385 lines
43 KiB
XML
385 lines
43 KiB
XML
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<mods:titleInfo id="A3B36A77270AA09DD3288ACEE4E315BB">
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<mods:title id="E383D4944AB138E41DEF03DC0A2AE2EC">The identity of Chelodina oblonga Gray 1841 (Testudines: Chelidae) reassessed</mods:title>
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<mods:name id="CA224090CB4AB24F2A7C3B789B70DF99" type="personal">
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<mods:namePart id="D313F3E73695E435982349C4B59D3E78">Shea, Glenn</mods:namePart>
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<mods:affiliation id="B2D5ABADC6B70650CB11C80872707878">Sydney School of Veterinary Science, B 01, University of Sydney, NSW 2006 Australia & Australian Museum Research Institute, Australian Museum, 1 William St, Sydney, NSW 2010, Australia</mods:affiliation>
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<mods:namePart id="DE4566866BF486ECA8DD3A49CACF9206">Thomson, Scott</mods:namePart>
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<mods:affiliation id="17BA67F44F719366AA5F3210C44D099C">Chelonian Research Institute, 402 South Central Avenue, Oviedo, 32765, Florida, USA</mods:affiliation>
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<mods:namePart id="797DE5FD88CE9BA56F79065B4A088806">Georges, Arthur</mods:namePart>
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<mods:affiliation id="A881B2B511DCD32E52032C97962C134C">Institute for Applied Ecology, University of Canberra, ACT 2601 Australia</mods:affiliation>
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<mods:title id="B78B944AAF54A15AE1A7BFB059825767">Zootaxa</mods:title>
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<mods:date id="D4FACFA96653BF692488564133B5AADF">2020</mods:date>
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<mods:number id="744B08A9423E2DE4E2E48F51E6FA3ED3">2020-05-20</mods:number>
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<mods:number id="827C35A92B91C2A6A18D49CEDFAB612F">4779</mods:number>
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<mods:identifier id="5EF440096CE53C897765707791584896" type="DOI">10.11646/zootaxa.4779.3.9</mods:identifier>
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<treatment id="03E82A6F917D2851FF43FC06044522F7" ID-DOI="http://doi.org/10.5281/zenodo.4328850" ID-GBIF-Taxon="166097415" ID-Zenodo-Dep="4328850" LSID="urn:lsid:plazi:treatment:03E82A6F917D2851FF43FC06044522F7" httpUri="http://treatment.plazi.org/id/03E82A6F917D2851FF43FC06044522F7" lastPageId="14" lastPageNumber="433" pageId="13" pageNumber="432">
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<subSection id="E2CE8292917D2852FF43FC06052524B8" box="[151,1132,910,937]" pageId="13" pageNumber="432" type="multiple">
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<subSubSection id="C35BC8F2917D2852FF43FC06052524B8" box="[151,1132,910,937]" pageId="13" pageNumber="432" type="nomenclature">
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<paragraph id="8BFE9B79917D2852FF43FC06052524B8" blockId="13.[151,1132,910,937]" box="[151,1132,910,937]" pageId="13" pageNumber="432">
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<heading id="D0B62C15917D2852FF43FC06052524B8" bold="true" box="[151,1132,910,937]" fontSize="11" level="2" pageId="13" pageNumber="432">
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<emphasis id="B935476B917D2852FF43FC06052524B8" bold="true" box="[151,1132,910,937]" pageId="13" pageNumber="432">
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3.4. Synonymy of
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<taxonomicName id="4C41E0FA917D2852FEA2FC06031424B9" authorityName="J. Gray" authorityYear="1841" box="[374,605,910,937]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="432" phylum="Chordata" rank="species" species="oblonga">
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<emphasis id="B935476B917D2852FEA2FC06031424B9" bold="true" box="[374,605,910,937]" italics="true" pageId="13" pageNumber="432">Chelodina oblonga</emphasis>
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</taxonomicName>
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, and redescriptions of the
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<typeStatus id="54FA25DB917D2852FC7CFC07029724B9" box="[936,990,911,937]" pageId="13" pageNumber="432">type</typeStatus>
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specimens.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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</subSection>
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<subSection id="E2CE8292917D2852FF43FC5E03EC2257" pageId="13" pageNumber="432" type="multiple">
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<subSubSection id="C35BC8F2917D2852FF43FC5E03EC2257" pageId="13" pageNumber="432" type="reference_group">
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<paragraph id="8BFE9B79917D2852FF43FC5E002C24E0" blockId="13.[151,357,982,1008]" box="[151,357,982,1008]" pageId="13" pageNumber="432">
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<emphasis id="B935476B917D2852FF43FC5E002C24E0" bold="true" box="[151,357,982,1008]" pageId="13" pageNumber="432">3.4.1. Synonymy</emphasis>
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</paragraph>
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<paragraph id="8BFE9B79917D2852FF43FB9203192323" blockId="13.[151,890,1050,1351]" box="[151,592,1050,1075]" pageId="13" pageNumber="432">
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<taxonomicName id="4C41E0FA917D2852FF43FB9203512323" authority="J. Gray 1841: 434" authorityName="J. Gray" authorityPageNumber="434" authorityYear="1841" box="[151,536,1050,1075]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="432" phylum="Chordata" rank="species" species="oblonga">
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<emphasis id="B935476B917D2852FF43FB9200132322" box="[151,346,1050,1074]" italics="true" pageId="13" pageNumber="432">Chelodina oblonga</emphasis>
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J.
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<bibRefCitation id="EFD0E688917D2852FEA3FB9303512323" author="Gray, J. E." box="[375,536,1051,1075]" pageId="13" pageNumber="432" pagination="422 - 449" refId="ref12110" refString="Gray, J. E. (1841) A Catalogue of the Species of Reptiles and Amphibia hitherto described as inhabiting Australia, with a description of some New Species from Western Australia, and some remarks on their geographical distribution. In: Grey, G. Journals of Two Expeditions of Discovery in North-west and Western Australia, during the years 1837, 38, and 39, Under the Authority of Her Majesty's Government. Vol. 2. T. and W. Boone, London, 28 pp., 8 pls. [pp. 422 - 449, pls. 1 - 8]" type="book chapter" year="1841">Gray 1841: 434</bibRefCitation>
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</taxonomicName>
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, 446.
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</paragraph>
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<paragraph id="8BFE9B79917D2852FF43FBB1021A2341" blockId="13.[151,890,1050,1351]" box="[151,851,1080,1105]" pageId="13" pageNumber="432">
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<taxonomicName id="4C41E0FA917D2852FF43FBB100B22341" authority="Gray 1856 b: 267" authorityName="Gray" authorityPageNumber="267" authorityYear="1856" box="[151,507,1080,1105]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="432" phylum="Chordata" rank="species" species="colliei">
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<emphasis id="B935476B917D2852FF43FBB1000F2341" box="[151,326,1081,1105]" italics="true" pageId="13" pageNumber="432">Chelodina colliei</emphasis>
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<bibRefCitation id="EFD0E688917D2852FE99FBB100B22341" author="Gray, J. E." box="[333,507,1080,1105]" pageId="13" pageNumber="432" pagination="197 - 202" refId="ref12446" refString="Gray, J. E. (1856 b) On some new species of freshwater tortoises from North America, Ceylon and Australia, in the collection of the British Museum. Proceedings of the Zoological Society of London, 23 (296 - 297), 197 - 202. [" 1855 "] *" type="journal article" year="1856">Gray 1856b: 267</bibRefCitation>
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</taxonomicName>
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. Synonymy by
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<bibRefCitation id="EFD0E688917D2852FD4DFBB002192341" author="Boulenger, G. A." box="[665,848,1080,1105]" pageId="13" pageNumber="432" refId="ref11069" refString="Boulenger, G. A. (1889) Catalogue of the Chelonians, Rhynchocephalians, and Crocodiles in the British Museum (Natural History). Trustees of the British Museum, London, x + 311 pp., pls. I-VI. https: // doi. org / 10.5962 / bhl. title. 13541" type="book" year="1889">Boulenger (1889)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BFE9B79917D2852FF43FBD003942360" blockId="13.[151,890,1050,1351]" box="[151,733,1111,1136]" pageId="13" pageNumber="432">
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<emphasis id="B935476B917D2852FF43FBD000FD2360" box="[151,436,1112,1136]" italics="true" pageId="13" pageNumber="432">
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<taxonomicName id="4C41E0FA917D2852FF43FBD000DE2360" authorityName="Wells & Wellington" authorityYear="1985" box="[151,407,1112,1136]" class="Testudines" family="Chelidae" genus="Macrochelodina" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurodira" pageId="13" pageNumber="432" phylum="Chordata" rank="species" species="oblonga">Macrochelodina oblonga</taxonomicName>
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—
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</emphasis>
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<bibRefCitation id="EFD0E688917D2852FE6EFBDF03902360" author="Wells, R. W. & Wellington, C. R." box="[442,729,1111,1136]" pageId="13" pageNumber="432" pagination="1 - 61" refId="ref15188" refString="Wells, R. W. & Wellington, C. R. (1985) A classification of the Amphibia and Reptilia of Australia. Australian Journal of Herpetology, Supplementary Series 1, 1 - 61." type="book chapter" year="1985">Wells & Wellington 1985: 9</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BFE9B79917D2852FF43FBFE0245239F" blockId="13.[151,890,1050,1351]" box="[151,780,1142,1167]" pageId="13" pageNumber="432">
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<emphasis id="B935476B917D2852FF43FBFE00E9239E" box="[151,416,1142,1166]" italics="true" pageId="13" pageNumber="432">
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<taxonomicName id="4C41E0FA917D2852FF43FBFE00CB239E" authorityName="Mccord & Josephouni" authorityYear="2007" box="[151,386,1142,1166]" class="Reptilia" family="Chelidae" genus="Macrodiremys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="432" phylum="Chordata" rank="species" species="oblonga">Macrodiremys oblonga</taxonomicName>
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—
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</emphasis>
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<bibRefCitation id="EFD0E688917D2852FE72FBFE0241239F" author="McCord, W. P. & Joseph-Ouni, M." box="[422,776,1142,1167]" pageId="13" pageNumber="432" pagination="56 - 64" refId="ref13631" refString="McCord, W. P. & Joseph-Ouni, M. (2007 a) A new genus of Australian longneck turtle (Testudines: Chelidae) and a new species of Macrochelodina from the Kimberley region of Western Australia (Australia). Reptilia, 55, 56 - 64." type="journal article" year="2007">McCord & Joseph-Ouni 2007a: 57</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BFE9B79917D2852FF43FB1D020623BD" blockId="13.[151,890,1050,1351]" box="[151,847,1172,1197]" pageId="13" pageNumber="432">
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<taxonomicName id="4C41E0FA917D2852FF43FB1D00A623BD" authorityName="Gray" authorityYear="1856" box="[151,495,1173,1197]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="432" phylum="Chordata" rank="species" species="colliei" subGenus="Macrodiremys">
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<emphasis id="B935476B917D2852FF43FB1D004823BD" box="[151,257,1173,1197]" italics="true" pageId="13" pageNumber="432">Chelodina</emphasis>
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(
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<emphasis id="B935476B917D2852FEC4FB1D00EB23BD" box="[272,418,1173,1197]" italics="true" pageId="13" pageNumber="432">Macrodiremys</emphasis>
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||
)
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<emphasis id="B935476B917D2852FE64FB1D00A623BD" box="[432,495,1173,1197]" italics="true" pageId="13" pageNumber="432">colliei</emphasis>
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</taxonomicName>
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—
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<bibRefCitation id="EFD0E688917D2852FDC1FB1D020223BD" author="Georges, A. & Thomson, S." box="[533,843,1172,1197]" pageId="13" pageNumber="432" pagination="1 - 37" refId="ref11886" refString="Georges, A. & Thomson, S. (2010) Diversity of Australasian freshwater turtles, with an annotated synonymy and keys to species. Zootaxa, 2496 (1), 1 - 37. https: // doi. org / 10.11646 / zootaxa. 2496.1.1" type="journal article" year="2010">Georges & Thomson 2010: 19</bibRefCitation>
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||
.
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</paragraph>
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||
<paragraph id="8BFE9B79917D2852FF43FB3C023323DC" blockId="13.[151,890,1050,1351]" box="[151,890,1203,1228]" pageId="13" pageNumber="432">
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||
<taxonomicName id="4C41E0FA917D2852FF43FB3C00A623DC" authorityName="Gray" authorityYear="1856" box="[151,495,1204,1228]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="432" phylum="Chordata" rank="species" species="colliei" subGenus="Macrodiremys">
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||
<emphasis id="B935476B917D2852FF43FB3C004823DC" box="[151,257,1204,1228]" italics="true" pageId="13" pageNumber="432">Chelodina</emphasis>
|
||
(
|
||
<emphasis id="B935476B917D2852FEC4FB3C00EB23DC" box="[272,418,1204,1228]" italics="true" pageId="13" pageNumber="432">Macrodiremys</emphasis>
|
||
)
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<emphasis id="B935476B917D2852FE64FB3C00A623DC" box="[432,495,1204,1228]" italics="true" pageId="13" pageNumber="432">colliei</emphasis>
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||
</taxonomicName>
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||
— van Dijk
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<emphasis id="B935476B917D2852FDA0FB3D03E923DC" box="[628,672,1204,1228]" italics="true" pageId="13" pageNumber="432">et al</emphasis>
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||
. (TTWG) 2014: 430.
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||
</paragraph>
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||
<paragraph id="8BFE9B79917D2852FF43FB5A036323FB" blockId="13.[151,890,1050,1351]" box="[151,554,1234,1259]" pageId="13" pageNumber="432">
|
||
<taxonomicName id="4C41E0FA917D2852FF43FB5A000F23FA" authorityName="Gray" authorityYear="1856" box="[151,326,1234,1258]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="432" phylum="Chordata" rank="species" species="colliei">
|
||
<emphasis id="B935476B917D2852FF43FB5A000F23FA" box="[151,326,1234,1258]" italics="true" pageId="13" pageNumber="432">Chelodina colliei</emphasis>
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||
</taxonomicName>
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||
— Cogger 2014: 243.
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</paragraph>
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||
<paragraph id="8BFE9B79917D2852FF43FB7902232219" blockId="13.[151,890,1050,1351]" box="[151,874,1264,1289]" pageId="13" pageNumber="432">
|
||
<taxonomicName id="4C41E0FA917D2852FF43FB7900A62219" authorityName="Gray" authorityYear="1856" box="[151,495,1265,1289]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="432" phylum="Chordata" rank="species" species="colliei" subGenus="Macrodiremys">
|
||
<emphasis id="B935476B917D2852FF43FB7900482219" box="[151,257,1265,1289]" italics="true" pageId="13" pageNumber="432">Chelodina</emphasis>
|
||
(
|
||
<emphasis id="B935476B917D2852FEC4FB7900EB2219" box="[272,418,1265,1289]" italics="true" pageId="13" pageNumber="432">Macrodiremys</emphasis>
|
||
)
|
||
<emphasis id="B935476B917D2852FE64FB7900A62219" box="[432,495,1265,1289]" italics="true" pageId="13" pageNumber="432">colliei</emphasis>
|
||
</taxonomicName>
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||
<emphasis id="B935476B917D2852FE22FB7A03442219" box="[502,525,1266,1289]" italics="true" pageId="13" pageNumber="432">—</emphasis>
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||
Rhodin
|
||
<emphasis id="B935476B917D2852FDB0FB7A03D92219" box="[612,656,1265,1289]" italics="true" pageId="13" pageNumber="432">et al</emphasis>
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. (TTWG) 2017: 193.
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||
</paragraph>
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<paragraph id="8BFE9B79917D2852FF43FA9803232238" blockId="13.[151,890,1050,1351]" box="[151,618,1295,1320]" pageId="13" pageNumber="432">
|
||
<taxonomicName id="4C41E0FA917D2852FF43FA98000F2238" authorityName="Gray" authorityYear="1856" box="[151,326,1296,1320]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="432" phylum="Chordata" rank="species" species="colliei">
|
||
<emphasis id="B935476B917D2852FF43FA98000F2238" box="[151,326,1296,1320]" italics="true" pageId="13" pageNumber="432">Chelodina colliei</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation id="EFD0E688917D2852FEB8FA87032F2238" author="Kehlmaier, C. & Zhang, X. & Georges, A. & Campbell, P. D. & Thomson, S. & Fritz, U." box="[364,614,1295,1320]" pageId="13" pageNumber="432" pagination="5841" refId="ref13343" refString="Kehlmaier, C., Zhang, X., Georges, A., Campbell, P. D., Thomson, S. & Fritz, U. (2019) Mitogenomics of historical type specimens of Australasian turtles: clarification of taxonomic confusion and old mitochondrial introgression. Nature: Scientific Reports, 9, 5841. https: // doi. org / 10.1038 / s 41598 - 019 - 42310 - x" type="journal article" year="2019">
|
||
Kehlmaier
|
||
<emphasis id="B935476B917D2852FE0AFA9903432238" box="[478,522,1296,1320]" italics="true" pageId="13" pageNumber="432">et al</emphasis>
|
||
. 2019: 6
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFE9B79917D2852FF43FAA603EC2257" blockId="13.[151,890,1050,1351]" box="[151,677,1326,1351]" pageId="13" pageNumber="432">
|
||
<taxonomicName id="4C41E0FA917D2852FF43FAA603512256" authorityName="J. Gray" authorityYear="1841" box="[151,536,1326,1351]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="432" phylum="Chordata" rank="species" species="oblonga" subGenus="Macrochelodina">
|
||
<emphasis id="B935476B917D2852FF43FAA600482256" box="[151,257,1326,1350]" italics="true" pageId="13" pageNumber="432">Chelodina</emphasis>
|
||
(
|
||
<emphasis id="B935476B917D2852FEC4FAA600FF2256" box="[272,438,1326,1350]" italics="true" pageId="13" pageNumber="432">Macrochelodina</emphasis>
|
||
)
|
||
<emphasis id="B935476B917D2852FE11FAA603512256" box="[453,536,1326,1350]" italics="true" pageId="13" pageNumber="432">oblonga</emphasis>
|
||
</taxonomicName>
|
||
— this study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
<subSection id="E2CE8292917D2851FF43FA06044522F7" lastPageId="14" lastPageNumber="433" pageId="13" pageNumber="432" type="multiple">
|
||
<subSubSection id="C35BC8F2917D2852FF43FA0602B322E1" pageId="13" pageNumber="432" type="materials_examined">
|
||
<paragraph id="8BFE9B79917D2852FF43FA06031722B8" blockId="13.[151,606,1422,1449]" box="[151,606,1422,1449]" pageId="13" pageNumber="432">
|
||
<emphasis id="B935476B917D2852FF43FA06031722B8" bold="true" box="[151,606,1422,1449]" pageId="13" pageNumber="432">
|
||
3.4.2. Description of
|
||
<typeStatus id="54FA25DB917D2852FE4DFA07008622B9" box="[409,463,1423,1449]" pageId="13" pageNumber="432">type</typeStatus>
|
||
specimens:
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFE9B79917D2852FF43FA5F02B322E1" blockId="13.[151,1437,1493,2025]" box="[151,1018,1493,1521]" pageId="13" pageNumber="432">
|
||
<typeStatus id="54FA25DB917D2852FF43FA5F004222E1" box="[151,267,1495,1521]" pageId="13" pageNumber="432" type="lectotype">Lectotype</typeStatus>
|
||
: BMNH 1947.3.5.89 (
|
||
<figureCitation id="137A87FC917D2852FDD1FA5E030922E0" box="[517,576,1494,1520]" captionStart="FIGURE 1" captionStartId="3.[151,250,1945,1970]" captionTargetBox="[357,1230,179,1918]" captionTargetId="figure@3.[357,1230,179,1918]" captionTargetPageId="3" captionText="FIGURE 1. Ventral, left lateral and dorsal views of the lectotype of Chelodina oblonga Gray (BMNH 1947.3.5.89). The lateral image is slightly dorsally angled, affecting the profile of the carapace. (Image courtesy of P. Campbell, BMNH)." figureDoi="http://doi.org/10.5281/zenodo.5918350" httpUri="https://zenodo.org/record/5918350/files/figure.png" pageId="13" pageNumber="432">Fig 1</figureCitation>
|
||
). Carapace length (straight) =
|
||
<quantity id="4CB9369C917D2852FC4DFA5E02BE22E1" box="[921,1015,1494,1521]" metricMagnitude="-1" metricUnit="m" metricValue="1.72" pageId="13" pageNumber="432" unit="cm" value="17.2">17.2 cm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35BC8F2917D2852FF13FA7204D02024" pageId="13" pageNumber="432" type="description">
|
||
<paragraph id="8BFE9B79917D2852FF13FA7204D02024" blockId="13.[151,1437,1493,2025]" pageId="13" pageNumber="432">
|
||
Description. Plastron with three inked markings, on the left humeral scute is the BMNH number “1947.3.5.89”, on the right humeral is the number “74a” Also on the bridge area of the right abdominal is the original number “40.12.9.81”; the latter part of the number is less clear. Dry mounted specimen, entire with the limbs extended. Neck broken, posterior part of neck skin collapsed inside shell. Carapace oval, slightly elongated from above, slightly furrowed along midline, dark brown with a reticulated pattern. 5
|
||
<superScript id="7C343631917D2852FC99F90102132187" attach="left" box="[845,858,1673,1687]" fontSize="6" pageId="13" pageNumber="432">th</superScript>
|
||
vertebral scute divided along midline; 2
|
||
<superScript id="7C343631917D2852FAF5F90104782187" attach="left" box="[1313,1329,1673,1687]" fontSize="6" pageId="13" pageNumber="432">nd</superScript>
|
||
marginal scute twice the length of first marginal, all other marginals normal. Plastron narrow and slightly trapezoidal, yellow-orange and highly reticulated. Legs all overstuffed; 4 claws on front and rear feet. Poor condition of neck makes determination of skin characters uncertain, however skin just behind the head finely reticulate with no tubercles observable. Head with artificial eyes inserted, and poorly preserved, though some osseous features visible externally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35BC8F2917D2851FF13F8B602BC242B" lastPageId="14" lastPageNumber="433" pageId="13" pageNumber="432" type="diagnosis">
|
||
<paragraph id="8BFE9B79917D2851FF13F8B602BC242B" blockId="13.[151,1437,1493,2025]" lastBlockId="14.[151,1437,152,1511]" lastPageId="14" lastPageNumber="433" pageId="13" pageNumber="432">
|
||
<materialsCitation id="3B299124917D2851FF13F8B602B8242B" ID-GBIF-Occurrence="2824704301" collectorName="McCord & Joseph-Ouni" country="Australia" lastPageId="14" lastPageNumber="433" location="Frontal region" pageId="13" pageNumber="432" specimenCount="6" typeStatus="lectotype">
|
||
Diagnostic comparison. The extreme narrowing and anterior elongation of the shell typical of the south-western Australian species is not obvious; however, carapace shape varies ontogenetically in this species, as apparent from a large series of specimens ranging from hatching to adult examined at the Chelonian Research Institute (CRI). The
|
||
<typeStatus id="54FA25DB917D2852FF43F82201B720D4" box="[151,254,1962,1988]" pageId="13" pageNumber="432" type="lectotype">lectotype</typeStatus>
|
||
fits within the range of carapace shapes of a subadult of this taxon. Plastron length/width at level of axillary notch ratio (P L/W) 2.63, which is comparable with specimens of known Perth provenance (CRI 3129: P L/W = 2.49; CRI 5070: P L/W = 2.56; CRI 4632: P L/W = 2.48), while species of
|
||
<taxonomicName id="4C41E0FA917E2851FBC6FF11045F27A3" authority="" authorityName="Shea & Thomson & Georges" authorityYear="2020" box="[1042,1302,153,179]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="subGenus" subGenus="Chelydera">
|
||
<emphasis id="B935476B917E2851FBC6FF1105CE27A3" box="[1042,1159,153,179]" italics="true" pageId="14" pageNumber="433">Chelodina</emphasis>
|
||
(
|
||
<emphasis id="B935476B917E2851FB4CFF11044427A3" box="[1176,1293,153,179]" italics="true" pageId="14" pageNumber="433">Chelydera</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
range from 1.88 (
|
||
<taxonomicName id="4C41E0FA917E2851FF0CFF35001A27C7" authorityName="Gray" authorityYear="1857" box="[216,339,189,215]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="expansa">
|
||
<emphasis id="B935476B917E2851FF0CFF35001A27C7" box="[216,339,189,215]" italics="true" pageId="14" pageNumber="433">C. expansa</emphasis>
|
||
</taxonomicName>
|
||
CRI 2397) to 2.10 (
|
||
<taxonomicName id="4C41E0FA917E2851FDE6FF3502CE27C7" box="[562,903,189,215]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="subSpecies" species="burrungandjii" subSpecies="wolloyarrina">
|
||
<emphasis id="B935476B917E2851FDE6FF3502CE27C7" box="[562,903,189,215]" italics="true" pageId="14" pageNumber="433">C. burrungandjii wolloyarrina</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<collectorName id="26B4FEAF917E2851FC41FF3502BB27C7" box="[917,1010,189,215]" pageId="14" pageNumber="433">McCord</collectorName>
|
||
&
|
||
<collectorName id="26B4FEAF917E2851FBC1FF3505EC27C7" box="[1045,1189,189,215]" pageId="14" pageNumber="433">Joseph-Ouni</collectorName>
|
||
, 2009).
|
||
<location id="8E9ECDA2917E2851FB2AFF3504D427C7" LSID="urn:lsid:plazi:treatment:03E82A6F917D2851FF43FC06044522F7:8E9ECDA2917E2851FB2AFF3504D427C7" box="[1278,1437,189,215]" country="Australia" name="Frontal region" pageId="14" pageNumber="433">Frontal region</location>
|
||
of
|
||
<typeStatus id="54FA25DB917E2851FF61FF69005527EB" box="[181,284,225,251]" pageId="14" pageNumber="433" type="lectotype">lectotype</typeStatus>
|
||
skull is narrower than in members of
|
||
<taxonomicName id="4C41E0FA917E2851FD12FF69028327EB" authority="(Chelodina)" authorityName="Fitzinger" authorityYear="1826" baseAuthorityName="Chelodina" box="[710,970,225,251]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="subGenus" subGenus="Chelodina">
|
||
<emphasis id="B935476B917E2851FD12FF69027227EB" box="[710,827,225,251]" italics="true" pageId="14" pageNumber="433">Chelodina</emphasis>
|
||
(
|
||
<emphasis id="B935476B917E2851FC9FFF69028927EB" box="[843,960,225,251]" italics="true" pageId="14" pageNumber="433">Chelodina</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
; however, this character is even narrower in
|
||
<taxonomicName id="4C41E0FA917E2851FF61FE8D00F0260F" authority="" authorityName="Shea & Thomson & Georges" authorityYear="2020" box="[181,441,261,287]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="subGenus" subGenus="Chelydera">
|
||
<emphasis id="B935476B917E2851FF61FE8D0063260F" box="[181,298,261,287]" italics="true" pageId="14" pageNumber="433">Chelodina</emphasis>
|
||
(
|
||
<emphasis id="B935476B917E2851FEE8FE8D00F8260F" box="[316,433,261,287]" italics="true" pageId="14" pageNumber="433">Chelydera</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
species from northern
|
||
<collectingCountry id="F356DBE9917E2851FD14FE8D0263260F" box="[704,810,261,287]" name="Australia" pageId="14" pageNumber="433">Australia</collectingCountry>
|
||
, due to the anterior rotation and enlargement of the orbits. Postorbitals appear to be narrow and arched, similar to
|
||
<taxonomicName id="4C41E0FA917E2851FC95FEA1050E2653" authority="(Chelodina)" authorityName="Fitzinger" authorityYear="1826" baseAuthorityName="Chelodina" box="[833,1095,297,323]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="subGenus" subGenus="Chelodina">
|
||
<emphasis id="B935476B917E2851FC95FEA102FF2653" box="[833,950,297,323]" italics="true" pageId="14" pageNumber="433">Chelodina</emphasis>
|
||
(
|
||
<emphasis id="B935476B917E2851FC13FEA105752653" box="[967,1084,297,323]" italics="true" pageId="14" pageNumber="433">Chelodina</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
, but differing from
|
||
<taxonomicName id="4C41E0FA917E2851FAFCFEA100572677" authority="" authorityName="Shea & Thomson & Georges" authorityYear="2020" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="subGenus" subGenus="Chelydera">
|
||
<emphasis id="B935476B917E2851FAFCFEA104D42653" box="[1320,1437,297,323]" italics="true" pageId="14" pageNumber="433">Chelodina</emphasis>
|
||
(
|
||
<emphasis id="B935476B917E2851FF75FEC5005F2677" box="[161,278,333,359]" italics="true" pageId="14" pageNumber="433">Chelydera</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
which has laterally flattened postorbitals. Posterior part of skull either poorly preserved or damaged, making outline of posterior parietal region difficult to ascertain. The skulls of both subspecies of
|
||
<taxonomicName id="4C41E0FA917E2851FB3BFEF9004426BF" authorityName="Thomson, Kennett & Georges" authorityYear="2000" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="burrungandjii">
|
||
<emphasis id="B935476B917E2851FB3BFEF9004426BF" italics="true" pageId="14" pageNumber="433">Chelodina burrungandjii</emphasis>
|
||
</taxonomicName>
|
||
are dorsoventrally flattened, a shape not present in this specimen. Further, the skull of
|
||
<taxonomicName id="4C41E0FA917E2851FB24FE1D00F826C3" authority="burrungandjii" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="subSpecies" species="burrungandjii" subSpecies="burrungandjii">
|
||
<emphasis id="B935476B917E2851FB24FE1D00F826C3" italics="true" pageId="14" pageNumber="433">Chelodina burrungandjii burrungandjii</emphasis>
|
||
</taxonomicName>
|
||
is laterally broadened and the palate anteroposteriorly elongated (
|
||
<bibRefCitation id="EFD0E688917E2851FB44FE31043426C3" author="Thomson, S. & Kennett, R. & Georges, A." box="[1168,1405,441,467]" pageId="14" pageNumber="433" pagination="675 - 685" refId="ref14808" refString="Thomson, S., Kennett, R. & Georges, A. (2000) A new species of long-necked turtle (Testudines: Chelidae) from the Arnhem Land Plateau, Northern Territory, Australia. Chelonian Conservation and Biology, 3 (4), 675 - 685." type="journal article" year="2000">
|
||
Thomson
|
||
<emphasis id="B935476B917E2851FAD7FE32047D26C3" box="[1283,1332,441,467]" italics="true" pageId="14" pageNumber="433">et al</emphasis>
|
||
. 2000
|
||
</bibRefCitation>
|
||
), a configuration also not present in the
|
||
<typeStatus id="54FA25DB917E2851FDFEFE5503DD26E7" box="[554,660,477,503]" pageId="14" pageNumber="433" type="lectotype">lectotype</typeStatus>
|
||
. The anterior bridge struts of the
|
||
<taxonomicName id="4C41E0FA917E2851FBD4FE55044B26E7" authority="" authorityName="Shea & Thomson & Georges" authorityYear="2020" box="[1024,1282,477,503]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="subGenus" subGenus="Chelydera">
|
||
<emphasis id="B935476B917E2851FBD4FE55053C26E7" box="[1024,1141,477,503]" italics="true" pageId="14" pageNumber="433">Chelodina</emphasis>
|
||
(
|
||
<emphasis id="B935476B917E2851FB50FE5505B026E7" box="[1156,1273,477,503]" italics="true" pageId="14" pageNumber="433">Chelydera</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
species are in general enlarged and encroach to over halfway across the first pleural—this is most extensive in
|
||
<taxonomicName id="4C41E0FA917E2851FB12FD8904D5250B" authorityName="Gray" authorityYear="1857" box="[1222,1436,513,539]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="expansa">
|
||
<emphasis id="B935476B917E2851FB12FD8904D5250B" box="[1222,1436,513,539]" italics="true" pageId="14" pageNumber="433">Chelodina expansa</emphasis>
|
||
</taxonomicName>
|
||
and least in
|
||
<taxonomicName id="4C41E0FA917E2851FEF6FDAD00A4252F" authorityName="Ogilby" authorityYear="1890" box="[290,493,549,575]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="rugosa">
|
||
<emphasis id="B935476B917E2851FEF6FDAD00A4252F" box="[290,493,549,575]" italics="true" pageId="14" pageNumber="433">Chelodina rugosa</emphasis>
|
||
</taxonomicName>
|
||
; however this character is absent in the
|
||
<typeStatus id="54FA25DB917E2851FC6FFDAD056C252F" box="[955,1061,549,575]" pageId="14" pageNumber="433" type="lectotype">lectotype</typeStatus>
|
||
.
|
||
<taxonomicName id="4C41E0FA917E2851FBE5FDAD0402252F" authorityName="Thomson, Kennett & Georges" authorityYear="2000" box="[1073,1355,549,575]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="burrungandjii">
|
||
<emphasis id="B935476B917E2851FBE5FDAD0402252F" box="[1073,1355,549,575]" italics="true" pageId="14" pageNumber="433">Chelodina burrungandjii</emphasis>
|
||
</taxonomicName>
|
||
has an expanded posterior lobe of the plastron at the level of the femoral scutes (
|
||
<bibRefCitation id="EFD0E688917E2851FC1AFDC105F22573" author="Thomson, S. & Kennett, R. & Georges, A." box="[974,1211,585,611]" pageId="14" pageNumber="433" pagination="675 - 685" refId="ref14808" refString="Thomson, S., Kennett, R. & Georges, A. (2000) A new species of long-necked turtle (Testudines: Chelidae) from the Arnhem Land Plateau, Northern Territory, Australia. Chelonian Conservation and Biology, 3 (4), 675 - 685." type="journal article" year="2000">
|
||
Thomson
|
||
<emphasis id="B935476B917E2851FB94FDC205382573" box="[1088,1137,585,611]" italics="true" pageId="14" pageNumber="433">et al</emphasis>
|
||
. 2000
|
||
</bibRefCitation>
|
||
; McCord & Joseph- Ouni 2007a), a feature not present in the
|
||
<typeStatus id="54FA25DB917E2851FD8AFDE5038C2597" box="[606,709,621,647]" pageId="14" pageNumber="433" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C41E0FA917E2851FD3CFDE5022D2597" authorityName="J. Gray" authorityYear="1841" box="[744,868,621,647]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="oblonga">
|
||
<emphasis id="B935476B917E2851FD3CFDE5022D2597" box="[744,868,621,647]" italics="true" pageId="14" pageNumber="433">C. oblonga</emphasis>
|
||
</taxonomicName>
|
||
, which has a more evenly tapered posterior lobe of the plastron characteristic of the south-western species (
|
||
<bibRefCitation id="EFD0E688917E2851FCD3FD1902DC25BB" author="Goode, J." box="[775,917,657,683]" pageId="14" pageNumber="433" refId="ref11997" refString="Goode, J. (1967) Freshwater tortoises of Australia and New Guinea (in the family Chelidae). Lansdowne Press, Melbourne, x + 154 pp." type="book" year="1967">Goode 1967</bibRefCitation>
|
||
; Cogger 2017). The combination of characters above clearly assigns the
|
||
<typeStatus id="54FA25DB917E2851FE6DFD3D036925DF" box="[441,544,693,719]" pageId="14" pageNumber="433" type="lectotype">lectotype</typeStatus>
|
||
to the Oblong Turtle. Thomson (2000) identified the presence of insertions for an expanded muscular system on the ventral surface of the carapace, only found in the members of the
|
||
<taxonomicName id="4C41E0FA917E2851FAFCFD5104D425E3" authorityName="Fitzinger" authorityYear="1826" box="[1320,1437,729,755]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="genus">
|
||
<emphasis id="B935476B917E2851FAFCFD5104D425E3" box="[1320,1437,729,755]" italics="true" pageId="14" pageNumber="433">Chelodina</emphasis>
|
||
</taxonomicName>
|
||
B group. It is probable that this was an artefact of preservation as the muscles themselves were long gone, and this character should be considered to have been incorrectly codified at that time
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35BC8F2917E2851FF13FCCD051E244F" box="[199,1111,836,864]" pageId="14" pageNumber="433" type="materials_examined">
|
||
<paragraph id="8BFE9B79917E2851FF13FCCD051E244F" blockId="14.[151,1437,152,1511]" box="[199,1111,836,864]" pageId="14" pageNumber="433">
|
||
<typeStatus id="54FA25DB917E2851FF13FCCD002D244F" box="[199,356,837,863]" pageId="14" pageNumber="433" type="paralectotype">Paralectotype</typeStatus>
|
||
: OUMNH 02584 (
|
||
<figureCitation id="137A87FC917E2851FDE0FCCD03EB244F" box="[564,674,837,863]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="11.[151,250,1082,1107]" captionStartId-1="12.[151,250,1945,1970]" captionTargetBox-0="[189,1398,381,1057]" captionTargetBox-1="[282,1304,181,1919]" captionTargetId-0="figure@11.[189,1398,381,1057]" captionTargetId-1="figure@12.[282,1304,181,1920]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURE 3. Dorsal view of the paralectotype of Chelodina oblonga Gray (OUMNH 02584). (Image courtesy of K. Child, OUMNH)." captionText-1="FIGURE 4. Ventral, right lateral and dorsal views of the shell of the paralectotype of Chelodina oblonga Gray (OUMNH 02584). (Images courtesy of K. Child, OUMNH)." figureDoi-0="http://doi.org/10.5281/zenodo.5918354" figureDoi-1="http://doi.org/10.5281/zenodo.5918356" httpUri-0="https://zenodo.org/record/5918354/files/figure.png" httpUri-1="https://zenodo.org/record/5918356/files/figure.png" pageId="14" pageNumber="433">Figs. 3–4</figureCitation>
|
||
). Carapace length (straight) =
|
||
<quantity id="4CB9369C917E2851FC22FCCC051D244F" box="[1014,1108,836,863]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" pageId="14" pageNumber="433" unit="cm" value="27.5">27.5 cm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35BC8F2917E2851FF13FCE1055B236F" pageId="14" pageNumber="433" type="description">
|
||
<paragraph id="8BFE9B79917E2851FF13FCE1055B236F" blockId="14.[151,1437,152,1511]" pageId="14" pageNumber="433">
|
||
Description. Plastron with three labels and markings: “Ref. 2584” inked onto left abdominal scute; “
|
||
<taxonomicName id="4C41E0FA917E2851FAFCFCE1034624B7" authority="Bell Coll Rev F. W. Hope" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="oblonga">
|
||
<emphasis id="B935476B917E2851FAFCFCE101BA24B7" italics="true" pageId="14" pageNumber="433">Chelodina oblonga</emphasis>
|
||
Bell Coll Rev F.W. Hope
|
||
</taxonomicName>
|
||
” on a label stuck to the left abdominal and pectoral scutes, and “a” in a small label on the left abdominal scute. Dry mounted specimen, entire, with neck and limbs extended. Deep reticulated pattern on the shell typical of the Perth species. 4
|
||
<superScript id="7C343631917E2851FDA5FC5C033724F2" attach="left" box="[625,638,980,994]" fontSize="6" pageId="14" pageNumber="433">th</superScript>
|
||
vertebral scute and left 3
|
||
<superScript id="7C343631917E2851FC4BFC5C02E424F2" attach="left" box="[927,941,980,994]" fontSize="6" pageId="14" pageNumber="433">rd</superScript>
|
||
costal scute divided; nuchal scute wide; 2
|
||
<superScript id="7C343631917E2851FA58FC5C04D524F2" attach="none" box="[1420,1436,980,994]" fontSize="6" pageId="14" pageNumber="433">nd</superScript>
|
||
marginal longer than 1
|
||
<superScript id="7C343631917E2851FE4EFC7000EC2316" attach="none" box="[410,421,1016,1030]" fontSize="6" pageId="14" pageNumber="433">st</superScript>
|
||
; supracaudal scute is offset and deformed; asymmetry of marginal length on the left and right side, though 11 are present on each side; lateral marginals reduced in lateral view. Bridge small, narrow. Plastron dark yellow with no deformities in the scutes. Neck and rear legs slightly overstuffed. 4 claws on front and rear feet. Dorsal surface of neck dark grey, skin finely reticulated, lacking tubercles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35BC8F2917E2851FF13FB01044522F7" pageId="14" pageNumber="433" type="diagnosis">
|
||
<paragraph id="8BFE9B79917E2851FF13FB01044522F7" blockId="14.[151,1437,152,1511]" pageId="14" pageNumber="433">
|
||
Diagnostic comparison. Carapace elongate and narrow in dorsal view, dorsoventrally flattened in lateral view, medium brown; this combination of features is unique to the Oblong Turtle, the shell shape features are only prominent in adults. Bridge small, narrow. Plastron long, narrow (plastral length/width 2.51, markedly higher than any
|
||
<taxonomicName id="4C41E0FA917E2851FF43FB7D00D0221F" authority="" authorityName="Shea & Thomson & Georges" authorityYear="2020" box="[151,409,1269,1295]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="subGenus" subGenus="Chelydera">
|
||
<emphasis id="B935476B917E2851FF43FB7D0045221F" box="[151,268,1269,1295]" italics="true" pageId="14" pageNumber="433">Chelodina</emphasis>
|
||
(
|
||
<emphasis id="B935476B917E2851FEC8FB7D00D8221F" box="[284,401,1269,1295]" italics="true" pageId="14" pageNumber="433">Chelydera</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
species). Some features of skull shape are visible due to shrinking of the skin over the skull. Frontal bone and postorbital region narrow, however the frontal is wider than in
|
||
<taxonomicName id="4C41E0FA917E2851FBC0FA9104512223" authority="" authorityName="Shea & Thomson & Georges" authorityYear="2020" box="[1044,1304,1305,1331]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="subGenus" subGenus="Chelydera">
|
||
<emphasis id="B935476B917E2851FBC0FA9105C02223" box="[1044,1161,1305,1331]" italics="true" pageId="14" pageNumber="433">Chelodina</emphasis>
|
||
(
|
||
<emphasis id="B935476B917E2851FB4CFA9104442223" box="[1176,1293,1305,1331]" italics="true" pageId="14" pageNumber="433">Chelydera</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
; postorbital highly arched as against flat for
|
||
<taxonomicName id="4C41E0FA917E2851FDD4FAB5024C2247" authority="" authorityName="Shea & Thomson & Georges" authorityYear="2020" box="[512,773,1341,1367]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="433" phylum="Chordata" rank="subGenus" subGenus="Chelydera">
|
||
<emphasis id="B935476B917E2851FDD4FAB5033C2247" box="[512,629,1341,1367]" italics="true" pageId="14" pageNumber="433">Chelodina</emphasis>
|
||
(
|
||
<emphasis id="B935476B917E2851FD51FAB503B32247" box="[645,762,1341,1367]" italics="true" pageId="14" pageNumber="433">Chelydera</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
, see above. Posterior part of parietal elongated and narrow, beginning to expand anteriorly halfway towards orbits, giving a diamond shaped anterior parietal region. Parietal arches, dorsal to quadrates, do not reach supraoccipital, but form cristae that arch dorsomedially. While this specimen has not been genotyped, we believe that morphologically its identification as an Oblong Turtle is without question and genotyping would be for curatorial reasons only, as this specimen is not a name bearing
|
||
<typeStatus id="54FA25DB917E2851FB00FA45044122F7" box="[1236,1288,1485,1511]" pageId="14" pageNumber="433">type</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |