171 lines
19 KiB
XML
171 lines
19 KiB
XML
<document id="0A69342F9D44A741E00051290741AEC8" ID-DOI="10.5281/zenodo.13203160" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203160" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio & Müller, Patrick" docDate="2022" docId="03DAD52BFF85FFC8FF0DBE635B00A1F0" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Sanaungulus nilsi Fanti & Müller, 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="365" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="363" updateTime="1722697113453" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:mods id="90B42062EB11ACB27EE09DDC47D1D373" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4FBFFF7765F64D89AC43708AF0D2DFAF">
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<mods:title id="9731C021C7536D1C9BB2515735FAB215">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:name id="C2960BC177D7B04FC59E332459FBA5E3" type="personal">
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</mods:role>
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<mods:namePart id="0057477C9E89B946243E9A3DEB6F33E0">Fanti, Fabrizio</mods:namePart>
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<mods:name id="F528F49AE5C85C651871F4AA0F3BC379" type="personal">
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<mods:roleTerm id="F6CA747848852EEB99554E9CB78E97C6">Author</mods:roleTerm>
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<mods:namePart id="0956D1789B4239B406132906B37880DB">Müller, Patrick</mods:namePart>
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<mods:typeOfResource id="097DAA3EA99C8D930099C9E336C6B047">text</mods:typeOfResource>
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<mods:titleInfo id="C4CFDC0D83265285D4546732E649A16F">
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<mods:title id="BF230D255CE4C369FE43EFA1563D6F0F">Baltic Journal of Coleopterology</mods:title>
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</mods:titleInfo>
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<mods:part id="2CDC36004C96A2B6F32BB67C85A2EE83">
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<mods:date id="89D3A5528E91E264D9B1AB8E509BC283">2022</mods:date>
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<mods:detail id="C51254A62E4276C67D55CDB1056BACC5" type="volume">
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<mods:number id="FBA0E5B961D6D51898B4FBCD2C031596">22</mods:number>
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<mods:detail id="2F49A06CE4D15FF74A24ABB21C588FF7" type="issue">
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<mods:number id="3B2723AC84C61349FF9DB5DD66AC0954">2</mods:number>
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<mods:start id="B330B5B98C5A376384D172EDCE2BF67C">331</mods:start>
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<mods:classification id="B9D2B75E87FF1A8404F95D2A6C3F7034">journal article</mods:classification>
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<mods:identifier id="9C21679B5D7F86CB6AFD30CEA476C643" type="DOI">10.5281/zenodo.13203160</mods:identifier>
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<mods:identifier id="73485726C07442152B3B3ACC71341D93" type="ISSN">1407-8619</mods:identifier>
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<mods:identifier id="A2E7C307419D33C53C6EE7F2E18F6925" type="Zenodo-Dep">13203160</mods:identifier>
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</mods:mods>
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<treatment id="03DAD52BFF85FFC8FF0DBE635B00A1F0" LSID="urn:lsid:plazi:treatment:03DAD52BFF85FFC8FF0DBE635B00A1F0" httpUri="http://treatment.plazi.org/id/03DAD52BFF85FFC8FF0DBE635B00A1F0" lastPageId="34" lastPageNumber="365" pageId="32" pageNumber="363">
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<subSubSection id="C36937B6FF85FFCAFF0DBE635EFDA219" box="[151,443,849,873]" pageId="32" pageNumber="363" type="nomenclature">
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<paragraph id="8BCC643DFF85FFCAFF0DBE635EFDA219" blockId="32.[151,443,849,904]" box="[151,443,849,873]" pageId="32" pageNumber="363">
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<heading id="D084D351FF85FFCAFF0DBE635EFDA219" bold="true" box="[151,443,849,873]" fontSize="10" level="2" pageId="32" pageNumber="363" reason="4">
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<emphasis id="B907B82FFF85FFCAFF0DBE635EFDA219" bold="true" box="[151,443,849,873]" pageId="32" pageNumber="363">
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<taxonomicName id="4C731FBEFF85FFCAFF0DBE635E26A219" authorityName="Fanti & Müller" authorityYear="2022" box="[151,352,849,873]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="363" phylum="Arthropoda" rank="species" species="nilsi" status="sp. nov.">
