429 lines
67 KiB
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429 lines
67 KiB
XML
<document id="3A2FCF8C51BB1047CEF601A06570BC0A" ID-CLB-Dataset="3948" ID-DOI="10.5852/ejt.2021.775.1551" ID-GBIF-Dataset="cc10c087-bec3-4947-81d7-b3e5580a5c94" ID-ISSN="2118-9773" ID-Zenodo-Dep="5589689" ID-ZooBank="976D7020-5904-4951-97CE-B4FE58DA12A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634822039012" checkinUser="felipe" docAuthor="González-Alvarado, Arturo & Vaz-De, Fernando Z." docDate="2021" docId="03F48795DC57FFBEA63AA65BFC04FE94" docLanguage="en" docName="ejt-775_86-106.pdf" docOrigin="European Journal of Taxonomy 775" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Deltochilum jocelynae González-Alvarado & Vaz-De 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="99" masterDocId="FFCDFFEDDC5DFFB3A45AA753FFCDFFC7" masterDocTitle="Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum)" masterLastPageNumber="106" masterPageNumber="86" pageNumber="96" updateTime="1699464201688" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9427DBEF2D8CE3A9D390E98534ADE42C">Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum)</mods:title>
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<mods:namePart id="AEE1DA7BB497F002687F86C1E13CF1DF">González-Alvarado, Arturo</mods:namePart>
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<mods:affiliation id="0F448C6D56DE07254B3F31923295452B">Programa de Pós-graduação em Ecologia e Conservação da Biodiversidade, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="EEC6B97DE1B3F274A97452C866977882" type="email">fagakorn@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B906E0DBBEADB0E1D3181FA3481F1738">Vaz-De, Fernando Z.</mods:namePart>
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<mods:affiliation id="D5D2E0889BF17BF6FEA3C707FF2AF431">Instituto de Biociências, Departamento de Biologia e Zoologia, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="0B63797815AA51F9ACD58992FD8021D7" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:title id="7ADC36559AB42B091ED70F7C332E866A">European Journal of Taxonomy</mods:title>
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<mods:date id="DCA186175852102B7C204F9F91A217F3">2021</mods:date>
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<treatment id="03F48795DC57FFBEA63AA65BFC04FE94" ID-DOI="http://doi.org/10.5281/zenodo.5589703" ID-GBIF-Taxon="189151489" ID-Zenodo-Dep="5589703" LSID="urn:lsid:plazi:treatment:03F48795DC57FFBEA63AA65BFC04FE94" httpUri="http://treatment.plazi.org/id/03F48795DC57FFBEA63AA65BFC04FE94" lastPageId="13" lastPageNumber="99" pageId="10" pageNumber="96">
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<paragraph id="8BE23683DC57FFB9A63AA65BFC19FEE4" blockId="10.[363,1225,264,362]" box="[608,980,264,291]" pageId="10" pageNumber="96">
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<heading id="D0AA81EFDC57FFB9A63AA65BFC19FEE4" box="[608,980,264,291]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="96" reason="2">
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<taxonomicName id="4C5D4D00DC57FFB9A63AA65BFCBFFEE4" ID-CoL="8SSPY" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[608,882,264,291]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="96" phylum="Arthropoda" rank="species" species="jocelynae" status="sp. nov.">
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<emphasis id="B929EA91DC57FFB9A63AA65BFCBFFEE4" bold="true" box="[608,882,264,291]" italics="true" pageId="10" pageNumber="96">Deltochilum jocelynae</emphasis>
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<taxonomicNameLabel id="A21A57EADC57FFB9A723A65AFC19FEE4" box="[889,980,265,291]" pageId="10" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C3476508DC57FFB9A531A678FB9DFEAE" pageId="10" pageNumber="96" type="description">
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<paragraph id="8BE23683DC57FFB9A5B9A61CFB9DFEAE" blockId="10.[363,1225,264,362]" box="[483,1104,335,362]" pageId="10" pageNumber="96">
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<figureCitation id="13662A06DC57FFB9A5B9A61CFD8CFEAE" box="[483,577,335,361]" captionStart="Fig" captionStartId="3.[189,232,1892,1918]" captionTargetBox="[222,1362,786,1773]" captionTargetId="figure-203@3.[219,1368,777,1781]" captionTargetPageId="3" captionText="Fig. 1. Habitus of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589691" httpUri="https://zenodo.org/record/5589691/files/figure.png" pageId="10" pageNumber="96">Figs 1B</figureCitation>
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,
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<figureCitation id="13662A06DC57FFB9A614A61CFDBEFEAE" box="[590,627,335,361]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[237,1351,525,1826]" captionTargetId="figure-106@4.[237,1351,525,1826]" captionTargetPageId="4" captionText="Fig. 2. Head of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589695" httpUri="https://zenodo.org/record/5589695/files/figure.png" pageId="10" pageNumber="96">2B</figureCitation>
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,
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<figureCitation id="13662A06DC57FFB9A625A61CFD0CFEAE" box="[639,705,335,361]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="10" pageNumber="96">3B, F</figureCitation>
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,
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<figureCitation id="13662A06DC57FFB9A697A61CFD3CFEAE" box="[717,753,335,361]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="10" pageNumber="96">4B</figureCitation>
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,
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<figureCitation id="13662A06DC57FFB9A6A4A61CFCEFFEAD" box="[766,802,335,362]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="10" pageNumber="96">5B</figureCitation>
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,
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<figureCitation id="13662A06DC57FFB9A774A61CFC9EFEAE" box="[814,851,335,361]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[230,1363,815,1803]" captionTargetId="figure-152@8.[220,1368,814,1813]" captionTargetPageId="8" captionText="Fig. 6. Ventral view of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589707" httpUri="https://zenodo.org/record/5589707/files/figure.png" pageId="10" pageNumber="96">6B</figureCitation>
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,
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<figureCitation id="13662A06DC57FFB9A705A61CFC6CFEAE" box="[863,929,335,361]" captionStart="Fig" captionStartId="9.[189,232,1822,1848]" captionTargetBox="[206,1383,744,1716]" captionTargetId="figure-193@9.[203,1383,744,1716]" captionTargetPageId="9" captionText="Fig. 7.Aedeagus of members of thegilli species group.A–E. Lateral view.F–G. Dorsal view of paramera. Arrows show the apical-dorsal notches of the paramera. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov. CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Holotype of D. jocelynae sp. nov. G. Holotype of D. nonstriatum sp. nov., CMNEN WDS00041746." figureDoi="http://doi.org/10.5281/zenodo.5589711" httpUri="https://zenodo.org/record/5589711/files/figure.png" pageId="10" pageNumber="96">7B, F</figureCitation>
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,
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<figureCitation id="13662A06DC57FFB9A7F7A61CFC1FFEAE" box="[941,978,335,361]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[233,1362,650,1777]" captionTargetId="figure-181@10.[218,1370,644,1787]" captionTargetPageId="10" captionText="Fig. 8. Endophallus of members of thegilli species group.Arrows show the scales on the sub-medial area of the endophallus. A. Elongate scales. C. Regular scales. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589713" httpUri="https://zenodo.org/record/5589713/files/figure.png" pageId="10" pageNumber="96">8B</figureCitation>
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,
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<figureCitation id="13662A06DC57FFB9A784A61CFBEAFEAE" box="[990,1063,335,361]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[268,1315,559,1863]" captionTargetId="figure-129@12.[268,1320,553,1873]" captionTargetPageId="12" captionText="Fig. 9. Habitus A, C and ventral view B, D of the only two females known of the gilli species group. A–B. Paratype of Deltochilum gilli González-Alvarado & Vaz-de-Mello, 2021. C–D. Paratype D. jocelynae sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589715" httpUri="https://zenodo.org/record/5589715/files/figure.png" pageId="10" pageNumber="96">9C–D</figureCitation>
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,
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<figureCitation id="13662A06DC57FFB9A069A61CFB9DFEAE" box="[1075,1104,335,361]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[336,1251,578,1870]" captionTargetId="figure-135@14.[336,1251,578,1870]" captionTargetPageId="14" captionText="Fig. 10. Distribution map of thegilli species group. Blue circle =Deltochilum gilli González-Alvarado & Vaz-de-Mello, 2021; yellow square = D. jocelynae sp. nov.; red star = D. nonstriatum sp. nov.; green triangle = D. quasistriatum sp. nov.; white eight-point star = D. tenuistriatum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589717" httpUri="https://zenodo.org/record/5589717/files/figure.png" pageId="10" pageNumber="96">10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3476508DC57FFB9A4E7A6C3FAC2FD84" pageId="10" pageNumber="96" type="diagnosis">
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<paragraph id="8BE23683DC57FFB9A4E7A6C3FEFBFE6D" blockId="10.[189,310,400,426]" box="[189,310,400,426]" pageId="10" pageNumber="96">
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<heading id="D0AA81EFDC57FFB9A4E7A6C3FEFBFE6D" bold="true" box="[189,310,400,426]" fontSize="11" level="3" pageId="10" pageNumber="96" reason="3">
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<emphasis id="B929EA91DC57FFB9A4E7A6C3FEFBFE6D" bold="true" box="[189,310,400,426]" pageId="10" pageNumber="96">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE23683DC57FFB9A4E7A6ECFAC2FD84" blockId="10.[188,1401,447,579]" pageId="10" pageNumber="96">
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Close to
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<taxonomicName id="4C5D4D00DC57FFB9A576A693FE12FE1E" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[300,479,447,473]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="96" phylum="Arthropoda" rank="species" species="nonstriatum" status="sp. nov.">
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<emphasis id="B929EA91DC57FFB9A576A693FEF2FE1E" box="[300,319,448,473]" italics="true" pageId="10" pageNumber="96">D</emphasis>
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||
.
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<emphasis id="B929EA91DC57FFB9A50AA693FE12FE1E" box="[336,479,448,473]" italics="true" pageId="10" pageNumber="96">nonstriatum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21A57EADC57FFB9A5B6A6ECFD85FE1E" box="[492,584,447,473]" pageId="10" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
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by having striae I–VII inconspicuous (
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<figureCitation id="13662A06DC57FFB9A045A6ECFB69FE1E" box="[1055,1188,447,473]" captionStart="Fig" captionStartId="3.[189,232,1892,1918]" captionTargetBox="[222,1362,786,1773]" captionTargetId="figure-203@3.[219,1368,777,1781]" captionTargetPageId="3" captionText="Fig. 1. Habitus of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589691" httpUri="https://zenodo.org/record/5589691/files/figure.png" pageId="10" pageNumber="96">Figs 1B–C</figureCitation>
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,
|
||
<figureCitation id="13662A06DC57FFB9A0E8A6ECFB35FE1E" box="[1202,1272,447,473]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="10" pageNumber="96">4B–C</figureCitation>
|
||
) including apically (
|
||
<figureCitation id="13662A06DC57FFB9A575A6B1FE7CFE3A" box="[303,433,482,509]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="10" pageNumber="96">Fig. 5B–C</figureCitation>
|
||
). However, it can be distinguished by presenting the smallest and most disperse interstrial punctures (
|
||
<figureCitation id="13662A06DC57FFB9A5EEA556FDC0FDD8" box="[436,525,517,543]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="10" pageNumber="96">Fig. 4B</figureCitation>
|
||
) as well as most disperse punctures on head frons (
|
||
<figureCitation id="13662A06DC57FFB9A000A556FB7FFDD8" box="[1114,1202,517,543]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[237,1351,525,1826]" captionTargetId="figure-106@4.[237,1351,525,1826]" captionTargetPageId="4" captionText="Fig. 2. Head of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589695" httpUri="https://zenodo.org/record/5589695/files/figure.png" pageId="10" pageNumber="96">Fig. 2B</figureCitation>
|
||
) and on pronotal disc (
|
||
<figureCitation id="13662A06DC57FFB9A4A1A57BFEB8FD84" box="[251,373,552,579]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="10" pageNumber="96">Fig. 3B, F</figureCitation>
|
||
) and finally, by the well-defined shiny points on the pronotal disc (
|
||
<figureCitation id="13662A06DC57FFB9A0DCA57BFACCFD84" box="[1158,1281,552,579]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="10" pageNumber="96">Fig. 3B, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF22660BDC57FFB9A4E7A012FA98F82E" ID-DOI="http://doi.org/10.5281/zenodo.5589713" ID-Zenodo-Dep="5589713" httpUri="https://zenodo.org/record/5589713/files/figure.png" pageId="10" pageNumber="96" startId="10.[189,232,1857,1883]" targetBox="[233,1362,650,1777]" targetPageId="10">
|
||
<paragraph id="8BE23683DC57FFB9A4E7A012FA98F82E" blockId="10.[187,1399,1857,2025]" pageId="10" pageNumber="96">
|
||
<emphasis id="B929EA91DC57FFB9A4E7A012FEC4F89C" bold="true" box="[189,265,1857,1883]" pageId="10" pageNumber="96">Fig. 8.</emphasis>
|
||
Endophallus of members of the
|
||
<taxonomicName id="4C5D4D00DC57FFB9A621A012FD66F89C" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[635,683,1857,1883]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="96" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC57FFB9A621A012FD66F89C" box="[635,683,1857,1883]" italics="true" pageId="10" pageNumber="96">gilli</emphasis>
|
||
</taxonomicName>
|
||
species group.Arrows show the scales on the sub-medial area of the endophallus.
|
||
<emphasis id="B929EA91DC57FFB9A5FDA037FE71F8B9" bold="true" box="[423,444,1892,1918]" pageId="10" pageNumber="96">A</emphasis>
|
||
. Elongate scales.
|
||
<emphasis id="B929EA91DC57FFB9A6C8A037FD6AF8B9" bold="true" box="[658,679,1892,1918]" pageId="10" pageNumber="96">C</emphasis>
|
||
. Regular scales.
|
||
<emphasis id="B929EA91DC57FFB9A72BA037FC4BF8B9" bold="true" box="[881,902,1892,1918]" pageId="10" pageNumber="96">A</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC57FFB9A076A036FB3BF8B8" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[1068,1270,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="96" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC57FFB9A076A036FB3BF8B8" box="[1068,1270,1893,1919]" italics="true" pageId="10" pageNumber="96">Deltochilum gilli</emphasis>
|
||
</taxonomicName>
|
||
González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888.
|
||
<emphasis id="B929EA91DC57FFB9A75FA0DBFCD5F865" bold="true" box="[773,792,1928,1954]" pageId="10" pageNumber="96">B</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC57FFB9A7E5A0DAFB9BF865" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[959,1110,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="96" phylum="Arthropoda" rank="species" species="jocelynae" status="sp. nov.">
|
||
<emphasis id="B929EA91DC57FFB9A7E5A0DAFB9BF865" box="[959,1110,1928,1954]" italics="true" pageId="10" pageNumber="96">D. jocelynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC57FFB9A007A0DBFB71F865" box="[1117,1212,1928,1954]" pageId="10" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CNCI 379889.
|
||
<emphasis id="B929EA91DC57FFB9A4E4A0F8FF1FF802" bold="true" box="[190,210,1963,1989]" pageId="10" pageNumber="96">C</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC57FFB9A524A0FFFDF8F802" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[382,565,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="96" phylum="Arthropoda" rank="species" species="nonstriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC57FFB9A524A0FFFE5EF802" box="[382,403,1964,1989]" italics="true" pageId="10" pageNumber="96">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC57FFB9A5FCA0FFFDF8F802" box="[422,565,1964,1989]" italics="true" pageId="10" pageNumber="96">nonstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC57FFB9A61AA0FFFD6DF801" box="[576,672,1964,1990]" pageId="10" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CMNEN WSD00041746.
|
||
<emphasis id="B929EA91DC57FFB9A7B1A0F8FBCDF802" bold="true" box="[1003,1024,1963,1989]" pageId="10" pageNumber="96">D</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC57FFB9A0F6A0FFFABBF802" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[1196,1398,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="96" phylum="Arthropoda" rank="species" species="quasistriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC57FFB9A0F6A0FFFB0CF802" box="[1196,1217,1964,1989]" italics="true" pageId="10" pageNumber="96">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC57FFB9A089A0FFFABBF802" box="[1235,1398,1964,1989]" italics="true" pageId="10" pageNumber="96">quasistriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC57FFB9A4E7A09CFED7F82E" box="[189,282,1999,2025]" pageId="10" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CMNEN WSD00041744.
|
||
<emphasis id="B929EA91DC57FFB9A606A09DFDBDF82F" bold="true" box="[604,624,1998,2024]" pageId="10" pageNumber="96">E</emphasis>
|
||
. Holotype
|
||
<taxonomicName id="4C5D4D00DC57FFB9A6A9A09CFC78F82F" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[755,949,1999,2025]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="96" phylum="Arthropoda" rank="species" species="tenuistriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC57FFB9A6A9A09CFCC5F82F" box="[755,776,1999,2024]" italics="true" pageId="10" pageNumber="96">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC57FFB9A74DA09CFC78F82F" box="[791,949,1999,2024]" italics="true" pageId="10" pageNumber="96">tenuistriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC57FFB9A7E7A09CFBD7F82E" box="[957,1050,1999,2025]" pageId="10" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CMNEN WSD00041745.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3476508DC56FFB8A4E7A65AFAF1FE96" box="[189,1340,265,338]" pageId="11" pageNumber="97" type="etymology">
|
||
<paragraph id="8BE23683DC56FFB8A4E7A65AFE89FEE4" blockId="11.[189,324,265,291]" box="[189,324,265,291]" pageId="11" pageNumber="97">
|
||
<heading id="D0AA81EFDC56FFB8A4E7A65AFE89FEE4" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="11" pageNumber="97" reason="3">
|
||
<emphasis id="B929EA91DC56FFB8A4E7A65AFE89FEE4" bold="true" box="[189,324,265,291]" pageId="11" pageNumber="97">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE23683DC56FFB8A4E7A66BFAF1FE96" blockId="11.[189,1340,311,338]" box="[189,1340,311,338]" pageId="11" pageNumber="97">A patronym, noun in the genitive case, for Jocelyn Gill. See also the “Acknowledgments” section.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3476508DC56FFB8A4E7A629FC26FD70" pageId="11" pageNumber="97" type="materials_examined">
|
||
<paragraph id="8BE23683DC56FFB8A4E7A629FEA1FE54" blockId="11.[189,364,377,404]" box="[189,364,377,404]" pageId="11" pageNumber="97">
|
||
<heading id="D0AA81EFDC56FFB8A4E7A629FEA1FE54" bold="true" box="[189,364,377,404]" fontSize="11" level="3" pageId="11" pageNumber="97" reason="3">
|
||
<emphasis id="B929EA91DC56FFB8A4E7A629FEA1FE54" bold="true" box="[189,364,377,404]" pageId="11" pageNumber="97">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE23683DC56FFB8A4B0A6FAFDB7FDE9" blockId="11.[189,1399,425,558]" pageId="11" pageNumber="97">
|
||
<materialsCitation id="3B353CDEDC56FFB8A4B0A6FAFDB7FDE9" ID-GBIF-Occurrence="3391047302" collectedFrom="primary forest" collectingDate="2004-11-04" collectingDateMax="2004-11-08" collectingDateMin="2004-11-04" collectingMethod="pitfall trap" collectionCode="BDGC" collectorName="B. Hubley" country="Guyana" elevation="1411" latitude="5.0926113" location="District" longLatPrecision="1" longitude="-59.842888" municipality="Mount Wokomung" pageId="11" pageNumber="97" specimenCode="CNC 379889" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="B929EA91DC56FFB8A4B0A6FAFE91FE04" bold="true" box="[234,348,425,451]" pageId="11" pageNumber="97">
|
||
<typeStatus id="54E68821DC56FFB8A4B0A6FAFE91FE04" box="[234,348,425,451]" pageId="11" pageNumber="97" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F34A7613DC56FFB8A4E7A69FFEFAFE21" box="[189,311,460,486]" name="Guyana" pageId="11" pageNumber="97">GUYANA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D5BFD0ADC56FFB8A50EA69FFE48FE20" box="[340,389,460,487]" pageId="11" pageNumber="97" type="male">1 ♂</specimenCount>
|
||
;
|
||
<location id="8E826058DC56FFB8A5C9A69FFE26FE21" LSID="urn:lsid:plazi:treatment:03F48795DC57FFBEA63AA65BFC04FE94:8E826058DC56FFB8A5C9A69FFE26FE21" box="[403,491,460,486]" country="Guyana" latitude="5.0926113" longLatPrecision="1" longitude="-59.842888" municipality="Mount Wokomung" name="District" pageId="11" pageNumber="97">District</location>
|
||
8,
|
||
<collectingMunicipality id="6B86ACF9DC56FFB8A649A69EFD3BFE21" box="[531,758,460,487]" pageId="11" pageNumber="97">Mount Wokomung</collectingMunicipality>
|
||
;
|
||
<geoCoordinate id="EE695044DC56FFB8A759A698FC6FFE20" box="[771,930,459,487]" degrees="5" direction="north" minutes="05" orientation="latitude" pageId="11" pageNumber="97" precision="1" seconds="33.4" value="5.0926113">5°05′33.4″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695044DC56FFB8A7EAA698FBAEFE20" box="[944,1123,459,487]" degrees="59" direction="west" minutes="50" orientation="longitude" pageId="11" pageNumber="97" precision="1" seconds="34.4" value="-59.842888">59°50′34.4″ W</geoCoordinate>
|
||
; alt.
|
||
<quantity id="4CA59B66DC56FFB8A0C5A69FFB36FE20" box="[1183,1275,460,487]" metricMagnitude="3" metricUnit="m" metricValue="1.411" pageId="11" pageNumber="97" unit="m" value="1411.0">
|
||
<elevation id="0070D1B0DC56FFB8A0C5A69FFB36FE20" box="[1183,1275,460,487]" metricMagnitude="3" metricUnit="m" metricValue="1.411" pageId="11" pageNumber="97" unit="m" value="1411.0">1411 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="FFE31043DC56FFB8A153A69FFF36FDCD" pageId="11" pageNumber="97" value="2004-11-04" valueMax="2004-11-08" valueMin="2004-11-04">
|
||
<collectingDate id="EFA7E9ABDC56FFB8A153A69FFF36FDCD" pageId="11" pageNumber="97" value="2004-11-04" valueMax="2004-11-08" valueMin="2004-11-04">4–8 Nov. 2004</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26A85355DC56FFB8A55DA6A3FEB2FDCD" box="[263,383,496,522]" pageId="11" pageNumber="97">B. Hubley</collectorName>
|
||
leg.;
|
||
<collectingMethod id="521C4E94DC56FFB8A598A6A3FDF0FDCD" box="[450,573,496,522]" pageId="11" pageNumber="97">pitfall trap</collectingMethod>
|
||
(human dung),
|
||
<collectedFrom id="3841757EDC56FFB8A6A2A6A3FC6EFDCD" box="[760,931,496,522]" pageId="11" pageNumber="97">primary forest</collectedFrom>
|
||
; ROM 2004526;
|
||
<collectionCode id="ED4CAE46DC56FFB8A035A6A3FB09FDCD" box="[1135,1220,496,522]" pageId="11" pageNumber="97">BDGC</collectionCode>
|
||
; [aedeagus and endophallus extracted];
|
||
<specimenCode id="DBFB9EF8DC56FFB8A58DA547FDBBFDE9" box="[471,630,532,558]" pageId="11" pageNumber="97">CNC 379889</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BE23683DC56FFB8A4B0A505FC26FD70" blockId="11.[189,1399,598,696]" pageId="11" pageNumber="97">
|
||
<materialsCitation id="3B353CDEDC56FFB8A4B0A505FCE2FD54" ID-GBIF-Occurrence="3391047301" collectedFrom="primary forest" collectingDate="2004-11-04" collectingDateMax="2004-11-08" collectingDateMin="2004-11-04" collectingMethod="pitfall trap" collectionCode="BDGC" collectorName="B. Hubley" country="Guyana" elevation="1411" latitude="5.0926113" location="GUYANA" longLatPrecision="1" longitude="-59.842888" municipality="Mount Wokomung" pageId="11" pageNumber="97" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<emphasis id="B929EA91DC56FFB8A4B0A505FEAAFDB7" bold="true" box="[234,359,598,624]" pageId="11" pageNumber="97">
|
||
<typeStatus id="54E68821DC56FFB8A4B0A505FEAAFDB7" box="[234,359,598,624]" pageId="11" pageNumber="97" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F34A7613DC56FFB8A4E7A52AFEF4FD54" box="[189,313,633,659]" name="Guyana" pageId="11" pageNumber="97">GUYANA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D5BFD0ADC56FFB8A50DA52AFE48FD51" box="[343,389,633,662]" pageId="11" pageNumber="97" type="male">
|
||
1
|
||
<emphasis id="B929EA91DC56FFB8A535A529FE48FD51" bold="true" box="[367,389,634,662]" pageId="11" pageNumber="97">♂</emphasis>
|
||
</specimenCount>
|
||
; same data as for holotype;
|
||
<collectionCode id="ED4CAE46DC56FFB8A684A52AFCE2FD54" box="[734,815,633,659]" pageId="11" pageNumber="97">BDGC</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B353CDEDC56FFB8A717A52AFC2BFD70" ID-GBIF-Occurrence="3391047304" collectedFrom="primary forest" collectingDate="2004-10-21" collectingDateMax="2004-10-26" collectingDateMin="2004-10-21" collectingMethod="pitfall trap" collectionCode="BDGC" collectorName="B. Hubley" country="Guyana" elevation="698" latitude="5.1314445" location="District" longLatPrecision="1" longitude="-59.80872" municipality="Mount Wokomung" pageId="11" pageNumber="97" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="9D5BFD0ADC56FFB8A717A52AFCB1FD51" box="[845,892,633,662]" pageId="11" pageNumber="97" type="female">
|
||
1
|
||
<emphasis id="B929EA91DC56FFB8A73CA529FCB1FD51" bold="true" box="[870,892,634,662]" pageId="11" pageNumber="97">♀</emphasis>
|
||
</specimenCount>
|
||
; same data as for holotype;
|
||
<geoCoordinate id="EE695044DC56FFB8A08FA524FABFFD53" box="[1237,1394,631,660]" degrees="5" direction="north" minutes="07" orientation="latitude" pageId="11" pageNumber="97" precision="1" seconds="53.2" value="5.1314445">5º07′53.2″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE695044DC56FFB8A4E7A5C8FEA1FD70" box="[189,364,667,695]" degrees="59" direction="west" minutes="48" orientation="longitude" pageId="11" pageNumber="97" precision="1" seconds="31.4" value="-59.80872">59º48′31.4″ W</geoCoordinate>
|
||
; alt.
|
||
<quantity id="4CA59B66DC56FFB8A5FEA5CEFE3CFD70" box="[420,497,669,695]" metricMagnitude="2" metricUnit="m" metricValue="6.9799999999999995" pageId="11" pageNumber="97" unit="m" value="698.0">
|
||
<elevation id="0070D1B0DC56FFB8A5FEA5CEFE3CFD70" box="[420,497,669,695]" metricMagnitude="2" metricUnit="m" metricValue="6.9799999999999995" pageId="11" pageNumber="97" unit="m" value="698.0">698 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="FFE31043DC56FFB8A5A7A5CEFD08FD70" box="[509,709,669,695]" pageId="11" pageNumber="97" value="2004-10-21" valueMax="2004-10-26" valueMin="2004-10-21">
|
||
<collectingDate id="EFA7E9ABDC56FFB8A5A7A5CEFD08FD70" box="[509,709,669,695]" pageId="11" pageNumber="97" value="2004-10-21" valueMax="2004-10-26" valueMin="2004-10-21">21–26 Oct. 2004</collectingDate>
|
||
</date>
|
||
; ROM 2004509;
|
||
<collectionCode id="ED4CAE46DC56FFB8A7CBA5CEFC2BFD70" box="[913,998,669,695]" pageId="11" pageNumber="97">BDGC</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3476508DC56FFB8A4E7A58DFCE9F8F7" pageId="11" pageNumber="97" type="description">
|
||
<paragraph id="8BE23683DC56FFB8A4E7A58DFE83FD3F" blockId="11.[189,334,734,760]" box="[189,334,734,760]" pageId="11" pageNumber="97">
|
||
<heading id="D0AA81EFDC56FFB8A4E7A58DFE83FD3F" bold="true" box="[189,334,734,760]" fontSize="11" level="3" pageId="11" pageNumber="97" reason="3">
|
||
<emphasis id="B929EA91DC56FFB8A4E7A58DFE83FD3F" bold="true" box="[189,334,734,760]" pageId="11" pageNumber="97">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE23683DC56FFB8A4E7A45DFC38FCB7" blockId="11.[189,1398,781,880]" pageId="11" pageNumber="97">
|
||
MEASUREMENTS AND COLOR.
|
||
<typeStatus id="54E68821DC56FFB8A652A45DFDB8FCEF" box="[520,629,782,808]" pageId="11" pageNumber="97" type="holotype">Holotype</typeStatus>
|
||
male, length
|
||
<quantity id="4CA59B66DC56FFB8A744A45DFCB3FCEF" box="[798,894,782,808]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="11" pageNumber="97" unit="mm" value="8.1">8.1 mm</quantity>
|
||
, humeral width
|
||
<quantity id="4CA59B66DC56FFB8A01EA45EFB69FCEF" box="[1092,1188,781,808]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="11" pageNumber="97" unit="mm" value="5.1">5.1 mm</quantity>
|
||
. Dark green with some red reflections dorsally (
|
||
<figureCitation id="13662A06DC56FFB8A67DA462FD4EFC8C" box="[551,643,817,843]" captionStart="Fig" captionStartId="3.[189,232,1892,1918]" captionTargetBox="[222,1362,786,1773]" captionTargetId="figure-203@3.[219,1368,777,1781]" captionTargetPageId="3" captionText="Fig. 1. Habitus of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589691" httpUri="https://zenodo.org/record/5589691/files/figure.png" pageId="11" pageNumber="97">Fig. 1B</figureCitation>
|
||
). Black ventrally, with shiny red reflections on anterior area of metaventral process, meso- and metafemora and ventrite VI (
|
||
<figureCitation id="13662A06DC56FFB8A7D1A406FC2AFCB7" box="[907,999,853,880]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[230,1363,815,1803]" captionTargetId="figure-152@8.[220,1368,814,1813]" captionTargetPageId="8" captionText="Fig. 6. Ventral view of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589707" httpUri="https://zenodo.org/record/5589707/files/figure.png" pageId="11" pageNumber="97">Fig. 6B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE23683DC56FFB8A4E7A4CBFDA9FC3E" blockId="11.[189,1398,919,1017]" pageId="11" pageNumber="97">
|
||
HEAD (
|
||
<figureCitation id="13662A06DC56FFB8A554A4C4FEA7FC76" box="[270,362,919,946]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[237,1351,525,1826]" captionTargetId="figure-106@4.[237,1351,525,1826]" captionTargetPageId="4" captionText="Fig. 2. Head of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589695" httpUri="https://zenodo.org/record/5589695/files/figure.png" pageId="11" pageNumber="97">Fig. 2B</figureCitation>
|
||
). Dorsal inter-ocular distance approximately seven times width of one eye. Punctures on frons separated by less than one diameter of each puncture. Punctures on head disc separated by less than one diameter of each puncture.
|
||
</paragraph>
|
||
<paragraph id="8BE23683DC56FFB8A4E7A372FBDCFB99" blockId="11.[188,1398,1056,1118]" pageId="11" pageNumber="97">
|
||
PRONOTUM (
|
||
<figureCitation id="13662A06DC56FFB8A51DA373FE0DFBFC" box="[327,448,1056,1083]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="11" pageNumber="97">Fig. 3B, F</figureCitation>
|
||
). Medial angle slightly projected. Punctures on the disc separated by less than one diameter. Shiny points on disc well-defined and contiguous to punctures.
|
||
</paragraph>
|
||
<paragraph id="8BE23683DC56FFB8A4E7A3D5FC8CFAE8" blockId="11.[189,1400,1158,1327]" pageId="11" pageNumber="97">
|
||
ELYTRA (
|
||
<figureCitation id="13662A06DC56FFB8A57DA3D5FE44FB67" box="[295,393,1158,1184]" captionStart="Fig" captionStartId="3.[189,232,1892,1918]" captionTargetBox="[222,1362,786,1773]" captionTargetId="figure-203@3.[219,1368,777,1781]" captionTargetPageId="3" captionText="Fig. 1. Habitus of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589691" httpUri="https://zenodo.org/record/5589691/files/figure.png" pageId="11" pageNumber="97">Figs 1B</figureCitation>
|
||
,
|
||
<figureCitation id="13662A06DC56FFB8A5C0A3D5FE0DFB67" box="[410,448,1158,1184]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="11" pageNumber="97">4B</figureCitation>
|
||
). Carina of the ninth interstria reaching middle of elytral length. Striae I–VIII inconspicuous including apically (
|
||
<figureCitation id="13662A06DC56FFB8A60EA3FAFD7DFB03" box="[596,688,1193,1220]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="11" pageNumber="97">Fig. 5B</figureCitation>
|
||
), only in some parts striae slightly visible and very narrow. Punctures of second interstria on disc separated by one or less than one diameter, on third a slightly disperse. Punctures of third interstria on disc occupying about
|
||
<emphasis id="B929EA91DC56FFB8A7C3A3A2FC76FACC" bold="true" box="[921,955,1265,1291]" pageId="11" pageNumber="97">1/14</emphasis>
|
||
<superScript id="7C289BCBDC56FFB8A7E1A3A3FC05FB39" attach="left" box="[955,968,1264,1278]" fontSize="6" pageId="11" pageNumber="97">th</superScript>
|
||
of the distance between striae II and III. Apical tubercles on interstriae III, V–VII (
|
||
<figureCitation id="13662A06DC56FFB8A68DA247FCF9FAE8" box="[727,820,1300,1327]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="11" pageNumber="97">Fig. 5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE23683DC56FFB8A4E7A204FDA9FA1C" blockId="11.[189,1399,1366,1500]" pageId="11" pageNumber="97">
|
||
ABDOMEN (
|
||
<figureCitation id="13662A06DC56FFB8A564A205FE56FAB6" box="[318,411,1366,1393]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[230,1363,815,1803]" captionTargetId="figure-152@8.[220,1368,814,1813]" captionTargetPageId="8" captionText="Fig. 6. Ventral view of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589707" httpUri="https://zenodo.org/record/5589707/files/figure.png" pageId="11" pageNumber="97">Fig. 6B</figureCitation>
|
||
). Width of expansion of the ventrite I, on ventrite III, subequal to distance between clypeal teeth; expansion reaching distal margin of ventrite IV. Margins of expansion between ventrites II–IV forming an acute angle. Apex of expansion truncate. Basal area of expansion with punctures separated by less than one diameter.
|
||
</paragraph>
|
||
<paragraph id="8BE23683DC56FFB8A4E7A157FCB3F9A2" blockId="11.[189,1398,1539,1637]" pageId="11" pageNumber="97">LEGS. Apex of mesotibia on ventral-internal margin with a small spatulate expansion. Expansion of metafemur 1.8 × wider than the width of metafemur basal to expansion. Internal margin of metatibia with large tubercles, occupying almost all metatibial length.</paragraph>
|
||
<paragraph id="8BE23683DC56FFB8A4E7A1DEFABAF90D" blockId="11.[188,1399,1677,1739]" pageId="11" pageNumber="97">
|
||
PYGIDIUM (
|
||
<figureCitation id="13662A06DC56FFB8A560A1DEFE58F96F" box="[314,405,1677,1704]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="11" pageNumber="97">Fig. 5B</figureCitation>
|
||
). Most of the punctures separated by less than one diameter; punctures basally denser than punctures on disc. Discal punctures occupying approximately
|
||
<emphasis id="B929EA91DC56FFB8A7E5A1E3FC2EF90D" bold="true" box="[959,995,1712,1738]" pageId="11" pageNumber="97">1/30</emphasis>
|
||
<superScript id="7C289BCBDC56FFB8A7B9A1E3FC3DF979" attach="left" box="[995,1008,1712,1726]" fontSize="6" pageId="11" pageNumber="97">th</superScript>
|
||
the width on middle of pygidium.
|
||
</paragraph>
|
||
<paragraph id="8BE23683DC56FFB8A4E1A1A1FCE9F8F7" blockId="11.[187,1398,1778,1841]" pageId="11" pageNumber="97">
|
||
G ENITALIA (
|
||
<figureCitation id="13662A06DC56FFB8A519A1A1FE6FF8CA" box="[323,418,1778,1805]" captionStart="Fig" captionStartId="9.[189,232,1822,1848]" captionTargetBox="[206,1383,744,1716]" captionTargetId="figure-193@9.[203,1383,744,1716]" captionTargetPageId="9" captionText="Fig. 7.Aedeagus of members of thegilli species group.A–E. Lateral view.F–G. Dorsal view of paramera. Arrows show the apical-dorsal notches of the paramera. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov. CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Holotype of D. jocelynae sp. nov. G. Holotype of D. nonstriatum sp. nov., CMNEN WDS00041746." figureDoi="http://doi.org/10.5281/zenodo.5589711" httpUri="https://zenodo.org/record/5589711/files/figure.png" pageId="11" pageNumber="97">Figs 7B</figureCitation>
|
||
,
|
||
<figureCitation id="13662A06DC56FFB8A5F5A1A1FE19F8CB" box="[431,468,1778,1804]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[233,1362,650,1777]" captionTargetId="figure-181@10.[218,1370,644,1787]" captionTargetPageId="10" captionText="Fig. 8. Endophallus of members of thegilli species group.Arrows show the scales on the sub-medial area of the endophallus. A. Elongate scales. C. Regular scales. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589713" httpUri="https://zenodo.org/record/5589713/files/figure.png" pageId="11" pageNumber="97">8B</figureCitation>
|
||
). Aedeagus as described in the
|
||
<taxonomicName id="4C5D4D00DC56FFB8A71FA1A1FCB8F8CB" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[837,885,1778,1804]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC56FFB8A71FA1A1FCB8F8CB" box="[837,885,1778,1804]" italics="true" pageId="11" pageNumber="97">gilli</emphasis>
|
||
</taxonomicName>
|
||
species group. Medial endophallite slightly sinuate. Sub-medial area of endophallus with scales.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3476508DC56FFBFA4E7A00BFD80FE77" lastPageId="12" lastPageNumber="98" pageId="11" pageNumber="97" type="discussion">
|
||
<paragraph id="8BE23683DC56FFB8A4E7A00BFEE2F8B5" blockId="11.[189,303,1880,1906]" box="[189,303,1880,1906]" pageId="11" pageNumber="97">
|
||
<heading id="D0AA81EFDC56FFB8A4E7A00BFEE2F8B5" bold="true" box="[189,303,1880,1906]" fontSize="11" level="3" pageId="11" pageNumber="97" reason="3">
|
||
<emphasis id="B929EA91DC56FFB8A4E7A00BFEE2F8B5" bold="true" box="[189,303,1880,1906]" pageId="11" pageNumber="97">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE23683DC56FFBFA4E7A0D4FD80FE77" blockId="11.[189,1399,1927,2024]" lastBlockId="12.[188,1399,265,432]" lastPageId="12" lastPageNumber="98" pageId="11" pageNumber="97">
|
||
Only
|
||
<specimenCount id="9D5BFD0ADC56FFB8A55EA0D4FE08F866" box="[260,453,1927,1953]" pageId="11" pageNumber="97" type="generic">three specimens</specimenCount>
|
||
are known for this species, all teneral. The
|
||
<typeStatus id="54E68821DC56FFB8A7AFA0D4FB91F866" box="[1013,1116,1927,1953]" pageId="11" pageNumber="97" type="holotype">holotype</typeStatus>
|
||
does not appear to be teneral externally, however, the aedeagus is poorly sclerotised (
|
||
<figureCitation id="13662A06DC56FFB8A7F3A0F8FBCBF802" box="[937,1030,1963,1989]" captionStart="Fig" captionStartId="9.[189,232,1822,1848]" captionTargetBox="[206,1383,744,1716]" captionTargetId="figure-193@9.[203,1383,744,1716]" captionTargetPageId="9" captionText="Fig. 7.Aedeagus of members of thegilli species group.A–E. Lateral view.F–G. Dorsal view of paramera. Arrows show the apical-dorsal notches of the paramera. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov. CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Holotype of D. jocelynae sp. nov. G. Holotype of D. nonstriatum sp. nov., CMNEN WDS00041746." figureDoi="http://doi.org/10.5281/zenodo.5589711" httpUri="https://zenodo.org/record/5589711/files/figure.png" pageId="11" pageNumber="97">Fig. 7B</figureCitation>
|
||
). The
|
||
<typeStatus id="54E68821DC56FFB8A00BA0F8FB0CF802" box="[1105,1217,1963,1989]" pageId="11" pageNumber="97" type="paratype">paratypes</typeStatus>
|
||
differ from the
|
||
<typeStatus id="54E68821DC56FFB8A4E4A09DFEE8F82F" box="[190,293,1998,2024]" pageId="11" pageNumber="97" type="holotype">holotype</typeStatus>
|
||
by the sexual dimorphism and by having the elytra and the pygidium less sclerotised, with those structures light brown (
|
||
<figureCitation id="13662A06DC51FFBFA644A65AFD6FFEE4" box="[542,674,265,291]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[268,1315,559,1863]" captionTargetId="figure-129@12.[268,1320,553,1873]" captionTargetPageId="12" captionText="Fig. 9. Habitus A, C and ventral view B, D of the only two females known of the gilli species group. A–B. Paratype of Deltochilum gilli González-Alvarado & Vaz-de-Mello, 2021. C–D. Paratype D. jocelynae sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589715" httpUri="https://zenodo.org/record/5589715/files/figure.png" pageId="12" pageNumber="98">Fig. 9C–D</figureCitation>
|
||
). It appears that this species is sympatric with
|
||
<taxonomicName id="4C5D4D00DC51FFBFA080A659FAFDFEE4" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[1242,1328,265,292]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="98" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC51FFBFA080A659FB22FEE4" box="[1242,1263,266,291]" italics="true" pageId="12" pageNumber="98">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC51FFBFA15AA65AFAFDFEE4" box="[1280,1328,265,291]" italics="true" pageId="12" pageNumber="98">gilli</emphasis>
|
||
</taxonomicName>
|
||
since both species were collected a few kilometres apart (see
|
||
<figureCitation id="13662A06DC51FFBFA736A67FFC0BFE81" box="[876,966,300,326]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[336,1251,578,1870]" captionTargetId="figure-135@14.[336,1251,578,1870]" captionTargetPageId="14" captionText="Fig. 10. Distribution map of thegilli species group. Blue circle =Deltochilum gilli González-Alvarado & Vaz-de-Mello, 2021; yellow square = D. jocelynae sp. nov.; red star = D. nonstriatum sp. nov.; green triangle = D. quasistriatum sp. nov.; white eight-point star = D. tenuistriatum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589717" httpUri="https://zenodo.org/record/5589717/files/figure.png" pageId="12" pageNumber="98">Fig. 10</figureCitation>
|
||
, red square).
|
||
<taxonomicName id="4C5D4D00DC51FFBFA02AA67FFAF3FE81" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[1136,1342,300,326]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="98" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC51FFBFA02AA67FFAF3FE81" box="[1136,1342,300,326]" italics="true" pageId="12" pageNumber="98">Deltochilum gilli</emphasis>
|
||
</taxonomicName>
|
||
was collected almost 200 meters higher (altitude) than
|
||
<taxonomicName id="4C5D4D00DC51FFBFA77EA603FC70FEAE" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[804,957,335,362]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="98" phylum="Arthropoda" rank="species" species="jocelynae" status="sp. nov.">
|
||
<emphasis id="B929EA91DC51FFBFA77EA603FCF4FEAE" box="[804,825,336,361]" italics="true" pageId="12" pageNumber="98">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC51FFBFA711A61CFC70FEAE" box="[843,957,335,361]" italics="true" pageId="12" pageNumber="98">jocelynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC51FFBFA792A603FBECFEAD" box="[968,1057,336,362]" pageId="12" pageNumber="98" rank="species">sp. nov</taxonomicNameLabel>
|
||
.. However, both species are easily separated via the elytra;
|
||
<taxonomicName id="4C5D4D00DC51FFBFA645A620FDBEFE4B" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[543,627,370,397]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="98" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC51FFBFA645A620FDF9FE4B" box="[543,564,371,396]" italics="true" pageId="12" pageNumber="98">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC51FFBFA619A621FDBEFE4B" box="[579,627,370,396]" italics="true" pageId="12" pageNumber="98">gilli</emphasis>
|
||
</taxonomicName>
|
||
has conspicuous striae (
|
||
<figureCitation id="13662A06DC51FFBFA7D6A621FC24FE4A" box="[908,1001,370,397]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="12" pageNumber="98">Fig. 4A</figureCitation>
|
||
), whereas these are inconspicuous in
|
||
<taxonomicName id="4C5D4D00DC51FFBFA481A6C4FEBCFE77" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[219,369,406,432]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="98" phylum="Arthropoda" rank="species" species="jocelynae" status="sp. nov.">
|
||
<emphasis id="B929EA91DC51FFBFA481A6C4FF3DFE77" box="[219,240,407,432]" italics="true" pageId="12" pageNumber="98">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC51FFBFA4A5A6C5FEBCFE77" box="[255,369,406,432]" italics="true" pageId="12" pageNumber="98">jocelynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC51FFBFA522A6C5FE1EFE77" box="[376,467,406,432]" pageId="12" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13662A06DC51FFBFA5B8A6C5FDF2FE77" box="[482,575,406,432]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="12" pageNumber="98">Fig. 4B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF22660BDC51FFBFA4E7A0DBFE78F82E" ID-DOI="http://doi.org/10.5281/zenodo.5589715" ID-Zenodo-Dep="5589715" httpUri="https://zenodo.org/record/5589715/files/figure.png" pageId="12" pageNumber="98" startId="12.[189,232,1928,1954]" targetBox="[268,1315,559,1863]" targetPageId="12">
|
||
<paragraph id="8BE23683DC51FFBFA4E7A0DBFE78F82E" blockId="12.[189,1399,1928,2025]" pageId="12" pageNumber="98">
|
||
<emphasis id="B929EA91DC51FFBFA4E7A0DBFEC0F865" bold="true" box="[189,269,1928,1954]" pageId="12" pageNumber="98">Fig. 9.</emphasis>
|
||
Habitus A, C and ventral view B, D of the only two females known of the
|
||
<taxonomicName id="4C5D4D00DC51FFBFA0CDA0DBFB0BF865" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[1175,1222,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="98" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC51FFBFA0CDA0DBFB0BF865" box="[1175,1222,1928,1954]" italics="true" pageId="12" pageNumber="98">gilli</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
<emphasis id="B929EA91DC51FFBFA4E4A0F8FF38F802" bold="true" box="[190,245,1963,1990]" pageId="12" pageNumber="98">A–B</emphasis>
|
||
. Paratype of
|
||
<taxonomicName id="4C5D4D00DC51FFBFA5F4A0F8FB64F802" authority="Gonzalez-Alvarado & Vaz-de-Mello, 2021" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[430,1193,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="98" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC51FFBFA5F4A0F8FD4FF802" box="[430,642,1963,1989]" italics="true" pageId="12" pageNumber="98">Deltochilum gilli</emphasis>
|
||
<bibRefCitation id="EFCC4B72DC51FFBFA6C9A0F8FB64F802" author="Gonzalez-Alvarado A. & Vaz-de-Mello F. Z." box="[659,1193,1963,1989]" pageId="12" pageNumber="98" pagination="0244657" refId="ref7969" refString="Gonzalez-Alvarado A. & Vaz-de-Mello F. Z. 2021. Towards a comprehensive taxonomic revision of the Neotropical dung beetle subgenus Deltochilum (Deltohyboma) Lane, 1946 (Coleoptera: Scarabaeidae: Scarabaeinae): Division into species groups. PLOS ONE 16: e 0244657. https: // doi. org / 10.1371 / journal. pone. 0244657" type="journal article" year="2021">González-Alvarado & Vaz-de-Mello, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B929EA91DC51FFBFA0E4A0F8FB3AF802" bold="true" box="[1214,1271,1963,1990]" pageId="12" pageNumber="98">C–D</emphasis>
|
||
. Paratype
|
||
<taxonomicName id="4C5D4D00DC51FFBFA4E7A09CFE9EF82F" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[189,339,1998,2024]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="98" phylum="Arthropoda" rank="species" species="jocelynae" status="sp. nov.">
|
||
<emphasis id="B929EA91DC51FFBFA4E7A09CFE9EF82F" box="[189,339,1998,2024]" italics="true" pageId="12" pageNumber="98">D. jocelynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC51FFBFA500A09CFE78F82E" box="[346,437,1999,2025]" pageId="12" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3476508DC50FFBEA4E7A65AFC04FE94" pageId="13" pageNumber="99" type="distribution">
|
||
<paragraph id="8BE23683DC50FFBEA4E7A65AFE79FEE5" blockId="13.[189,436,264,291]" box="[189,436,264,291]" pageId="13" pageNumber="99">
|
||
<heading id="D0AA81EFDC50FFBEA4E7A65AFE79FEE5" bold="true" box="[189,436,264,291]" fontSize="11" level="3" pageId="13" pageNumber="99" reason="3">
|
||
<emphasis id="B929EA91DC50FFBEA4E7A65AFE79FEE5" bold="true" box="[189,436,264,291]" pageId="13" pageNumber="99">Known distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE23683DC50FFBEA4E7A66BFC04FE94" blockId="13.[189,969,312,339]" box="[189,969,312,339]" pageId="13" pageNumber="99">
|
||
<collectingCountry id="F34A7613DC50FFBEA4E7A66BFEF0FE95" box="[189,317,312,338]" name="Guyana" pageId="13" pageNumber="99">GUYANA</collectingCountry>
|
||
. District 8, Mount Wokomung (
|
||
<figureCitation id="13662A06DC50FFBEA6EBA66BFCC5FE95" box="[689,776,312,338]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[336,1251,578,1870]" captionTargetId="figure-135@14.[336,1251,578,1870]" captionTargetPageId="14" captionText="Fig. 10. Distribution map of thegilli species group. Blue circle =Deltochilum gilli González-Alvarado & Vaz-de-Mello, 2021; yellow square = D. jocelynae sp. nov.; red star = D. nonstriatum sp. nov.; green triangle = D. quasistriatum sp. nov.; white eight-point star = D. tenuistriatum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589717" httpUri="https://zenodo.org/record/5589717/files/figure.png" pageId="13" pageNumber="99">Fig. 10</figureCitation>
|
||
, yellow square).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |