166 lines
19 KiB
XML
166 lines
19 KiB
XML
<document id="23C4352824B1EA450AF2A6F4F875A31B" ID-DOI="10.5281/zenodo.13203160" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203160" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio & Müller, Patrick" docDate="2022" docId="03DAD52BFFBEFFF6FFEBBFEA5B10A181" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Sanaungulus myanmaricus Fanti & Müller, 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="359" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="358" updateTime="1722697113453" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:mods id="BCC30B8B2FE3ABE63672389884C124FF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C25F0B959C2EF127DEA72532EE37F63D">
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<mods:title id="54179A39079529FCA43C93D43BF421EE">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="326DD492FEB2BB7394C4478FEC736B2F">Fanti, Fabrizio</mods:namePart>
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<mods:roleTerm id="E83A71D14E8AEFE46460C9114D527762">Author</mods:roleTerm>
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<mods:namePart id="1365C106774577E7582EBE81C6BEDA01">Müller, Patrick</mods:namePart>
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<mods:typeOfResource id="45A07709BFDFAFF595CD6641E958FBC3">text</mods:typeOfResource>
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<mods:titleInfo id="B051924C1828735BA2AB49BE46ABAC4B">
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<mods:title id="6E0D01B0106DDFAF3B9511936FF1AA33">Baltic Journal of Coleopterology</mods:title>
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</mods:titleInfo>
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<mods:part id="EB8BAA7D50B4CDB562579B07F54E9C1C">
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<mods:date id="4E7C8011EA4F6DDE2EA597FC72A8AF36">2022</mods:date>
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<mods:detail id="D5C51EFDD8B3BDB2B1A0E478F821D924" type="volume">
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<mods:number id="DDC1B58CF37AED218668C7DF0990A395">22</mods:number>
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<mods:detail id="4484B43B36A98B6B299FF1EAF076C081" type="issue">
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<mods:number id="E8E87D9B35475A1D6DAA377A14C0732B">2</mods:number>
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<mods:start id="C8C290A0D9CBA1C7B81E0779739100D0">331</mods:start>
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<mods:classification id="9AC909BF5A66EFCE7CDC6CF934F399FF">journal article</mods:classification>
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<mods:identifier id="D9665F9BCAEB411AA2AEA31CFBA519CB" type="DOI">10.5281/zenodo.13203160</mods:identifier>
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<mods:identifier id="B7C45126ABB1B8A186E4F336B759CB3F" type="ISSN">1407-8619</mods:identifier>
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<treatment id="03DAD52BFFBEFFF6FFEBBFEA5B10A181" LSID="urn:lsid:plazi:treatment:03DAD52BFFBEFFF6FFEBBFEA5B10A181" httpUri="http://treatment.plazi.org/id/03DAD52BFFBEFFF6FFEBBFEA5B10A181" lastPageId="28" lastPageNumber="359" pageId="27" pageNumber="358">
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<subSubSection id="C36937B6FFBEFFF1FFEBBFEA5EBAA39F" box="[113,508,727,752]" pageId="27" pageNumber="358" type="nomenclature">
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<paragraph id="8BCC643DFFBEFFF1FFEBBFEA5EBAA39F" blockId="27.[113,508,727,784]" box="[113,508,727,752]" pageId="27" pageNumber="358">
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<heading id="D084D351FFBEFFF1FFEBBFEA5EBAA39F" bold="true" box="[113,508,727,752]" fontSize="10" level="2" pageId="27" pageNumber="358" reason="4">
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<emphasis id="B907B82FFFBEFFF1FFEBBFEA5EBAA39F" bold="true" box="[113,508,727,752]" pageId="27" pageNumber="358">
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<taxonomicName id="4C731FBEFFBEFFF1FFEBBFEA5EE7A380" authorityName="Fanti & Müller" authorityYear="2022" box="[113,417,728,752]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="358" phylum="Arthropoda" rank="species" species="myanmaricus" status="sp. nov.">
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<emphasis id="B907B82FFFBEFFF1FFEBBFEA5EE7A380" bold="true" box="[113,417,728,752]" italics="true" pageId="27" pageNumber="358">Sanaungulus myanmaricus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FFBEFFF1FE3DBFE55EBAA39F" box="[423,508,727,751]" pageId="27" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36937B6FFBEFFF1FFEBBFCB5F88A260" box="[113,206,761,784]" pageId="27" pageNumber="358" type="description">
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||
<paragraph id="8BCC643DFFBEFFF1FFEBBFCB5F88A260" blockId="27.[113,508,727,784]" box="[113,206,761,784]" pageId="27" pageNumber="358">
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(
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<figureCitation id="134878B8FFBEFFF1FFE3BFCB5F80A260" box="[121,198,761,784]" captionStart="Fig" captionStartId="27.[113,152,1665,1688]" captionTargetBox="[113,1134,1022,1634]" captionTargetId="figure-363@27.[113,1134,1022,1634]" captionTargetPageId="27" captionText="Fig. 15. Sanaungulus myanmaricus sp. nov. in Burmese amber. Holotype, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13203209" httpUri="https://zenodo.org/record/13203209/files/figure.png" pageId="27" pageNumber="358">Fig. 15</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFBEFFF1FFEBBE025CE4A058" pageId="27" pageNumber="358" type="materials_examined">
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<paragraph id="8BCC643DFFBEFFF1FFEBBE025EFBA2BA" blockId="27.[113,605,816,970]" pageId="27" pageNumber="358">
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<materialsCitation id="3B1B6E60FFBEFFF1FFEBBE025EFBA2BA" collectionCode="BUB" county="Probable" location="Catalog" municipality="Burmese" pageId="27" pageNumber="358" specimenCode="BUB4166" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
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<emphasis id="B907B82FFFBEFFF1FFEBBE025FA6A238" bold="true" box="[113,224,816,840]" pageId="27" pageNumber="358">
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<typeStatus id="54C8DA9FFFBEFFF1FFEBBE025F9DA238" box="[113,219,816,840]" pageId="27" pageNumber="358" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectingCounty id="62AD1CB1FFBEFFF1FF7DBE025E00A237" box="[231,326,816,839]" pageId="27" pageNumber="358">Probable</collectingCounty>
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male, adult specimen in a
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<collectingMunicipality id="6BA8FE47FFBEFFF1FFEBBE635F8BA218" box="[113,205,849,872]" pageId="27" pageNumber="358">Burmese</collectingMunicipality>
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(
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<collectingRegion id="49B7AADFFFBEFFF1FF43BE635E61A218" box="[217,295,849,872]" country="Myanmar" name="Kachin" pageId="27" pageNumber="358">Kachin</collectingRegion>
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) amber piece:
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<location id="8EAC32E6FFBEFFF1FE23BE635D4FA218" LSID="urn:lsid:plazi:treatment:03DAD52BFFBEFFF6FFEBBFEA5B10A181:8EAC32E6FFBEFFF1FE23BE635D4FA218" box="[441,521,849,872]" county="Probable" municipality="Burmese" name="Catalog" pageId="27" pageNumber="358" stateProvince="Kachin">Catalog</location>
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number SNHM
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<quantity id="4C8BC9D8FFBEFFF1FF50BE405E67A2F9" box="[202,289,882,905]" metricMagnitude="2" metricUnit="m" metricValue="1.526286" pageId="27" pageNumber="358" unit="in" value="6009.0">6009 in</quantity>
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the
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<location id="8EAC32E6FFBEFFF1FEC0BE405E8FA2DA" LSID="urn:lsid:plazi:treatment:03DAD52BFFBEFFF6FFEBBFEA5B10A181:8EAC32E6FFBEFFF1FEC0BE405E8FA2DA" county="Probable" municipality="Burmese" name="Staatliches Naturhistorisches Museum Braunschweig" pageId="27" pageNumber="358" stateProvince="Kachin">Staatliches Naturhistorisches Museum Braunschweig</location>
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(ex
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<location id="8EAC32E6FFBEFFF1FD9ABEA15D1AA2DA" LSID="urn:lsid:plazi:treatment:03DAD52BFFBEFFF6FFEBBFEA5B10A181:8EAC32E6FFBEFFF1FD9ABEA15D1AA2DA" box="[512,604,915,938]" county="Probable" municipality="Burmese" name="Muller's" pageId="27" pageNumber="358" stateProvince="Kachin">Müller’s</location>
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collection number:
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<specimenCode id="DBD5CC46FFBEFFF1FEDABE815EF7A2BA" box="[320,433,947,970]" collectionCode="BUB" pageId="27" pageNumber="358">BUB4166</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BCC643DFFBEFFF1FD19BDA85C93A1A2" blockId="27.[643,1134,154,210]" pageId="27" pageNumber="358">
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<emphasis id="B907B82FFFBEFFF1FD19BDA85C5AA1C2" bold="true" box="[643,796,154,178]" pageId="27" pageNumber="358">Type locality.</emphasis>
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<collectingCountry id="F36424ADFFBEFFF1FCBEBDA95CD6A1C2" box="[804,912,155,178]" name="Myanmar" pageId="27" pageNumber="358">Myanmar</collectingCountry>
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:
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<collectingRegion id="49B7AADFFFBEFFF1FC01BDA95B63A1C2" box="[923,1061,155,178]" country="Myanmar" name="Kachin" pageId="27" pageNumber="358">Kachin state</collectingRegion>
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, Myitkyina District, Hukawng Valley.
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</paragraph>
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<paragraph id="8BCC643DFFBEFFF1FD19BDC35CE4A058" blockId="27.[643,1134,241,296]" pageId="27" pageNumber="358">
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<emphasis id="B907B82FFFBEFFF1FD19BDC35C50A079" bold="true" box="[643,790,241,265]" pageId="27" pageNumber="358">Type horizon.</emphasis>
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Lowermost Cenomanian (98.79 ± 0.62 Ma), midCretaceous.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFBEFFF1FD19BC7A5C62A30F" pageId="27" pageNumber="358" type="diagnosis">
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<paragraph id="8BCC643DFFBEFFF1FD19BC7A5C62A30F" blockId="27.[643,1134,328,639]" pageId="27" pageNumber="358">
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<emphasis id="B907B82FFFBEFFF1FD19BC7A5C3CA010" bold="true" box="[643,890,328,352]" pageId="27" pageNumber="358">Differential diagnosis.</emphasis>
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<taxonomicName id="4C731FBEFFBEFFF1FCE4BC7A5DFEA00F" authorityName="Fanti & Müller" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="358" phylum="Arthropoda" rank="species" species="myanmaricus" status="sp. nov.">
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<emphasis id="B907B82FFFBEFFF1FCE4BC7A5DFEA00F" italics="true" pageId="27" pageNumber="358">Sanaungulus myanmaricus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FFBEFFF1FD5ABC5A5C55A00F" box="[704,787,360,383]" pageId="27" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
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is characterized by a very elongated pronotum with the surface slightly undulate, a feature present only in
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<taxonomicName id="4C731FBEFFBEFFF1FC11BC9A5B0FA0CF" authorityName="Fanti & Müller" authorityYear="2022" box="[907,1097,424,447]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="358" phylum="Arthropoda" rank="species" species="nilsi" status="sp. nov.">
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<emphasis id="B907B82FFFBEFFF1FC11BC9A5B0FA0CF" box="[907,1097,424,447]" italics="true" pageId="27" pageNumber="358">Sanaungulus nilsi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FFBEFFF1FBD5BC9A5DF6A0AF" pageId="27" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="4C731FBEFFBEFFF1FD7DBCFA5CB0A0AF" authorityName="Fanti & Müller" authorityYear="2022" box="[743,1014,456,479]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="358" phylum="Arthropoda" rank="species" species="kachinensis" status="sp. nov.">
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<emphasis id="B907B82FFFBEFFF1FD7DBCFA5CB0A0AF" box="[743,1014,456,479]" italics="true" pageId="27" pageNumber="358">Sanaungulus kachinensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FFBEFFF1FC64BCFA5B14A0AF" box="[1022,1106,456,479]" pageId="27" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
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<taxonomicName id="4C731FBEFFBEFFF1FBC0BCFA5DF6A08F" authorityName="Fanti & Müller" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="358" phylum="Arthropoda" rank="species" species="nilsi">
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<emphasis id="B907B82FFFBEFFF1FBC0BCFA5DF6A08F" italics="true" pageId="27" pageNumber="358">S. nilsi</emphasis>
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</taxonomicName>
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has a pronotum more concave in the middle and more bordered at the margins (especially at the posterior margin), while
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<taxonomicName id="4C731FBEFFBEFFF1FC4FBF1A5B28A34F" authorityName="Fanti & Müller" authorityYear="2022" box="[981,1134,552,575]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="358" phylum="Arthropoda" rank="species" species="kachinensis">
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<emphasis id="B907B82FFFBEFFF1FC4FBF1A5B28A34F" box="[981,1134,552,575]" italics="true" pageId="27" pageNumber="358">S. kachinensis</emphasis>
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</taxonomicName>
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has a pronotum that is shorter and with a transversal incision.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFBEFFF6FD19BFAC5D1CA482" lastPageId="28" lastPageNumber="359" pageId="27" pageNumber="358" type="description">
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<paragraph id="8BCC643DFFBEFFF1FD19BFAC5DBBA266" blockId="27.[643,1134,670,790]" pageId="27" pageNumber="358">
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<emphasis id="B907B82FFFBEFFF1FD19BFAC5C48A3C6" bold="true" box="[643,782,670,694]" pageId="27" pageNumber="358">Description.</emphasis>
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Adult, winged, slender. Probable male, defined on the basis of slender abdomen and long antennae. Body length
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<quantity id="4C8BC9D8FFBEFFF1FC40BFED5B69A386" box="[986,1071,735,758]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="27" pageNumber="358" unit="mm" value="3.2">3.2 mm</quantity>
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. Entirely brown.
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</paragraph>
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<paragraph id="8BCC643DFFBEFFF6FD19BE075D1CA482" blockId="27.[643,1135,821,972]" lastBlockId="28.[151,644,155,1522]" lastPageId="28" lastPageNumber="359" pageId="27" pageNumber="358">Head transverse, constricted (triangularshaped) behind the eyes, equipped with pubescence and punctation. Eyes prominent, large, roundish, inserted in the upper and lateral part of the head. Mandibles elongated, falciform. Maxillary palps 4 segmented with the last palpomere securiform. Labial palps 3 segmented, with the last palpomere elongated and strongly securiform.Antennae 11 segmented, pectinate, surpassing half of elytra and reaching half of abdomen, antennal insertion in the eyes proximity; scape slender, elongated, slightly enlarged apically; antennomere II short, swollen apically; antennomere III elongated, filiform, about 1.8 times longer than second; antennomere IV longer and stouter than previous one; antennomeres V–IX extremely long, each equipped near the base with a long antennal process which is very long and rounded apically; antennomere X shorter than previous ones, equipped with an antennal process shorter than others and slightly squadrate at the base; antennomere XI short and robust, with rounded apex; all the antennomeres pubescent. Pronotum very elongated, narrower than head, anterior margin undulate, posterior margin extend backward in the middle and slightly bordered, sides straight and bordered, corners obtuse, pronotum disc irregular with small concavity and drafts. Scutellum triangularshaped. Elytra short which reveals various abdominal segments, at humeri wider than pronotum, posteriorly parallelsided and slightly dehiscent apically, rounded apexes, surface pubescent. Metathoracic wings transparent, slightly longer than elytra. Sternum elongated, sternites transverse, last abdominal segment wide and with rounded apex. Legs thin, extremely long, pubescent; coxae massive; trochanters elongated, with rounded apex; femora almost straight, cylindrical, slightly sturdier than tibiae; tibiae, thin, cylindrical, extremely long, with a spur near the apex, pro and mesotibiae shorter than pro and mesofemora, metatibiae as long as metafemora. Tarsal formula 5 5 5; first tarsomere long, about 2.0 times longer than second; third tarsomere short, triangularshaped; fourth strongly bilobed; fifth tarsomere extremely elongated, thin, flat and curved; claws simple, long, pointed, without denticle at the base.</paragraph>
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</subSubSection>
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||
<caption id="DF0C34B5FFBEFFF1FFEBBBB35B4CA7E8" ID-DOI="http://doi.org/10.5281/zenodo.13203209" ID-Zenodo-Dep="13203209" box="[113,1034,1665,1688]" httpUri="https://zenodo.org/record/13203209/files/figure.png" pageId="27" pageNumber="358" startId="27.[113,152,1665,1688]" targetBox="[113,1134,1022,1634]" targetPageId="27" targetType="figure">
|
||
<paragraph id="8BCC643DFFBEFFF1FFEBBBB35B4CA7E8" blockId="27.[113,1034,1665,1688]" box="[113,1034,1665,1688]" pageId="27" pageNumber="358">
|
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Fig. 15.
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<taxonomicName id="4C731FBEFFBEFFF1FF50BBB35EACA7E8" authorityName="Fanti & Müller" authorityYear="2022" box="[202,490,1665,1688]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="358" phylum="Arthropoda" rank="species" species="myanmaricus" status="sp. nov.">
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<emphasis id="B907B82FFFBEFFF1FF50BBB35EACA7E8" box="[202,490,1665,1688]" italics="true" pageId="27" pageNumber="358">Sanaungulus myanmaricus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2340554FFBEFFF1FE6ABBB35D05A7E8" box="[496,579,1665,1688]" pageId="27" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
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in Burmese amber. Holotype, lateral view.
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</paragraph>
|
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</caption>
|
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<subSubSection id="C36937B6FFB9FFF6FF0DBB235E85A7EF" pageId="28" pageNumber="359" type="etymology">
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<paragraph id="8BCC643DFFB9FFF6FF0DBB235DC2A738" blockId="28.[151,644,1553,1608]" pageId="28" pageNumber="359">
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<emphasis id="B907B82FFFB9FFF6FF0DBB235E5EA759" bold="true" box="[151,280,1553,1577]" pageId="28" pageNumber="359">Etymology.</emphasis>
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Derived from the toponym
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<collectingCountry id="F36424ADFFB9FFF6FDA6BB235F86A738" name="Myanmar" pageId="28" pageNumber="359">Myanmar</collectingCountry>
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(nation where the amber mines are located).
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</paragraph>
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<paragraph id="8BCC643DFFB9FFF6FF0DBB5A5E85A7EF" blockId="28.[151,643,1640,1695]" pageId="28" pageNumber="359">
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<emphasis id="B907B82FFFB9FFF6FF0DBB5A5E7DA7F0" bold="true" box="[151,315,1640,1664]" pageId="28" pageNumber="359">Syninclusions.</emphasis>
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Air bubbles, detritus, stellate hairs, an unidentified insect.
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="C36937B6FFB9FFF6FD32BDA85B10A181" pageId="28" pageNumber="359" type="discussion">
|
||
<paragraph id="8BCC643DFFB9FFF6FD32BDA85B10A181" blockId="28.[680,1172,154,241]" pageId="28" pageNumber="359">
|
||
<emphasis id="B907B82FFFB9FFF6FD32BDA85C52A1C2" bold="true" box="[680,788,154,178]" pageId="28" pageNumber="359">Remarks.</emphasis>
|
||
Piece of clear, golden amber that measures 18x11x
|
||
<quantity id="4C8BC9D8FFB9FFF6FCAEBD885C30A1A1" box="[820,886,186,209]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="28" pageNumber="359" unit="mm" value="4.0">4 mm</quantity>
|
||
. The inclusion is complete but well visible only laterally (one side).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |