo.4590519" httpUri="https://zenodo.org/record/4590519/files/figure.png" pageId="8" pageNumber="2507">Figures 1</figureCitation>
and
<figureCitation id="13662A09FFA8451119A3FEC5FC4C1454" box="[897,967,274,298]" captionStart="Figures 1921" captionStartId="7.[146,222,1493,1515]" captionTargetBox="[146,1104,1034,1457]" captionTargetId="figure-61@7.[146,1104,1034,1457]" captionTargetPageId="7" captionText="Figures 1921. Pronotum and elytra. (1920) Neochthebius granulosus; (21) N. vandykei." figureDoi="http://doi.org/10.5281/zenodo.4590527" httpUri="https://zenodo.org/record/4590527/files/figure.png" pageId="8" pageNumber="2507">1920</figureCitation>
) about 1.2 times wider than long; widest in front of middle; moderately narrowed apically, strongly narrowed basally. Elytra (
<figureCitation id="13662A09FFA8451118A0FE84FCAD1415" box="[642,806,339,363]" captionStart="Figures 1921" captionStartId="7.[146,222,1493,1515]" captionTargetBox="[146,1104,1034,1457]" captionTargetId="figure-61@7.[146,1104,1034,1457]" captionTargetPageId="7" captionText="Figures 1921. Pronotum and elytra. (1920) Neochthebius granulosus; (21) N. vandykei." figureDoi="http://doi.org/10.5281/zenodo.4590527" httpUri="https://zenodo.org/record/4590527/files/figure.png" pageId="8" pageNumber="2507">Figures 1920</figureCitation>
) about 1.5 times longer than wide. Abdomen (
<figureCitation id="13662A09FFA845111BBCFEA4FD8414F5" box="[414,527,371,395]" captionStart="Figures 2225" captionStartId="9.[146,222,1476,1498]" captionTargetBox="[146,1104,165,1440]" captionTargetId="figure-53@9.[146,1104,165,1440]" captionTargetPageId="9" captionText="Figures 2225. (2223) pro- and mesoventrite; (2425) metaventrite and abdomen. (22 and 24) Neochthebius granulosus; (23 and 25) N. vandykei." figureDoi="http://doi.org/10.5281/zenodo.4590529" httpUri="https://zenodo.org/record/4590529/files/figure.png" pageId="8" pageNumber="2507">Figure 24</figureCitation>
) strongly imbricate; last abdominal segment of female with a row of 712 stout spines apically (
<figureCitation id="13662A09FFA845111925FE44FC2514D5" box="[775,942,403,427]" captionStart="Figures 1518" captionStartId="7.[146,222,922,944]" captionTargetBox="[146,1104,165,885]" captionTargetId="figure-39@7.[146,1104,165,885]" captionTargetPageId="7" captionText="Figures 1518. (1516) head; (1718) female last abdominal segment, ventral aspect. (15 and 1718) Neochthebius granulosus; (16) N. vandykei." figureDoi="http://doi.org/10.5281/zenodo.4590525" httpUri="https://zenodo.org/record/4590525/files/figure.png" pageId="8" pageNumber="2507">Figures 1718</figureCitation>
). Aedeagus as in
<figureCitation id="13662A09FFA845111AF7FE63FED514B2" box="[213,350,436,460]" captionStart="Figures 38" captionStartId="4.[213,289,1532,1554]" captionTargetBox="[292,1092,165,1495]" captionTargetId="figure-0@4.[292,1092,165,1495]" captionTargetPageId="4" captionText="Figures 38. Neochthebius granulosus. (3) antenna, dorsal aspect; (4) labrum, dorsal aspect; (5) mandible, dorsal aspect; (6) maxilla, ventral aspect; (7) aedeagus, dorsal aspect; (8) aedeagus, lateral aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590523" httpUri="https://zenodo.org/record/4590523/files/figure.png" pageId="8" pageNumber="2507">Figures 78</figureCitation>
. Parameres as in
<figureCitation id="13662A09FFA845111813FE63FD3114B2" box="[561,698,436,460]" captionStart="Figures 38" captionStartId="4.[213,289,1532,1554]" captionTargetBox="[292,1092,165,1495]" captionTargetId="figure-0@4.[292,1092,165,1495]" captionTargetPageId="4" captionText="Figures 38. Neochthebius granulosus. (3) antenna, dorsal aspect; (4) labrum, dorsal aspect; (5) mandible, dorsal aspect; (6) maxilla, ventral aspect; (7) aedeagus, dorsal aspect; (8) aedeagus, lateral aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590523" httpUri="https://zenodo.org/record/4590523/files/figure.png" pageId="8" pageNumber="2507">Figures 78</figureCitation>
, apex with very short setae ventrally in lateral aspect.
</paragraph>
<paragraph id="8BE2368CFFA845121AF7FDF2FC9714DD" blockId="8.[213,1170,549,1706]" lastBlockId="11.[146,1103,168,419]" lastPageId="11" lastPageNumber="2510" pageId="8" pageNumber="2507">
<emphasis id="B929EA9EFFA845111AF7FDF2FEA81743" box="[213,291,549,573]" italics="true" pageId="8" pageNumber="2507">Larvae</emphasis>
(
<emphasis id="B929EA9EFFA845111B19FDF2FE3A1743" box="[315,433,549,573]" italics="true" pageId="8" pageNumber="2507">last instar</emphasis>
). Length
<quantity id="4CA59B69FFA845111817FDF2FD361743" box="[565,701,549,573]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="8" pageNumber="2507" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.52.7 mm</quantity>
. Body (
<figureCitation id="13662A09FFA84511190AFDF2FC1C1743" box="[808,919,549,573]" captionStart="Figure 26" captionStartId="10.[213,280,1607,1629]" captionTargetBox="[465,918,165,1571]" captionTargetId="figure-0@10.[465,918,165,1571]" captionTargetPageId="10" captionText="Figure 26. Larva habitus of Neochthebius granulosus. Note: Scale50.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4590531" httpUri="https://zenodo.org/record/4590531/files/figure.png" pageId="8" pageNumber="2507">Figure 26</figureCitation>
) pale, dark brown, antennae and urogomphi darker; elongate, parallel-sided. Head (
<figureCitation id="13662A09FFA8451119FCFD92FBC61723" box="[990,1101,581,605]" captionStart="Figures 2731" captionStartId="12.[213,289,1471,1493]" captionTargetBox="[292,1092,165,1431]" captionTargetId="figure-29@12.[292,1092,165,1434]" captionTargetPageId="12" captionText="Figures 2731. Larva of Neochthebius granulosus. (27) head, dorsal aspect; (28) labrum, dorsal aspect; (29) mandible, dorsal aspect; (30) maxilla, ventral aspect; (31) antenna, ventral aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590533" httpUri="https://zenodo.org/record/4590533/files/figure.png" pageId="8" pageNumber="2507">Figure 27</figureCitation>
) subquadrate, lateral margins broadly rounded; widest at middle, 1.5 times wider than long; chaetotaxy as in
<figureCitation id="13662A09FFA845111829FD51FDF017E0" box="[523,635,646,670]" captionStart="Figures 2731" captionStartId="12.[213,289,1471,1493]" captionTargetBox="[292,1092,165,1431]" captionTargetId="figure-29@12.[292,1092,165,1434]" captionTargetPageId="12" captionText="Figures 2731. Larva of Neochthebius granulosus. (27) head, dorsal aspect; (28) labrum, dorsal aspect; (29) mandible, dorsal aspect; (30) maxilla, ventral aspect; (31) antenna, ventral aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590533" httpUri="https://zenodo.org/record/4590533/files/figure.png" pageId="8" pageNumber="2507">Figure 27</figureCitation>
; frontoclypeal suture indistinctly present; epicranial stem relatively short; V-shaped frontal arms long, reached nearly to antennae; neck absent. Clypeus (
<figureCitation id="13662A09FFA845111856FD10FD6817A1" box="[628,739,711,735]" captionStart="Figures 2731" captionStartId="12.[213,289,1471,1493]" captionTargetBox="[292,1092,165,1431]" captionTargetId="figure-29@12.[292,1092,165,1434]" captionTargetPageId="12" captionText="Figures 2731. Larva of Neochthebius granulosus. (27) head, dorsal aspect; (28) labrum, dorsal aspect; (29) mandible, dorsal aspect; (30) maxilla, ventral aspect; (31) antenna, ventral aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590533" httpUri="https://zenodo.org/record/4590533/files/figure.png" pageId="8" pageNumber="2507">Figure 27</figureCitation>
) truncate, transverse, clypeolabral suture straight; three distinct setae (Cl1Cl3) present on each side. Five stemmata present (
<figureCitation id="13662A09FFA845111B1BFCDFFE20165E" box="[313,427,776,800]" captionStart="Figures 2731" captionStartId="12.[213,289,1471,1493]" captionTargetBox="[292,1092,165,1431]" captionTargetId="figure-29@12.[292,1092,165,1434]" captionTargetPageId="12" captionText="Figures 2731. Larva of Neochthebius granulosus. (27) head, dorsal aspect; (28) labrum, dorsal aspect; (29) mandible, dorsal aspect; (30) maxilla, ventral aspect; (31) antenna, ventral aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590533" httpUri="https://zenodo.org/record/4590533/files/figure.png" pageId="8" pageNumber="2507">Figure 27</figureCitation>
). Antenna (
<figureCitation id="13662A09FFA845111816FCDFFD29165E" box="[564,674,776,800]" captionStart="Figures 2731" captionStartId="12.[213,289,1471,1493]" captionTargetBox="[292,1092,165,1431]" captionTargetId="figure-29@12.[292,1092,165,1434]" captionTargetPageId="12" captionText="Figures 2731. Larva of Neochthebius granulosus. (27) head, dorsal aspect; (28) labrum, dorsal aspect; (29) mandible, dorsal aspect; (30) maxilla, ventral aspect; (31) antenna, ventral aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590533" httpUri="https://zenodo.org/record/4590533/files/figure.png" pageId="8" pageNumber="2507">Figure 31</figureCitation>
) with three antennomeres; antennomere 1 widest; 2 sub-truncate, longest, 2.0 times longer than wide, 2.4 times longer than 3, four long setae present on apex; sensorium IIS1 present on apex of 2, as long as antennomere 3, IIS 2 minute; antennomere 3 short with four long and two short setae, 1.6 times longer than wide. Labrum (
<figureCitation id="13662A09FFA8451118FDFC5EFCC516DF" box="[735,846,905,929]" captionStart="Figures 2731" captionStartId="12.[213,289,1471,1493]" captionTargetBox="[292,1092,165,1431]" captionTargetId="figure-29@12.[292,1092,165,1434]" captionTargetPageId="12" captionText="Figures 2731. Larva of Neochthebius granulosus. (27) head, dorsal aspect; (28) labrum, dorsal aspect; (29) mandible, dorsal aspect; (30) maxilla, ventral aspect; (31) antenna, ventral aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590533" httpUri="https://zenodo.org/record/4590533/files/figure.png" pageId="8" pageNumber="2507">Figure 28</figureCitation>