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<emphasis id="B907B82FFF85FFCAFF0DBE635E26A219" bold="true" box="[151,352,849,873]" italics="true" pageId="32" pageNumber="363">Sanaungulus nilsi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FF85FFCAFEFCBE635EFDA219" box="[358,443,849,873]" pageId="32" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FF85FFCAFF0DBE435FB2A2F8" box="[151,244,881,904]" pageId="32" pageNumber="363" type="description">
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<paragraph id="8BCC643DFF85FFCAFF0DBE435FB2A2F8" blockId="32.[151,443,849,904]" box="[151,244,881,904]" pageId="32" pageNumber="363">
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(
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<figureCitation id="134878B8FF85FFCAFF05BE435FAAA2F8" box="[159,236,881,904]" captionStart="Fig" captionStartId="33.[113,152,1409,1432]" captionTargetBox="[113,1134,151,1376]" captionTargetId="figure-28@33.[113,1134,151,1376]" captionTargetPageId="33" captionText="Fig. 18. Sanaungulus nilsi sp. nov. in Burmese amber: A: Holotype, lateral view; B: Holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13203215" httpUri="https://zenodo.org/record/13203215/files/figure.png" pageId="32" pageNumber="363">Fig. 18</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FF85FFCAFF0DBE9A5EF0A59F" pageId="32" pageNumber="363" type="materials_examined">
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<paragraph id="8BCC643DFF85FFCAFF0DBE9A5D02A530" blockId="32.[151,643,936,1088]" pageId="32" pageNumber="363">
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<materialsCitation id="3B1B6E60FF85FFCAFF0DBE9A5D02A530" collectingDate="2021-12" collectionCode="BUB" county="Female" location="Catalog" municipality="Burmese" pageId="32" pageNumber="363" specimenCode="BUB4393" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
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<emphasis id="B907B82FFF85FFCAFF0DBE9A5E42A2B0" bold="true" box="[151,260,936,960]" pageId="32" pageNumber="363">
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<typeStatus id="54C8DA9FFF85FFCAFF0DBE9A5FB8A2B0" box="[151,254,936,960]" pageId="32" pageNumber="363" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectingCounty id="62AD1CB1FF85FFCAFE9DBE9A5E10A2CF" box="[263,342,936,959]" pageId="32" pageNumber="363">Female</collectingCounty>
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, adult specimen in a
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<collectingMunicipality id="6BA8FE47FF85FFCAFDBCBE9A5DC5A2CF" box="[550,643,936,959]" pageId="32" pageNumber="363">Burmese</collectingMunicipality>
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(
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<collectingRegion id="49B7AADFFF85FFCAFF05BEFA5FA9A2AF" box="[159,239,968,991]" country="Myanmar" name="Kachin" pageId="32" pageNumber="363">Kachin</collectingRegion>
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) amber piece:
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<location id="8EAC32E6FF85FFCAFE12BEFA5E9DA2AF" LSID="urn:lsid:plazi:treatment:03DAD52BFF85FFC8FF0DBE635B00A1F0:8EAC32E6FF85FFCAFE12BEFA5E9DA2AF" box="[392,475,968,991]" county="Female" municipality="Burmese" name="Catalog" pageId="32" pageNumber="363" stateProvince="Kachin">Catalog</location>
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number SNSBBSPG
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<date id="FFCD42FDFF85FFCAFF79BEDB5E01A570" box="[227,327,1001,1024]" pageId="32" pageNumber="363" value="2021-12">
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<collectingDate id="EF89BB15FF85FFCAFF79BEDB5E01A570" box="[227,327,1001,1024]" pageId="32" pageNumber="363" value="2021-12">2021 XII</collectingDate>
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</date>
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<quantity id="4C8BC9D8FF85FFCAFECABEDB5ECEA570" box="[336,392,1001,1024]" metricMagnitude="0" metricUnit="m" metricValue="1.6256" pageId="32" pageNumber="363" unit="in" value="64.0">64 in</quantity>
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the
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<location id="8EAC32E6FF85FFCAFE21BEDB5E40A550" LSID="urn:lsid:plazi:treatment:03DAD52BFF85FFC8FF0DBE635B00A1F0:8EAC32E6FF85FFCAFE21BEDB5E40A550" county="Female" municipality="Burmese" name="Bayerische Staatssammlung" pageId="32" pageNumber="363" stateProvince="Kachin">Bayerische Staatssammlung</location>
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für
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<location id="8EAC32E6FF85FFCAFEA8B93B5E83A550" LSID="urn:lsid:plazi:treatment:03DAD52BFF85FFC8FF0DBE635B00A1F0:8EAC32E6FF85FFCAFEA8B93B5E83A550" box="[306,453,1033,1056]" county="Female" municipality="Burmese" name="Palaontologie" pageId="32" pageNumber="363" stateProvince="Kachin">Paläontologie</location>
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und
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<location id="8EAC32E6FF85FFCAFE63B93B5D1DA550" LSID="urn:lsid:plazi:treatment:03DAD52BFF85FFC8FF0DBE635B00A1F0:8EAC32E6FF85FFCAFE63B93B5D1DA550" box="[505,603,1033,1056]" county="Female" municipality="Burmese" name="Geologie" pageId="32" pageNumber="363" stateProvince="Kachin">Geologie</location>
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(ex
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<location id="8EAC32E6FF85FFCAFF0DB91B5FB4A530" LSID="urn:lsid:plazi:treatment:03DAD52BFF85FFC8FF0DBE635B00A1F0:8EAC32E6FF85FFCAFF0DB91B5FB4A530" box="[151,242,1065,1088]" county="Female" municipality="Burmese" name="Muller's" pageId="32" pageNumber="363" stateProvince="Kachin">Müller’s</location>
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collection number:
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<specimenCode id="DBD5CC46FF85FFCAFE5DB91B5D7EA530" box="[455,568,1065,1088]" collectionCode="BUB" pageId="32" pageNumber="363">BUB4393</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BCC643DFF85FFCAFF0DB9525EACA5E7" blockId="32.[151,643,1120,1175]" pageId="32" pageNumber="363">
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<emphasis id="B907B82FFF85FFCAFF0DB9525E76A508" bold="true" box="[151,304,1120,1144]" pageId="32" pageNumber="363">Type locality.</emphasis>
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<collectingCountry id="F36424ADFF85FFCAFEA2B9525EE2A507" box="[312,420,1120,1143]" name="Myanmar" pageId="32" pageNumber="363">Myanmar</collectingCountry>
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:
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<collectingRegion id="49B7AADFFF85FFCAFE2AB9525D7CA507" box="[432,570,1120,1143]" country="Myanmar" name="Kachin" pageId="32" pageNumber="363">Kachin state</collectingRegion>
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, Myitkyina District, Hukawng Valley.
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</paragraph>
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<paragraph id="8BCC643DFF85FFCAFF0DB9855EF0A59F" blockId="32.[151,643,1207,1263]" pageId="32" pageNumber="363">
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<emphasis id="B907B82FFF85FFCAFF0DB9855E6CA5BF" bold="true" box="[151,298,1207,1231]" pageId="32" pageNumber="363">Type horizon.</emphasis>
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Lowermost Cenomanian (98.79 ± 0.62 Ma), midCretaceous.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FF85FFCAFF0DB83C5FA4A4F6" pageId="32" pageNumber="363" type="diagnosis">
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<paragraph id="8BCC643DFF85FFCAFF0DB83C5FA4A4F6" blockId="32.[151,643,1294,1414]" pageId="32" pageNumber="363">
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<emphasis id="B907B82FFF85FFCAFF0DB83C5ED2A456" bold="true" box="[151,404,1294,1318]" pageId="32" pageNumber="363">Differential diagnosis.</emphasis>
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<taxonomicName id="4C731FBEFF85FFCAFE06B83D5D1AA456" authorityName="Fanti & Müller" authorityYear="2022" box="[412,604,1295,1318]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="363" phylum="Arthropoda" rank="species" species="nilsi" status="sp. nov.">
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<emphasis id="B907B82FFF85FFCAFE06B83D5D1AA456" box="[412,604,1295,1318]" italics="true" pageId="32" pageNumber="363">Sanaungulus nilsi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FF85FFCAFDFEB83C5F83A436" pageId="32" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
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differs from other species of
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<taxonomicName id="4C731FBEFF85FFCAFE63B81D5DC5A436" authorityName="Fanti, Damgaard & Ellenberger" authorityYear="2018" box="[505,643,1327,1350]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="363" phylum="Arthropoda" rank="genus">
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<emphasis id="B907B82FFF85FFCAFE63B81D5DC5A436" box="[505,643,1327,1350]" italics="true" pageId="32" pageNumber="363">Sanaungulus</emphasis>
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</taxonomicName>
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with filiform antennae by its very elongated pronotum.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FF85FFCBFF0DB8945DF9A1F0" lastPageId="33" lastPageNumber="364" pageId="32" pageNumber="363" type="description">
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<paragraph id="8BCC643DFF85FFCAFF0DB8945EEFA76E" blockId="32.[151,643,1446,1566]" pageId="32" pageNumber="363">
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<emphasis id="B907B82FFF85FFCAFF0DB8945E65A4CE" bold="true" box="[151,291,1446,1470]" pageId="32" pageNumber="363">Description.</emphasis>
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Adult, winged, slender. Female, defined on the basis of short antennae and large abdomen. Body length
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<quantity id="4C8BC9D8FF85FFCAFE10B8D55E9BA48E" box="[394,477,1511,1534]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="32" pageNumber="363" unit="mm" value="3.3">3.3 mm</quantity>
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. Entirely brown with testaceous abdomen.
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</paragraph>
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<paragraph id="8BCC643DFF85FFCAFF0DBB0C5C40A7E2" blockId="32.[151,643,1598,1685]" lastBlockId="32.[680,1172,155,1682]" pageId="32" pageNumber="363">Head rounded, slightly constricted (triangularshaped) behind the eyes, equipped with dense pubescence and shallow punctation. Eyes prominent, large, roundish, inserted in the upper and lateral part of the head. Mandibles elongated, falciform, without teeth. Maxillary palps 4 segmented with the last palpomere very elongated and securiform. Labial palps 3 segmented, with the last palpomere elongated and securiform. Antennae 11segmented, filiform, robust, short, reaching half of elytra and the first abdominal segment, antennal insertion in the eyes proximity; scape robust, clubshaped; antennomeres II–III short, subequal, thinner than first; antennomere IV elongated, the longest; antennomeres V–X slightly shorter than antennomere IV, subequal, with antennomeres V– VIII sturdier than others; antennomere XI filiform, longer than previous ones, enlarged in the middle, rounded at apex; all the antennomeres with short pubescence. Pronotum very elongated, subrectangular, as wide as head, concave in the middle, anterior margin straight, posterior margin slightly undulated and strongly bordered, sides straight and bordered, corners obtuse, pronotum disc concave and pubescent. Scutellum triangularshaped. Elytra short which reveals various abdominal segments, at humeri slightly wider than pronotum, posteriorly parallelsided and slighty dehiscent apically, rounded apexes, surface pubescent and with a very small rugosity. Metathoracic wings longer than elytra, slightly surpassing the half of abdomen. Sternum elongated with posterior margin rounded, sternites large and transverse, last tergite elongated with rounded apex and narrower than penultimate tergite which is large and transverse, last sternite wide and enlarged at sides and slightly pointed apically. Legs thin, relatively long, pubescent; coxae massive; trochanters elongated, with rounded apex; femora slightly curved, more robust than tibiae, slightly flat; tibiae flattened, thin, with spurs near the apex, protibiae as long as profemora, mesotibiae slightly longer than mesofemora, metatibiae longer than metafemora. Tarsal formula 5 5 5; first tarsomere long, about 0.4 times longer than second; third tarsomere short and slightly bilobed; fourth strongly bilobed; fifth tarsomere elongated, thin, slightly flat and curved; claws simple, without denticle at the base.</paragraph>
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<paragraph id="8BCC643DFF84FFCBFDBABD5F5DF9A1F0" blockId="33.[544,703,109,128]" box="[544,703,109,128]" pageId="33" pageNumber="364">
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<emphasis id="B907B82FFF84FFCBFDBABD5F5DF9A1F0" box="[544,703,109,128]" italics="true" pageId="33" pageNumber="364">Fanti F., Müller P.</emphasis>
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</paragraph>
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||
</subSubSection>
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||
<caption id="DF0C34B5FF84FFCBFFEBB8B35FBCA4C8" ID-DOI="http://doi.org/10.5281/zenodo.13203215" ID-Zenodo-Dep="13203215" httpUri="https://zenodo.org/record/13203215/files/figure.png" pageId="33" pageNumber="364" startId="33.[113,152,1409,1432]" targetBox="[113,1134,151,1376]" targetPageId="33" targetType="figure">
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<paragraph id="8BCC643DFF84FFCBFFEBB8B35FBCA4C8" blockId="33.[113,1134,1409,1464]" pageId="33" pageNumber="364">
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Fig. 18.
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<taxonomicName id="4C731FBEFF84FFCBFF56B8B35ECAA4E8" authorityName="Fanti & Müller" authorityYear="2022" box="[204,396,1409,1432]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="364" phylum="Arthropoda" rank="species" species="nilsi" status="sp. nov.">
|
||
<emphasis id="B907B82FFF84FFCBFF56B8B35ECAA4E8" box="[204,396,1409,1432]" italics="true" pageId="33" pageNumber="364">Sanaungulus nilsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2340554FF84FFCBFE0EB8B35EA1A4E8" box="[404,487,1409,1432]" pageId="33" pageNumber="364" rank="species">sp. nov.</taxonomicNameLabel>
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in Burmese amber: A: Holotype, lateral view; B: Holotype, ventral view.
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</paragraph>
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</caption>
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<subSubSection id="C36937B6FF84FFCBFFEBB8DD5ECDA7F1" pageId="33" pageNumber="364" type="etymology">
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||
<paragraph id="8BCC643DFF84FFCBFFEBB8DD5EF3A758" blockId="33.[113,604,1519,1576]" pageId="33" pageNumber="364">
|
||
<emphasis id="B907B82FFF84FFCBFFEBB8DD5FA9A777" bold="true" box="[113,239,1519,1543]" pageId="33" pageNumber="364">Etymology.</emphasis>
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Named in honor of Nils Müller, son of the second author (Patrick).
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</paragraph>
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<paragraph id="8BCC643DFF84FFCBFFEBBB7A5ECDA7F1" blockId="33.[113,605,1608,1665]" pageId="33" pageNumber="364">
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<emphasis id="B907B82FFF84FFCBFFEBBB7A5E5EA710" bold="true" box="[113,280,1608,1632]" pageId="33" pageNumber="364">Syninclusions.</emphasis>
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<taxonomicName id="4C731FBEFF84FFCBFEB8BB7B5E81A710" box="[290,455,1609,1632]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="364" phylum="Arthropoda" rank="family">Cecidomyiidae</taxonomicName>
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(leg), cicada (leg), air bubbles, detritus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FF84FFC8FD19B8DD5B00A1F0" lastPageId="34" lastPageNumber="365" pageId="33" pageNumber="364" type="discussion">
|
||
<paragraph id="8BCC643DFF84FFCBFD19B8DD5DFBA739" blockId="33.[643,1134,1519,1609]" pageId="33" pageNumber="364">
|
||
<emphasis id="B907B82FFF84FFCBFD19B8DD5DB4A777" bold="true" box="[643,754,1519,1543]" pageId="33" pageNumber="364">Remarks.</emphasis>
|
||
Piece of clear, golden amber that measures 19x7.5x
|
||
<quantity id="4C8BC9D8FF84FFCBFCA1BB235CCAA758" box="[827,908,1553,1576]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="33" pageNumber="364" unit="mm" value="3.3">3.3 mm</quantity>
|
||
. The inclusion is complete.
|
||
</paragraph>
|
||
<paragraph id="8BCC643DFF87FFC8FF7CBD5F5B00A1F0" blockId="34.[230,1094,109,128]" box="[230,1094,109,128]" pageId="34" pageNumber="365">
|
||
<emphasis id="B907B82FFF87FFC8FF7CBD5F5B00A1F0" box="[230,1094,109,128]" italics="true" pageId="34" pageNumber="365">
|
||
Fossil
|
||
<taxonomicName id="4C731FBEFF87FFC8FEBBBD5F5ECBA1F0" authorityName="Imhoff" authorityYear="1856" box="[289,397,109,128]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="365" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
|
||
from the Cretaceous Burmese (
|
||
<collectingRegion id="49B7AADFFF87FFC8FD3ABD5F5D99A1F0" box="[672,735,109,128]" country="Myanmar" name="Kachin" pageId="34" pageNumber="365">Kachin</collectingRegion>
|
||
) amber of the Patrick Müller collection..
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |