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<document id="C20E4F33B41B49DE849D5CBF42C22FB3" ID-DOI="10.5281/zenodo.13203160" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203160" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio &amp; Müller, Patrick" docDate="2022" docId="03DAD52BFFA6FFEFFD19BC725FE2A406" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Burmomiles volpei Fanti &amp; Müller, 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="336" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="334" updateTime="1722697113453" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="2EA7120D84EF7887A74667629B1B5EB5">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="F2EE1E39C66883D941D750A1528FF75C">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="EAE63186AC7B9C975C5800A528C3088A">Müller, Patrick</mods:namePart>
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<mods:date id="B0289862FD6A92C4976E5D8BA7463F3E">2022</mods:date>
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<treatment id="03DAD52BFFA6FFEFFD19BC725FE2A406" LSID="urn:lsid:plazi:treatment:03DAD52BFFA6FFEFFD19BC725FE2A406" httpUri="http://treatment.plazi.org/id/03DAD52BFFA6FFEFFD19BC725FE2A406" lastPageId="5" lastPageNumber="336" pageId="3" pageNumber="334">
<subSubSection id="C36937B6FFA6FFE9FD19BC725CE8A028" box="[643,942,320,344]" pageId="3" pageNumber="334" type="nomenclature">
<paragraph id="8BCC643DFFA6FFE9FD19BC725CE8A028" blockId="3.[643,942,320,375]" box="[643,942,320,344]" pageId="3" pageNumber="334">
<heading id="D084D351FFA6FFE9FD19BC725CE8A028" bold="true" box="[643,942,320,344]" fontSize="10" level="2" pageId="3" pageNumber="334" reason="4">
<emphasis id="B907B82FFFA6FFE9FD19BC725CE8A028" bold="true" box="[643,942,320,344]" pageId="3" pageNumber="334">
<taxonomicName id="4C731FBEFFA6FFE9FD19BC725C17A028" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[643,849,320,344]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="species" species="volpei" status="sp. nov.">
<emphasis id="B907B82FFFA6FFE9FD19BC725C17A028" bold="true" box="[643,849,320,344]" italics="true" pageId="3" pageNumber="334">Burmomiles volpei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFA6FFE9FCC2BC725CE8A028" box="[856,942,320,344]" pageId="3" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA6FFE9FD19BC525D94A007" box="[643,722,352,375]" pageId="3" pageNumber="334" type="description">
<paragraph id="8BCC643DFFA6FFE9FD19BC525D94A007" blockId="3.[643,942,320,375]" box="[643,722,352,375]" pageId="3" pageNumber="334">
(
<figureCitation id="134878B8FFA6FFE9FD11BC525D8CA007" box="[651,714,352,375]" captionStart="Fig" captionStartId="4.[151,190,1639,1662]" captionTargetBox="[151,1172,397,1610]" captionTargetId="figure-135@4.[151,1172,397,1610]" captionTargetPageId="4" captionText="Fig. 2. Burmomiles volpei sp. nov. in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13203166" httpUri="https://zenodo.org/record/13203166/files/figure.png" pageId="3" pageNumber="334">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA6FFE9FD19BCA55CE4A3AA" pageId="3" pageNumber="334" type="materials_examined">
<paragraph id="8BCC643DFFA6FFE9FD19BCA55B69A35E" blockId="3.[643,1134,407,558]" pageId="3" pageNumber="334">
<materialsCitation id="3B1B6E60FFA6FFE9FD19BCA55B69A35E" collectingDate="2021-12" collectionCode="BUB" county="Male" location="Catalog" municipality="Burmese" pageId="3" pageNumber="334" specimenCode="BUB4027" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFA6FFE9FD19BCA55DB7A0DF" bold="true" box="[643,753,407,431]" pageId="3" pageNumber="334">
<typeStatus id="54C8DA9FFFA6FFE9FD19BCA55DABA0DF" box="[643,749,407,431]" pageId="3" pageNumber="334" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFA6FFE9FD6DBCA55C76A0DE" box="[759,816,407,430]" pageId="3" pageNumber="334">Male</collectingCounty>
, adult specimen in a
<collectingMunicipality id="6BA8FE47FFA6FFE9FB95BCA55B28A0DE" box="[1039,1134,407,430]" pageId="3" pageNumber="334">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFA6FFE9FD10BC855D9CA0BE" box="[650,730,439,462]" country="Myanmar" name="Kachin" pageId="3" pageNumber="334">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFA6FFE9FCE9BC855C80A0BE" LSID="urn:lsid:plazi:treatment:03DAD52BFFA6FFEFFD19BC725FE2A406:8EAC32E6FFA6FFE9FCE9BC855C80A0BE" box="[883,966,439,462]" county="Male" municipality="Burmese" name="Catalog" pageId="3" pageNumber="334" stateProvince="Kachin">Catalog</location>
number SNSB­BSPG
<date id="FFCD42FDFFA6FFE9FD54BCE55C74A09E" box="[718,818,471,494]" pageId="3" pageNumber="334" value="2021-12">
<collectingDate id="EF89BB15FFA6FFE9FD54BCE55C74A09E" box="[718,818,471,494]" pageId="3" pageNumber="334" value="2021-12">2021 XII</collectingDate>
</date>
<quantity id="4C8BC9D8FFA6FFE9FCA1BCE55C32A09E" box="[827,884,471,494]" metricMagnitude="0" metricUnit="m" metricValue="1.8796" pageId="3" pageNumber="334" unit="in" value="74.0">74 in</quantity>
the
<location id="8EAC32E6FFA6FFE9FC3CBCE55DB4A37E" LSID="urn:lsid:plazi:treatment:03DAD52BFFA6FFEFFD19BC725FE2A406:8EAC32E6FFA6FFE9FC3CBCE55DB4A37E" county="Male" municipality="Burmese" name="Bayerische Staatssammlung" pageId="3" pageNumber="334" stateProvince="Kachin">Bayerische Staatssammlung</location>
für
<location id="8EAC32E6FFA6FFE9FC87BCC55CF6A37E" LSID="urn:lsid:plazi:treatment:03DAD52BFFA6FFEFFD19BC725FE2A406:8EAC32E6FFA6FFE9FC87BCC55CF6A37E" box="[797,944,503,526]" county="Male" municipality="Burmese" name="Palaontologie" pageId="3" pageNumber="334" stateProvince="Kachin">Paläontologie</location>
und
<location id="8EAC32E6FFA6FFE9FC7EBCC55B00A37E" LSID="urn:lsid:plazi:treatment:03DAD52BFFA6FFEFFD19BC725FE2A406:8EAC32E6FFA6FFE9FC7EBCC55B00A37E" box="[996,1094,503,526]" county="Male" municipality="Burmese" name="Geologie" pageId="3" pageNumber="334" stateProvince="Kachin">Geologie</location>
(ex
<location id="8EAC32E6FFA6FFE9FD19BF255D99A35E" LSID="urn:lsid:plazi:treatment:03DAD52BFFA6FFEFFD19BC725FE2A406:8EAC32E6FFA6FFE9FD19BF255D99A35E" box="[643,735,535,558]" county="Male" municipality="Burmese" name="Muller's" pageId="3" pageNumber="334" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFA6FFE9FC29BF255B62A35E" box="[947,1060,535,558]" collectionCode="BUB" pageId="3" pageNumber="334">BUB4027</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFA6FFE9FD19BF7F5C93A3F4" blockId="3.[643,1134,589,644]" pageId="3" pageNumber="334">
<emphasis id="B907B82FFFA6FFE9FD19BF7F5C5AA315" bold="true" box="[643,796,589,613]" pageId="3" pageNumber="334">Type locality.</emphasis>
<collectingCountry id="F36424ADFFA6FFE9FCBEBF7F5CD6A314" box="[804,912,589,612]" name="Myanmar" pageId="3" pageNumber="334">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFA6FFE9FC01BF7F5B63A314" box="[923,1061,589,612]" country="Myanmar" name="Kachin" pageId="3" pageNumber="334">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFA6FFE9FD19BF915CE4A3AA" blockId="3.[643,1134,675,730]" pageId="3" pageNumber="334">
<emphasis id="B907B82FFFA6FFE9FD19BF915C50A3CB" bold="true" box="[643,790,675,699]" pageId="3" pageNumber="334">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA6FFE9FD19BFCA5B0DA580" pageId="3" pageNumber="334" type="diagnosis">
<paragraph id="8BCC643DFFA6FFE9FD19BFCA5B0DA580" blockId="3.[643,1134,760,1264]" pageId="3" pageNumber="334">
<emphasis id="B907B82FFFA6FFE9FD19BFCA5C38A260" bold="true" box="[643,894,760,784]" pageId="3" pageNumber="334">Differential diagnosis.</emphasis>
Head rounded behind the eyes, short elytra that do not reach the last abdominal segments and which are equipped with punctuation gathered in striae, plus a pronotum rounded at the sides, make this new species belong to the genus
<taxonomicName id="4C731FBEFFA6FFE9FC30BEAB5B6DA2C0" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[938,1067,921,944]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFA6FFE9FC30BEAB5B6DA2C0" box="[938,1067,921,944]" italics="true" pageId="3" pageNumber="334">Burmomiles</emphasis>
</taxonomicName>
(
<taxonomicName id="4C731FBEFFA6FFE9FBA0BEAB5D98A2A0" authorityName="Imhoff" authorityYear="1856" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
).
<taxonomicName id="4C731FBEFFA6FFE9FD6BBE8B5CFAA2A0" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[753,956,953,976]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="species" species="volpei" status="sp. nov.">
<emphasis id="B907B82FFFA6FFE9FD6BBE8B5CFAA2A0" box="[753,956,953,976]" italics="true" pageId="3" pageNumber="334">Burmomiles volpei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFA6FFE9FC5CBE8B5B5DA2A0" box="[966,1051,953,976]" pageId="3" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
is very similar to
<taxonomicName id="4C731FBEFFA6FFE9FD6DBEEB5C9CA560" authority="Fanti, Damgaard &amp; Ellenberger, 2018" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="species" species="willerslevorum">
<emphasis id="B907B82FFFA6FFE9FD6DBEEB5B60A280" box="[759,1062,985,1008]" italics="true" pageId="3" pageNumber="334">Burmomiles willerslevorum</emphasis>
<bibRefCitation id="EFE219CCFFA6FFE9FBAABEEB5C9CA560" author="Fanti F. &amp; Damgaard A. L. &amp; Ellenberger S." pageId="3" pageNumber="334" pagination="170 - 177" refId="ref20442" refString="Fanti F., Damgaard A. L., Ellenberger S. 2018. Two new genera of Cantharidae from Burmese amber of the Hukawng Valley (Insecta, Coleoptera). Cretaceous Research 86: 170 - 177." type="journal article" year="2018">Fanti, Damgaard &amp; Ellenberger, 2018</bibRefCitation>
</taxonomicName>
, but it differs based on the slightly longer elytra, and by its pronotum less protruded at the sides and less protruded in the middle of the anterior margin (Fanti et al. 2018). Instead,
<taxonomicName id="4C731FBEFFA6FFE9FC0FB94B5CD6A5C0" authority="Fanti &amp; Damgaard, 2019" authorityName="Fanti &amp; Damgaard" authorityYear="2019" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="species" species="blixenae">
<emphasis id="B907B82FFFA6FFE9FC0FB94B5B2BA5E0" box="[917,1133,1145,1168]" italics="true" pageId="3" pageNumber="334">Burmomiles blixenae</emphasis>
<bibRefCitation id="EFE219CCFFA6FFE9FD19B9AB5CD6A5C0" author="Fanti F. &amp; Damgaard A. L." box="[643,912,1177,1200]" pageId="3" pageNumber="334" pagination="101 - 125" refId="ref20361" refString="Fanti F., Damgaard A. L. 2019. New soldier beetles (Cantharidae) from Baltic, Burmese and Dominican ambers of the Anders Damgaard amber collection. Baltic Journal of Coleopterology 19 (2): 101 - 125." type="journal article" year="2019">Fanti &amp; Damgaard, 2019</bibRefCitation>
</taxonomicName>
has a pronotum that is more bordered and shorter elytra with a different sculpture (
<bibRefCitation id="EFE219CCFFA6FFE9FCADB9EB5B06A580" author="Fanti F. &amp; Damgaard A. L." box="[823,1088,1241,1264]" pageId="3" pageNumber="334" pagination="101 - 125" refId="ref20361" refString="Fanti F., Damgaard A. L. 2019. New soldier beetles (Cantharidae) from Baltic, Burmese and Dominican ambers of the Anders Damgaard amber collection. Baltic Journal of Coleopterology 19 (2): 101 - 125." type="journal article" year="2019">Fanti &amp; Damgaard 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA6FFEFFD19B83C5E25A542" lastPageId="5" lastPageNumber="336" pageId="3" pageNumber="334" type="description">
<paragraph id="8BCC643DFFA6FFE9FD19B83C5C2CA415" blockId="3.[643,1134,1294,1381]" pageId="3" pageNumber="334">
<emphasis id="B907B82FFFA6FFE9FD19B83C5C48A456" bold="true" box="[643,782,1294,1318]" pageId="3" pageNumber="334">Description.</emphasis>
Adult, alate, rather robust. Male, for the shape of last sternite. Entirely dark brown. Body length:
<quantity id="4C8BC9D8FFA6FFE9FC88B87C5C21A415" box="[786,871,1358,1381]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="334" unit="mm" value="3.5">3.5 mm</quantity>
.
</paragraph>
<paragraph id="8BCC643DFFA6FFEFFD19B8B75E25A542" blockId="3.[643,1134,1413,1692]" lastBlockId="5.[113,605,155,1074]" lastPageId="5" lastPageNumber="336" pageId="3" pageNumber="334">Head rounded behind the eyes, partially covered by the pronotum, surface with sparce punctation. Eyes very prominent, rounded, inserted laterally to the head. Mandibles falciform. Maxillary palps 4­ segmented, with the last palpomere securiform. Labial palps 3­ segmented with the last palpomere securiform. Antennae 11­ segmented, pectinate, inserted in the frons, relatively long, surpassing the half of the elytra; scape club­shaped, not particularly robust, slightly enlarged apically; antennomere II very short, globular; antennomere III short, slightly longer than second, triangular­shaped; antennomeres IVIX subequal, robust, swollen, over two times longer than antennomeres IIIII together, each antennomere with a long process inserted near the base; antennomere X leaner than previous ones but subequal in length, with a long process near the base; antennomere XI elongate, sturdier, squadrate apically; all antennomeres with setae.Pronotum transverse, wider than head, anterior margin rounded and slightly bordered, posterior margin almost straight and bordered, sides rather curved, surface almost flat with pubescence. Scutellum triangular­shaped, elongated, with a subtruncate apex. Elytra wider than pronotum, robust, rounded at apex, not covering the last abdominal segment (and very slightly the penultimate), slightly wider at humeri, surface presenting a coarse and very wide punctation gathered in evident striae visible for the entire elytral length. Metathoracic wings slightly infuscate, longer than elytra. Sternum elongate and robust. Sternites transverse, last tergite as a caudal appendage which is relative short and rounded apically, last sternite narrower than last tergite. Legs long and slender with short and fitted pubescence; coxae short, robust; trochanters elongated with rounded apex; femora curved, sturdier than tibiae; tibiae thin, elongated, cylindrical, longer than femora, with a long spur at apex. Tarsal formula 5­ 5­ 5; first tarsomere elongated, about 1.4 times longer than second; second tarsomere elongated (pro­ and mesotarsomere triangular­shaped); third tarsomere triangular, shorter than previous one; fourth tarsomere lobated at corners; fifth tarsomere very elongated, thin, curved, slightly enlarged apically; claws simple, very long, apparently without tooth at the base (with the base not enlarged).</paragraph>
</subSubSection>
<caption id="DF0C34B5FFA1FFEEFF0DBB555E59A7EE" ID-DOI="http://doi.org/10.5281/zenodo.13203166" ID-Zenodo-Dep="13203166" httpUri="https://zenodo.org/record/13203166/files/figure.png" pageId="4" pageNumber="335" startId="4.[151,190,1639,1662]" targetBox="[151,1172,397,1610]" targetPageId="4" targetType="figure">
<paragraph id="8BCC643DFFA1FFEEFF0DBB555E59A7EE" blockId="4.[151,1172,1639,1694]" pageId="4" pageNumber="335">
Fig. 2.
<taxonomicName id="4C731FBEFFA1FFEEFF7CBB5A5EF6A70F" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[230,432,1640,1663]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="volpei" status="sp. nov.">
<emphasis id="B907B82FFFA1FFEEFF7CBB5A5EF6A70F" box="[230,432,1640,1663]" italics="true" pageId="4" pageNumber="335">Burmomiles volpei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFA1FFEEFE22BB555D4AA70E" box="[440,524,1639,1662]" pageId="4" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FFA0FFEFFFEBB9635EBFA5AF" pageId="5" pageNumber="336" type="etymology">
<paragraph id="8BCC643DFFA0FFEFFFEBB9635FAAA5D8" blockId="5.[113,605,1105,1192]" pageId="5" pageNumber="336">
<emphasis id="B907B82FFFA0FFEFFFEBB9635FB4A519" bold="true" box="[113,242,1105,1129]" pageId="5" pageNumber="336">Etymology.</emphasis>
Named in honor of Felice Volpe (Montella, Avellino,
<collectingCountry id="F36424ADFFA0FFEFFECEB9435ECFA5F8" box="[340,393,1137,1160]" name="Italy" pageId="5" pageNumber="336">Italy</collectingCountry>
), dear friend of the first author.
</paragraph>
<paragraph id="8BCC643DFFA0FFEFFFEBB9FA5EBFA5AF" blockId="5.[113,505,1224,1248]" box="[113,505,1224,1248]" pageId="5" pageNumber="336">
<emphasis id="B907B82FFFA0FFEFFFEBB9FA5E52A590" bold="true" box="[113,276,1224,1248]" pageId="5" pageNumber="336">Syninclusions.</emphasis>
Air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA0FFEFFFEBB9CC5FE2A406" pageId="5" pageNumber="336" type="discussion">
<paragraph id="8BCC643DFFA0FFEFFFEBB9CC5FE2A406" blockId="5.[113,605,1278,1398]" pageId="5" pageNumber="336">
<emphasis id="B907B82FFFA0FFEFFFEBB9CC5F9BA466" bold="true" box="[113,221,1278,1302]" pageId="5" pageNumber="336">Remarks.</emphasis>
Piece of rounded, clear, golden amber that measures 12x12x
<quantity id="4C8BC9D8FFA0FFEFFEFAB82D5EE7A446" box="[352,417,1311,1334]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="336" unit="mm" value="3.0">3 mm</quantity>
. To the inclusion lacks almost completely the legs of the right side.
</paragraph>
</subSubSection>
</treatment>
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