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<document id="9A82B35572D7FED5974446352604B7CF" ID-CLB-Dataset="4014" ID-DOI="10.11646/zootaxa.5051.1.8" ID-GBIF-Dataset="1694bae6-36b6-43dd-b533-b40f545aa291" ID-ISSN="1175-5326" ID-Zenodo-Dep="5564091" ID-ZooBank="0B868A45-6804-4EC2-A6F7-4CB521BD7B79" IM.bibliography_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634024638876" checkinUser="plazi" docAuthor="Garlitska, Lesya &amp; Chertoprud, Elena S." docDate="2021" docId="03DA2D322039FFDEFF0C161CD530FBBA" docLanguage="en" docName="zootaxa.5051.1.8.pdf" docOrigin="Zootaxa 5051 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Danielssenia spitsbergensis Gee &amp; Huys 1994" docType="treatment" docVersion="7" lastPageNumber="97" masterDocId="FFE3554A2038FFDDFF9B130BD777FFC7" masterDocTitle="Two rare species of Danielsseniinae (Copepoda: Harpacticoida: Pseudotachidiidae) from the Kara and East Siberian Seas, with description of their previously unknown sexes" masterLastPageNumber="100" masterPageNumber="94" pageNumber="95" updateTime="1699459631203" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="98C50A620C987EC009F15BBA9669A230">Two rare species of Danielsseniinae (Copepoda: Harpacticoida: Pseudotachidiidae) from the Kara and East Siberian Seas, with description of their previously unknown sexes</mods:title>
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<mods:namePart id="E39204E7E3D190A99F4FE0C003F08372">Garlitska, Lesya</mods:namePart>
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<mods:affiliation id="06F7C03E38B121CA17DF9CB17CBE4877">P. P. Shirshov Institute of Oceanology, Russian Academy of Sciences, 36, Nakhimovsky Prospect, 117997 Moscow, Russian Federation. garlitska @ gmail. com; https: // orcid. org / 0000 - 0002 - 1317 - 729 X</mods:affiliation>
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<mods:namePart id="14CC22E03DE15ADD2D8528E253237CCC">Chertoprud, Elena S.</mods:namePart>
<mods:affiliation id="B7CF6318BB47448F26B36A077E0F7A0C">M. V. Lomonosov Moscow State University, Leninskie Gory, Moscow 119991, Russian Federation. horsax @ yandex. ru &amp; A. N. Severtsov Institute of Ecology &amp; Evolution, Leninsky Pr., 33, Moscow 119071, Russian Federation.</mods:affiliation>
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<treatment id="03DA2D322039FFDEFF0C161CD530FBBA" ID-DOI="http://doi.org/10.5281/zenodo.5564059" ID-GBIF-Taxon="188909462" ID-Zenodo-Dep="5564059" LSID="urn:lsid:plazi:treatment:03DA2D322039FFDEFF0C161CD530FBBA" httpUri="http://treatment.plazi.org/id/03DA2D322039FFDEFF0C161CD530FBBA" lastPageId="3" lastPageNumber="97" pageId="1" pageNumber="95">
<subSubSection id="C369CFAF2039FFDCFF0C161CD5BAFAF6" box="[151,717,1303,1330]" pageId="1" pageNumber="95" type="nomenclature">
<paragraph id="8BCC9C242039FFDCFF0C161CD5BAFAF6" blockId="1.[151,717,1303,1365]" box="[151,717,1303,1330]" pageId="1" pageNumber="95">
<heading id="D0842B482039FFDCFF0C161CD5BAFAF6" bold="true" box="[151,717,1303,1330]" fontSize="11" level="1" pageId="1" pageNumber="95" reason="1">
<taxonomicName id="4C73E7A72039FFDCFF0C161CD5BAFAF6" authority="Gee &amp; Huys, 1994" authorityName="Gee &amp; Huys" authorityYear="1994" box="[151,717,1303,1330]" class="Hexanauplia" family="Pseudotachidiidae" genus="Danielssenia" kingdom="Animalia" order="Harpacticoida" pageId="1" pageNumber="95" phylum="Arthropoda" rank="species" species="spitsbergensis">
<emphasis id="B90740362039FFDCFF0C161CD5BAFAF6" bold="true" box="[151,717,1303,1330]" pageId="1" pageNumber="95">