) more or less semicircular; five labral marginal setae (Lm1Lm5) present on each side, Lm2 furcated twice; two labral dorsal setae (Ld1Ld2), one campaniform sensilla (C2). Mandibles (
<figureCitation id="13662A09FFA845111AFEFC3DFEC7117C" box="[220,332,1002,1026]" captionStart="Figures 2731" captionStartId="12.[213,289,1471,1493]" captionTargetBox="[292,1092,165,1431]" captionTargetId="figure-29@12.[292,1092,165,1434]" captionTargetPageId="12" captionText="Figures 2731. Larva of Neochthebius granulosus. (27) head, dorsal aspect; (28) labrum, dorsal aspect; (29) mandible, dorsal aspect; (30) maxilla, ventral aspect; (31) antenna, ventral aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590533" httpUri="https://zenodo.org/record/4590533/files/figure.png" pageId="8" pageNumber="2507">Figure 29</figureCitation>
) more or less symmetrical, triangular; base broad, apical half thin, moderately incurved; apex blunt; incisor edge serrate; prostheca fixed, hyaline, with serrate process at apex; mola well developed, tuberculate; with a distinct tooth above mola; two mandibula setae (M1M2) and two campaniform sensillae (C1C2) present. Maxilla as in
<figureCitation id="13662A09FFA845111867FBBBFD3F11FA" box="[581,692,1132,1156]" captionStart="Figures 2731" captionStartId="12.[213,289,1471,1493]" captionTargetBox="[292,1092,165,1431]" captionTargetId="figure-29@12.[292,1092,165,1434]" captionTargetPageId="12" captionText="Figures 2731. Larva of Neochthebius granulosus. (27) head, dorsal aspect; (28) labrum, dorsal aspect; (29) mandible, dorsal aspect; (30) maxilla, ventral aspect; (31) antenna, ventral aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590533" httpUri="https://zenodo.org/record/4590533/files/figure.png" pageId="8" pageNumber="2507">Figure 30</figureCitation>
; palpifer with crescent sclerite at base of maxillary palpus with one seta (Pf1); cardo triangular with one seta (Cdo1); stipes with three lateral setae (Stp1Stp3) and one campaniform sensilla (C1); five lacinial setae (La1, La3La6) present; maxillary palpi with three palpomeres, tapered apically; palpomere 1 about 0.8 times longer than wide, two campaniform sensillae (C1C2) present; palpomere 2 sub-quadrate, 1.4 times longer than wide, two long setae (Pm1Pm2) and one campaniform sensilla (C1) present; 3 slender, longest, 4.2 times longer than wide, apical third abruptly narrowed, sensorial digitiform appendage (SD) present. Labium indistinct; labial palpi with two palpomeres, small, indistinct; ligula pointed apically. Pronotum (
<figureCitation id="13662A09FFA8451119FEFA58FBC010D9" box="[988,1099,1423,1447]" captionStart="Figures 3234" captionStartId="13.[146,222,1536,1558]" captionTargetBox="[225,1025,165,1499]" captionTargetId="figure-0@13.[225,1025,165,1499]" captionTargetPageId="13" captionText="Figures 3234. Larva of Neochthebius granulosus. (32) pronotum, dorsal aspect; (33) mesonotum, dorsal aspect; (34) metanotum, dorsal aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590535" httpUri="https://zenodo.org/record/4590535/files/figure.png" pageId="8" pageNumber="2507">Figure 32</figureCitation>
) with margins rounded; 1.7 times wider than long; widest at middle; four anterior (A1 A4), three lateral (L1L3), four posterior (P1P4), eight additional setae present on each side as in
<figureCitation id="13662A09FFA845111B8CFA27FD951376" box="[430,542,1520,1544]" captionStart="Figures 3234" captionStartId="13.[146,222,1536,1558]" captionTargetBox="[225,1025,165,1499]" captionTargetId="figure-0@13.[225,1025,165,1499]" captionTargetPageId="13" captionText="Figures 3234. Larva of Neochthebius granulosus. (32) pronotum, dorsal aspect; (33) mesonotum, dorsal aspect; (34) metanotum, dorsal aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590535" httpUri="https://zenodo.org/record/4590535/files/figure.png" pageId="8" pageNumber="2507">Figure 32</figureCitation>
; five campaniform sensillae (C1C5) present on each side. Mesonotum (
<figureCitation id="13662A09FFA845111B99F9C6FDA01357" box="[443,555,1553,1577]" captionStart="Figures 3234" captionStartId="13.[146,222,1536,1558]" captionTargetBox="[225,1025,165,1499]" captionTargetId="figure-0@13.[225,1025,165,1499]" captionTargetPageId="13" captionText="Figures 3234. Larva of Neochthebius granulosus. (32) pronotum, dorsal aspect; (33) mesonotum, dorsal aspect; (34) metanotum, dorsal aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590535" httpUri="https://zenodo.org/record/4590535/files/figure.png" pageId="8" pageNumber="2507">Figure 33</figureCitation>
) rounded laterally, sharply narrowed posteriorly, moderately narrowed anteriorly; 2.0 times wider than long; widest near apex; four minute anterior setae (A1A4), three lateral (L1L3), four posterior (P1P4), five additional setae present on each side as in
<figureCitation id="13662A09FFA8451118F5F9A5FCCC13F4" box="[727,839,1650,1674]" captionStart="Figures 3234" captionStartId="13.[146,222,1536,1558]" captionTargetBox="[225,1025,165,1499]" captionTargetId="figure-0@13.[225,1025,165,1499]" captionTargetPageId="13" captionText="Figures 3234. Larva of Neochthebius granulosus. (32) pronotum, dorsal aspect; (33) mesonotum, dorsal aspect; (34) metanotum, dorsal aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590535" httpUri="https://zenodo.org/record/4590535/files/figure.png" pageId="8" pageNumber="2507">Figure 33</figureCitation>
; five campaniform sensillae present on each side. Metanotum (
<figureCitation id="13662A09FFA845111882F945FC8413D4" box="[672,783,1682,1706]" captionStart="Figures 3234" captionStartId="13.[146,222,1536,1558]" captionTargetBox="[225,1025,165,1499]" captionTargetId="figure-0@13.[225,1025,165,1499]" captionTargetPageId="13" captionText="Figures 3234. Larva of Neochthebius granulosus. (32) pronotum, dorsal aspect; (33) mesonotum, dorsal aspect; (34) metanotum, dorsal aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590535" httpUri="https://zenodo.org/record/4590535/files/figure.png" pageId="8" pageNumber="2507">Figure 34</figureCitation>
) similar to mesonotum. Legs (
<figureCitation id="13662A09FFA945101AB8F9E7FE821336" box="[154,265,1584,1608]" captionStart="Figures 3536" captionStartId="14.[213,289,792,814]" captionTargetBox="[213,1171,165,753]" captionTargetId="figure-251@14.[213,1171,165,756]" captionTargetPageId="14" captionText="Figures 3536. Larva of Neochthebius granulosus. (35) mesoleg, dorsal aspect; (36) urogomphus, dorsal aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590537" httpUri="https://zenodo.org/record/4590537/files/figure.png" pageId="9" pageNumber="2508">Figure 35</figureCitation>
) long; all very similar in shape and distribution of setal patterns; trochanter large, triangular; femur transverse; tibia slender, narrowed apically, one stout seta present at inner middle dorsally, a small tooth present at inner margin of basal fourth with one pretarsal seta; tarsungulus with apex sharply pointed ventrally. Abdomen (
<figureCitation id="13662A09FFAB45121BD2FF7FFDEB15BE" box="[496,608,168,192]" captionStart="Figure 26" captionStartId="10.[213,280,1607,1629]" captionTargetBox="[465,918,165,1571]" captionTargetId="figure-0@10.[465,918,165,1571]" captionTargetPageId="10" captionText="Figure 26. Larva habitus of Neochthebius granulosus. Note: Scale50.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4590531" httpUri="https://zenodo.org/record/4590531/files/figure.png" pageId="11" pageNumber="2510">Figure 26</figureCitation>
) with 10 segments; parallel-sided in basal half, slightly narrowed posteriorly; tergites IVI expanded posteriorly and suddenly narrowed; VII parallel; VIIIIX slightly narrowed posteriorly; X strongly narrowed posteriorly, apical margin straight; two small anal hooks recurved ventrally. Urogomphi (
<figureCitation id="13662A09FFAB45121831FEFDFD09143C" box="[531,642,298,322]" captionStart="Figures 3536" captionStartId="14.[213,289,792,814]" captionTargetBox="[213,1171,165,753]" captionTargetId="figure-251@14.[213,1171,165,756]" captionTargetPageId="14" captionText="Figures 3536. Larva of Neochthebius granulosus. (35) mesoleg, dorsal aspect; (36) urogomphus, dorsal aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590537" httpUri="https://zenodo.org/record/4590537/files/figure.png" pageId="11" pageNumber="2510">Figure 36</figureCitation>
) triangular, two articled (URIURII); apical fifth strongly narrowed and parallel; apex with a long seta (
<collectingCountry id="F34A761CFFAB4512198FFE9DFC5E141C" box="[941,981,330,354]" name="American Samoa" pageId="11" pageNumber="2510">AS</collectingCountry>
), six setae (U1U6) present. Abdominal sternites well sclerotized, form single plates. Respiratory system peripneustic and spiracles annular.
</paragraph>
<caption id="DF226604FFA945101AB0FA13FD1F1089" ID-DOI="http://doi.org/10.5281/zenodo.4590529" ID-Zenodo-Dep="4590529" httpUri="https://zenodo.org/record/4590529/files/figure.png" pageId="9" pageNumber="2508" startId="9.[146,222,1476,1498]" targetBox="[146,1104,165,1440]" targetPageId="9">
<paragraph id="8BE2368CFFA945101AB0FA13FD1F1089" blockId="9.[146,1103,1476,1527]" pageId="9" pageNumber="2508">
Figures 2225. (2223) pro- and mesoventrite; (2425) metaventrite and abdomen. (22 and 24)
<taxonomicName id="4C5D4D0FFFA945101AB0FA35FE021089" authorityName=": Perkins" authorityYear="1980" box="[146,393,1506,1527]" class="Insecta" family="Hydraenidae" genus="Neochthebius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2508" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis id="B929EA9EFFA945101AB0FA35FE021089" box="[146,393,1506,1527]" italics="true" pageId="9" pageNumber="2508">Neochthebius granulosus</emphasis>
</taxonomicName>
; (23 and 25)
<taxonomicName id="4C5D4D0FFFA945101834FA35FD051089" baseAuthorityName="Knisch" baseAuthorityYear="1924" box="[534,654,1506,1527]" class="Insecta" family="Hydraenidae" genus="Neochthebius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2508" phylum="Arthropoda" rank="species" species="vandykei">
<emphasis id="B929EA9EFFA945101834FA35FD051089" box="[534,654,1506,1527]" italics="true" pageId="9" pageNumber="2508">N. vandykei</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF226604FFAA45131AF7F990FC6B1323" ID-DOI="http://doi.org/10.5281/zenodo.4590531" ID-Zenodo-Dep="4590531" box="[213,992,1607,1629]" httpUri="https://zenodo.org/record/4590531/files/figure.png" pageId="10" pageNumber="2509" startId="10.[213,280,1607,1629]" targetBox="[465,918,165,1571]" targetPageId="10">
<paragraph id="8BE2368CFFAA45131AF7F990FC6B1323" blockId="10.[213,992,1607,1629]" box="[213,992,1607,1629]" pageId="10" pageNumber="2509">
Figure 26. Larva habitus of
<taxonomicName id="4C5D4D0FFFAA45131BDDF990FD7D1322" authorityName=": Perkins" authorityYear="1980" box="[511,758,1607,1628]" class="Insecta" family="Hydraenidae" genus="Neochthebius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2509" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis id="B929EA9EFFAA45131BDDF990FD7D1322" box="[511,758,1607,1628]" italics="true" pageId="10" pageNumber="2509">Neochthebius granulosus</emphasis>
</taxonomicName>
. Note: Scale50.5 mm.
</paragraph>
</caption>
<paragraph id="8BE2368CFFAB45121AB0FE0BFEFA148A" blockId="11.[146,369,476,500]" box="[146,369,476,500]" pageId="11" pageNumber="2510">
<emphasis id="B929EA9EFFAB45121AB0FE0BFEFA148A" box="[146,369,476,500]" italics="true" pageId="11" pageNumber="2510">Materials examined</emphasis>
</paragraph>
<paragraph id="8BE2368CFFAB45121AB0FDD2FF5C169C" blockId="11.[146,1103,516,994]" pageId="11" pageNumber="2510">
<emphasis id="B929EA9EFFAB45121AB0FDD2FF511763" box="[146,218,517,541]" italics="true" pageId="11" pageNumber="2510">Adults</emphasis>
.
<materialsCitation id="3B353CD1FFAB45121ACEFDD3FF58169C" collectingDate="1991-07-06" collectingDateMax="2005-06-11" collectingDateMin="1991-07-06" collectorName="K. - J. Ahn &amp; J. - Y. Lyu &amp; H. - J. Kim &amp; M. - J. Jeon &amp; S. - J. Park &amp; C. - W. Shin &amp; Is. Baekryeongdo &amp; City &amp; Geumgab &amp; Imhoe-myeon &amp; Jindo &amp; Myeongsasimri &amp; Sinji-myeon &amp; Wando &amp; J. - H. Ahn &amp; Ilchulbong &amp; Seongsan-eup &amp; Namjeju-gun &amp; S. - I. Lee &amp; D. - H. Lee" location="Geoje Island" pageId="11" pageNumber="2510" specimenCount="12" stateProvince="Gyeongbuk">
Ten specimens, Gugae,
<collectingRegion id="4999F86EFFAB45121820FDD3FD591762" box="[514,722,516,540]" country="South Korea" name="Gyeongbuk" pageId="11" pageNumber="2510">Gyeongbuk Prov.</collectingRegion>
,
<date id="FFE3104CFFAB451218C2FDD3FCE91762" box="[736,866,516,540]" pageId="11" pageNumber="2510" value="1991-07-06">
<collectingDate id="EFA7E9A4FFAB451218C2FDD3FCE91762" box="[736,866,516,540]" pageId="11" pageNumber="2510" value="1991-07-06">6 VII 1991</collectingDate>
</date>
,
<collectorName id="26A8535AFFAB45121952FDD3FC631762" box="[880,1000,516,540]" pageId="11" pageNumber="2510">K.-J. Ahn</collectorName>
,
<emphasis id="B929EA9EFFAB451219D5FDD2FB9A1763" box="[1015,1041,517,541]" italics="true" pageId="11" pageNumber="2510">ex</emphasis>
rock crevice; seven specimens, Yongyeon,
<collectingRegion id="4999F86EFFAB45121819FDF2FC841743" box="[571,783,549,573]" country="South Korea" name="Gyeongnam" pageId="11" pageNumber="2510">Gyeongnam Prov.</collectingRegion>
,
<date id="FFE3104CFFAB45121939FDF2FC2E1743" box="[795,933,549,573]" pageId="11" pageNumber="2510" value="1991-07-20">
<collectingDate id="EFA7E9A4FFAB45121939FDF2FC2E1743" box="[795,933,549,573]" pageId="11" pageNumber="2510" value="1991-07-20">20 VII 1991</collectingDate>
</date>
,
<collectorName id="26A8535AFFAB45121990FDF2FBA31743" box="[946,1064,549,573]" pageId="11" pageNumber="2510">K.-J. Ahn</collectorName>
,
<emphasis id="B929EA9EFFAB45121E16FDF2FBC51743" box="[1076,1102,549,573]" italics="true" pageId="11" pageNumber="2510">ex</emphasis>
rock crevice; two specimens, Gabae-ri, Dongbu-myeon,
<location id="8E826057FFAB4512193DFD92FC3E1723" LSID="urn:lsid:plazi:treatment:03F4879AFFA745121BE2F9FCFF6F13D4:8E826057FFAB4512193DFD92FC3E1723" box="[799,949,581,605]" name="Geoje Island" pageId="11" pageNumber="2510" stateProvince="Gyeongbuk">Geoje Island</location>
,
<collectingRegion id="4999F86EFFAB451219E1FD92FF581700" country="South Korea" name="Gyeongnam" pageId="11" pageNumber="2510">Gyeongnam Prov.</collectingRegion>
,
<collectorName id="26A8535AFFAB45121AFCFDB1FEC11700" box="[222,330,614,638]" pageId="11" pageNumber="2510">J.-Y. Lyu</collectorName>
and
<collectorName id="26A8535AFFAB45121BA0FDB1FE731700" box="[386,504,614,638]" pageId="11" pageNumber="2510">H.-J. Kim</collectorName>
,
<date id="FFE3104CFFAB45121821FDB1FDF61700" box="[515,637,614,638]" pageId="11" pageNumber="2510" value="1998-07-03">
<collectingDate id="EFA7E9A4FFAB45121821FDB1FDF61700" box="[515,637,614,638]" pageId="11" pageNumber="2510" value="1998-07-03">3 VII 1998</collectingDate>
</date>
,
<emphasis id="B929EA9EFFAB451218AAFDB1FD291700" box="[648,674,614,638]" italics="true" pageId="11" pageNumber="2510">ex</emphasis>
empty barnacles; one specimen, same as former except
<date id="FFE3104CFFAB45121B74FD51FE5417E0" box="[342,479,646,670]" pageId="11" pageNumber="2510" value="1999-07-30">
<collectingDate id="EFA7E9A4FFAB45121B74FD51FE5417E0" box="[342,479,646,670]" pageId="11" pageNumber="2510" value="1999-07-30">30 VII 1999</collectingDate>
</date>
,
<collectorName id="26A8535AFFAB45121BC9FD51FDEB17E0" box="[491,608,646,670]" pageId="11" pageNumber="2510">K.-J. Ahn</collectorName>
,
<emphasis id="B929EA9EFFAB4512184EFD51FD0D17E0" box="[620,646,646,670]" italics="true" pageId="11" pageNumber="2510">ex</emphasis>
on rocks;
<specimenCount id="9D5BFD05FFAB451218DCFD51FC1D17E0" box="[766,918,646,670]" pageId="11" pageNumber="2510" type="generic">12 specimens</specimenCount>
, same as former except
<date id="FFE3104CFFAB45121AC3FD71FED017C0" box="[225,347,678,702]" pageId="11" pageNumber="2510" value="2000-07-01">
<collectingDate id="EFA7E9A4FFAB45121AC3FD71FED017C0" box="[225,347,678,702]" pageId="11" pageNumber="2510" value="2000-07-01">1 VII 2000</collectingDate>
</date>
,
<collectorName id="26A8535AFFAB45121B44FD71FE5117C0" box="[358,474,678,702]" pageId="11" pageNumber="2510">K.-J. Ahn</collectorName>
,
<collectorName id="26A8535AFFAB45121BC7FD71FDD317C0" box="[485,600,678,702]" pageId="11" pageNumber="2510">H.-J. Kim</collectorName>
and
<collectorName id="26A8535AFFAB451218B2FD71FC8017C0" box="[656,779,678,702]" pageId="11" pageNumber="2510">M.-J. Jeon</collectorName>
,
<emphasis id="B929EA9EFFAB45121934FD70FCBB17C1" box="[790,816,679,703]" italics="true" pageId="11" pageNumber="2510">ex</emphasis>
barnacles; one specimen, same as former except
<date id="FFE3104CFFAB45121BBDFD10FDA917A1" box="[415,546,711,735]" pageId="11" pageNumber="2510" value="2001-06-28">
<collectingDate id="EFA7E9A4FFAB45121BBDFD10FDA917A1" box="[415,546,711,735]" pageId="11" pageNumber="2510" value="2001-06-28">28 VI 2001</collectingDate>
</date>
,
<collectorName id="26A8535AFFAB45121812FD10FD2A17A1" box="[560,673,711,735]" pageId="11" pageNumber="2510">S.-J. Park</collectorName>
and
<collectorName id="26A8535AFFAB451218FDFD10FCE817A1" box="[735,867,711,735]" pageId="11" pageNumber="2510">C.-W. Shin</collectorName>
,
<emphasis id="B929EA9EFFAB45121953FD10FC0017A1" box="[881,907,711,735]" italics="true" pageId="11" pageNumber="2510">ex</emphasis>
under seaweeds; seven specimens, same as former except
<date id="FFE3104CFFAB451218A9FD30FC911781" box="[651,794,743,767]" pageId="11" pageNumber="2510" value="2001-06-30">
<collectingDate id="EFA7E9A4FFAB451218A9FD30FC911781" box="[651,794,743,767]" pageId="11" pageNumber="2510" value="2001-06-30">30 VI 2001</collectingDate>
</date>
,
<collectorName id="26A8535AFFAB4512190CFD30FC231781" box="[814,936,743,767]" pageId="11" pageNumber="2510">S.-J. Park</collectorName>
,
<emphasis id="B929EA9EFFAB4512199EFD30FC5D1781" box="[956,982,743,767]" italics="true" pageId="11" pageNumber="2510">ex</emphasis>
inside of barnacles; two specimens,
<collectorName id="26A8535AFFAB45121BE7FCD0FD191661" box="[453,658,775,799]" pageId="11" pageNumber="2510">Is. Baekryeongdo</collectorName>
,
<collectingRegion id="4999F86EFFAB45121882FCD0FD771661" box="[672,764,775,799]" country="South Korea" name="Incheon" pageId="11" pageNumber="2510">Incheon</collectingRegion>
<collectorName id="26A8535AFFAB45121924FCD0FCBC1661" box="[774,823,775,799]" pageId="11" pageNumber="2510">City</collectorName>
<collectingRegion id="4999F86EFFAB45121962FCD0FC731661" box="[832,1016,775,799]" country="South Korea" name="Gyeonggi" pageId="11" pageNumber="2510">Gyeonggi Prov.</collectingRegion>
,
<date id="FFE3104CFFAB45121E24FCD0FF41163E" pageId="11" pageNumber="2510" value="2000-08-07">
<collectingDate id="EFA7E9A4FFAB45121E24FCD0FF41163E" pageId="11" pageNumber="2510" value="2000-08-07">7 VIII 2000</collectingDate>
</date>
,
<collectorName id="26A8535AFFAB45121AF9FCFFFEE9163E" box="[219,354,808,832]" pageId="11" pageNumber="2510">C.-W. Shin</collectorName>
,
<emphasis id="B929EA9EFFAB45121B51FCFFFE06163E" box="[371,397,808,832]" italics="true" pageId="11" pageNumber="2510">ex</emphasis>
on rocks; nine specimens,
<collectorName id="26A8535AFFAB451218F5FCFFFCCD163E" box="[727,838,808,832]" pageId="11" pageNumber="2510">Geumgab</collectorName>
beach,
<collectorName id="26A8535AFFAB4512198BFCFFFBC1163E" box="[937,1098,808,832]" pageId="11" pageNumber="2510">Imhoe-myeon</collectorName>
,
<collectorName id="26A8535AFFAB45121AB0FC9FFF5F161E" box="[146,212,840,864]" pageId="11" pageNumber="2510">Jindo</collectorName>
,
<collectingRegion id="4999F86EFFAB45121AFDFC9FFE07161E" box="[223,396,840,864]" country="South Korea" name="Jeonnam" pageId="11" pageNumber="2510">Jeonnam Prov.</collectingRegion>
,
<date id="FFE3104CFFAB45121BB5FC9FFDA2161E" box="[407,553,840,864]" pageId="11" pageNumber="2510" value="2001-08-21">
<collectingDate id="EFA7E9A4FFAB45121BB5FC9FFDA2161E" box="[407,553,840,864]" pageId="11" pageNumber="2510" value="2001-08-21">21 VIII 2001</collectingDate>
</date>
,
<collectorName id="26A8535AFFAB45121816FC9FFD2E161E" box="[564,677,840,864]" pageId="11" pageNumber="2510">S.-J. Park</collectorName>
,
<emphasis id="B929EA9EFFAB45121892FC9FFD41161E" box="[688,714,840,864]" italics="true" pageId="11" pageNumber="2510">ex</emphasis>
on the barnacles; eight specimens,
<collectorName id="26A8535AFFAB45121AB0FCBEFECB16FF" box="[146,320,873,897]" pageId="11" pageNumber="2510">Myeongsasimri</collectorName>
beach,
<collectorName id="26A8535AFFAB45121B80FCBEFDA616FF" box="[418,557,873,897]" pageId="11" pageNumber="2510">Sinji-myeon</collectorName>
,
<collectorName id="26A8535AFFAB4512181CFCBEFD1F16FF" box="[574,660,873,897]" pageId="11" pageNumber="2510">Wando</collectorName>
,
<collectingRegion id="4999F86EFFAB45121887FCBEFCD316FF" box="[677,856,873,897]" country="South Korea" name="Jeonnam" pageId="11" pageNumber="2510">Jeonnam Prov.</collectingRegion>
,
<date id="FFE3104CFFAB4512194AFCBEFB8E16FF" box="[872,1029,873,897]" pageId="11" pageNumber="2510" value="2001-08-23">
<collectingDate id="EFA7E9A4FFAB4512194AFCBEFB8E16FF" box="[872,1029,873,897]" pageId="11" pageNumber="2510" value="2001-08-23">23 VIII 2001</collectingDate>
</date>
,
<collectorName id="26A8535AFFAB45121E34FCBEFF4F16DF" pageId="11" pageNumber="2510">K.-J. Ahn</collectorName>
and
<collectorName id="26A8535AFFAB45121ADDFC5EFEFE16DF" box="[255,373,905,929]" pageId="11" pageNumber="2510">J.-H. Ahn</collectorName>
; three specimens,
<collectorName id="26A8535AFFAB45121867FC5EFD4816DF" box="[581,707,905,929]" pageId="11" pageNumber="2510">Ilchulbong</collectorName>
,
<collectorName id="26A8535AFFAB451218F2FC5EFCFB16DF" box="[720,880,905,929]" pageId="11" pageNumber="2510">Seongsan-eup</collectorName>
,
<collectorName id="26A8535AFFAB4512195EFC5EFB9E16DF" box="[892,1045,905,929]" pageId="11" pageNumber="2510">Namjeju-gun</collectorName>
,
<collectingRegion id="4999F86EFFAB45121E03FC5EFF5816BF" country="South Korea" name="Jeju" pageId="11" pageNumber="2510">Jeju Prov.</collectingRegion>
,
<date id="FFE3104CFFAB45121AFDFC7EFED516BF" box="[223,350,937,961]" pageId="11" pageNumber="2510" value="2005-06-11">
<collectingDate id="EFA7E9A4FFAB45121AFDFC7EFED516BF" box="[223,350,937,961]" pageId="11" pageNumber="2510" value="2005-06-11">11 VI 2005</collectingDate>
</date>
,
<collectorName id="26A8535AFFAB45121B48FC7EFE4516BF" box="[362,462,937,961]" pageId="11" pageNumber="2510">S.-I. Lee</collectorName>
,
<collectorName id="26A8535AFFAB45121BFBFC7EFDC716BF" box="[473,588,937,961]" pageId="11" pageNumber="2510">S.-J. Park</collectorName>
,
<collectorName id="26A8535AFFAB4512187AFC7EFD4616BF" box="[600,717,937,961]" pageId="11" pageNumber="2510">K.-J. Ahn</collectorName>
,
<collectorName id="26A8535AFFAB451218FBFC7EFCD816BF" box="[729,851,937,961]" pageId="11" pageNumber="2510">M.-J. Jeon</collectorName>
and
<collectorName id="26A8535AFFAB451219AFFC7EFB8816BF" box="[909,1027,937,961]" pageId="11" pageNumber="2510">D.-H. Lee</collectorName>
,
<emphasis id="B929EA9EFFAB45121E2DFC7DFBA216BC" box="[1039,1065,938,962]" italics="true" pageId="11" pageNumber="2510">ex</emphasis>
on rocks
</materialsCitation>
.
</paragraph>
<paragraph id="8BE2368CFFAB45121AB0FBCCFD2F112D" blockId="11.[146,1103,1050,1107]" pageId="11" pageNumber="2510">
<emphasis id="B929EA9EFFAB45121AB0FBCCFF6B114D" box="[146,224,1051,1075]" italics="true" pageId="11" pageNumber="2510">Larvae</emphasis>
.
<materialsCitation id="3B353CD1FFAB45121AD2FBCDFD2B112D" collectingDate="1998-06-30" collectorName="K. - J. Ahn &amp; J. - H. Ahn" country="South Korea" county="Geoje Island" location="Six" municipality="Dongbu-myeon" pageId="11" pageNumber="2510" specimenCount="1" stateProvince="Gyeongnam">
<location id="8E826057FFAB45121AD2FBCDFE98114C" LSID="urn:lsid:plazi:treatment:03F4879AFFA745121BE2F9FCFF6F13D4:8E826057FFAB45121AD2FBCDFE98114C" box="[240,275,1050,1074]" country="South Korea" county="Geoje Island" municipality="Dongbu-myeon" name="Six" pageId="11" pageNumber="2510" stateProvince="Gyeongnam">Six</location>
specimens,
<location id="8E826057FFAB45121BBFFBCDFD8E114C" LSID="urn:lsid:plazi:treatment:03F4879AFFA745121BE2F9FCFF6F13D4:8E826057FFAB45121BBFFBCDFD8E114C" box="[413,517,1050,1074]" country="South Korea" county="Geoje Island" municipality="Dongbu-myeon" name="Gabae-ri" pageId="11" pageNumber="2510" stateProvince="Gyeongnam">Gabae-ri</location>
,
<collectingMunicipality id="6B86ACF6FFAB45121830FBCDFD43114C" box="[530,712,1050,1074]" pageId="11" pageNumber="2510">Dongbu-myeon</collectingMunicipality>
,
<collectingCounty id="62834E00FFAB451218F6FBCDFCE2114C" box="[724,873,1050,1074]" pageId="11" pageNumber="2510">Geoje Island</collectingCounty>
,
<collectingRegion id="4999F86EFFAB45121954FBCDFBC1114C" box="[886,1098,1050,1074]" country="South Korea" name="Gyeongnam" pageId="11" pageNumber="2510">Gyeongnam Prov.</collectingRegion>
,
<collectingCountry id="F34A761CFFAB45121AB0FBECFF57112D" box="[146,220,1083,1107]" name="South Korea" pageId="11" pageNumber="2510">Korea</collectingCountry>
,
<date id="FFE3104CFFAB45121AC8FBECFEE7112D" box="[234,364,1083,1107]" pageId="11" pageNumber="2510" value="1998-06-30">
<collectingDate id="EFA7E9A4FFAB45121AC8FBECFEE7112D" box="[234,364,1083,1107]" pageId="11" pageNumber="2510" value="1998-06-30">30 VI 1998</collectingDate>
</date>
,
<collectorName id="26A8535AFFAB45121B5BFBECFE66112D" box="[377,493,1083,1107]" pageId="11" pageNumber="2510">K.-J. Ahn</collectorName>
and
<collectorName id="26A8535AFFAB4512180BFBECFD2B112D" box="[553,672,1083,1107]" pageId="11" pageNumber="2510">J.-H. Ahn</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE2368CFFAB45121AB0FB5BFE9311DA" blockId="11.[146,280,1164,1188]" box="[146,280,1164,1188]" pageId="11" pageNumber="2510">
<emphasis id="B929EA9EFFAB45121AB0FB5BFE9311DA" box="[146,280,1164,1188]" italics="true" pageId="11" pageNumber="2510">Distribution</emphasis>
</paragraph>
<paragraph id="8BE2368CFFAB45121AB0FB63FEE911B2" blockId="11.[146,354,1204,1228]" box="[146,354,1204,1228]" pageId="11" pageNumber="2510">
<collectingCountry id="F34A761CFFAB45121AB0FB63FF5111B2" box="[146,218,1204,1228]" name="South Korea" pageId="11" pageNumber="2510">Korea</collectingCountry>
and
<collectingCountry id="F34A761CFFAB45121B34FB63FED611B2" box="[278,349,1204,1228]" name="Japan" pageId="11" pageNumber="2510">Japan</collectingCountry>
.
</paragraph>
<paragraph id="8BE2368CFFAB45121AB0FAD2FF7E1063" blockId="11.[146,245,1285,1309]" box="[146,245,1285,1309]" pageId="11" pageNumber="2510">
<emphasis id="B929EA9EFFAB45121AB0FAD2FF7E1063" box="[146,245,1285,1309]" italics="true" pageId="11" pageNumber="2510">Remarks</emphasis>
</paragraph>
<paragraph id="8BE2368CFFAB45121AB0FAF9FF6F13D4" blockId="11.[146,1103,1326,1706]" pageId="11" pageNumber="2510">
<taxonomicName id="4C5D4D0FFFAB45121AB0FAF9FE3A1038" authorityName=": Perkins" authorityYear="1980" box="[146,433,1326,1350]" class="Insecta" family="Hydraenidae" genus="Neochthebius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2510" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis id="B929EA9EFFAB45121AB0FAF9FE3A1038" box="[146,433,1326,1350]" italics="true" pageId="11" pageNumber="2510">Neochthebius granulosus</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="4C5D4D0FFFAB451218BFFAF9FCA01038" baseAuthorityName="Knisch" baseAuthorityYear="1924" box="[669,811,1326,1350]" class="Insecta" family="Hydraenidae" genus="Neochthebius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2510" phylum="Arthropoda" rank="species" species="vandykei">
<emphasis id="B929EA9EFFAB451218BFFAF9FCA01038" box="[669,811,1326,1350]" italics="true" pageId="11" pageNumber="2510">N. vandykei</emphasis>
</taxonomicName>
in body shape and the structure of the aedeagus. However, it can be distinguished from the latter by the characters in the key. This species shows geographical variation: specimens from eastern costal regions of
<collectingCountry id="F34A761CFFAB45121B97FA58FE7510D9" box="[437,510,1423,1447]" name="South Korea" pageId="11" pageNumber="2510">Korea</collectingCountry>
and
<collectingCountry id="F34A761CFFAB4512181BFA58FDF510D9" box="[569,638,1423,1447]" name="Japan" pageId="11" pageNumber="2510">Japan</collectingCountry>
have strongly imbricate antennomere 2 and maxillary palpomere 3 (Figures 10 and 13), pronotum with three distinct impressions on disk (
<figureCitation id="13662A09FFAB45121BAFFA07FE761096" box="[397,509,1488,1512]" captionStart="Figures 1921" captionStartId="7.[146,222,1493,1515]" captionTargetBox="[146,1104,1034,1457]" captionTargetId="figure-61@7.[146,1104,1034,1457]" captionTargetPageId="7" captionText="Figures 1921. Pronotum and elytra. (1920) Neochthebius granulosus; (21) N. vandykei." figureDoi="http://doi.org/10.5281/zenodo.4590527" httpUri="https://zenodo.org/record/4590527/files/figure.png" pageId="11" pageNumber="2510">Figure 20</figureCitation>
) and distal lobe of median lobe weakly sinuate in the lateral aspect. In contrast, specimens from southern costal regions and
<collectingRegion id="4999F86EFFAB45121E02FA27FBC51376" box="[1056,1102,1520,1544]" country="South Korea" name="Jeju" pageId="11" pageNumber="2510">Jeju</collectingRegion>
Island of
<collectingCountry id="F34A761CFFAB45121B20F9C6FEC01357" box="[258,331,1553,1577]" name="South Korea" pageId="11" pageNumber="2510">Korea</collectingCountry>
have weakly imbricate antennomere 2 and maxillary palpomere 3 (Figures 9 and 12), pronotum with indistinct impressions on disk (
<figureCitation id="13662A09FFAB451219BEF9E6FB861337" box="[924,1037,1585,1609]" captionStart="Figures 1921" captionStartId="7.[146,222,1493,1515]" captionTargetBox="[146,1104,1034,1457]" captionTargetId="figure-61@7.[146,1104,1034,1457]" captionTargetPageId="7" captionText="Figures 1921. Pronotum and elytra. (1920) Neochthebius granulosus; (21) N. vandykei." figureDoi="http://doi.org/10.5281/zenodo.4590527" httpUri="https://zenodo.org/record/4590527/files/figure.png" pageId="11" pageNumber="2510">Figure 19</figureCitation>
), and distal lobe of median lobe strongly sinuate in the lateral aspect (
<figureCitation id="13662A09FFAB45121987F986FB831317" box="[933,1032,1617,1641]" captionStart="Figures 38" captionStartId="4.[213,289,1532,1554]" captionTargetBox="[292,1092,165,1495]" captionTargetId="figure-0@4.[292,1092,165,1495]" captionTargetPageId="4" captionText="Figures 38. Neochthebius granulosus. (3) antenna, dorsal aspect; (4) labrum, dorsal aspect; (5) mandible, dorsal aspect; (6) maxilla, ventral aspect; (7) aedeagus, dorsal aspect; (8) aedeagus, lateral aspect. Note: Scales50.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590523" httpUri="https://zenodo.org/record/4590523/files/figure.png" pageId="11" pageNumber="2510">Figure 8</figureCitation>
). The number of stout spines in the female last abdominal segment varies from seven to twelve.
</paragraph>
</subSubSection>
</treatment>
</document>