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<document id="4F14832ED0C2D84611C7A26310C5D38C" ID-CLB-Dataset="291528" ID-DOI="10.11646/zootaxa.5428.2.5" ID-GBIF-Dataset="b0a98796-768e-43e2-be6b-1f79c24b9227" ID-ISSN="1175-5326" ID-Zenodo-Dep="10847635" ID-ZooBank="135D8285-BEC0-467E-BCD5-4DA116C6DFBC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1711009984219" checkinUser="plazi" docAuthor="Janák, Jiří &amp; Baňař, Petr" docDate="2024" docId="03E8307A6653027B9992FAACC5B9FCF3" docLanguage="en" docName="zootaxa.5428.2.5.pdf" docOrigin="Zootaxa 5428 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5428.2.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paulianidia densepunctata Janak &amp; Banar 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="256" masterDocId="FFD14802665202789905FFF3C74BFFCA" masterDocTitle="Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="268" masterPageNumber="253" pageNumber="254" updateTime="1711133149338" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6FC804B7712C30D0F34355210543651F">Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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<mods:namePart id="06F24B49F43DD1E9C76ECB66DC452125">Janák, Jiří</mods:namePart>
<mods:affiliation id="B8FF9445CC632665A19F3448C49A1DA0">CZ- 417 62 Rtyně nad Bílinou, č. 4, Czech Republic</mods:affiliation>
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</mods:role>
<mods:namePart id="A81FA61831242C8C7F95A28DB59E0F00">Baňař, Petr</mods:namePart>
<mods:affiliation id="19B84EBB99300FF113A56B7B7049D9EF">Moravian Museum, Department of Entomology, Hviezdoslavova 29 a, Brno, CZ- 627 00, Czech Republic</mods:affiliation>
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<mods:title id="FFD762040363348F101514FD5099E70E">Zootaxa</mods:title>
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<mods:date id="4594C1B4456D0FBDDBC652B372CC4BDD">2024</mods:date>
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<mods:number id="D51C13ACCD2841DD07B1F47A3A004BD4">2024-03-21</mods:number>
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<mods:classification id="4D39D094CAB9EBDEEC6DF6C42F101E2C">journal article</mods:classification>
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<treatment id="03E8307A6653027B9992FAACC5B9FCF3" ID-DOI="http://doi.org/10.5281/zenodo.10847569" ID-Zenodo-Dep="10847569" LSID="urn:lsid:plazi:treatment:03E8307A6653027B9992FAACC5B9FCF3" httpUri="http://treatment.plazi.org/id/03E8307A6653027B9992FAACC5B9FCF3" lastPageId="3" lastPageNumber="256" pageId="1" pageNumber="254">
<subSubSection id="C35BD2E7665302799992FAACC45BFAB3" box="[151,784,1374,1402]" pageId="1" pageNumber="254" type="nomenclature">
<paragraph id="8BFE816C665302799992FAACC45BFAB3" blockId="1.[151,784,1374,1437]" box="[151,784,1374,1402]" pageId="1" pageNumber="254">
<heading id="D0B63600665302799992FAACC45BFAB3" bold="true" box="[151,784,1374,1402]" fontSize="11" level="1" pageId="1" pageNumber="254" reason="1">
<emphasis id="B9355D7E665302799992FAACC45BFAB3" bold="true" box="[151,784,1374,1402]" pageId="1" pageNumber="254">
<taxonomicName id="4C41FAEF665302799992FAACC5EDFAB3" authority="Janak &amp; Banar" authorityName="Janak &amp; Banar" authorityYear="2024" box="[151,678,1374,1402]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="254" phylum="Arthropoda" rank="species" species="densepunctata" status="sp. nov.">
<emphasis id="B9355D7E665302799992FAACC691FAB0" bold="true" box="[151,474,1375,1402]" italics="true" pageId="1" pageNumber="254">Paulianidia densepunctata</emphasis>
Janák &amp; Baňař
</taxonomicName>
,
<taxonomicNameLabel id="A206E005665302799BB6FAACC45BFAB3" box="[691,784,1375,1401]" pageId="1" pageNumber="254" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35BD2E7665302799992FA70C600FA57" box="[151,331,1411,1437]" pageId="1" pageNumber="254" type="description">
<paragraph id="8BFE816C665302799992FA70C600FA57" blockId="1.[151,784,1374,1437]" box="[151,331,1411,1437]" pageId="1" pageNumber="254">
(
<figureCitation id="137A9DE966530279999AFA70C640FA57" box="[159,267,1411,1437]" captionStart="FIGURES 1–18" captionStartId="2.[151,264,1696,1720]" captionTargetBox="[176,1412,191,1653]" captionTargetId="figure-34@2.[163,1424,181,1672]" captionTargetPageId="2" captionText="FIGURES 1–18. 1–8: Paulianidia densepunctata sp. nov., holotype: 1, habitus; 2, forebody; 3, outer part of left elytron; 4–6, aedeagus (4, ventral; 5, 1/3 lateral; 6, lateral); 7, male sternite VII; 8, male sternite VIII. 9–18: P. hova Lecoq, paratypes: 9–14, 16–17; specimen from Ranomafana: 15, 17; 9, habitus; 10, forebody; 11–15, aedeagus (11, 14–15, ventral; 12, 1/3 lateral; 13, lateral); 16, male sternite VII; 17, posterior part of male sternite VII; 18, male sternite VIII. Scales 1 mm: 1–2, 9–10; 0.5 mm: 4–6, 7–8, 11–18; scale 0.1 mm: 3." figureDoi="http://doi.org/10.5281/zenodo.10847640" httpUri="https://zenodo.org/record/10847640/files/figure.png" pageId="1" pageNumber="254">Figs. 1–8</figureCitation>
,
<figureCitation id="137A9DE9665302799812FA70C609FA57" box="[279,322,1411,1437]" captionStart="FIGURES 116–122" captionStartId="12.[151,264,1780,1804]" captionTargetBox="[180,1408,194,1743]" captionTargetId="figure-34@12.[167,1421,181,1756]" captionText="FIGURES 116–122. 116–120: Paulianidia steineri Lecoq, Andasibe: 116–118, aedeagus (116, ventral; 117, 1/3 lateral; 118, lateral); 119, male sternite VII; 120, male sternite VIII. Scale 0.5 mm: 116–120. 121: Distribution of P. densepunctata sp. nov., (rhombus), P. valohoaka sp. nov. (star), P. tanala sp. nov. and P. vohimana sp. nov. (rectangle). 122: Distribution of Paulianidia species in Madagascar." figureDoi="http://doi.org/10.5281/zenodo.10854991" httpUri="https://zenodo.org/record/10854991/files/figure.png" pageId="1" pageNumber="254">121</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35BD2E7665302799992FA38C237F99B" pageId="1" pageNumber="254" type="materials_examined">
<paragraph id="8BFE816C665302799992FA38C47AFA2C" blockId="1.[151,1437,1483,2013]" box="[151,817,1483,1510]" pageId="1" pageNumber="254">
<materialsCitation id="3B298B31665302799992FA38C47AFA2C" box="[151,817,1483,1510]" country="Madagascar" location="Ambohitantely Special Reserve" pageId="1" pageNumber="254" specimenCount="1" typeStatus="holotype">
<emphasis id="B9355D7E665302799992FA38C67DFA2F" bold="true" box="[151,310,1483,1509]" pageId="1" pageNumber="254">
<typeStatus id="54FA3FCE665302799992FA38C799FA2F" box="[151,210,1483,1509]" pageId="1" pageNumber="254">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F356C1FC665302799839FA38C68EFA2F" box="[316,453,1483,1509]" name="Madagascar" pageId="1" pageNumber="254">Madagascar</collectingCountry>
,
<location id="8E9ED7B76653027998CAFA38C47AFA2C" LSID="urn:lsid:plazi:treatment:03E8307A6653027B9992FAACC5B9FCF3:8E9ED7B76653027998CAFA38C47AFA2C" box="[463,817,1483,1510]" country="Madagascar" name="Ambohitantely Special Reserve" pageId="1" pageNumber="254">Ambohitantely Special Reserve</location>
</materialsCitation>
</paragraph>
<paragraph id="8BFE816C6653027999C2FA1CC237F99B" blockId="1.[151,1437,1483,2013]" pageId="1" pageNumber="254">
<emphasis id="B9355D7E6653027999C2FA1CC635F9C3" bold="true" box="[199,382,1519,1545]" pageId="1" pageNumber="254">Type material.</emphasis>
<materialsCitation id="3B298B3166530279988BFA1CC6E8F99B" collectingDate="2011-11" collectorName="L. S. Rahanitriniaina" country="Madagascar" elevation="1497" latitude="-18.181166" location="MADAGASCAR" longLatPrecision="1" longitude="47.28958" pageId="1" pageNumber="254" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B9355D7E66530279988BFA1CC6B0F9C3" bold="true" box="[398,507,1519,1545]" pageId="1" pageNumber="254">
<typeStatus id="54FA3FCE66530279988BFA1CC6B0F9C3" box="[398,507,1519,1545]" pageId="1" pageNumber="254" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D474AE5665302799B0EFA03C568F9C0" box="[523,547,1520,1546]" count="1" pageId="1" pageNumber="254" type="male">♂</specimenCount>
:
<emphasis id="B9355D7E665302799B3DFA1CC44EF9C3" bold="true" box="[568,773,1519,1545]" pageId="1" pageNumber="254">
<collectingCountry id="F356C1FC665302799B3DFA1CC44EF9C3" box="[568,773,1519,1545]" name="Madagascar" pageId="1" pageNumber="254">MADAGASCAR</collectingCountry>
</emphasis>
: “ABT/
<date id="FFFFA7AC665302799A6FFA1CC493F9C3" box="[874,984,1519,1545]" pageId="1" pageNumber="254" value="2011-11">
<collectingDate id="EFBB5E44665302799A6FFA1CC493F9C3" box="[874,984,1519,1545]" pageId="1" pageNumber="254" value="2011-11">Nov.2011</collectingDate>
</date>
/11 MADAGASCAR, AMBOHITANTELY Spec. Res.;
<quantity id="4CB92C89665302799819F9E0C63FF9E4" box="[284,372,1555,1582]" metricMagnitude="3" metricUnit="m" metricValue="1.4969999999999999" pageId="1" pageNumber="254" unit="m" value="1497.0">
<elevation id="006C665F665302799819F9E0C63FF9E4" box="[284,372,1555,1582]" metricMagnitude="3" metricUnit="m" metricValue="1.4969999999999999" pageId="1" pageNumber="254" unit="m" value="1497.0">1497 m</elevation>
</quantity>
,
<geoCoordinate id="EE75E7AB665302799885F9E0C551F9E4" box="[384,538,1555,1582]" degrees="18" direction="south" minutes="10" orientation="latitude" pageId="1" pageNumber="254" precision="1" seconds="52.2" value="-18.181166">S18°10’52.2”</geoCoordinate>
<geoCoordinate id="EE75E7AB665302799B1FF9E7C5F8F9E4" box="[538,691,1555,1582]" degrees="47" direction="east" minutes="17" orientation="longitude" pageId="1" pageNumber="254" precision="1" seconds="22.5" value="47.28958">E 47°17’22.5</geoCoordinate>
; 18.xi.” // “2011; sifting forest litter, Winkler app. extraction;
<collectorName id="26B4E4BA665302799C68F9E0C60FF99B" pageId="1" pageNumber="254">L.S. Rahanitriniaina</collectorName>
lgt.” // “
</materialsCitation>
<materialsCitation id="3B298B316653027998A6F9C4C233F99B" box="[419,1400,1591,1618]" collectionCode="MMBC" country="Madagascar" determinerName="Janak &amp; Banar" pageId="1" pageNumber="254" specimenCount="1" typeStatus="holotype">
<typeStatus id="54FA3FCE6653027998A6F9C4C507F99B" box="[419,588,1591,1617]" pageId="1" pageNumber="254" type="holotype">HOLOTYPUS</typeStatus>
<taxonomicName id="4C41FAEF665302799B56F9C4C43CF99B" authority="Janák &amp; Baňař, 2024" authorityName="Janák &amp; Baňař" authorityYear="2024" box="[595,887,1591,1617]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="254" phylum="Arthropoda" rank="species" species="densepunctata" status="sp. nov.">Paulianidia densepunctata</taxonomicName>
<taxonomicNameLabel id="A206E005665302799A7BF9CBC49CF998" box="[894,983,1592,1618]" pageId="1" pageNumber="254" rank="species">sp. nov.</taxonomicNameLabel>
,
<determinerName id="7E226DA8665302799AE7F9C4C369F99B" box="[994,1058,1591,1617]" pageId="1" pageNumber="254">Janák</determinerName>
&amp;
<determinerName id="7E226DA8665302799D40F9C4C3C3F99B" box="[1093,1160,1591,1617]" pageId="1" pageNumber="254">Baňař</determinerName>
det. 2023” (
<collectionCode id="ED5019A9665302799C16F9C4C23AF99B" box="[1299,1393,1591,1617]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="1" pageNumber="254">MMBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35BD2E76653027B99C2F9A8C6DAFE67" lastPageId="3" lastPageNumber="256" pageId="1" pageNumber="254" type="description">
<paragraph id="8BFE816C6653027999C2F9A8C5BAF953" blockId="1.[151,1437,1483,2013]" pageId="1" pageNumber="254">
<emphasis id="B9355D7E6653027999C2F9A8C61AF9BF" bold="true" box="[199,337,1627,1653]" pageId="1" pageNumber="254">Description</emphasis>
: Body length
<quantity id="4CB92C896653027998F1F9A8C506F9BC" box="[500,589,1627,1654]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="1" pageNumber="254" unit="mm" value="6.4">6.4 mm</quantity>
, forebody length
<quantity id="4CB92C89665302799A11F9A8C425F9BC" box="[788,878,1627,1654]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="254" unit="mm" value="3.6">3.6 mm</quantity>
. Dark brown, abdomen dark reddish brown, legs, mouthparts and antennomeres reddish brown (
<figureCitation id="137A9DE9665302799B9BF98CC5ABF953" box="[670,736,1663,1689]" captionStart="FIGURES 1–18" captionStartId="2.[151,264,1696,1720]" captionTargetBox="[176,1412,191,1653]" captionTargetId="figure-34@2.[163,1424,181,1672]" captionTargetPageId="2" captionText="FIGURES 1–18. 1–8: Paulianidia densepunctata sp. nov., holotype: 1, habitus; 2, forebody; 3, outer part of left elytron; 4–6, aedeagus (4, ventral; 5, 1/3 lateral; 6, lateral); 7, male sternite VII; 8, male sternite VIII. 9–18: P. hova Lecoq, paratypes: 9–14, 16–17; specimen from Ranomafana: 15, 17; 9, habitus; 10, forebody; 11–15, aedeagus (11, 14–15, ventral; 12, 1/3 lateral; 13, lateral); 16, male sternite VII; 17, posterior part of male sternite VII; 18, male sternite VIII. Scales 1 mm: 1–2, 9–10; 0.5 mm: 4–6, 7–8, 11–18; scale 0.1 mm: 3." figureDoi="http://doi.org/10.5281/zenodo.10847640" httpUri="https://zenodo.org/record/10847640/files/figure.png" pageId="1" pageNumber="254">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BFE816C6653027999C2F950C35FF8E3" blockId="1.[151,1437,1483,2013]" pageId="1" pageNumber="254">
Head (
<figureCitation id="137A9DE9665302799816F950C61CF977" box="[275,343,1699,1725]" captionStart="FIGURES 1–18" captionStartId="2.[151,264,1696,1720]" captionTargetBox="[176,1412,191,1653]" captionTargetId="figure-34@2.[163,1424,181,1672]" captionTargetPageId="2" captionText="FIGURES 1–18. 1–8: Paulianidia densepunctata sp. nov., holotype: 1, habitus; 2, forebody; 3, outer part of left elytron; 4–6, aedeagus (4, ventral; 5, 1/3 lateral; 6, lateral); 7, male sternite VII; 8, male sternite VIII. 9–18: P. hova Lecoq, paratypes: 9–14, 16–17; specimen from Ranomafana: 15, 17; 9, habitus; 10, forebody; 11–15, aedeagus (11, 14–15, ventral; 12, 1/3 lateral; 13, lateral); 16, male sternite VII; 17, posterior part of male sternite VII; 18, male sternite VIII. Scales 1 mm: 1–2, 9–10; 0.5 mm: 4–6, 7–8, 11–18; scale 0.1 mm: 3." figureDoi="http://doi.org/10.5281/zenodo.10847640" httpUri="https://zenodo.org/record/10847640/files/figure.png" pageId="1" pageNumber="254">Fig. 2</figureCitation>
) moderately convex, a quarter longer than wide (W/L = 1.23). Surface with moderately fine and very dense punctures and with very small interstices. Eyes small, temples about 1.4 times as long as eyes (R = 1.42). Temples markedly roundly narrowed posteriorly. Antennae long and slender, fifth antennomere about 1.3 times as long as wide (L/W = 1.32), tenth antennomere slightly transverse (L/W = 0.89).
</paragraph>
<paragraph id="8BFE816C6653027999C2F8C7C7BFF817" blockId="1.[151,1437,1483,2013]" pageId="1" pageNumber="254">
Pronotum (
<figureCitation id="137A9DE9665302799840F8C0C6C2F887" box="[325,393,1843,1869]" captionStart="FIGURES 1–18" captionStartId="2.[151,264,1696,1720]" captionTargetBox="[176,1412,191,1653]" captionTargetId="figure-34@2.[163,1424,181,1672]" captionTargetPageId="2" captionText="FIGURES 1–18. 1–8: Paulianidia densepunctata sp. nov., holotype: 1, habitus; 2, forebody; 3, outer part of left elytron; 4–6, aedeagus (4, ventral; 5, 1/3 lateral; 6, lateral); 7, male sternite VII; 8, male sternite VIII. 9–18: P. hova Lecoq, paratypes: 9–14, 16–17; specimen from Ranomafana: 15, 17; 9, habitus; 10, forebody; 11–15, aedeagus (11, 14–15, ventral; 12, 1/3 lateral; 13, lateral); 16, male sternite VII; 17, posterior part of male sternite VII; 18, male sternite VIII. Scales 1 mm: 1–2, 9–10; 0.5 mm: 4–6, 7–8, 11–18; scale 0.1 mm: 3." figureDoi="http://doi.org/10.5281/zenodo.10847640" httpUri="https://zenodo.org/record/10847640/files/figure.png" pageId="1" pageNumber="254">Fig. 2</figureCitation>
) egg-shaped, moderately convex, with shallow basal impressions, about 1.1 times as long as wide (L/W = 1.13) and slightly narrower than head (PW/HW = 0.94). Sides slightly roundly widened posteriorly. Surface with moderately fine and very dense punctures and with very narrow, hardly visible interstices, similar as on head, punctation less dense along very narrow, in anterior part of pronotum at most one puncture wide unpunctured midline.
</paragraph>
<caption id="DF3ED1E46650027A9992F953C6A4F882" ID-DOI="http://doi.org/10.5281/zenodo.10847640" ID-Zenodo-Dep="10847640" httpUri="https://zenodo.org/record/10847640/files/figure.png" pageId="2" pageNumber="255" startId="2.[151,264,1696,1720]" targetBox="[176,1412,191,1653]" targetPageId="2" targetType="figure">
<paragraph id="8BFE816C6650027A9992F953C6A4F882" blockId="2.[151,1437,1696,1864]" pageId="2" pageNumber="255">
<emphasis id="B9355D7E6650027A9992F953C609F972" bold="true" box="[151,322,1696,1720]" pageId="2" pageNumber="255">FIGURES 1–18</emphasis>
. 1–8:
<taxonomicName id="4C41FAEF6650027A9887F953C5D8F972" authority="Janák &amp; Baňař, 2024" authorityName="Janák &amp; Baňař" authorityYear="2024" box="[386,659,1696,1720]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="255" phylum="Arthropoda" rank="species" species="densepunctata" status="sp. nov.">
<emphasis id="B9355D7E6650027A9887F953C5D8F972" box="[386,659,1696,1720]" italics="true" pageId="2" pageNumber="255">Paulianidia densepunctata</emphasis>
</taxonomicName>
<emphasis id="B9355D7E6650027A9B9FF953C5A0F972" bold="true" box="[666,747,1696,1720]" pageId="2" pageNumber="255">
<taxonomicNameLabel id="A206E0056650027A9B9FF953C5A0F972" box="[666,747,1696,1720]" pageId="2" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype: 1, habitus; 2, forebody; 3, outer part of left elytron; 4–6, aedeagus (4, ventral; 5, 1/3 lateral; 6, lateral); 7, male sternite VII; 8, male sternite VIII. 9–18:
<taxonomicName id="4C41FAEF6650027A9D5DF936C3A0F916" authority="Lecoq" authorityName="Lecoq" authorityYear="2008" box="[1112,1259,1732,1756]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="255" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="B9355D7E6650027A9D5DF936C3E9F916" box="[1112,1186,1732,1756]" italics="true" pageId="2" pageNumber="255">P. hova</emphasis>
Lecoq
</taxonomicName>
, paratypes: 9–14, 16–17; specimen from Ranomafana: 15, 17; 9, habitus; 10, forebody; 11–15, aedeagus (11, 14–15, ventral; 12, 1/3 lateral; 13, lateral); 16, male sternite VII; 17, posterior part of male sternite VII; 18, male sternite VIII. Scales 1 mm: 1–2, 9–10; 0.5 mm: 4–6, 7–8, 11–18; scale 0.1 mm: 3.
</paragraph>
</caption>
<paragraph id="8BFE816C6650027A99C2F867C41CF83C" blockId="2.[151,1436,1940,2038]" pageId="2" pageNumber="255">
Elytra (
<figureCitation id="137A9DE96650027A981FF867C616F864" box="[282,349,1940,1966]" captionStart="FIGURES 1–18" captionStartId="2.[151,264,1696,1720]" captionTargetBox="[176,1412,191,1653]" captionTargetId="figure-34@2.[163,1424,181,1672]" captionTargetPageId="2" captionText="FIGURES 1–18. 1–8: Paulianidia densepunctata sp. nov., holotype: 1, habitus; 2, forebody; 3, outer part of left elytron; 4–6, aedeagus (4, ventral; 5, 1/3 lateral; 6, lateral); 7, male sternite VII; 8, male sternite VIII. 9–18: P. hova Lecoq, paratypes: 9–14, 16–17; specimen from Ranomafana: 15, 17; 9, habitus; 10, forebody; 11–15, aedeagus (11, 14–15, ventral; 12, 1/3 lateral; 13, lateral); 16, male sternite VII; 17, posterior part of male sternite VII; 18, male sternite VIII. Scales 1 mm: 1–2, 9–10; 0.5 mm: 4–6, 7–8, 11–18; scale 0.1 mm: 3." figureDoi="http://doi.org/10.5281/zenodo.10847640" httpUri="https://zenodo.org/record/10847640/files/figure.png" pageId="2" pageNumber="255">Fig. 2</figureCitation>
) elongate with very slightly rounded sides, distinct depression along suture in first quarter behind scutellum, markedly flattened along suture, about 1.1 times longer than wide (R = 1.15). Surface with raised, dense, granular sculpture. Interstices shining, without microsculpture.
</paragraph>
<paragraph id="8BFE816C6651027B99C2FF64C473FF33" blockId="3.[151,1437,151,825]" pageId="3" pageNumber="256">
Abdomen (
<figureCitation id="137A9DE96651027B9842FF64C6C1FF7B" box="[327,394,151,177]" captionStart="FIGURES 1–18" captionStartId="2.[151,264,1696,1720]" captionTargetBox="[176,1412,191,1653]" captionTargetId="figure-34@2.[163,1424,181,1672]" captionTargetPageId="2" captionText="FIGURES 1–18. 1–8: Paulianidia densepunctata sp. nov., holotype: 1, habitus; 2, forebody; 3, outer part of left elytron; 4–6, aedeagus (4, ventral; 5, 1/3 lateral; 6, lateral); 7, male sternite VII; 8, male sternite VIII. 9–18: P. hova Lecoq, paratypes: 9–14, 16–17; specimen from Ranomafana: 15, 17; 9, habitus; 10, forebody; 11–15, aedeagus (11, 14–15, ventral; 12, 1/3 lateral; 13, lateral); 16, male sternite VII; 17, posterior part of male sternite VII; 18, male sternite VIII. Scales 1 mm: 1–2, 9–10; 0.5 mm: 4–6, 7–8, 11–18; scale 0.1 mm: 3." figureDoi="http://doi.org/10.5281/zenodo.10847640" httpUri="https://zenodo.org/record/10847640/files/figure.png" pageId="3" pageNumber="256">Fig. 1</figureCitation>
) slightly widened to segment V, very finely and densely punctured. Interstices on average about 1.5 times larger than diameter of punctures, surface less shining, with distinct microsculpture of transverse fields. Tergite VII with distinct membranous fringe apically.
</paragraph>
<paragraph id="8BFE816C6651027B99C2FEF0C67DFE43" blockId="3.[151,1437,151,825]" pageId="3" pageNumber="256">
Male. Posterior margin of sternite VII with very short almost invisible middle projection (
<figureCitation id="137A9DE96651027B9DBCFEF0C3B6FED7" box="[1209,1277,259,285]" captionStart="FIGURES 1–18" captionStartId="2.[151,264,1696,1720]" captionTargetBox="[176,1412,191,1653]" captionTargetId="figure-34@2.[163,1424,181,1672]" captionTargetPageId="2" captionText="FIGURES 1–18. 1–8: Paulianidia densepunctata sp. nov., holotype: 1, habitus; 2, forebody; 3, outer part of left elytron; 4–6, aedeagus (4, ventral; 5, 1/3 lateral; 6, lateral); 7, male sternite VII; 8, male sternite VIII. 9–18: P. hova Lecoq, paratypes: 9–14, 16–17; specimen from Ranomafana: 15, 17; 9, habitus; 10, forebody; 11–15, aedeagus (11, 14–15, ventral; 12, 1/3 lateral; 13, lateral); 16, male sternite VII; 17, posterior part of male sternite VII; 18, male sternite VIII. Scales 1 mm: 1–2, 9–10; 0.5 mm: 4–6, 7–8, 11–18; scale 0.1 mm: 3." figureDoi="http://doi.org/10.5281/zenodo.10847640" httpUri="https://zenodo.org/record/10847640/files/figure.png" pageId="3" pageNumber="256">Fig. 7</figureCitation>
), sternite VIII with long triangular emargination in posterior third (
<figureCitation id="137A9DE96651027B9BDBFED4C46AFE8B" box="[734,801,295,321]" captionStart="FIGURES 1–18" captionStartId="2.[151,264,1696,1720]" captionTargetBox="[176,1412,191,1653]" captionTargetId="figure-34@2.[163,1424,181,1672]" captionTargetPageId="2" captionText="FIGURES 1–18. 1–8: Paulianidia densepunctata sp. nov., holotype: 1, habitus; 2, forebody; 3, outer part of left elytron; 4–6, aedeagus (4, ventral; 5, 1/3 lateral; 6, lateral); 7, male sternite VII; 8, male sternite VIII. 9–18: P. hova Lecoq, paratypes: 9–14, 16–17; specimen from Ranomafana: 15, 17; 9, habitus; 10, forebody; 11–15, aedeagus (11, 14–15, ventral; 12, 1/3 lateral; 13, lateral); 16, male sternite VII; 17, posterior part of male sternite VII; 18, male sternite VIII. Scales 1 mm: 1–2, 9–10; 0.5 mm: 4–6, 7–8, 11–18; scale 0.1 mm: 3." figureDoi="http://doi.org/10.5281/zenodo.10847640" httpUri="https://zenodo.org/record/10847640/files/figure.png" pageId="3" pageNumber="256">Fig. 8</figureCitation>
). Aedeagus (
<figureCitation id="137A9DE96651027B9AB7FED4C350FE8B" box="[946,1051,295,321]" captionStart="FIGURES 1–18" captionStartId="2.[151,264,1696,1720]" captionTargetBox="[176,1412,191,1653]" captionTargetId="figure-34@2.[163,1424,181,1672]" captionTargetPageId="2" captionText="FIGURES 1–18. 1–8: Paulianidia densepunctata sp. nov., holotype: 1, habitus; 2, forebody; 3, outer part of left elytron; 4–6, aedeagus (4, ventral; 5, 1/3 lateral; 6, lateral); 7, male sternite VII; 8, male sternite VIII. 9–18: P. hova Lecoq, paratypes: 9–14, 16–17; specimen from Ranomafana: 15, 17; 9, habitus; 10, forebody; 11–15, aedeagus (11, 14–15, ventral; 12, 1/3 lateral; 13, lateral); 16, male sternite VII; 17, posterior part of male sternite VII; 18, male sternite VIII. Scales 1 mm: 1–2, 9–10; 0.5 mm: 4–6, 7–8, 11–18; scale 0.1 mm: 3." figureDoi="http://doi.org/10.5281/zenodo.10847640" httpUri="https://zenodo.org/record/10847640/files/figure.png" pageId="3" pageNumber="256">Figs. 4–6</figureCitation>
) moderately long and slender, less than
<quantity id="4CB92C896651027B99D5FEB8C658FEAF" box="[208,275,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="256" unit="mm" value="1.0">1 mm</quantity>
long (AL =
<quantity id="4CB92C896651027B98A0FEB8C544FEAF" box="[421,527,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="3" pageNumber="256" unit="mm" value="0.91">0.91 mm</quantity>
). Median lobe in ventral view with wide middle part and moderately wide subrounded apex.
</paragraph>
<paragraph id="8BFE816C6651027B99C2FE60C6DAFE67" blockId="3.[151,1437,151,825]" box="[199,401,403,429]" pageId="3" pageNumber="256">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C35BD2E76651027B99C2FE45C44AFD63" pageId="3" pageNumber="256" type="diagnosis">
<paragraph id="8BFE816C6651027B99C2FE45C44AFD63" blockId="3.[151,1437,151,825]" pageId="3" pageNumber="256">
<emphasis id="B9355D7E6651027B99C2FE45C684FE1A" bold="true" box="[199,463,438,464]" pageId="3" pageNumber="256">Differential diagnosis.</emphasis>
The new species is arranged in a group of the species with raised, granular punctured elytra together with
<taxonomicName id="4C41FAEF6651027B9876FE28C517FE3F" authority="Lecoq, 2008" authorityName="Lecoq" authorityYear="2008" box="[371,604,475,501]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B9355D7E6651027B9876FE28C681FE3E" box="[371,458,475,500]" italics="true" pageId="3" pageNumber="256">P. nigra</emphasis>
<bibRefCitation id="EFD0FC9D6651027B98CAFE28C517FE3F" author="Lecoq, J. C." box="[463,604,475,501]" pageId="3" pageNumber="256" pagination="579 - 603" refId="ref12275" refString="Lecoq, J. C. (2008) Revision du genre Paulianidia Jarrige, 1978 de Madagascar (Insecta, Coleoptera, Staphylinidae, Paederinae). Zoosystema, 30 (3), 579 - 603." type="journal article" year="2008">Lecoq, 2008</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C41FAEF6651027B9B60FE28C43BFE3F" authority="Lecoq, 2008" authorityName="Lecoq" authorityYear="2008" box="[613,880,475,501]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="marojejy">
<emphasis id="B9355D7E6651027B9B60FE28C5AAFE3E" box="[613,737,475,500]" italics="true" pageId="3" pageNumber="256">P. marojejy</emphasis>
<bibRefCitation id="EFD0FC9D6651027B9BE3FE28C43BFE3F" author="Lecoq, J. C." box="[742,880,475,501]" pageId="3" pageNumber="256" pagination="579 - 603" refId="ref12275" refString="Lecoq, J. C. (2008) Revision du genre Paulianidia Jarrige, 1978 de Madagascar (Insecta, Coleoptera, Staphylinidae, Paederinae). Zoosystema, 30 (3), 579 - 603." type="journal article" year="2008">Lecoq, 2008</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C41FAEF6651027B9AA7FE28C393FE3F" authority="Jarrige, 1978" authorityName="Jarrige" authorityYear="1978" box="[930,1240,475,501]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="densipennis">
<emphasis id="B9355D7E6651027B9AA7FE28C30BFE3F" box="[930,1088,475,501]" italics="true" pageId="3" pageNumber="256">P. densipennis</emphasis>
<bibRefCitation id="EFD0FC9D6651027B9D40FE28C393FE3F" author="Jarrige, J." box="[1093,1240,475,501]" pageId="3" pageNumber="256" pagination="245 - 296" refId="ref12166" refString="Jarrige, J. (1978) Contribution a l'etude des Coleoptera Brachelytra Staphylinidae du massif de l'Andringitra (Madagascar centre). Bulletin du Museum national d'histoire naturelle. Ser. 3. Zoologie, 353, 245 - 296." type="journal article" year="1978">Jarrige, 1978</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C41FAEF6651027B9DE4FE28C675FDD3" authority="Janak &amp; Banar" authorityName="Janak &amp; Banar" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="densepunctata" status="sp. nov.">
<emphasis id="B9355D7E6651027B9DE4FE28C2D7FE3F" box="[1249,1436,475,501]" italics="true" pageId="3" pageNumber="256">P. densepunctata</emphasis>
Janák &amp; Baňař
</taxonomicName>
,
<emphasis id="B9355D7E6651027B984DFE0CC6EAFDD3" bold="true" box="[328,417,511,537]" pageId="3" pageNumber="256">
<taxonomicNameLabel id="A206E0056651027B984DFE0CC6EAFDD3" box="[328,417,511,537]" pageId="3" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C41FAEF6651027B9B34FE0CC5E5FDD2" authorityName="Lecoq" authorityYear="2008" box="[561,686,511,536]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="marojejy">
<emphasis id="B9355D7E6651027B9B34FE0CC5E5FDD2" box="[561,686,511,536]" italics="true" pageId="3" pageNumber="256">P. marojejy</emphasis>
</taxonomicName>
by larger eyes and rounded temples, from
<taxonomicName id="4C41FAEF6651027B9D81FE0CC390FDD2" authorityName="Lecoq" authorityYear="2008" box="[1156,1243,511,536]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B9355D7E6651027B9D81FE0CC390FDD2" box="[1156,1243,511,536]" italics="true" pageId="3" pageNumber="256">P. nigra</emphasis>
</taxonomicName>
by much densely and coarsely punctate and less anteriorly narrowed pronotum, from
<taxonomicName id="4C41FAEF6651027B9A8AFDD0C364FDF7" authorityName="Jarrige" authorityYear="1978" box="[911,1071,547,573]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="densipennis">
<emphasis id="B9355D7E6651027B9A8AFDD0C364FDF7" box="[911,1071,547,573]" italics="true" pageId="3" pageNumber="256">P. densipennis</emphasis>
</taxonomicName>
(only in female hitherto known) by larger body, larger eyes and head slightly wider than pronotum. The aedeagus of the new species is slender than that of
<taxonomicName id="4C41FAEF6651027B99EEFD98C620FD4E" authorityName="Lecoq" authorityYear="2008" box="[235,363,619,644]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="marojejy">
<emphasis id="B9355D7E6651027B99EEFD98C620FD4E" box="[235,363,619,644]" italics="true" pageId="3" pageNumber="256">P. marojejy</emphasis>
</taxonomicName>
(cf.
<figureCitation id="137A9DE96651027B98A6FD98C6BEFD4F" box="[419,501,619,645]" captionStart="FIGURES 68–91" captionStartId="10.[151,264,1631,1655]" captionTargetBox="[172,1416,194,1594]" captionTargetId="figure-34@10.[158,1429,181,1607]" captionTargetPageId="10" captionText="FIGURES 68–91. 68–70: Paulianidia ikoka Lecoq, holotype: 68, aedeagus, ventral, transmitted light; 69, male sternite VII; 70, male sternite VIII. 71–72: P. itremo Lecoq, holotype: aedeagus ventral (71 ex Lecoq 2008: 3D). 73–77: P. ivohibensis Lecoq, Ranomafana: 73–75, aedeagus (73, ventral; 74, 1/3 lateral; 75, lateral); 76, male sternite VII; 77, male sternite VIII. 78–82: P. janaki Lecoq, paratype: 78–80, aedeagus (78, ventral; 79, 1/3 lateral; 80, lateral); 81, male sternite VII; 82, male sternite VIII. 83–84: P. marojejy Lecoq, holotype: 83, habitus; 84, aedeagus, ventral. 85: P. microptera Lecoq, holotype, aedeagus ventral. 86: P. minima Lecoq, holotype, habitus. 87–91: P. nigra Lecoq, holotype: 87–89, aedeagus (87, ventral; 88, 1/3 lateral; 89, lateral); 90, male sternite VII; 91, male sternite VIII. Scales 1 mm: 83, 86; 0.5 mm: 68–70, 71–72, 73–77, 78–80, 81–82, 84–85, 87–89, 90–91." figureDoi="http://doi.org/10.5281/zenodo.10847650" httpUri="https://zenodo.org/record/10847650/files/figure.png" pageId="3" pageNumber="256">Fig. 84</figureCitation>
) and not divided in two projections apically, slightly similar to
<taxonomicName id="4C41FAEF6651027B9DD6FD98C266FD4E" authorityName="Lecoq" authorityYear="2008" box="[1235,1325,619,644]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B9355D7E6651027B9DD6FD98C266FD4E" box="[1235,1325,619,644]" italics="true" pageId="3" pageNumber="256">P. nigra</emphasis>
</taxonomicName>
(cf.
<figureCitation id="137A9DE96651027B9C60FD98C7AAFD63" captionStart="FIGURES 68–91" captionStartId="10.[151,264,1631,1655]" captionTargetBox="[172,1416,194,1594]" captionTargetId="figure-34@10.[158,1429,181,1607]" captionTargetPageId="10" captionText="FIGURES 68–91. 68–70: Paulianidia ikoka Lecoq, holotype: 68, aedeagus, ventral, transmitted light; 69, male sternite VII; 70, male sternite VIII. 71–72: P. itremo Lecoq, holotype: aedeagus ventral (71 ex Lecoq 2008: 3D). 73–77: P. ivohibensis Lecoq, Ranomafana: 73–75, aedeagus (73, ventral; 74, 1/3 lateral; 75, lateral); 76, male sternite VII; 77, male sternite VIII. 78–82: P. janaki Lecoq, paratype: 78–80, aedeagus (78, ventral; 79, 1/3 lateral; 80, lateral); 81, male sternite VII; 82, male sternite VIII. 83–84: P. marojejy Lecoq, holotype: 83, habitus; 84, aedeagus, ventral. 85: P. microptera Lecoq, holotype, aedeagus ventral. 86: P. minima Lecoq, holotype, habitus. 87–91: P. nigra Lecoq, holotype: 87–89, aedeagus (87, ventral; 88, 1/3 lateral; 89, lateral); 90, male sternite VII; 91, male sternite VIII. Scales 1 mm: 83, 86; 0.5 mm: 68–70, 71–72, 73–77, 78–80, 81–82, 84–85, 87–89, 90–91." figureDoi="http://doi.org/10.5281/zenodo.10847650" httpUri="https://zenodo.org/record/10847650/files/figure.png" pageId="3" pageNumber="256">Figs. 87–91</figureCitation>
), but much more slender and in details different.
</paragraph>
</subSubSection>
<subSubSection id="C35BD2E76651027B99C2FD40C439FD07" box="[199,882,691,717]" pageId="3" pageNumber="256" type="etymology">
<paragraph id="8BFE816C6651027B99C2FD40C439FD07" blockId="3.[151,1437,151,825]" box="[199,882,691,717]" pageId="3" pageNumber="256">
<emphasis id="B9355D7E6651027B99C2FD40C605FD07" bold="true" box="[199,334,691,717]" pageId="3" pageNumber="256">Etymology.</emphasis>
The name refers to the densely punctured elytra.
</paragraph>
</subSubSection>
<subSubSection id="C35BD2E76651027B99C2FD25C5B9FCF3" pageId="3" pageNumber="256" type="distribution">
<paragraph id="8BFE816C6651027B99C2FD25C31DFCDF" blockId="3.[151,1437,151,825]" pageId="3" pageNumber="256">
<emphasis id="B9355D7E6651027B99C2FD25C62AFD3A" bold="true" box="[199,353,726,752]" pageId="3" pageNumber="256">Distribution.</emphasis>
<taxonomicName id="4C41FAEF6651027B986AFD24C427FD3B" authority="Janak &amp; Banar" authorityName="Janak &amp; Banar" authorityYear="2024" box="[367,876,727,753]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="densepunctata" status="sp. nov.">
<emphasis id="B9355D7E6651027B986AFD24C5EFFD3B" box="[367,676,727,753]" italics="true" pageId="3" pageNumber="256">Paulianidia densepunctata</emphasis>
Janák &amp; Baňař
</taxonomicName>
,
<emphasis id="B9355D7E6651027B9A7BFD24C4ABFD3B" bold="true" box="[894,992,727,753]" pageId="3" pageNumber="256">
<taxonomicNameLabel id="A206E0056651027B9A7BFD24C4ABFD3B" box="[894,992,727,753]" pageId="3" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is currently recorded only from the Ambohitantely Special Reserve in the northern central part of
<collectingCountry id="F356C1FC6651027B9A54FD08C493FCDF" box="[849,984,763,789]" name="Madagascar" pageId="3" pageNumber="256">Madagascar</collectingCountry>
(
<figureCitation id="137A9DE96651027B9AE2FD08C302FCDF" box="[999,1097,763,789]" captionStart="FIGURES 116–122" captionStartId="12.[151,264,1780,1804]" captionTargetBox="[180,1408,194,1743]" captionTargetId="figure-34@12.[167,1421,181,1756]" captionText="FIGURES 116–122. 116–120: Paulianidia steineri Lecoq, Andasibe: 116–118, aedeagus (116, ventral; 117, 1/3 lateral; 118, lateral); 119, male sternite VII; 120, male sternite VIII. Scale 0.5 mm: 116–120. 121: Distribution of P. densepunctata sp. nov., (rhombus), P. valohoaka sp. nov. (star), P. tanala sp. nov. and P. vohimana sp. nov. (rectangle). 122: Distribution of Paulianidia species in Madagascar." figureDoi="http://doi.org/10.5281/zenodo.10854991" httpUri="https://zenodo.org/record/10854991/files/figure.png" pageId="3" pageNumber="256">Fig. 121</figureCitation>
).
</paragraph>
<paragraph id="8BFE816C6651027B99C2FCEDC5B9FCF3" blockId="3.[151,1437,151,825]" box="[199,754,798,825]" pageId="3" pageNumber="256">
<emphasis id="B9355D7E6651027B99C2FCEDC600FCF2" bold="true" box="[199,331,798,824]" pageId="3" pageNumber="256">Bionomics.</emphasis>
The
<typeStatus id="54FA3FCE6651027B9881FCECC6FFFCF3" box="[388,436,799,825]" pageId="3" pageNumber="256">type</typeStatus>
was sifted from forest litter.
</paragraph>
</subSubSection>
</treatment>
</document>
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