164 lines
19 KiB
XML
164 lines
19 KiB
XML
<document id="8E46D934B79E3A93E27577A2152D7F26" ID-DOI="10.5281/zenodo.13203160" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203160" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio & Müller, Patrick" docDate="2022" docId="03DAD52BFFA4FFE9FD19BFD85C59A062" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Burmomiles parisii Fanti & Müller, 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="334" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="332" updateTime="1722697113453" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:titleInfo id="7B1429D0B5E6CBC7845606B8D8336438">
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<mods:title id="41427696499B0812E5BD5B7870E7F4F4">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="41C446530183F5C5C3FA9AAAF1E753A4">Fanti, Fabrizio</mods:namePart>
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<mods:roleTerm id="26FC0D1C8E8B05CAA2FC20DE88D633DD">Author</mods:roleTerm>
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<mods:namePart id="29890B2C80CB240CCB578BC47C493283">Müller, Patrick</mods:namePart>
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<mods:typeOfResource id="DC1561BEC13D0F5D608ED787B759C2CF">text</mods:typeOfResource>
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<mods:title id="836F23D679304315C8D9042A23BBAC31">Baltic Journal of Coleopterology</mods:title>
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</mods:titleInfo>
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<mods:part id="CEC8309105BB54EFC464BEAEC6BCECC4">
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<mods:date id="A3A43C70B68ED1B8F4034331035203C5">2022</mods:date>
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<mods:detail id="C8614E965EFF981F01B3519B21493227" type="volume">
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<mods:number id="E4AD73BEDA4E05682174D7971617F879">22</mods:number>
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<mods:number id="ED072861A6F171EAEDDC7624826D212F">2</mods:number>
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<mods:start id="4BE709B184031FA3D03EFF62D1E96DAC">331</mods:start>
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<mods:identifier id="74B1F78EAA73EE43723ECD8E17D53C37" type="DOI">10.5281/zenodo.13203160</mods:identifier>
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<mods:identifier id="713F6382D1CFAD2F659B2ECCC5EA36FA" type="ISSN">1407-8619</mods:identifier>
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<treatment id="03DAD52BFFA4FFE9FD19BFD85C59A062" LSID="urn:lsid:plazi:treatment:03DAD52BFFA4FFE9FD19BFD85C59A062" httpUri="http://treatment.plazi.org/id/03DAD52BFFA4FFE9FD19BFD85C59A062" lastPageId="3" lastPageNumber="334" pageId="1" pageNumber="332">
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<subSubSection id="C36937B6FFA4FFEBFD19BFD85CF4A272" box="[643,946,746,770]" pageId="1" pageNumber="332" type="nomenclature">
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<paragraph id="8BCC643DFFA4FFEBFD19BFD85CF4A272" blockId="1.[643,946,746,801]" box="[643,946,746,770]" pageId="1" pageNumber="332">
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<heading id="D084D351FFA4FFEBFD19BFD85CF4A272" bold="true" box="[643,946,746,770]" fontSize="10" level="2" pageId="1" pageNumber="332" reason="4">
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<emphasis id="B907B82FFFA4FFEBFD19BFD85CF4A272" bold="true" box="[643,946,746,770]" pageId="1" pageNumber="332">
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<taxonomicName id="4C731FBEFFA4FFEBFD19BFD85C10A272" authorityName="Fanti & Müller" authorityYear="2022" box="[643,854,746,770]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="species" species="parisii" status="sp. nov.">
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<emphasis id="B907B82FFFA4FFEBFD19BFD85C10A272" bold="true" box="[643,854,746,770]" italics="true" pageId="1" pageNumber="332">Burmomiles parisii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FFA4FFEBFCC7BFD85CF4A272" box="[861,946,746,770]" pageId="1" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFA4FFEBFD19BE385D95A251" box="[643,723,778,801]" pageId="1" pageNumber="332" type="description">
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<paragraph id="8BCC643DFFA4FFEBFD19BE385D95A251" blockId="1.[643,946,746,801]" box="[643,723,778,801]" pageId="1" pageNumber="332">
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(
