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<document id="B372DEA120FA33A1AC1E92A0CC7025A2" ID-DOI="10.5281/zenodo.158751" ID-GBIF-Dataset="dbf79923-ab26-419c-a759-a47cc13bebfc" ID-ISSN="1175­5326" ID-Zenodo-Dep="158751" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461035716970" checkinUser="plazi" docAuthor="Webb, Donald W. &amp; Metz, Mark A." docDate="2004" docId="03F48790FF9A12171637FDDDD43CB8A4" docLanguage="en" docName="zt00757.pdf" docOrigin="Zootaxa 757" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Nigranitida irwini Webb, spec. nov." docType="treatment" docVersion="8" lastPageNumber="12" masterDocId="FFCDFFE8FF93121B173FFFE9D72DBA62" masterDocTitle="The South American genus Nigranitida Metz (Diptera: Therevidae: Therevinae) with the description of three new species" masterLastPageNumber="16" masterPageNumber="1" pageNumber="10" updateTime="1698574591596" updateUser="plazi">
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<mods:title id="10E44DBB11918E9BD7DE056126335776">The South American genus Nigranitida Metz (Diptera: Therevidae: Therevinae) with the description of three new species</mods:title>
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<mods:namePart id="796C1DB69709DBC653FC5143D98AD729">Webb, Donald W.</mods:namePart>
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<mods:namePart id="D34E989AE50B0DFF80CE69217D64EAB1">Metz, Mark A.</mods:namePart>
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<mods:date id="C164F332DD58E2B571036FF84866B863">2004</mods:date>
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<treatment id="03F48790FF9A12171637FDDDD43CB8A4" ID-DOI="http://doi.org/10.5281/zenodo.6272755" ID-GBIF-Taxon="119608952" ID-Zenodo-Dep="6272755" LSID="urn:lsid:plazi:treatment:03F48790FF9A12171637FDDDD43CB8A4" httpUri="http://treatment.plazi.org/id/03F48790FF9A12171637FDDDD43CB8A4" lastPageId="12" lastPageNumber="12" pageId="9" pageNumber="10">
<subSubSection id="C347650DFF9A12121637FDDDD59EB82F" box="[264,691,563,590]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8BE23686FF9A12121637FDDDD59EB82F" blockId="9.[264,691,563,590]" box="[264,691,563,590]" pageId="9" pageNumber="10">
<heading id="D0AA81EAFF9A12121637FDDDD59EB82F" bold="true" box="[264,691,563,590]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="B929EA94FF9A12121637FDDDD59EB82F" bold="true" box="[264,691,563,590]" pageId="9" pageNumber="10">
<taxonomicName id="4C5D4D05FF9A12121637FDDDD519B82F" ID-CoL="6RY8Z" authority="Webb" authorityName="Webb" box="[264,564,563,590]" class="Insecta" family="Therevidae" genus="Nigranitida" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="irwini" status="sp. nov.">
<emphasis id="B929EA94FF9A12121637FDDDD6CEB82C" bold="true" box="[264,483,564,590]" italics="true" pageId="9" pageNumber="10">Nigranitida irwini</emphasis>
Webb
</taxonomicName>
<taxonomicNameLabel id="A21A57EFFF9A12121504FDDAD59EB82F" box="[571,691,563,589]" pageId="9" pageNumber="10" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C347650DFF9A12121637FD97D586B8A3" pageId="9" pageNumber="10" type="etymology">
<paragraph id="8BE23686FF9A12121637FD97D586B8A3" blockId="9.[264,1326,638,1905]" pageId="9" pageNumber="10">
<emphasis id="B929EA94FF9A12121637FD97D6B5B8FA" bold="true" box="[264,408,638,664]" pageId="9" pageNumber="10">Etymology:</emphasis>
This species is named after Dr. Michael E. Irwin, who was involved in the collection this species in
<collectingCountry id="F34A7616FF9A12121511FD4ED58AB8A3" box="[558,679,679,705]" name="Argentina" pageId="9" pageNumber="10">Argentina</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C347650DFF9A12121607FD27D208BE4B" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="8BE23686FF9A12121607FD27D208BE4B" blockId="9.[264,1326,638,1905]" pageId="9" pageNumber="10">
<emphasis id="B929EA94FF9A12121607FD27D69AB88A" bold="true" box="[312,439,718,744]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
Like
<taxonomicName id="4C5D4D05FF9A1212153FFD26D5CAB88A" box="[512,743,719,744]" class="Insecta" family="Therevidae" genus="Nigranitida" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="B929EA94FF9A1212153FFD26D5CAB88A" box="[512,743,719,744]" italics="true" pageId="9" pageNumber="10">Nigranitida costata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5D4D05FF9A1212141AFD26D4A6B88A" box="[805,907,719,744]" class="Insecta" family="Therevidae" genus="Nigranitida" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gibba">
<emphasis id="B929EA94FF9A1212141AFD26D4A6B88A" box="[805,907,719,744]" italics="true" pageId="9" pageNumber="10">N. gibba</emphasis>
</taxonomicName>
in lacking setae on the katepisternum. Like
<taxonomicName id="4C5D4D05FF9A121216B0FD1ED6DAB972" box="[399,503,759,784]" class="Insecta" family="Therevidae" genus="Nigranitida" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gibba">
<emphasis id="B929EA94FF9A121216B0FD1ED6DAB972" box="[399,503,759,784]" italics="true" pageId="9" pageNumber="10">N. gibba</emphasis>
</taxonomicName>
in having the halter and maxillary palpus yellow and the posterior margin of the hypoproct ending distal to the apex of the cercus. It differs from
<taxonomicName id="4C5D4D05FF9A12121398FCF6D221B95A" box="[1191,1292,799,824]" class="Insecta" family="Therevidae" genus="Nigranitida" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gibba">
<emphasis id="B929EA94FF9A12121398FCF6D221B95A" box="[1191,1292,799,824]" italics="true" pageId="9" pageNumber="10">N. gibba</emphasis>
</taxonomicName>
in having the male setae on the mesonotum dark brown intermixed with brownish gold setae; the gonostylus lacking a basodorsal lobe and lateral pointed flange; the anterior margin of the male ventral apodeme being deeply emarginate and lacking a pair of short, ventromedial projections; and the male ejaculatory apodeme extending beyond the anterior margin of the dorsal apodeme. The female frons is dark reddish brown, glossy with silver pruinescence ventrally and gray pruinescence along the lateral margin and on the ocellar tubercle.
</paragraph>
</subSubSection>
<subSubSection id="C347650DFF9A12101607FBDFD61BBCB4" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="description">
<paragraph id="8BE23686FF9A12121607FBDFD47BBE33" blockId="9.[264,1326,638,1905]" box="[312,854,1078,1105]" pageId="9" pageNumber="10">
<emphasis id="B929EA94FF9A12121607FBDFD5B3BE32" bold="true" box="[312,670,1078,1104]" pageId="9" pageNumber="10">
Description of
<typeStatus id="54E68824FF9A121216D0FBDFD576BE32" box="[495,603,1078,1104]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
male
</emphasis>
(MEI 106759).
</paragraph>
<paragraph id="8BE23686FF9A12121607FBB6D599BE1B" blockId="9.[264,1326,638,1905]" box="[312,692,1119,1145]" pageId="9" pageNumber="10">Morphological variation (n = 2).</paragraph>
<paragraph id="8BE23686FF9A12121607FB6ED426BEC3" blockId="9.[264,1326,638,1905]" box="[312,779,1159,1185]" pageId="9" pageNumber="10">
Body length
<quantity id="4CA59B63FF9A121216F0FB6ED504BEC3" box="[463,553,1159,1185]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="10" unit="mm" value="6.5">6.5 mm</quantity>
(6.56.7,
<quantity id="4CA59B63FF9A1212159FFB6ED5D2BEC3" box="[672,767,1159,1185]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="9" pageNumber="10" unit="mm" value="6.6">6.6 mm</quantity>
).
</paragraph>
<paragraph id="8BE23686FF9A12121607FB47D32DBC53" blockId="9.[264,1326,638,1905]" pageId="9" pageNumber="10">
<emphasis id="B929EA94FF9A12121607FB47D6ACBEAA" bold="true" box="[312,385,1198,1224]" pageId="9" pageNumber="10">Head.</emphasis>
Length
<quantity id="4CA59B63FF9A121216CEFB46D54EBEAB" box="[497,611,1199,1225]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="9" pageNumber="10" unit="mm" value="0.92">0.92 mm</quantity>
. Frons pruinescence silver gray ventrally becoming dark brown dorsally. Antenna dark brown, pruinescence gray, Ant./HL 0.85 (0.760.85, 0.80); scape length
<quantity id="4CA59B63FF9A1212169BFB16D53FBF7B" box="[420,530,1279,1305]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="10" unit="mm" value="0.2">0.20 mm</quantity>
, length/width 1.8 (1.71.8, 1.7), scape/pedicel length 2.0 (1.72.0, 1.8), scape/flagellum width 0.79 (0.750.79, 0.77), setae dark brown; pedicel length
<quantity id="4CA59B63FF9A121213C6FACED61BBF0B" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="mm" value="0.1">0.10 mm</quantity>
(0.080.10,
<quantity id="4CA59B63FF9A121216F1FAA6D511BF0B" box="[462,572,1359,1385]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="10" unit="mm" value="0.09">0.09 mm</quantity>
), length/width 0.7; first flagellomere oval, tapered apically, flagellum length
<quantity id="4CA59B63FF9A1212168BFA9ED533BFF3" box="[436,542,1399,1425]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="9" pageNumber="10" unit="mm" value="0.48">0.48 mm</quantity>
(0.420.48,
<quantity id="4CA59B63FF9A12121589FA9ED409BFF3" box="[694,804,1399,1425]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="10" unit="mm" value="0.45">0.45 mm</quantity>
), length/width 3.4 (2.63.4, 3.0), flagellum/ scape length 2.4 (2.12.4, 2.3). Parafacial pruinescence silver gray. Maxillary palpus yellow, brown basally; length
<quantity id="4CA59B63FF9A1212157BFA2ED581BF83" box="[580,684,1479,1505]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="9" pageNumber="10" unit="mm" value="0.44">0.44 mm</quantity>
(0.380.44,
<quantity id="4CA59B63FF9A1212147FFA2ED481BF83" box="[832,940,1479,1505]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="9" pageNumber="10" unit="mm" value="0.41">0.41 mm</quantity>
), length/width 5.5 (5.56.3, 5.9); setae yellow. Genal setae white. Occipital setae white ventrally becoming appressed, lanceolate dorsolaterally; macrosetae dark reddish brown, scattered.
</paragraph>
<paragraph id="8BE23686FF9A12111607F9D7D6F5BB3C" blockId="9.[264,1326,638,1905]" lastBlockId="10.[264,1323,284,350]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="B929EA94FF9A12121607F9D7D6B1BC3A" bold="true" box="[312,412,1598,1624]" pageId="9" pageNumber="10">Thorax.</emphasis>
Macrosetae 3 np, 2 sa, 1 pa, 0 dc, 2 sc. Mesonotum dark brown, pruinescence gray; vittae dark brown separated by brownish gray pruinescent stripe; setae dark brown, erect intermixed with brownish gold, appressed, lanceolate setae. Pleuron dark brown, pruinescence white (occasionally glossy on anterodorsal third); setae white on propleuron, anepisternum, laterotergite, metanepisternum, and scutellum, absent on katepisternum.
<emphasis id="B929EA94FF9A12121637F8EED661BD42" box="[264,332,1799,1824]" italics="true" pageId="9" pageNumber="10">Wing.</emphasis>
Length
<quantity id="4CA59B63FF9A12121685F8EED536BD43" box="[442,539,1799,1825]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="10" unit="mm" value="5.5">5.5 mm</quantity>
(5.35.5,
<quantity id="4CA59B63FF9A1212159FF8EED429BD43" box="[672,772,1799,1825]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="9" pageNumber="10" unit="mm" value="5.4">5.4 mm</quantity>
), length/width 2.8 (2.82.9, 2.9); membrane opaque; veins pale brown; pterostigma dark brown. Halter pale yellow (dark brown on MEI 106756).
<emphasis id="B929EA94FF9A12121688F8BED6DBBD12" box="[439,502,1879,1904]" italics="true" pageId="9" pageNumber="10">Legs.</emphasis>
Femora dark reddish brown, pruinescence white; ventral setae white; av 6:2:7 (6:2:57), pv 0:0:3 (0:0:13). Tibiae dark yellowish brown, dark brown apically. Tarsi dark brown.
</paragraph>
<caption id="DF22660EFF9912111637F95BD614BD0B" httpUri="https://zenodo.org/record/158754/files/figure.png" pageId="10" pageNumber="11" targetBox="[301,1271,403,1713]" targetPageId="10">
<paragraph id="8BE23686FF9912111637F95BD614BD0B" blockId="10.[264,1323,1714,1897]" pageId="10" pageNumber="11">
<emphasis id="B929EA94FF9912111637F95BD6FDBCA8" bold="true" box="[264,464,1714,1738]" pageId="10" pageNumber="11">FIGURES 1927.</emphasis>
<taxonomicName id="4C5D4D05FF99121116E8F95BD5B1BCAB" box="[471,668,1714,1737]" class="Insecta" family="Therevidae" genus="Nigranitida" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="irwini" status="sp. nov.">
<emphasis id="B929EA94FF99121116E8F95BD5B1BCAB" box="[471,668,1714,1737]" italics="true" pageId="10" pageNumber="11">Nigranitida irwini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EFFF991211159CF95BD423BCAB" box="[675,782,1714,1737]" pageId="10" pageNumber="11" rank="species">spec. nov.</taxonomicNameLabel>
19. Tergite 8, epandrium, cerci, hypoproct, dorsal view (MEI 106758). 20. Sternite 8, hypandrium, gonocoxite ventral view. 21. Gonocoxite, hypandrium, dorsal view. 22. Gonocoxite, left lateral view. 23. Gonostylus, left lateral view. 24. Aedeagus, dorsal view (inset ejaculatory apodeme cross­section). 25. Aedeagus, ventral view. 26. Aedeagus, left lateral view. 27. Female internal reproductive organs (MEI 106761). Scale = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BE23686FF9812101607FEF2D3B7B9EC" blockId="11.[264,1324,283,1870]" pageId="11" pageNumber="12">
<emphasis id="B929EA94FF9812101607FEF2D694BB57" bold="true" box="[312,441,283,309]" pageId="11" pageNumber="12">Abdomen.</emphasis>
Cuticle black, tergites 13 with posterolateral area dark yellow, pruinescence white across posterior margin of tergite 1, in broad posteromedial area on tergites 2 5, over entire area of tergites 68; dorsal setae white, lanceolate, appressed; lateral setae white, erect.
<taxonomicName id="4C5D4D05FF981210169EFE7DD50DBBCF" box="[417,544,404,429]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="genus">
<emphasis id="B929EA94FF981210169EFE7DD50DBBCF" box="[417,544,404,429]" italics="true" pageId="11" pageNumber="12">Terminalia</emphasis>
</taxonomicName>
. Epandrium (
<figureCitation id="13662A03FF98121015FCFE7DD434BBCC" box="[707,793,404,430]" captionStart="FIGURES 19 27" captionStartId="10.[264,383,1714,1738]" captionTargetBox="[301,1271,403,1713]" captionTargetId="figure@10.[301,1285,402,1729]" captionTargetPageId="10" captionText="FIGURES 19 27. Nigranitida irwini spec. nov. 19. Tergite 8, epandrium, cerci, hypoproct, dorsal view (MEI 106758). 20. Sternite 8, hypandrium, gonocoxite ventral view. 21. Gonocoxite, hypandrium, dorsal view. 22. Gonocoxite, left lateral view. 23. Gonostylus, left lateral view. 24. Aedeagus, dorsal view (inset ejaculatory apodeme cross­section). 25. Aedeagus, ventral view. 26. Aedeagus, left lateral view. 27. Female internal reproductive organs (MEI 106761). Scale = 0.1 mm." httpUri="https://zenodo.org/record/158754/files/figure.png" pageId="11" pageNumber="12">Fig. 19</figureCitation>
) quadrate, anterior margin truncate, posterolateral margin tapering to thick point, setae dark yellow. Hypoproct (
<figureCitation id="13662A03FF9812101370FE55D385BBB4" box="[1103,1192,444,470]" captionStart="FIGURES 19 27" captionStartId="10.[264,383,1714,1738]" captionTargetBox="[301,1271,403,1713]" captionTargetId="figure@10.[301,1285,402,1729]" captionTargetPageId="10" captionText="FIGURES 19 27. Nigranitida irwini spec. nov. 19. Tergite 8, epandrium, cerci, hypoproct, dorsal view (MEI 106758). 20. Sternite 8, hypandrium, gonocoxite ventral view. 21. Gonocoxite, hypandrium, dorsal view. 22. Gonocoxite, left lateral view. 23. Gonostylus, left lateral view. 24. Aedeagus, dorsal view (inset ejaculatory apodeme cross­section). 25. Aedeagus, ventral view. 26. Aedeagus, left lateral view. 27. Female internal reproductive organs (MEI 106761). Scale = 0.1 mm." httpUri="https://zenodo.org/record/158754/files/figure.png" pageId="11" pageNumber="12">Fig. 19</figureCitation>
), posterior margin slightly emarginate, ending distal to cercus. Gonocoxite ventral view (
<figureCitation id="13662A03FF98121013F7FE0DD20EBB9C" box="[1224,1315,484,510]" captionStart="FIGURES 19 27" captionStartId="10.[264,383,1714,1738]" captionTargetBox="[301,1271,403,1713]" captionTargetId="figure@10.[301,1285,402,1729]" captionTargetPageId="10" captionText="FIGURES 19 27. Nigranitida irwini spec. nov. 19. Tergite 8, epandrium, cerci, hypoproct, dorsal view (MEI 106758). 20. Sternite 8, hypandrium, gonocoxite ventral view. 21. Gonocoxite, hypandrium, dorsal view. 22. Gonocoxite, left lateral view. 23. Gonostylus, left lateral view. 24. Aedeagus, dorsal view (inset ejaculatory apodeme cross­section). 25. Aedeagus, ventral view. 26. Aedeagus, left lateral view. 27. Female internal reproductive organs (MEI 106761). Scale = 0.1 mm." httpUri="https://zenodo.org/record/158754/files/figure.png" pageId="11" pageNumber="12">Fig. 20</figureCitation>
) quadrate, lateral margin tapered posteriorly, median margin emarginate then tapering posteriorly to short, thick point, setae dark yellow with setae in mediolateral concavity directed ventrally; dorsal view (
<figureCitation id="13662A03FF9812101543FDB5D5FDB814" box="[636,720,604,630]" captionStart="FIGURES 19 27" captionStartId="10.[264,383,1714,1738]" captionTargetBox="[301,1271,403,1713]" captionTargetId="figure@10.[301,1285,402,1729]" captionTargetPageId="10" captionText="FIGURES 19 27. Nigranitida irwini spec. nov. 19. Tergite 8, epandrium, cerci, hypoproct, dorsal view (MEI 106758). 20. Sternite 8, hypandrium, gonocoxite ventral view. 21. Gonocoxite, hypandrium, dorsal view. 22. Gonocoxite, left lateral view. 23. Gonostylus, left lateral view. 24. Aedeagus, dorsal view (inset ejaculatory apodeme cross­section). 25. Aedeagus, ventral view. 26. Aedeagus, left lateral view. 27. Female internal reproductive organs (MEI 106761). Scale = 0.1 mm." httpUri="https://zenodo.org/record/158754/files/figure.png" pageId="11" pageNumber="12">Fig. 21</figureCitation>
) gonocoxal bridge large, pointed; lateral view (
<figureCitation id="13662A03FF98121013C4FDB5D60AB8FC" captionStart="FIGURES 19 27" captionStartId="10.[264,383,1714,1738]" captionTargetBox="[301,1271,403,1713]" captionTargetId="figure@10.[301,1285,402,1729]" captionTargetPageId="10" captionText="FIGURES 19 27. Nigranitida irwini spec. nov. 19. Tergite 8, epandrium, cerci, hypoproct, dorsal view (MEI 106758). 20. Sternite 8, hypandrium, gonocoxite ventral view. 21. Gonocoxite, hypandrium, dorsal view. 22. Gonocoxite, left lateral view. 23. Gonostylus, left lateral view. 24. Aedeagus, dorsal view (inset ejaculatory apodeme cross­section). 25. Aedeagus, ventral view. 26. Aedeagus, left lateral view. 27. Female internal reproductive organs (MEI 106761). Scale = 0.1 mm." httpUri="https://zenodo.org/record/158754/files/figure.png" pageId="11" pageNumber="12">Fig. 22</figureCitation>
) oval, tapering apically to broad point. Gonostylus (
<figureCitation id="13662A03FF98121014A0FD6DD4DBB8FC" box="[927,1014,644,670]" captionStart="FIGURES 19 27" captionStartId="10.[264,383,1714,1738]" captionTargetBox="[301,1271,403,1713]" captionTargetId="figure@10.[301,1285,402,1729]" captionTargetPageId="10" captionText="FIGURES 19 27. Nigranitida irwini spec. nov. 19. Tergite 8, epandrium, cerci, hypoproct, dorsal view (MEI 106758). 20. Sternite 8, hypandrium, gonocoxite ventral view. 21. Gonocoxite, hypandrium, dorsal view. 22. Gonocoxite, left lateral view. 23. Gonostylus, left lateral view. 24. Aedeagus, dorsal view (inset ejaculatory apodeme cross­section). 25. Aedeagus, ventral view. 26. Aedeagus, left lateral view. 27. Female internal reproductive organs (MEI 106761). Scale = 0.1 mm." httpUri="https://zenodo.org/record/158754/files/figure.png" pageId="11" pageNumber="12">Fig. 23</figureCitation>
) lacking basodorsal lobe; ventral margin rounded, setae dark yellowish brown. Aedeagus with dorsal apodeme (
<figureCitation id="13662A03FF98121013C4FD45D60AB88C" captionStart="FIGURES 19 27" captionStartId="10.[264,383,1714,1738]" captionTargetBox="[301,1271,403,1713]" captionTargetId="figure@10.[301,1285,402,1729]" captionTargetPageId="10" captionText="FIGURES 19 27. Nigranitida irwini spec. nov. 19. Tergite 8, epandrium, cerci, hypoproct, dorsal view (MEI 106758). 20. Sternite 8, hypandrium, gonocoxite ventral view. 21. Gonocoxite, hypandrium, dorsal view. 22. Gonocoxite, left lateral view. 23. Gonostylus, left lateral view. 24. Aedeagus, dorsal view (inset ejaculatory apodeme cross­section). 25. Aedeagus, ventral view. 26. Aedeagus, left lateral view. 27. Female internal reproductive organs (MEI 106761). Scale = 0.1 mm." httpUri="https://zenodo.org/record/158754/files/figure.png" pageId="11" pageNumber="12">Fig. 24</figureCitation>
) quadrate, tapered posteriorly; ventral apodeme (
<figureCitation id="13662A03FF9812101457FD3DD492B88C" box="[872,959,724,750]" captionStart="FIGURES 19 27" captionStartId="10.[264,383,1714,1738]" captionTargetBox="[301,1271,403,1713]" captionTargetId="figure@10.[301,1285,402,1729]" captionTargetPageId="10" captionText="FIGURES 19 27. Nigranitida irwini spec. nov. 19. Tergite 8, epandrium, cerci, hypoproct, dorsal view (MEI 106758). 20. Sternite 8, hypandrium, gonocoxite ventral view. 21. Gonocoxite, hypandrium, dorsal view. 22. Gonocoxite, left lateral view. 23. Gonostylus, left lateral view. 24. Aedeagus, dorsal view (inset ejaculatory apodeme cross­section). 25. Aedeagus, ventral view. 26. Aedeagus, left lateral view. 27. Female internal reproductive organs (MEI 106761). Scale = 0.1 mm." httpUri="https://zenodo.org/record/158754/files/figure.png" pageId="11" pageNumber="12">Fig. 25</figureCitation>
), anterior margin deeply emarginate, tapered laterally to narrow point, ventral surface lacking pair of short, ventromedial projections; distiphallus (
<figureCitation id="13662A03FF981210150FFCCDD5A8B95C" box="[560,645,804,830]" captionStart="FIGURES 19 27" captionStartId="10.[264,383,1714,1738]" captionTargetBox="[301,1271,403,1713]" captionTargetId="figure@10.[301,1285,402,1729]" captionTargetPageId="10" captionText="FIGURES 19 27. Nigranitida irwini spec. nov. 19. Tergite 8, epandrium, cerci, hypoproct, dorsal view (MEI 106758). 20. Sternite 8, hypandrium, gonocoxite ventral view. 21. Gonocoxite, hypandrium, dorsal view. 22. Gonocoxite, left lateral view. 23. Gonostylus, left lateral view. 24. Aedeagus, dorsal view (inset ejaculatory apodeme cross­section). 25. Aedeagus, ventral view. 26. Aedeagus, left lateral view. 27. Female internal reproductive organs (MEI 106761). Scale = 0.1 mm." httpUri="https://zenodo.org/record/158754/files/figure.png" pageId="11" pageNumber="12">Fig. 24</figureCitation>
) tapered posteriorly, lateral view (
<figureCitation id="13662A03FF981210132AFCCDD340B95C" box="[1045,1133,804,830]" captionStart="FIGURES 19 27" captionStartId="10.[264,383,1714,1738]" captionTargetBox="[301,1271,403,1713]" captionTargetId="figure@10.[301,1285,402,1729]" captionTargetPageId="10" captionText="FIGURES 19 27. Nigranitida irwini spec. nov. 19. Tergite 8, epandrium, cerci, hypoproct, dorsal view (MEI 106758). 20. Sternite 8, hypandrium, gonocoxite ventral view. 21. Gonocoxite, hypandrium, dorsal view. 22. Gonocoxite, left lateral view. 23. Gonostylus, left lateral view. 24. Aedeagus, dorsal view (inset ejaculatory apodeme cross­section). 25. Aedeagus, ventral view. 26. Aedeagus, left lateral view. 27. Female internal reproductive organs (MEI 106761). Scale = 0.1 mm." httpUri="https://zenodo.org/record/158754/files/figure.png" pageId="11" pageNumber="12">Fig. 26</figureCitation>
), sinuate; ejaculatory apodeme (
<figureCitation id="13662A03FF98121016F0FCA5D508B904" box="[463,549,844,870]" captionStart="FIGURES 19 27" captionStartId="10.[264,383,1714,1738]" captionTargetBox="[301,1271,403,1713]" captionTargetId="figure@10.[301,1285,402,1729]" captionTargetPageId="10" captionText="FIGURES 19 27. Nigranitida irwini spec. nov. 19. Tergite 8, epandrium, cerci, hypoproct, dorsal view (MEI 106758). 20. Sternite 8, hypandrium, gonocoxite ventral view. 21. Gonocoxite, hypandrium, dorsal view. 22. Gonocoxite, left lateral view. 23. Gonostylus, left lateral view. 24. Aedeagus, dorsal view (inset ejaculatory apodeme cross­section). 25. Aedeagus, ventral view. 26. Aedeagus, left lateral view. 27. Female internal reproductive organs (MEI 106761). Scale = 0.1 mm." httpUri="https://zenodo.org/record/158754/files/figure.png" pageId="11" pageNumber="12">Fig. 24</figureCitation>
) with posterior half bulbous, anterior half quadrate, anterior margin slightly emarginate, extending beyond anterior margin of dorsal apodeme.
</paragraph>
<paragraph id="8BE23686FF9812101607FC75D512B9D4" blockId="11.[264,1324,283,1870]" box="[312,575,924,950]" pageId="11" pageNumber="12">Description of female.</paragraph>
<paragraph id="8BE23686FF9812101607FC2DD5C6B9BC" blockId="11.[264,1324,283,1870]" box="[312,747,964,990]" pageId="11" pageNumber="12">Similar to male except for following.</paragraph>
<paragraph id="8BE23686FF9812101607FC05D588BE64" blockId="11.[264,1324,283,1870]" box="[312,677,1004,1030]" pageId="11" pageNumber="12">Morphological variation (n=4).</paragraph>
<paragraph id="8BE23686FF9812101607FBFDD5BABE4C" blockId="11.[264,1324,283,1870]" box="[312,663,1044,1070]" pageId="11" pageNumber="12">Body length 6.87.2, 7.0 mm.</paragraph>
<paragraph id="8BE23686FF9812101607FBD5D3DEBFDC" blockId="11.[264,1324,283,1870]" pageId="11" pageNumber="12">
<emphasis id="B929EA94FF9812101607FBD5D6ACBE34" bold="true" box="[312,385,1084,1110]" pageId="11" pageNumber="12">Head.</emphasis>
Length 0.861.00,
<quantity id="4CA59B63FF9812101555FBD5D5FBBE34" box="[618,726,1084,1110]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="11" pageNumber="12" unit="mm" value="0.94">0.94 mm</quantity>
. Frons dark reddish brown, glossy with silver pruinescence ventrally and grayish black pruinescence laterally and on ocellar tubercle; setae variable, dark reddish brown, short, in single row dorsally along margin on eye (MEI 106760), dark brown short, scattered on ventral fourth (MEI
<date id="FFE31046FF98121014C5FB5DD378BEAC" box="[1018,1109,1204,1230]" pageId="11" pageNumber="12" value="34-12-11">121134</date>
), or absent (MEI 106761). Antenna varying from dark brown to dark yellowish brown; Ant./HL 0.690.85, 0.77; scape length 0.190.24,
<quantity id="4CA59B63FF98121015BDFAEDD5D8BF7C" box="[642,757,1284,1310]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="12" unit="mm" value="0.22">0.22 mm</quantity>
, length/width 1.62.0,
<quantity id="4CA59B63FF9812101328FAEDD354BF7C" box="[1047,1145,1284,1310]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="12" unit="mm" value="1.9">1.9 mm</quantity>
, scape/pedicel length, 2.43.0, 2.8, scape/flagellum width 0.71.0, 0.8; pedicel length
<quantity id="4CA59B63FF9812101361FAC5D3E1BF24" box="[1118,1228,1324,1350]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="12" unit="mm" value="0.08">0.08 mm</quantity>
, length/ width 0.7; flagellum length
<quantity id="4CA59B63FF9812101562FABDD5E0BF0C" box="[605,717,1364,1390]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="11" pageNumber="12" unit="mm" value="0.42">0.42 mm</quantity>
, length/width 2.53.5, 2.9, flagellum/scape length 1.82.2, 1.9. Maxillary palpus length 0.340.42,
<quantity id="4CA59B63FF9812101453FA95D4F3BFF4" box="[876,990,1404,1430]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="11" pageNumber="12" unit="mm" value="0.39">0.39 mm</quantity>
, length/width 4.87.0, 6.0. Occipital setae white ventrally becoming whitish pink, appressed, lanceolate dorsally.
</paragraph>
<paragraph id="8BE23686FF9812101607FA25D3C2BC6C" blockId="11.[264,1324,283,1870]" pageId="11" pageNumber="12">
<emphasis id="B929EA94FF9812101607FA25D6B1BF84" bold="true" box="[312,412,1484,1510]" pageId="11" pageNumber="12">Thorax.</emphasis>
Macrosetae 3 np, 12 sa, 1 pa, 0 dc, 2 sc.
<emphasis id="B929EA94FF98121014AFFA25D4F9BF87" box="[912,980,1484,1509]" italics="true" pageId="11" pageNumber="12">Wing.</emphasis>
Length 5.205.80,
<quantity id="4CA59B63FF9812101384FA25D20ABF84" box="[1211,1319,1484,1510]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="11" pageNumber="12" unit="mm" value="5.5">5.50 mm</quantity>
, length/width 2.93.3, 3.0.
<emphasis id="B929EA94FF9812101502FA1DD551BC6F" box="[573,636,1524,1549]" italics="true" pageId="11" pageNumber="12">Legs.</emphasis>
Anteroventral macrosetae 35:13:310, pv 0:0:14.
</paragraph>
<paragraph id="8BE23686FF9812101607F9F5D61BBCB4" blockId="11.[264,1324,283,1870]" pageId="11" pageNumber="12">
<emphasis id="B929EA94FF9812101607F9F5D694BC54" bold="true" box="[312,441,1564,1590]" pageId="11" pageNumber="12">Abdomen.</emphasis>
Cuticle dark reddish brown, glossy, posterolateral corners of tergites 27 slightly orange­brown; dorsal setae dark reddish brown, appressed with white, elongate, appressed setae across posterior margin of tergites 14.
<taxonomicName id="4C5D4D05FF98121014ABF985D33CBCE7" box="[916,1041,1644,1669]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="genus">
<emphasis id="B929EA94FF98121014ABF985D33CBCE7" box="[916,1041,1644,1669]" italics="true" pageId="11" pageNumber="12">Terminalia</emphasis>
</taxonomicName>
. Furca (
<figureCitation id="13662A03FF981210134AF985D3E1BCE4" box="[1141,1228,1644,1670]" captionStart="FIGURES 19 27" captionStartId="10.[264,383,1714,1738]" captionTargetBox="[301,1271,403,1713]" captionTargetId="figure@10.[301,1285,402,1729]" captionTargetPageId="10" captionText="FIGURES 19 27. Nigranitida irwini spec. nov. 19. Tergite 8, epandrium, cerci, hypoproct, dorsal view (MEI 106758). 20. Sternite 8, hypandrium, gonocoxite ventral view. 21. Gonocoxite, hypandrium, dorsal view. 22. Gonocoxite, left lateral view. 23. Gonostylus, left lateral view. 24. Aedeagus, dorsal view (inset ejaculatory apodeme cross­section). 25. Aedeagus, ventral view. 26. Aedeagus, left lateral view. 27. Female internal reproductive organs (MEI 106761). Scale = 0.1 mm." httpUri="https://zenodo.org/record/158754/files/figure.png" pageId="11" pageNumber="12">Fig. 27</figureCitation>
), length
<quantity id="4CA59B63FF9812101637F97DD658BCCC" box="[264,373,1684,1710]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="11" pageNumber="12" unit="mm" value="0.38">0.38 mm</quantity>
, width
<quantity id="4CA59B63FF98121016F2F97DD516BCCC" box="[461,571,1684,1710]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="11" pageNumber="12" unit="mm" value="0.28">0.28 mm</quantity>
; spermathecal sac elongate, lobate, spermatheca spherical (
<figureCitation id="13662A03FF98121013C3F97DD604BCB4" captionStart="FIGURES 19 27" captionStartId="10.[264,383,1714,1738]" captionTargetBox="[301,1271,403,1713]" captionTargetId="figure@10.[301,1285,402,1729]" captionTargetPageId="10" captionText="FIGURES 19 27. Nigranitida irwini spec. nov. 19. Tergite 8, epandrium, cerci, hypoproct, dorsal view (MEI 106758). 20. Sternite 8, hypandrium, gonocoxite ventral view. 21. Gonocoxite, hypandrium, dorsal view. 22. Gonocoxite, left lateral view. 23. Gonostylus, left lateral view. 24. Aedeagus, dorsal view (inset ejaculatory apodeme cross­section). 25. Aedeagus, ventral view. 26. Aedeagus, left lateral view. 27. Female internal reproductive organs (MEI 106761). Scale = 0.1 mm." httpUri="https://zenodo.org/record/158754/files/figure.png" pageId="11" pageNumber="12">Fig. 27</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C347650DFF9812171607F90DD43CB8A4" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="biology_ecology">
<paragraph id="8BE23686FF9812101607F90DD480BD44" blockId="11.[264,1324,283,1870]" pageId="11" pageNumber="12">
<emphasis id="B929EA94FF9812101607F90DD6B1BC9C" bold="true" box="[312,412,1764,1790]" pageId="11" pageNumber="12">Biology.</emphasis>
<taxonomicName id="4C5D4D05FF9812101699F90DD552BC9F" box="[422,639,1764,1789]" class="Insecta" family="Therevidae" genus="Nigranitida" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="B929EA94FF9812101699F90DD552BC9F" box="[422,639,1764,1789]" italics="true" pageId="11" pageNumber="12">Nigranitida irwini</emphasis>
</taxonomicName>
has been hand netted on inland dunes and a white sand area, and in Malaise traps from
<date id="FFE31046FF9812101543F8E5D484BD44" box="[636,937,1804,1830]" pageId="11" pageNumber="12">9 September to 22 March</date>
.
</paragraph>
<paragraph id="8BE23686FF9812171607F8DDD43CB8A4" blockId="11.[264,1324,283,1870]" lastBlockId="12.[264,1325,284,710]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="B929EA94FF9812101607F8DDD565BD2C" bold="true" box="[312,584,1844,1870]" pageId="11" pageNumber="12">Specimens Examined.</emphasis>
<emphasis id="B929EA94FF9812101570F8DDD428BD2F" box="[591,773,1844,1869]" italics="true" pageId="11" pageNumber="12">
<typeStatus id="54E68824FF9812101570F8DDD5ABBD2F" box="[591,646,1844,1869]" pageId="11" pageNumber="12">Type</typeStatus>
Specimen.
</emphasis>
The
<typeStatus id="54E68824FF9812101400F8DDD488BD2C" box="[831,933,1844,1870]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="4C5D4D05FF9812101334F8DDD204BD2C" authority="Webb" authorityName="Webb" box="[1035,1321,1844,1870]" class="Insecta" family="Therevidae" genus="Nigranitida" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="B929EA94FF9812101334F8DDD3CDBD2F" box="[1035,1248,1844,1869]" italics="true" pageId="11" pageNumber="12">Nigranitida irwini</emphasis>
Webb
</taxonomicName>
(MEI 106759) was collected in
<collectingCountry id="F34A7616FF9F121715ADFEF5D46CBB54" box="[658,833,284,310]" name="Argentina" pageId="12" pageNumber="13">ARGENTINA</collectingCountry>
, Catamarca Prov.,
<quantity id="4CA59B63FF9F12171316FEF5D355BB54" box="[1065,1144,284,310]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="12" pageNumber="13" unit="km" value="28.0">28 km</quantity>
SE Tinogasta,
<quantity id="4CA59B63FF9F12171637FEADD647BB3C" box="[264,362,324,350]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="13" unit="m" value="1100.0">1,100 m</quantity>
,
<date id="FFE31046FF9F12171649FEADD6D8BB3C" box="[374,501,324,350]" pageId="12" pageNumber="13" value="1997-10-17">17.X.1997</date>
, inland dunes, M.E. Irwin, F.D. Parker, S. Roig,
<geoCoordinate id="EE695041FF9F12171313FEADD382BB3C" box="[1068,1199,324,350]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="5" value="-28.245">28.2450º S</geoCoordinate>
.,
<geoCoordinate id="EE695041FF9F121713FEFEADD608BBE4" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="5" value="-67.4557">67.4557º W</geoCoordinate>
. and is deposited in the CAS (
<typeStatus id="54E68824FF9F121715B4FE85D5EABBE4" box="[651,711,364,390]" pageId="12" pageNumber="13">Type</typeStatus>
No. 18024).
<typeStatus id="54E68824FF9F1217145AFE85D321BBE4" box="[869,1036,364,390]" pageId="12" pageNumber="13" type="paratype">PARATYPES</typeStatus>
:
<emphasis id="B929EA94FF9F12171328FE82D206BBE7" bold="true" box="[1047,1323,363,389]" pageId="12" pageNumber="13">
<collectingCountry id="F34A7616FF9F12171328FE82D3E1BBE7" box="[1047,1228,363,389]" name="Argentina" pageId="12" pageNumber="13">ARGENTINA</collectingCountry>
. CATA­
</emphasis>
<emphasis id="B929EA94FF9F12171637FE7ED64CBBCE" bold="true" box="[264,353,407,428]" pageId="12" pageNumber="13">MARCA</emphasis>
. San Fernando, Rio Belen, [­27.307, ­66.894],
<quantity id="4CA59B63FF9F121714B4FE7DD4CBBBCC" box="[907,998,404,430]" metricMagnitude="3" metricUnit="m" metricValue="1.463" pageId="12" pageNumber="13" unit="m" value="1463.0">1463 m</quantity>
,
<date id="FFE31046FF9F121714CCFE7DD35FBBCC" box="[1011,1138,404,430]" pageId="12" pageNumber="13" value="1997-10-17">17.X.1997</date>
, M. E. Irwin, F. D. Parker, S. Roig,
<specimenCount id="9D5BFD0FFF9F121716CAFE55D570BBB4" box="[501,605,444,470]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
MEI 106761 (CAS).
<emphasis id="B929EA94FF9F1217145BFE52D4CDBBB7" bold="true" box="[868,992,443,469]" pageId="12" pageNumber="13">LA RIOJA</emphasis>
. General Lavalle,
<quantity id="4CA59B63FF9F121713FFFE55D221BBB4" box="[1216,1292,444,470]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="13" unit="km" value="10.0">10 km</quantity>
N Villa Union, [­29.222, ­68.246],
<quantity id="4CA59B63FF9F121715BDFE0DD5F1BB9C" box="[642,732,484,510]" metricMagnitude="3" metricUnit="m" metricValue="1.234" pageId="12" pageNumber="13" unit="m" value="1234.0">1234 m</quantity>
,
<date id="FFE31046FF9F121715D7FE0DD4BFBB9C" box="[744,914,484,510]" pageId="12" pageNumber="13" value="1997-10-13" valueMax="1997-10-15" valueMin="1997-10-13">1315.X.1997</date>
, M. E. Irwin, F. D. Parker, S. Roig,
<specimenCount id="9D5BFD0FFF9F12171637FDE5D675B844" box="[264,344,524,550]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
MEI 106758,
<specimenCount id="9D5BFD0FFF9F12171538FDE5D543B844" box="[519,622,524,550]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
106760 (INHS), Malaise trap.
<emphasis id="B929EA94FF9F121714DDFDE2D34FB847" bold="true" box="[994,1122,523,549]" pageId="12" pageNumber="13">MENDOZA</emphasis>
. Lavalle, Parque Provincial Telteca, [­32.382, ­68.055],
<date id="FFE31046FF9F121715E8FDDDD451B82C" box="[727,892,564,590]" pageId="12" pageNumber="13" value="1999-09-09" valueMax="1999-09-17" valueMin="1999-09-09">917.IX.1999</date>
, M. E. Irwin, F. D. Parker,
<specimenCount id="9D5BFD0FFF9F121713F9FDDDD200B82C" box="[1222,1325,564,590]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
MEI
<date id="FFE31046FF9F12171674FDB5D68FB814" box="[331,418,604,630]" pageId="12" pageNumber="13" value="34-12-11">121134</date>
(MEI), Malaise trap, dunes.
<emphasis id="B929EA94FF9F12171430FDB2D4A6B817" bold="true" box="[783,907,603,629]" pageId="12" pageNumber="13">TUCUMAN</emphasis>
. Casa del Campo, km 992 along Highway 40, [­26.547, ­66.023],
<quantity id="4CA59B63FF9F121715B2FD6DD5C5B8FC" box="[653,744,644,670]" metricMagnitude="3" metricUnit="m" metricValue="1.8039999999999998" pageId="12" pageNumber="13" unit="m" value="1804.0">1804 m</quantity>
,
<date id="FFE31046FF9F121715CAFD6DD459B8FC" box="[757,884,644,670]" pageId="12" pageNumber="13" value="1997-10-18">18.X.1997</date>
, M. E. Irwin, F. D. Parker, S. Roig,
<specimenCount id="9D5BFD0FFF9F12171224FD6DD675B8A4" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
MEI 106757 (MEI), white sand area.
</paragraph>
</subSubSection>
</treatment>
</document>