<emphasis id="B90740362039FFDCFF0C161CD694FAF5" bold="true" box="[151,483,1303,1330]" italics="true" pageId="1" pageNumber="95">Danielssenia spitsbergensis</emphasis>
<bibRefCitation id="EFE2E1D52039FFDCFE71161CD5BAFAF6" author="Gee, J. M. &amp; Huys, R." box="[490,717,1303,1329]" pageId="1" pageNumber="95" pagination="1007 - 1046" refId="ref3787" refString="Gee, J. M. &amp; Huys, R. (1994) Paranannopidae (Copepoda: Harpacticoida) from sublittoral soft sediments in Spitzbergen. Journal of natural History, 28, 1007 - 1046. https: // doi. org / 10.1080 / 00222939400770541" type="journal article" year="1994">Gee &amp; Huys, 1994</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="C369CFAF2039FFDEFF0C1630D209FE16" lastPageId="3" lastPageNumber="97" pageId="1" pageNumber="95" type="description">
<paragraph id="8BCC9C242039FFDCFF0C1630D79CFA92" blockId="1.[151,717,1303,1365]" box="[151,235,1339,1365]" pageId="1" pageNumber="95">
(
<figureCitation id="134880A12039FFDCFF3B1630D797FA92" box="[160,224,1339,1365]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[195,1395,192,1909]" captionTargetId="figure-17@2.[184,1403,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Danielssenia spitsbergensis Gee &amp; Huys, 1994, female:A, genital field; B, antennule; C, P2; D, P5." figureDoi="http://doi.org/10.5281/zenodo.5564093" httpUri="https://zenodo.org/record/5564093/files/figure.png" pageId="1" pageNumber="95">Fig. 1</figureCitation>
)
</paragraph>
<paragraph id="8BCC9C242039FFDCFF0C1688D5BCF9CE" blockId="1.[151,1437,1411,2014]" pageId="1" pageNumber="95">
<emphasis id="B90740362039FFDCFF0C1688D6EFFA5A" bold="true" box="[151,408,1411,1437]" pageId="1" pageNumber="95">Specimens examined.</emphasis>
Female dissected on four slides (collection numbers IORAS-Har199202), collected from
<quantity id="4C8B31C12039FFDCFF0C16ACD795FA05" box="[151,226,1447,1474]" metricMagnitude="2" metricUnit="m" metricValue="1.04" pageId="1" pageNumber="95" unit="m" value="104.0">104 m</quantity>
depth at location
<geoCoordinate id="EE47FAE32039FFDCFE2316ACD544FA06" box="[440,563,1447,1473]" degrees="78" direction="north" minutes="00" orientation="latitude" pageId="1" pageNumber="95" precision="15" seconds="9" value="78.0025">78°009”N</geoCoordinate>
<geoCoordinate id="EE47FAE32039FFDCFDA616ACD5C2FA06" box="[573,693,1447,1473]" degrees="87" direction="east" minutes="37" orientation="longitude" pageId="1" pageNumber="95" precision="15" seconds="3" value="87.6175">87°373”E</geoCoordinate>
(Kara Sea) and female partly dissected on one slide (collection number IORAS-Har203), collected from
<quantity id="4C8B31C12039FFDCFDFB16C0D5D1FA21" box="[608,678,1483,1510]" metricMagnitude="2" metricUnit="m" metricValue="3.35" pageId="1" pageNumber="95" unit="m" value="335.0">335 m</quantity>
depth at location
<geoCoordinate id="EE47FAE32039FFDCFCF216C0D492FA22" box="[873,997,1483,1509]" degrees="80" direction="north" minutes="00" orientation="latitude" pageId="1" pageNumber="95" precision="15" seconds="0" value="80.0">80°000”N</geoCoordinate>
<geoCoordinate id="EE47FAE32039FFDCFC7116C0D314FA21" box="[1002,1123,1483,1510]" degrees="85" direction="east" minutes="32" orientation="longitude" pageId="1" pageNumber="95" precision="15" seconds="0" value="85.53333">85°320”E</geoCoordinate>
(Kara Sea). Both specimens were collected in
<date id="FFCDBAE42039FFDCFEC616E4D560F9CD" box="[349,535,1519,1546]" pageId="1" pageNumber="95" value="2015-09">September 2015</date>
, leg. A. Udalov.
</paragraph>
<paragraph id="8BCC9C242039FFDCFF5C1518D638F95E" blockId="1.[151,1437,1411,2014]" pageId="1" pageNumber="95">
<emphasis id="B90740362039FFDCFF5C1518D6BBF9EA" bold="true" box="[199,460,1555,1581]" pageId="1" pageNumber="95">Description of female.</emphasis>
Total body length measured from tip of rostrum to posterior margin of caudal rami: 310 μm. Genital field (
<figureCitation id="134880A12039FFDCFEF7153CD6B3F996" box="[364,452,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[195,1395,192,1909]" captionTargetId="figure-17@2.[184,1403,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Danielssenia spitsbergensis Gee &amp; Huys, 1994, female:A, genital field; B, antennule; C, P2; D, P5." figureDoi="http://doi.org/10.5281/zenodo.5564093" httpUri="https://zenodo.org/record/5564093/files/figure.png" pageId="1" pageNumber="95">Fig. 1A</figureCitation>
) with small copulatory pore; short copulatory duct leading to seminal receptacle with paired anterior chambers extending to anterior margin of genital double-somite; vestigial P6 with one pinnate seta and one spinule.
</paragraph>
<paragraph id="8BCC9C242039FFDCFF5C15A8D3C9F8B6" blockId="1.[151,1437,1411,2014]" pageId="1" pageNumber="95">
Antennule (
<figureCitation id="134880A12039FFDCFED415A8D6D0F97A" box="[335,423,1699,1725]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[195,1395,192,1909]" captionTargetId="figure-17@2.[184,1403,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Danielssenia spitsbergensis Gee &amp; Huys, 1994, female:A, genital field; B, antennule; C, P2; D, P5." figureDoi="http://doi.org/10.5281/zenodo.5564093" httpUri="https://zenodo.org/record/5564093/files/figure.png" pageId="1" pageNumber="95">Fig. 1B</figureCitation>
) short, stout, four-segmented. Segment 1 largest, almost rectangular, with two rows of spinules on dorsal surface and a pinnate seta at distal corner. Segment 2 almost triangular, with eight setae (one pinnate seta proximally on anterior margin, five setae at anterior distal corner and two setae on posterior margin). Segment 3 triangular with eleven setae (five setae medially on anterior margin, and six setae and an aesthetasc at anterior distal corner). Segment 4 triangular with row of spinules on dorsal surface and seven setae at proximal margin (two long pinnate setae, two plane setae and three sort pinnate setae) and an aesthetasc.
</paragraph>
<paragraph id="8BCC9C242039FFDEFF5C1470D3A9FEDA" blockId="1.[151,1437,1411,2014]" lastBlockId="3.[151,1437,151,1149]" lastPageId="3" lastPageNumber="97" pageId="1" pageNumber="95">
P2 (
<figureCitation id="134880A12039FFDCFF6D1470D638F852" box="[246,335,1915,1941]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[195,1395,192,1909]" captionTargetId="figure-17@2.[184,1403,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Danielssenia spitsbergensis Gee &amp; Huys, 1994, female:A, genital field; B, antennule; C, P2; D, P5." figureDoi="http://doi.org/10.5281/zenodo.5564093" httpUri="https://zenodo.org/record/5564093/files/figure.png" pageId="1" pageNumber="95">Fig. 1C</figureCitation>
). Intercoxal sclerite well-developed, bearing two groups of setules at distal margin. Coxa almost rectangular with two groups of setules at proximal and distal outer corner of distal margin on anterior face. Basis with outer seta and row of spinules halfway down anterior margin. Both rami three-segmented, almost equal in length. Exopod-1 with rows of spinules on outer and distal margin, and one outer seta; exopod-2 with rows of spinules on outer and distal margin, one outer seta and one inner seta; exopod-3 only with outer row of spinules, and seven setae (223). Endopod-1 with outer rows of spinules and one inner seta; endopod-2 with outer row of spinules, long inner spinule and one inner seta; endopod-3 with outer rows of spinules and five setae (221).
</paragraph>
<caption id="DF0CCCAC203AFFDFFF0C149CD279F868" ID-DOI="http://doi.org/10.5281/zenodo.5564093" ID-Zenodo-Dep="5564093" box="[151,1294,1943,1968]" httpUri="https://zenodo.org/record/5564093/files/figure.png" pageId="2" pageNumber="96" startId="2.[151,250,1943,1967]" targetBox="[195,1395,192,1909]" targetPageId="2">
<paragraph id="8BCC9C24203AFFDFFF0C149CD279F868" blockId="2.[151,1294,1943,1968]" box="[151,1294,1943,1968]" pageId="2" pageNumber="96">
<emphasis id="B9074036203AFFDFFF0C149CD67AF868" bold="true" box="[151,269,1943,1967]" pageId="2" pageNumber="96">FIGURE 1</emphasis>
.
<taxonomicName id="4C73E7A7203AFFDFFE821493D58EF877" authority="Gee &amp; Huys, 1994" authorityName="Gee &amp; Huys" authorityYear="1994" box="[281,761,1944,1968]" class="Hexanauplia" family="Pseudotachidiidae" genus="Danielssenia" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="spitsbergensis">
<emphasis id="B9074036203AFFDFFE821493D544F877" box="[281,563,1944,1968]" italics="true" pageId="2" pageNumber="96">Danielssenia spitsbergensis</emphasis>
<bibRefCitation id="EFE2E1D5203AFFDFFDA21493D58EF877" author="Gee, J. M. &amp; Huys, R." box="[569,761,1944,1968]" pageId="2" pageNumber="96" pagination="1007 - 1046" refId="ref3787" refString="Gee, J. M. &amp; Huys, R. (1994) Paranannopidae (Copepoda: Harpacticoida) from sublittoral soft sediments in Spitzbergen. Journal of natural History, 28, 1007 - 1046. https: // doi. org / 10.1080 / 00222939400770541" type="journal article" year="1994">Gee &amp; Huys, 1994</bibRefCitation>
</taxonomicName>
, female: A, genital field; B, antennule; C, P2; D, P5.
</paragraph>
</caption>
<paragraph id="8BCC9C24203BFFDEFF5C122DD43AFE4E" blockId="3.[151,1437,151,1149]" pageId="3" pageNumber="97">
P5 (
<figureCitation id="134880A1203BFFDEFF6D122CD626FE86" box="[246,337,295,321]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[195,1395,192,1909]" captionTargetId="figure-17@2.[184,1403,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Danielssenia spitsbergensis Gee &amp; Huys, 1994, female:A, genital field; B, antennule; C, P2; D, P5." figureDoi="http://doi.org/10.5281/zenodo.5564093" httpUri="https://zenodo.org/record/5564093/files/figure.png" pageId="3" pageNumber="97">Fig. 1D</figureCitation>
). Element of each side not fused medially. Baseoendopod and exopod separate. Endopodal lobe with five pinnate setae; fourth seta the longest. Exopod slightly longer than wide with four setae; second seta the longest with plumose tip. Outer seta long and well developed.
</paragraph>
<paragraph id="8BCC9C24203BFFDEFF5C1299D209FE16" blockId="3.[151,1437,151,1149]" pageId="3" pageNumber="97">
<emphasis id="B9074036203BFFDEFF5C1299D63AFE6B" bold="true" box="[199,333,402,428]" pageId="3" pageNumber="97">Variability.</emphasis>
Only
<specimenCount id="9D7557AD203BFFDEFE0B1298D561FE6A" box="[400,534,403,429]" pageId="3" pageNumber="97" type="female">two females</specimenCount>
and
<specimenCount id="9D7557AD203BFFDEFDD21298D5CDFE6A" box="[585,698,403,429]" pageId="3" pageNumber="97" type="male">two males</specimenCount>
were collected and examined. No variable features or asymmetries were observed, and differences between male and female specimens are all part of a normal sexual dimorphism.
</paragraph>
</subSubSection>
<subSubSection id="C369CFAF203BFFDEFF5C12D0D530FBBA" pageId="3" pageNumber="97" type="discussion">
<paragraph id="8BCC9C24203BFFDEFF5C12D0D576FD36" blockId="3.[151,1437,151,1149]" pageId="3" pageNumber="97">
<emphasis id="B9074036203BFFDEFF5C12D0D64CFE32" bold="true" box="[199,315,475,501]" pageId="3" pageNumber="97">Remarks.</emphasis>
<taxonomicName id="4C73E7A7203BFFDEFEDE12D0D508FE32" authorityName="Gee &amp; Huys" authorityYear="1994" box="[325,639,475,501]" class="Hexanauplia" family="Pseudotachidiidae" genus="Danielssenia" kingdom="Animalia" order="Harpacticoida" pageId="3" pageNumber="97" phylum="Arthropoda" rank="species" species="spitsbergensis">
<emphasis id="B9074036203BFFDEFEDE12D0D508FE32" box="[325,639,475,501]" italics="true" pageId="3" pageNumber="97">Danielssenia spitsbergensis</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFE2E1D5203BFFDEFCC312D0D349FE32" author="Gee, J. M. &amp; Huys, R." box="[856,1086,475,501]" pageId="3" pageNumber="97" pagination="1007 - 1046" refId="ref3787" refString="Gee, J. M. &amp; Huys, R. (1994) Paranannopidae (Copepoda: Harpacticoida) from sublittoral soft sediments in Spitzbergen. Journal of natural History, 28, 1007 - 1046. https: // doi. org / 10.1080 / 00222939400770541" type="journal article" year="1994">Gee &amp; Huys (1994)</bibRefCitation>
from a single male specimen. It can easily be distinguished from other known species by the presence of only two outer spines on exopod-3 of P3 and P4. However, because of differences in the structure of the P2 endopod and P5, the species placed
<emphasis id="B9074036203BFFDEFADB1128D7B8FDA6" italics="true" pageId="3" pageNumber="97">incertae sedis</emphasis>
within
<taxonomicName id="4C73E7A7203BFFDEFEBE114CD6C0FDA6" authorityName="Boeck" authorityYear="1873" box="[293,439,583,609]" class="Hexanauplia" family="Pseudotachidiidae" genus="Danielssenia" kingdom="Animalia" order="Harpacticoida" pageId="3" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="B9074036203BFFDEFEBE114CD6C0FDA6" box="[293,439,583,609]" italics="true" pageId="3" pageNumber="97">Danielssenia</emphasis>
</taxonomicName>
pending the discovery of the female. A single female was also found and, according
<bibRefCitation id="EFE2E1D5203BFFDEFAF4114CD63FFD42" author="Gee, J. M. &amp; Huys, R." pageId="3" pageNumber="97" pagination="1007 - 1046" refId="ref3787" refString="Gee, J. M. &amp; Huys, R. (1994) Paranannopidae (Copepoda: Harpacticoida) from sublittoral soft sediments in Spitzbergen. Journal of natural History, 28, 1007 - 1046. https: // doi. org / 10.1080 / 00222939400770541" type="journal article" year="1994">Gee &amp; Huys (1994)</bibRefCitation>
, had only two outer spines on exopod-3 P3 and P4, but was lost before detailed inspection. The lack of some details of female morphology, particularly the segmentation of the female antennule and the detailed structure of the genital field and the seminal receptacle, made
<bibRefCitation id="EFE2E1D5203BFFDEFCDA11B8D36DFD0A" author="Gee, J. M. &amp; Huys, R." box="[833,1050,691,717]" pageId="3" pageNumber="97" pagination="1007 - 1046" refId="ref3787" refString="Gee, J. M. &amp; Huys, R. (1994) Paranannopidae (Copepoda: Harpacticoida) from sublittoral soft sediments in Spitzbergen. Journal of natural History, 28, 1007 - 1046. https: // doi. org / 10.1080 / 00222939400770541" type="journal article" year="1994">Gee &amp; Huys (1994)</bibRefCitation>
refrain from determining the exact taxonomic status of this species.
</paragraph>
<paragraph id="8BCC9C24203BFFDEFF5C11F0D530FBBA" blockId="3.[151,1437,151,1149]" pageId="3" pageNumber="97">
Our specimens exhibit all diagnostic features of the genus
<taxonomicName id="4C73E7A7203BFFDEFCF911F0D483FCD2" authorityName="Boeck" authorityYear="1873" box="[866,1012,763,789]" class="Hexanauplia" family="Pseudotachidiidae" genus="Danielssenia" kingdom="Animalia" order="Harpacticoida" pageId="3" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="B9074036203BFFDEFCF911F0D483FCD2" box="[866,1012,763,789]" italics="true" pageId="3" pageNumber="97">Danielssenia</emphasis>
</taxonomicName>
as redefined by
<bibRefCitation id="EFE2E1D5203BFFDEFB2811F0D2EEFCD2" author="Huys, R. &amp; Gee, J. M." box="[1203,1433,763,789]" pageId="3" pageNumber="97" pagination="45 - 81" refId="ref3861" refString="Huys, R. &amp; Gee, J. M. (1993) A revision of Danielssenia Boeck and Psammis Sars with the establishment of two new genera Archisenia and Bathypsammis (Harpacticoida: Paranannopidae). Bulletin of the Natural History Museum, London, Zoology, 59, 45 - 81." type="journal article" year="1993">Huys &amp; Gee (1993)</bibRefCitation>
: four-segmented antennule; one seta on exopod-1 of antennary exopod; structure of all mouthparts; absence of inner seta on exopod-1 of P2P4; P5 not fused medially; distinct P5 exopod and baseoendopod; five setae on P5 baseoendopod; and genital field with large copulatory pore and short copulatory duct leading to seminal receptacle with paired anterior chambers. Since our females have only two outer spines on exopod-3 of P3 and P4, this differentiating feature that is unique for
<taxonomicName id="4C73E7A7203BFFDEFDCF10A4D464FC0E" authorityName="Gee &amp; Huys" authorityYear="1994" box="[596,787,943,969]" class="Hexanauplia" family="Pseudotachidiidae" genus="Danielssenia" kingdom="Animalia" order="Harpacticoida" pageId="3" pageNumber="97" phylum="Arthropoda" rank="species" species="spitsbergensis">
<emphasis id="B9074036203BFFDEFDCF10A4D464FC0E" box="[596,787,943,969]" italics="true" pageId="3" pageNumber="97">D. spitsbergensis</emphasis>
</taxonomicName>
, should be included in species diagnosis. Examined males have the same features like in Gee &amp; Huys (1994) original description: (1) the mucroniform process on endopod-2 of P2 reaches to the distal margin of endopod-3; and (2) the exopod of the male P5 is fused to the baseoendopod. Although the confirmation of the deviating armature formula of the P3P4 exopod-
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both sexes would require an emendation of the generic diagnosis, we prefer to concur with
<bibRefCitation id="EFE2E1D5203BFFDEFCC71734D34AFB9E" author="Gee, J. M. &amp; Huys, R." box="[860,1085,1087,1113]" pageId="3" pageNumber="97" pagination="1007 - 1046" refId="ref3787" refString="Gee, J. M. &amp; Huys, R. (1994) Paranannopidae (Copepoda: Harpacticoida) from sublittoral soft sediments in Spitzbergen. Journal of natural History, 28, 1007 - 1046. https: // doi. org / 10.1080 / 00222939400770541" type="journal article" year="1994">Gee &amp; Huys (1994)</bibRefCitation>
in placing
<taxonomicName id="4C73E7A7203BFFDEFB271734D20AFB9E" authorityName="Gee &amp; Huys" authorityYear="1994" box="[1212,1405,1087,1113]" class="Hexanauplia" family="Pseudotachidiidae" genus="Danielssenia" kingdom="Animalia" order="Harpacticoida" pageId="3" pageNumber="97" phylum="Arthropoda" rank="species" species="spitsbergensis">
<emphasis id="B9074036203BFFDEFB271734D20AFB9E" box="[1212,1405,1087,1113]" italics="true" pageId="3" pageNumber="97">D. spitsbergensis</emphasis>
</taxonomicName>
as
<emphasis id="B9074036203BFFDEFF0C1768D6FDFBBA" box="[151,394,1123,1149]" italics="true" pageId="3" pageNumber="97">species incertae sedis</emphasis>
in
<taxonomicName id="4C73E7A7203BFFDEFE351768D537FBBA" authorityName="Boeck" authorityYear="1873" box="[430,576,1123,1149]" class="Hexanauplia" family="Pseudotachidiidae" genus="Danielssenia" kingdom="Animalia" order="Harpacticoida" pageId="3" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="B9074036203BFFDEFE351768D537FBBA" box="[430,576,1123,1149]" italics="true" pageId="3" pageNumber="97">Danielssenia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>