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<figureCitation id="134878B8FFA4FFEBFD11BE385D8EA251" box="[651,712,778,801]" captionStart="Fig" captionStartId="2.[151,190,1639,1662]" captionTargetBox="[151,1172,384,1609]" captionTargetId="figure-18@2.[151,1172,384,1609]" captionTargetPageId="2" captionText="Fig. 1. Burmomiles parisii sp. nov. in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13203164" httpUri="https://zenodo.org/record/13203164/files/figure.png" pageId="1" pageNumber="332">Fig. 1</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFA4FFEBFD19BE725CF1A5F5" pageId="1" pageNumber="332" type="materials_examined">
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<paragraph id="8BCC643DFFA4FFEBFD19BE725C46A2A8" blockId="1.[643,1135,832,984]" pageId="1" pageNumber="332">
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<materialsCitation id="3B1B6E60FFA4FFEBFD19BE725C46A2A8" collectionCode="BUB" county="Female" location="Catalog" municipality="Burmese" pageId="1" pageNumber="332" specimenCode="BUB4457" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
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<emphasis id="B907B82FFFA4FFEBFD19BE725DA9A228" bold="true" box="[643,751,832,856]" pageId="1" pageNumber="332">
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<typeStatus id="54C8DA9FFFA4FFEBFD19BE725DACA228" box="[643,746,832,856]" pageId="1" pageNumber="332" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectingCounty id="62AD1CB1FFA4FFEBFD69BE735C04A228" box="[755,834,833,856]" pageId="1" pageNumber="332">Female</collectingCounty>
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, adult specimen in a
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<collectingMunicipality id="6BA8FE47FFA4FFEBFB8BBE735B28A228" box="[1041,1134,833,856]" pageId="1" pageNumber="332">Burmese</collectingMunicipality>
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(
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<collectingRegion id="49B7AADFFFA4FFEBFD10BE535D9FA208" box="[650,729,865,888]" country="Myanmar" name="Kachin" pageId="1" pageNumber="332">Kachin</collectingRegion>
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) amber piece:
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<location id="8EAC32E6FFA4FFEBFCF4BE535C86A208" LSID="urn:lsid:plazi:treatment:03DAD52BFFA4FFE9FD19BFD85C59A062:8EAC32E6FFA4FFEBFCF4BE535C86A208" box="[878,960,865,888]" county="Female" municipality="Burmese" name="Catalog" pageId="1" pageNumber="332" stateProvince="Kachin">Catalog</location>
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number
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<quantity id="4C8BC9D8FFA4FFEBFB83BE535D8BA2E8" metricMagnitude="2" metricUnit="m" metricValue="1.525016" pageId="1" pageNumber="332" unit="in" value="6004.0">SNHM6004 in</quantity>
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the
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<location id="8EAC32E6FFA4FFEBFD69BEB35C5DA2C8" LSID="urn:lsid:plazi:treatment:03DAD52BFFA4FFE9FD19BFD85C59A062:8EAC32E6FFA4FFEBFD69BEB35C5DA2C8" county="Female" municipality="Burmese" name="Staatliches Naturhistorisches Museum Braunschweig" pageId="1" pageNumber="332" stateProvince="Kachin">Staatliches Naturhistorisches Museum Braunschweig</location>
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(ex
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<location id="8EAC32E6FFA4FFEBFCDDBE935CE4A2C8" LSID="urn:lsid:plazi:treatment:03DAD52BFFA4FFE9FD19BFD85C59A062:8EAC32E6FFA4FFEBFCDDBE935CE4A2C8" box="[839,930,929,952]" county="Female" municipality="Burmese" name="Muller's" pageId="1" pageNumber="332" stateProvince="Kachin">Müller’s</location>
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collection number:
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<specimenCode id="DBD5CC46FFA4FFEBFD19BEF35DB2A2A8" box="[643,756,961,984]" collectionCode="BUB" pageId="1" pageNumber="332">BUB4457</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BCC643DFFA4FFEBFD19BEC55C93A55E" blockId="1.[643,1134,1015,1070]" pageId="1" pageNumber="332">
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<emphasis id="B907B82FFFA4FFEBFD19BEC55C5AA57F" bold="true" box="[643,796,1015,1039]" pageId="1" pageNumber="332">Type locality.</emphasis>
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<collectingCountry id="F36424ADFFA4FFEBFCBEBEC55CD6A57E" box="[804,912,1015,1038]" name="Myanmar" pageId="1" pageNumber="332">Myanmar</collectingCountry>
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:
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<collectingRegion id="49B7AADFFFA4FFEBFC01BEC55B63A57E" box="[923,1061,1015,1038]" country="Myanmar" name="Kachin" pageId="1" pageNumber="332">Kachin state</collectingRegion>
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, Myitkyina District, Hukawng Valley.
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</paragraph>
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<paragraph id="8BCC643DFFA4FFEBFD19B97F5CF1A5F5" blockId="1.[643,1134,1101,1157]" pageId="1" pageNumber="332">
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<emphasis id="B907B82FFFA4FFEBFD19B97F5C5BA515" bold="true" box="[643,797,1101,1125]" pageId="1" pageNumber="332">Type horizon.</emphasis>
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Lowermost Cenomanian (98.79 ± 0.62 Ma), midCretaceous.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFA4FFEBFD19B9965C45A7EC" pageId="1" pageNumber="332" type="diagnosis">
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<paragraph id="8BCC643DFFA4FFEBFD19B9965C45A7EC" blockId="1.[643,1137,1188,1692]" pageId="1" pageNumber="332">
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<emphasis id="B907B82FFFA4FFEBFD19B9965C3FA5CC" bold="true" box="[643,889,1188,1212]" pageId="1" pageNumber="332">Differential diagnosis.</emphasis>
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The genus
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<taxonomicName id="4C731FBEFFA4FFEBFC74B9975B28A5CC" authorityName="Fanti, Damgaard & Ellenberger" authorityYear="2018" box="[1006,1134,1189,1212]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
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<emphasis id="B907B82FFFA4FFEBFC74B9975B28A5CC" box="[1006,1134,1189,1212]" italics="true" pageId="1" pageNumber="332">Burmomiles</emphasis>
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</taxonomicName>
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is characterized by short elytra that do not reach the last abdominal segments, tarsal formula 5 5 5, elytra with shallow but dense punctuation arranged in striae, a pronotum enlarged and rounded at sides, the head rounded, and the unequal maxillary palpomeres with the last segment securiform (Fanti et al. 2018), all features present in the new species described here. The new taxon differs from other
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<taxonomicName id="4C731FBEFFA4FFEBFC2BB8F75B77A4AC" authorityName="Fanti, Damgaard & Ellenberger" authorityYear="2018" box="[945,1073,1477,1500]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
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<emphasis id="B907B82FFFA4FFEBFC2BB8F75B77A4AC" box="[945,1073,1477,1500]" italics="true" pageId="1" pageNumber="332">Burmomiles</emphasis>
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</taxonomicName>
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by its much more transverse pronotum. Furthermore, the other
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<taxonomicName id="4C731FBEFFA4FFEBFD7FBB375C20A76C" authorityName="Fanti, Damgaard & Ellenberger" authorityYear="2018" box="[741,870,1541,1564]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
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<emphasis id="B907B82FFFA4FFEBFD7FBB375C20A76C" box="[741,870,1541,1564]" italics="true" pageId="1" pageNumber="332">Burmomiles</emphasis>
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</taxonomicName>
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have pectinate antennae.
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<taxonomicName id="4C731FBEFFA4FFEBFD19BB175B28A74C" authority="Yang, Geiser & Liu, 2021" authorityName="Yang, Geiser & Liu" authorityYear="2021" box="[643,1134,1573,1596]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="species" species="laticollis">
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<emphasis id="B907B82FFFA4FFEBFD19BB175C24A74C" box="[643,866,1573,1596]" italics="true" pageId="1" pageNumber="332">Burmomiles laticollis</emphasis>
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<bibRefCitation id="EFE219CCFFA4FFEBFCFCBB175B28A74C" author="Yang Y. & Zhao W. & Geiser M. F. & Zhang W. & Bai M. & Liu H." box="[870,1134,1573,1596]" pageId="1" pageNumber="332" refId="ref21251" refString="Yang Y., Zhao W., Geiser M. F., Zhang W., Bai M., Liu H. 2021. Review of the genus Burmomiles Fanti, Damgaard & Ellenberger from mid-Cretaceous Burmese amber (Coleoptera, Cantharidae). Cretaceous Research 128: 104961 [9 pp.]." type="book" year="2021">Yang, Geiser & Liu, 2021</bibRefCitation>
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</taxonomicName>
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has a similar pronotum, but it has shorter elytra (less punctated) and pectinate antennae (
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<bibRefCitation id="EFE219CCFFA4FFEBFBAFBB575DBEA7EC" author="Yang Y. & Zhao W. & Geiser M. F. & Zhang W. & Bai M. & Liu H." pageId="1" pageNumber="332" refId="ref21251" refString="Yang Y., Zhao W., Geiser M. F., Zhang W., Bai M., Liu H. 2021. Review of the genus Burmomiles Fanti, Damgaard & Ellenberger from mid-Cretaceous Burmese amber (Coleoptera, Cantharidae). Cretaceous Research 128: 104961 [9 pp.]." type="book" year="2021">Yang et al. 2021</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFA7FFE9FF0DBDA95E55A4B7" lastPageId="3" lastPageNumber="334" pageId="2" pageNumber="333" type="description">
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<paragraph id="8BCC643DFFA7FFE8FF0DBDA95D24A184" blockId="2.[151,644,155,244]" pageId="2" pageNumber="333">
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<emphasis id="B907B82FFFA7FFE8FF0DBDA95E65A1C3" bold="true" box="[151,291,155,179]" pageId="2" pageNumber="333">Description.</emphasis>
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Adult, alate, robust. Female, for the shape of last sternite and short antennae. Entirely dark brown. Body length: 3.0 mm.
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</paragraph>
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<paragraph id="8BCC643DFFA7FFE9FF0DBC265E55A4B7" blockId="2.[151,643,276,332]" lastBlockId="3.[113,606,155,1479]" lastPageId="3" lastPageNumber="334" pageId="2" pageNumber="333">Head rounded behind the eyes, partially covered by the pronotum, surface with shallow punctation. Eyes prominent, subelliptical, inserted laterally to the head. Mandibles falciform. Maxillary palps 4 segmented, with the last palpomere securiform. Labial palps 3 segmented with the last palpomere securiform. Antennae 11 segmented, filiform, inserted in the upper part of the head and not particularly close to the eyes, short, slightly surpassing the humeral zone of the elytra; scape clubshaped, very robust; antennomere II short, very wide and globular; antennomere III narrower than previous one and globular; antennomeres IV–X subequal, robust, swollen, longer than antennomeres II–III together; antennomere XI elongate, pointed apically; all antennomeres with setae. Pronotum very transverse, wider than head, anterior margin straight and slightly bordered, posterior margin almost straight, sides slightly curved and slightly bordered, surface completely flat with pubescence (just perceptible). Scutellum triangularshaped with rounded apex. Elytra wider than pronotum, robust and wide, rounded at apex, not covering the last abdominal segment, wider at humeri, surface presenting a coarse and wide punctation gathered in evident striae visible for the entire elytral length. Metathoracic wings covered by elytra, and not visible. Sternum subquadrate with almost straight posterior margin. Sternites transverse and wide, last two tergites as a caudal large appendage, last sternite rounded presenting two slender and divergent lobes (oviduct). Legs very robust and with short and fitted pubescence; coxae short, robust; trochanters elongated with rounded apex; femora slightly curved, sturdier than tibiae; tibiae thin, elongate, cylindrical and slightly flat apically, pro and mesotibiae as long as pro and mesofemora, metatibiae longer than metafemora, with a robust spur at apex. Tarsal formula 5 5 5; first tarsomere about 1.3 times longer than second; second tarsomere elongated; third tarsomere triangular, very slightly enlarged at corners; fourth tarsomere flat, wide, very slightly lobated at corners; fifth tarsomere very elongated, thin, slightly curved, slightly enlarged apically; claws simple, very long, apparently without tooth at the base (with the base slightly wider).</paragraph>
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</subSubSection>
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<caption id="DF0C34B5FFA7FFE8FF0DBB555E59A7EE" ID-DOI="http://doi.org/10.5281/zenodo.13203164" ID-Zenodo-Dep="13203164" httpUri="https://zenodo.org/record/13203164/files/figure.png" pageId="2" pageNumber="333" startId="2.[151,190,1639,1662]" targetBox="[151,1172,384,1609]" targetPageId="2" targetType="figure">
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||
<paragraph id="8BCC643DFFA7FFE8FF0DBB555E59A7EE" blockId="2.[151,1172,1639,1694]" pageId="2" pageNumber="333">
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||
Fig. 1.
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<taxonomicName id="4C731FBEFFA7FFE8FF7FBB555EF2A70E" authorityName="Fanti & Müller" authorityYear="2022" box="[229,436,1639,1662]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="333" phylum="Arthropoda" rank="species" species="parisii" status="sp. nov.">
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<emphasis id="B907B82FFFA7FFE8FF7FBB555EF2A70E" box="[229,436,1639,1662]" italics="true" pageId="2" pageNumber="333">Burmomiles parisii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FFA7FFE8FE26BB555D49A70E" box="[444,527,1639,1662]" pageId="2" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
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in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view.
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</paragraph>
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</caption>
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<subSubSection id="C36937B6FFA6FFE9FFEBB8D55E4CA7E7" pageId="3" pageNumber="334" type="etymology">
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<paragraph id="8BCC643DFFA6FFE9FFEBB8D55E4DA730" blockId="3.[113,605,1511,1600]" pageId="3" pageNumber="334">
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<emphasis id="B907B82FFFA6FFE9FFEBB8D55FA8A48F" bold="true" box="[113,238,1511,1535]" pageId="3" pageNumber="334">Etymology.</emphasis>
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Named in honor of Francesco Parisi (University of Florence,
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<collectingCountry id="F36424ADFFA6FFE9FEF4BB3A5EE7A76F" box="[366,417,1544,1567]" name="Italy" pageId="3" pageNumber="334">Italy</collectingCountry>
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), friend of the first author (Fanti).
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</paragraph>
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<paragraph id="8BCC643DFFA6FFE9FFEBBB525E4CA7E7" blockId="3.[113,605,1632,1687]" pageId="3" pageNumber="334">
|
||
<emphasis id="B907B82FFFA6FFE9FFEBBB525E48A708" bold="true" box="[113,270,1632,1656]" pageId="3" pageNumber="334">Syninclusions.</emphasis>
|
||
Air bubbles, detritus, Hymenoptera (
|
||
<quantity id="4C8BC9D8FFA6FFE9FF3DBBB25FB8A7E7" box="[167,254,1664,1687]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="334" unit="mm" value="0.7">0.7 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36937B6FFA6FFE9FD19BDA85C59A062" pageId="3" pageNumber="334" type="discussion">
|
||
<paragraph id="8BCC643DFFA6FFE9FD19BDA85C59A062" blockId="3.[643,1134,154,274]" pageId="3" pageNumber="334">
|
||
<emphasis id="B907B82FFFA6FFE9FD19BDA85DB4A1C2" bold="true" box="[643,754,154,178]" pageId="3" pageNumber="334">Remarks.</emphasis>
|
||
Piece of clear, golden amber that measures 11x10x
|
||
<quantity id="4C8BC9D8FFA6FFE9FCAFBD895CCEA1A2" box="[821,904,187,210]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="334" unit="mm" value="4.5">4.5 mm</quantity>
|
||
. The inclusion is complete, but the pronotum and ventral side are not clearly visible.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |