377 lines
59 KiB
XML
377 lines
59 KiB
XML
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<mods:title id="16CA24CBA2BEED899D48A97809D13B62">New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China</mods:title>
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<mods:namePart id="4A85880B36836AE9123968843B025C27">Hou, Yanmeng</mods:namePart>
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<mods:namePart id="8DE935CEED64CF9E329DAEE57DF8FD3E">Feng, Zegang</mods:namePart>
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<mods:namePart id="88FCD134A73E1FFDAEA2AF6B5541DFED">Zhang, Feng</mods:namePart>
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<mods:date id="4F63677BED55EA9B621AF666CB4C4957">2023</mods:date>
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<treatment id="03DA1A1EFF9FFFAEFF053F79E3C23D46" ID-DOI="http://doi.org/10.5281/zenodo.8090034" ID-GBIF-Taxon="210479509" ID-Zenodo-Dep="8090034" LSID="urn:lsid:plazi:treatment:03DA1A1EFF9FFFAEFF053F79E3C23D46" httpUri="http://treatment.plazi.org/id/03DA1A1EFF9FFFAEFF053F79E3C23D46" lastPageId="10" lastPageNumber="11" pageId="5" pageNumber="6">
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<subSubSection id="C369F883FF9FFFA1FF053F79E2233C03" box="[151,784,1312,1339]" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="8BCCAB08FF9FFFA1FF053F79E2233C03" blockId="5.[151,784,1312,1375]" box="[151,784,1312,1339]" pageId="5" pageNumber="6">
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<heading id="D0841C64FF9FFFA1FF053F79E2233C03" bold="true" box="[151,784,1312,1339]" fontSize="11" level="2" pageId="5" pageNumber="6" reason="4">
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<emphasis id="B907771AFF9FFFA1FF053F79E2233C03" bold="true" box="[151,784,1312,1339]" pageId="5" pageNumber="6">
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<taxonomicName id="4C73D08BFF9FFFA1FF053F79E0C03C00" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[151,499,1312,1339]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="biyunensis" status="sp. nov.">
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<emphasis id="B907771AFF9FFFA1FF053F79E0C03C00" bold="true" box="[151,499,1312,1339]" italics="true" pageId="5" pageNumber="6">Lagynochthonius biyunensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A234CA61FF9FFFA1FE693F78E36B3C00" box="[507,600,1313,1339]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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(ỠỄṄṁae)
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C369F883FF9FFFA1FF053F1DE1CA3C64" box="[151,249,1348,1375]" pageId="5" pageNumber="6" type="description">
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<paragraph id="8BCCAB08FF9FFFA1FF053F1DE1CA3C64" blockId="5.[151,784,1312,1375]" box="[151,249,1348,1375]" pageId="5" pageNumber="6">
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<figureCitation id="1348B78DFF9FFFA1FF053F1DE1CA3C64" box="[151,249,1348,1375]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="6.[151,250,1801,1825]" captionStartId-1="7.[151,250,781,805]" captionStartId-2="8.[151,250,1943,1967]" captionStartId-3="9.[151,250,1894,1918]" captionTargetBox-0="[151,1436,381,1777]" captionTargetBox-1="[159,1430,188,757]" captionTargetBox-2="[179,1408,196,1919]" captionTargetBox-3="[197,1391,181,1870]" captionTargetId-0="figure-132@6.[151,1436,381,1777]" captionTargetId-1="figure-18@7.[151,1436,181,757]" captionTargetId-2="figure-20@8.[179,1408,181,1919]" captionTargetId-3="figure-18@9.[197,1391,181,1870]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="FIGURE 2. Biyun Cave, type locality of Lagynochthonius biyunensis sp. nov., A. Entrance; B. Inside the cave entrance; C. Another entrance penetrated by a river; D. A narrow tunnel to the interior of the cave; E. Area where L. biyunensis sp. nov. specimens were collected (red arrows); F. Live male of L. biyunensis sp. nov. in its natural environment." captionText-1="FIGURE 3. Lagynochthonius biyunensis sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." captionText-2="FIGURE 4. Lagynochthonius biyunensis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." captionText-3="FIGURE 5. Lagynochthonius biyunensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg Ⅰ (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi-0="http://doi.org/10.5281/zenodo.8090052" figureDoi-1="http://doi.org/10.5281/zenodo.8090056" figureDoi-2="http://doi.org/10.5281/zenodo.8090058" figureDoi-3="http://doi.org/10.5281/zenodo.8090062" httpUri-0="https://zenodo.org/record/8090052/files/figure.png" httpUri-1="https://zenodo.org/record/8090056/files/figure.png" httpUri-2="https://zenodo.org/record/8090058/files/figure.png" httpUri-3="https://zenodo.org/record/8090062/files/figure.png" pageId="5" pageNumber="6">Figs 2–5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C369F883FF9FFFA1FF053FD4E3243F61" pageId="5" pageNumber="6" type="materials_examined">
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<paragraph id="8BCCAB08FF9FFFA1FF053FD4E3243F61" blockId="5.[151,1437,1420,2023]" pageId="5" pageNumber="6">
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<emphasis id="B907771AFF9FFFA1FF053FD4E07B3C9D" bold="true" box="[151,328,1420,1447]" pageId="5" pageNumber="6">Type material.</emphasis>
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<materialsCitation id="3B1BA155FF9FFFA1FEC03FD5E59E3CD5" ID-GBIF-Occurrence="4138993331" collectingDate="2019-08-05" collectionCode="MHBU-GZC" collectorName="Zegang Feng & Zhaoyi Li & Chen Zhang" country="China" county="Panzhou City" elevation="1496" latitude="25.77192" location="Nanmen Village" longLatPrecision="1" longitude="104.64" municipality="Chengguan Town" pageId="5" pageNumber="6" specimenCode="MHBU-GZC190805-01" specimenCount="1" stateProvince="Guizhou" typeStatus="holotype">
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<typeStatus id="54C815AAFF9FFFA1FEC03FD5E0893C9D" box="[338,442,1420,1446]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
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♂ (Ps.-
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):
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<collectingCountry id="F364EB98FF9FFFA1FCD23FD5E2B53C9D" box="[832,902,1420,1446]" name="China" pageId="5" pageNumber="6">China</collectingCountry>
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,
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<collectingRegion id="49B765EAFF9FFFA1FC063FD5E5563C9D" box="[916,1125,1420,1446]" country="China" name="Guizhou" pageId="5" pageNumber="6">Guizhou Province</collectingRegion>
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,
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<collectingCounty id="62ADD384FF9FFFA1FBE13FD5E43D3C9D" box="[1139,1294,1420,1446]" pageId="5" pageNumber="6">Panzhou City</collectingCounty>
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,
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<collectingMunicipality id="6BA83172FF9FFFA1FA8F3FD5E1EA3CF0" pageId="5" pageNumber="6">Chengguan Town</collectingMunicipality>
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,
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<location id="8EACFDD3FF9FFFA1FF703FE8E0AA3CF1" LSID="urn:lsid:plazi:treatment:03DA1A1EFF9FFFAEFF053F79E3C23D46:8EACFDD3FF9FFFA1FF703FE8E0AA3CF1" box="[226,409,1456,1483]" country="China" county="Panzhou City" latitude="25.77192" longLatPrecision="1" longitude="104.64" municipality="Chengguan Town" name="Nanmen Village" pageId="5" pageNumber="6" stateProvince="Guizhou">Nanmen Village</location>
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,
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<location id="8EACFDD3FF9FFFA1FE333FE9E3123CF1" LSID="urn:lsid:plazi:treatment:03DA1A1EFF9FFFAEFF053F79E3C23D46:8EACFDD3FF9FFFA1FE333FE9E3123CF1" box="[417,545,1456,1482]" country="China" county="Panzhou City" latitude="25.77192" longLatPrecision="1" longitude="104.64" municipality="Chengguan Town" name="Biyun Park" pageId="5" pageNumber="6" stateProvince="Guizhou">Biyun Park</location>
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,
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<location id="8EACFDD3FF9FFFA1FDBB3FE9E39D3CF0" LSID="urn:lsid:plazi:treatment:03DA1A1EFF9FFFAEFF053F79E3C23D46:8EACFDD3FF9FFFA1FDBB3FE9E39D3CF0" box="[553,686,1456,1483]" country="China" county="Panzhou City" latitude="25.77192" longLatPrecision="1" longitude="104.64" municipality="Chengguan Town" name="Biyun Cave" pageId="5" pageNumber="6" stateProvince="Guizhou">Biyun Cave</location>
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, under the detritus and on the walls in the deep zone [
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<geoCoordinate id="EE47CDCFFF9FFFA1FB783FEBE4AB3CF6" box="[1258,1432,1458,1485]" degrees="25" direction="north" minutes="46" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="18.91" value="25.77192">25°46′18.91″N</geoCoordinate>
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,
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<geoCoordinate id="EE47CDCFFF9FFFA1FF053F8FE0723CCA" box="[151,321,1494,1521]" degrees="104" direction="east" minutes="38" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="24.0" value="104.64">104°38′24.0″E</geoCoordinate>
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],
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<elevation id="005E4C3BFF9FFFA1FEC43F8DE0D83CD5" box="[342,491,1492,1519]" metricMagnitude="3" metricUnit="m" metricValue="1.496" pageId="5" pageNumber="6" unit="m" value="1496.0">
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a.s.l.
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</elevation>
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,
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<date id="FFCD8DC8FF9FFFA1FE6B3F8DE3973CD4" box="[505,676,1492,1519]" pageId="5" pageNumber="6" value="2019-08-05">
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<collectingDate id="EF897420FF9FFFA1FE6B3F8DE3973CD4" box="[505,676,1492,1519]" pageId="5" pageNumber="6" value="2019-08-05">5 August 2019</collectingDate>
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</date>
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,
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<collectorName id="2686CEDEFF9FFFA1FD233F8CE27A3CD4" box="[689,841,1493,1519]" pageId="5" pageNumber="6">Zegang Feng</collectorName>
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,
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<collectorName id="2686CEDEFF9FFFA1FCC43F8DE2F43CD5" box="[854,967,1492,1518]" pageId="5" pageNumber="6">Zhaoyi Li</collectorName>
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&
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<collectorName id="2686CEDEFF9FFFA1FC7D3F8DE5483CD5" box="[1007,1147,1492,1518]" pageId="5" pageNumber="6">Chen Zhang</collectorName>
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leg.
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</materialsCitation>
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<materialsCitation id="3B1BA155FF9FFFA1FB273F8CE20F3F28" ID-GBIF-Occurrence="4138993316" collectingDate="2022-08-02" collectionCode="MHBU-HBUARA" collectorName="Yanmeng Hou & Lu Zhang & Jianzhou Sun & Wenlong Fan" country="China" county="Panzhou City" elevation="1496" latitude="25.77192" location="Nanmen Village" longLatPrecision="1" longitude="104.64" municipality="Chengguan Town" pageId="5" pageNumber="6" specimenCode="MHBU-HBUARA#2022-498, 01, MSWU-HBUARA#2022-498, 05" specimenCount="4" specimenCount-female="4" stateProvince="Guizhou" typeStatus="paratype">
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<typeStatus id="54C815AAFF9FFFA1FB273F8CE4153CD4" box="[1205,1318,1493,1519]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D756081FF9FFFA1FAA13F8CE46C3CCA" box="[1331,1375,1493,1521]" pageId="5" pageNumber="6">4 ♂</specimenCount>
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(Ps.-
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-
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<specimenCode id="DBD50373FF9FFFA1FE603FA0E33F3F28" box="[498,524,1529,1555]" collectionCode="MHBU-HBUARA" pageId="5" pageNumber="6">01</specimenCode>
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–HBUARA#2022-498-04),
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</materialsCitation>
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<materialsCitation id="3B1BA155FF9FFFA1FCCE3FA0E3243F61" ID-GBIF-Occurrence="4138993304" collectingDate="2022-08-02" collectionCode="MSWU-HBUARA" collectorName="Yanmeng Hou & Lu Zhang & Jianzhou Sun & Wenlong Fan" country="China" county="Panzhou City" elevation="1496" latitude="25.77192" location="Nanmen Village" longLatPrecision="1" longitude="104.64" municipality="Chengguan Town" pageId="5" pageNumber="6" specimenCode="MHBU-HBUARA#2022-498, 01, MSWU-HBUARA#2022-498, 05" specimenCount="2" specimenCount-female="2" stateProvince="Guizhou" typeStatus="paratype">
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<specimenCount id="9D756081FF9FFFA1FCCE3FA0E2AD3F28" box="[860,926,1529,1555]" pageId="5" pageNumber="6" type="female">2 ♀</specimenCount>
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(Ps.-
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<specimenCode id="DBD50373FF9FFFA1FC783FA1E4763F28" box="[1002,1349,1528,1555]" collectionCode="MSWU-HBUARA" pageId="5" pageNumber="6">MSWU-HBUARA#2022-498</specimenCode>
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-
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<specimenCode id="DBD50373FF9FFFA1FADE3FA1E4543F28" box="[1356,1383,1528,1555]" collectionCode="MSWU-HBUARA" pageId="5" pageNumber="6">05</specimenCode>
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& HBUARA#2022-498-06), with the same location as the holotype,
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<date id="FFCD8DC8FF9FFFA1FC3B3C44E56A3F0C" box="[937,1113,1565,1591]" pageId="5" pageNumber="6" value="2022-08-02">
|
||
<collectingDate id="EF897420FF9FFFA1FC3B3C44E56A3F0C" box="[937,1113,1565,1591]" pageId="5" pageNumber="6" value="2022-08-02">2 August 2022</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="2686CEDEFF9FFFA1FBFB3C44E4213F0C" box="[1129,1298,1565,1591]" pageId="5" pageNumber="6">Yanmeng Hou</collectorName>
|
||
,
|
||
<collectorName id="2686CEDEFF9FFFA1FAB03C44E4AB3F0D" box="[1314,1432,1564,1591]" pageId="5" pageNumber="6">Lu Zhang</collectorName>
|
||
,
|
||
<collectorName id="2686CEDEFF9FFFA1FF053C19E01D3F60" box="[151,302,1600,1627]" pageId="5" pageNumber="6">Jianzhou Sun</collectorName>
|
||
&
|
||
<collectorName id="2686CEDEFF9FFFA1FEC33C19E0D53F60" box="[337,486,1600,1627]" pageId="5" pageNumber="6">Wenlong Fan</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C369F883FF9FFFA1FF553C3CE2073F44" box="[199,820,1636,1663]" pageId="5" pageNumber="6" type="etymology">
|
||
<paragraph id="8BCCAB08FF9FFFA1FF553C3CE2073F44" blockId="5.[151,1437,1420,2023]" box="[199,820,1636,1663]" pageId="5" pageNumber="6">
|
||
<emphasis id="B907771AFF9FFFA1FF553C3CE07D3F44" bold="true" box="[199,334,1637,1663]" pageId="5" pageNumber="6">Etymology.</emphasis>
|
||
Named after the
|
||
<typeStatus id="54C815AAFF9FFFA1FD823C3CE3733F44" box="[528,576,1637,1663]" pageId="5" pageNumber="6">type</typeStatus>
|
||
locality, Biyun Cave.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C369F883FF9FFFA1FF553CD1E2433E09" pageId="5" pageNumber="6" type="diagnosis">
|
||
<paragraph id="8BCCAB08FF9FFFA1FF553CD1E2433E09" blockId="5.[151,1437,1420,2023]" pageId="5" pageNumber="6">
|
||
<emphasis id="B907771AFF9FFFA1FF553CD1E0B73F9D" bold="true" box="[199,388,1672,1702]" pageId="5" pageNumber="6">
|
||
Diagnosis (♂
|
||
<specimenCount id="9D756081FF9FFFA1FECD3CD3E0473F9D" box="[351,372,1674,1702]" pageId="5" pageNumber="6" type="female">♀</specimenCount>
|
||
).
|
||
</emphasis>
|
||
<materialsCitation id="3B1BA155FF9FFFA1FE183CD1E25F3E09" ID-GBIF-Occurrence="4138993306" collectionCode="V" pageId="5" pageNumber="6" specimenCount="2" specimenCount-female="2">
|
||
Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, pointed, triangular; posterior margin of carapace with two setae; tergites Ⅰ–
|
||
<collectionCode id="ED6233CDFF9FFFA1FDA53C88E3783FD0" box="[567,587,1745,1771]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="6" type="Museum">Ⅴ</collectionCode>
|
||
each with four setae. Pedipalps slender, femur 8.06 (♂), 7.82–7.89 (
|
||
<specimenCount id="9D756081FF9FFFA1FAAF3C88E47C3FD0" box="[1341,1359,1745,1771]" pageId="5" pageNumber="6" type="female">♀</specimenCount>
|
||
) times longer than broad; chela 6.85–7.08 (♂), 7.27–7.43 (
|
||
<specimenCount id="9D756081FF9FFFA1FD763CACE3C53E34" box="[740,758,1781,1807]" pageId="5" pageNumber="6" type="female">♀</specimenCount>
|
||
) times longer than broad; chelal fingers without intercalary teeth; chemosensory setae (
|
||
<emphasis id="B907771AFF9FFFA1FE5E3D40E0D03E09" box="[460,483,1817,1842]" italics="true" pageId="5" pageNumber="6">sc</emphasis>
|
||
) present on dorsum of chelal hand
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C369F883FF9FFFAEFF553D65E3063D62" lastPageId="10" lastPageNumber="11" pageId="5" pageNumber="6" type="description">
|
||
<paragraph id="8BCCAB08FF9FFFA1FF553D65E2FF3E6C" blockId="5.[151,1437,1420,2023]" box="[199,972,1852,1879]" pageId="5" pageNumber="6">
|
||
<emphasis id="B907771AFF9FFFA1FF553D65E06A3E6D" bold="true" box="[199,345,1852,1878]" pageId="5" pageNumber="6">Description.</emphasis>
|
||
Males (
|
||
<typeStatus id="54C815AAFF9FFFA1FE213D65E3243E6D" box="[435,535,1852,1878]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54C815AAFF9FFFA1FDDF3D64E3893E6C" box="[589,698,1853,1879]" pageId="5" pageNumber="6" type="paratype">paratypes</typeStatus>
|
||
) (
|
||
<figureCitation id="1348B78DFF9FFFA1FD423D65E2143E6C" box="[720,807,1852,1879]" captionStart="FIGURE 2" captionStartId="6.[151,250,1801,1825]" captionTargetBox="[151,1436,381,1777]" captionTargetId="figure-132@6.[151,1436,381,1777]" captionTargetPageId="6" captionText="FIGURE 2. Biyun Cave, type locality of Lagynochthonius biyunensis sp. nov., A. Entrance; B. Inside the cave entrance; C. Another entrance penetrated by a river; D. A narrow tunnel to the interior of the cave; E. Area where L. biyunensis sp. nov. specimens were collected (red arrows); F. Live male of L. biyunensis sp. nov. in its natural environment." figureDoi="http://doi.org/10.5281/zenodo.8090052" httpUri="https://zenodo.org/record/8090052/files/figure.png" pageId="5" pageNumber="6">Figs 2F</figureCitation>
|
||
,
|
||
<figureCitation id="1348B78DFF9FFFA1FCA03D64E2653E6C" box="[818,854,1853,1879]" captionStart="FIGURE 3" captionStartId="7.[151,250,781,805]" captionTargetBox="[159,1430,188,757]" captionTargetId="figure-18@7.[151,1436,181,757]" captionTargetPageId="7" captionText="FIGURE 3. Lagynochthonius biyunensis sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8090056" httpUri="https://zenodo.org/record/8090056/files/figure.png" pageId="5" pageNumber="6">3A</figureCitation>
|
||
,
|
||
<figureCitation id="1348B78DFF9FFFA1FCF13D64E2903E6C" box="[867,931,1853,1879]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[179,1408,196,1919]" captionTargetId="figure-20@8.[179,1408,181,1919]" captionTargetPageId="8" captionText="FIGURE 4. Lagynochthonius biyunensis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.8090058" httpUri="https://zenodo.org/record/8090058/files/figure.png" pageId="5" pageNumber="6">4A–F</figureCitation>
|
||
,
|
||
<figureCitation id="1348B78DFF9FFFA1FC3C3D65E28D3E6C" box="[942,958,1852,1879]" captionStart="FIGURE 5" captionStartId="9.[151,250,1894,1918]" captionTargetBox="[197,1391,181,1870]" captionTargetId="figure-18@9.[197,1391,181,1870]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius biyunensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg Ⅰ (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.8090062" httpUri="https://zenodo.org/record/8090062/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCCAB08FF9FFFA1FF553D39E5DC3E41" blockId="5.[151,1437,1420,2023]" box="[199,1263,1888,1915]" pageId="5" pageNumber="6">Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</paragraph>
|
||
<paragraph id="8BCCAB08FF9FFFA2FF553DDDE249385E" blockId="5.[151,1437,1420,2023]" lastBlockId="6.[151,1437,151,357]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||
Cephalothorax (
|
||
<figureCitation id="1348B78DFF9FFFA1FEED3DDDE0ED3EA4" box="[383,478,1924,1951]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[179,1408,196,1919]" captionTargetId="figure-20@8.[179,1408,181,1919]" captionTargetPageId="8" captionText="FIGURE 4. Lagynochthonius biyunensis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.8090058" httpUri="https://zenodo.org/record/8090058/files/figure.png" pageId="5" pageNumber="6">Figs 4C</figureCitation>
|
||
,
|
||
<figureCitation id="1348B78DFF9FFFA1FE7C3DDDE3273EA4" box="[494,532,1924,1951]" captionStart="FIGURE 5" captionStartId="9.[151,250,1894,1918]" captionTargetBox="[197,1391,181,1870]" captionTargetId="figure-18@9.[197,1391,181,1870]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius biyunensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg Ⅰ (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.8090062" httpUri="https://zenodo.org/record/8090062/files/figure.png" pageId="5" pageNumber="6">5A</figureCitation>
|
||
): carapace 1.00 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome pointed and small, triangular; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, Ⅰ 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa Ⅰ with small, rounded anteromedial process; coxae II with ten terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (
|
||
<figureCitation id="1348B78DFF9CFFA2FF7E3B12E077385E" box="[236,324,330,357]" captionStart="FIGURE 5" captionStartId="9.[151,250,1894,1918]" captionTargetBox="[197,1391,181,1870]" captionTargetId="figure-18@9.[197,1391,181,1870]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius biyunensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg Ⅰ (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.8090062" httpUri="https://zenodo.org/record/8090062/files/figure.png" pageId="6" pageNumber="7">Fig. 5C</figureCitation>
|
||
); intercoxal tubercle absent; without sub-oral seta.
|
||
</paragraph>
|
||
<caption id="DF0CFB80FF9CFFA2FF053D50E5883E52" ID-DOI="http://doi.org/10.5281/zenodo.8090052" ID-Zenodo-Dep="8090052" httpUri="https://zenodo.org/record/8090052/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,250,1801,1825]" targetBox="[151,1436,381,1777]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8BCCAB08FF9CFFA2FF053D50E5883E52" blockId="6.[151,1437,1801,1897]" pageId="6" pageNumber="7">
|
||
<emphasis id="B907771AFF9CFFA2FF053D50E0263E1A" bold="true" box="[151,277,1801,1825]" pageId="6" pageNumber="7">FIGURE 2.</emphasis>
|
||
Biyun Cave, type locality of
|
||
<taxonomicName id="4C73D08BFF9CFFA2FDD83D50E25F3E1A" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[586,876,1801,1825]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="biyunensis" status="sp. nov.">
|
||
<emphasis id="B907771AFF9CFFA2FDD83D50E25F3E1A" box="[586,876,1801,1825]" italics="true" pageId="6" pageNumber="7">Lagynochthonius biyunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B907771AFF9CFFA2FCE63D50E2F53E1A" bold="true" box="[884,966,1801,1825]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="A234CA61FF9CFFA2FCE63D50E2F53E1A" box="[884,966,1801,1825]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, A. Entrance; B. Inside the cave entrance; C. Another entrance penetrated by a river; D. A narrow tunnel to the interior of the cave; E. Area where
|
||
<taxonomicName id="4C73D08BFF9CFFA2FB283D77E4723E7E" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[1210,1345,1837,1861]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="biyunensis" status="sp. nov.">
|
||
<emphasis id="B907771AFF9CFFA2FB283D77E4723E7E" box="[1210,1345,1837,1861]" italics="true" pageId="6" pageNumber="7">L. biyunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B907771AFF9CFFA2FADB3D74E4AF3E7E" bold="true" box="[1353,1436,1837,1861]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="A234CA61FF9CFFA2FADB3D74E4AF3E7E" box="[1353,1436,1837,1861]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
specimens were collected (red arrows); F. Live male of
|
||
<taxonomicName id="4C73D08BFF9CFFA2FD593D0BE2633E52" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[715,848,1873,1897]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="biyunensis" status="sp. nov.">
|
||
<emphasis id="B907771AFF9CFFA2FD593D0BE2633E52" box="[715,848,1873,1897]" italics="true" pageId="6" pageNumber="7">L. biyunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B907771AFF9CFFA2FCC53D08E2943E52" bold="true" box="[855,935,1873,1897]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="A234CA61FF9CFFA2FCC53D08E2943E52" box="[855,935,1873,1897]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in its natural environment.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0CFB80FF9DFFA3FF053954E1EB3A72" ID-DOI="http://doi.org/10.5281/zenodo.8090056" ID-Zenodo-Dep="8090056" httpUri="https://zenodo.org/record/8090056/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,781,805]" targetBox="[159,1430,188,757]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8BCCAB08FF9DFFA3FF053954E1EB3A72" blockId="7.[151,1436,781,841]" pageId="7" pageNumber="8">
|
||
<emphasis id="B907771AFF9DFFA3FF053954E0213A1E" bold="true" box="[151,274,781,805]" pageId="7" pageNumber="8">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C73D08BFF9DFFA3FE8A3954E30B3A1E" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[280,568,781,805]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="biyunensis" status="sp. nov.">
|
||
<emphasis id="B907771AFF9DFFA3FE8A3954E30B3A1E" box="[280,568,781,805]" italics="true" pageId="7" pageNumber="8">Lagynochthonius biyunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B907771AFF9DFFA3FDAF3954E3BE3A1E" bold="true" box="[573,653,781,805]" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="A234CA61FF9DFFA3FDAF3954E3BE3A1E" box="[573,653,781,805]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCCAB08FF9DFFA3FF5539C2E36E3D89" blockId="7.[151,1437,923,1922]" pageId="7" pageNumber="8">
|
||
Chelicera (
|
||
<figureCitation id="1348B78DFF9DFFA3FED239C2E0AE3A8E" box="[320,413,923,949]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[179,1408,196,1919]" captionTargetId="figure-20@8.[179,1408,181,1919]" captionTargetPageId="8" captionText="FIGURE 4. Lagynochthonius biyunensis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.8090058" httpUri="https://zenodo.org/record/8090058/files/figure.png" pageId="7" pageNumber="8">Figs 4D</figureCitation>
|
||
,
|
||
<figureCitation id="1348B78DFF9DFFA3FE3539C2E0F83A8D" box="[423,459,923,950]" captionStart="FIGURE 5" captionStartId="9.[151,250,1894,1918]" captionTargetBox="[197,1391,181,1870]" captionTargetId="figure-18@9.[197,1391,181,1870]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius biyunensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg Ⅰ (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.8090062" httpUri="https://zenodo.org/record/8090062/files/figure.png" pageId="7" pageNumber="8">5B</figureCitation>
|
||
): large, about as long as carapace, 2.40–2.47 times longer than broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 15–17 teeth, distal one largest; movable finger with 21–22 retrorse contiguous small teeth; galea absent (
|
||
<figureCitation id="1348B78DFF9DFFA3FBAD3E72E5A93D7D" box="[1087,1178,1067,1094]" captionStart="FIGURE 5" captionStartId="9.[151,250,1894,1918]" captionTargetBox="[197,1391,181,1870]" captionTargetId="figure-18@9.[197,1391,181,1870]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius biyunensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg Ⅰ (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.8090062" httpUri="https://zenodo.org/record/8090062/files/figure.png" pageId="7" pageNumber="8">Fig. 5B</figureCitation>
|
||
). Serrula exterior with 22–23 blades and serrula interior with 14–15 blades. Rallum with nine blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (
|
||
<figureCitation id="1348B78DFF9DFFA3FE643ECEE3633D89" box="[502,592,1175,1202]" captionStart="FIGURE 5" captionStartId="9.[151,250,1894,1918]" captionTargetBox="[197,1391,181,1870]" captionTargetId="figure-18@9.[197,1391,181,1870]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius biyunensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg Ⅰ (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.8090062" httpUri="https://zenodo.org/record/8090062/files/figure.png" pageId="7" pageNumber="8">Fig. 5D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCCAB08FF9DFFA3FF553EE2E00A3FC9" blockId="7.[151,1437,923,1922]" pageId="7" pageNumber="8">
|
||
Pedipalp (
|
||
<figureCitation id="1348B78DFF9DFFA3FEAB3EE2E0E43DED" box="[313,471,1211,1238]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[179,1408,196,1919]" captionTargetId="figure-20@8.[179,1408,181,1919]" captionTargetPageId="8" captionText="FIGURE 4. Lagynochthonius biyunensis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.8090058" httpUri="https://zenodo.org/record/8090058/files/figure.png" pageId="7" pageNumber="8">Figs 4A–B, E</figureCitation>
|
||
,
|
||
<figureCitation id="1348B78DFF9DFFA3FE763EE2E3193DED" box="[484,554,1211,1238]" captionStart="FIGURE 5" captionStartId="9.[151,250,1894,1918]" captionTargetBox="[197,1391,181,1870]" captionTargetId="figure-18@9.[197,1391,181,1870]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius biyunensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg Ⅰ (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.8090062" httpUri="https://zenodo.org/record/8090062/files/figure.png" pageId="7" pageNumber="8">5E–G</figureCitation>
|
||
): long and slender, trochanter 1.20–1.26, femur 8.06, patella 2.35, chela 6.85– 7.08, hand 2.70–2.81 times longer than broad; femur 2.74 times longer than patella; movable chelal finger 1.49–1.52 times longer than hand and 0.59–0.60 times longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (
|
||
<figureCitation id="1348B78DFF9DFFA3FE7B3F7EE3763C7A" box="[489,581,1319,1345]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[179,1408,196,1919]" captionTargetId="figure-20@8.[179,1408,181,1919]" captionTargetPageId="8" captionText="FIGURE 4. Lagynochthonius biyunensis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.8090058" httpUri="https://zenodo.org/record/8090058/files/figure.png" pageId="7" pageNumber="8">Figs 4E</figureCitation>
|
||
,
|
||
<figureCitation id="1348B78DFF9DFFA3FDC03F7EE3473C79" box="[594,628,1319,1346]" captionStart="FIGURE 5" captionStartId="9.[151,250,1894,1918]" captionTargetBox="[197,1391,181,1870]" captionTargetId="figure-18@9.[197,1391,181,1870]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius biyunensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg Ⅰ (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.8090062" httpUri="https://zenodo.org/record/8090062/files/figure.png" pageId="7" pageNumber="8">5E</figureCitation>
|
||
). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria,
|
||
<emphasis id="B907771AFF9DFFA3FAC43F36E45F3CB2" box="[1366,1388,1391,1417]" italics="true" pageId="7" pageNumber="8">ib</emphasis>
|
||
and
|
||
<emphasis id="B907771AFF9DFFA3FF053FCAE18B3C96" box="[151,184,1427,1453]" italics="true" pageId="7" pageNumber="8">isb</emphasis>
|
||
situated close together, submedially on dorsum of chelal hand;
|
||
<emphasis id="B907771AFF9DFFA3FC113FCAE2AE3C96" box="[899,925,1427,1453]" italics="true" pageId="7" pageNumber="8">eb</emphasis>
|
||
,
|
||
<emphasis id="B907771AFF9DFFA3FC3E3FCAE2E23C96" box="[940,977,1427,1453]" italics="true" pageId="7" pageNumber="8">esb</emphasis>
|
||
and
|
||
<emphasis id="B907771AFF9DFFA3FB9A3FCDE5113C96" box="[1032,1058,1428,1453]" italics="true" pageId="7" pageNumber="8">ist</emphasis>
|
||
forming a straight oblique row at base of fixed chelal finger;
|
||
<emphasis id="B907771AFF9DFFA3FE5F3FE1E0EE3CEA" box="[461,477,1464,1489]" italics="true" pageId="7" pageNumber="8">it</emphasis>
|
||
slightly distal to
|
||
<emphasis id="B907771AFF9DFFA3FD303FE1E3F23CEA" box="[674,705,1464,1489]" italics="true" pageId="7" pageNumber="8">est</emphasis>
|
||
, situated subdistally;
|
||
<emphasis id="B907771AFF9DFFA3FC253FE1E2F83CEA" box="[951,971,1464,1489]" italics="true" pageId="7" pageNumber="8">et</emphasis>
|
||
slightly near to tip of fixed chelal finger, very close to chelal teeth;
|
||
<emphasis id="B907771AFF9DFFA3FE2F3F82E0E43CCE" box="[445,471,1499,1525]" italics="true" pageId="7" pageNumber="8">dx</emphasis>
|
||
situated distal to
|
||
<emphasis id="B907771AFF9DFFA3FD0C3F85E3813CCE" box="[670,690,1500,1525]" italics="true" pageId="7" pageNumber="8">et</emphasis>
|
||
;
|
||
<emphasis id="B907771AFF9DFFA3FD533F82E3E93CCE" box="[705,730,1499,1525]" italics="true" pageId="7" pageNumber="8">sb</emphasis>
|
||
situated midway between
|
||
<emphasis id="B907771AFF9DFFA3FB973F82E5203CCE" box="[1029,1043,1499,1525]" italics="true" pageId="7" pageNumber="8">b</emphasis>
|
||
and
|
||
<emphasis id="B907771AFF9DFFA3FBD83F85E56E3CCE" box="[1098,1117,1500,1525]" italics="true" pageId="7" pageNumber="8">st</emphasis>
|
||
;
|
||
<emphasis id="B907771AFF9DFFA3FBFE3F82E5493CCE" box="[1132,1146,1499,1525]" italics="true" pageId="7" pageNumber="8">b</emphasis>
|
||
and
|
||
<emphasis id="B907771AFF9DFFA3FB223F85E58B3CCE" box="[1200,1208,1500,1525]" italics="true" pageId="7" pageNumber="8">t</emphasis>
|
||
situated subdistally,
|
||
<emphasis id="B907771AFF9DFFA3FF053C59E1AC3F22" box="[151,159,1536,1561]" italics="true" pageId="7" pageNumber="8">t</emphasis>
|
||
situated distal to
|
||
<emphasis id="B907771AFF9DFFA3FEF83C59E0493F22" box="[362,378,1536,1561]" italics="true" pageId="7" pageNumber="8">it</emphasis>
|
||
;
|
||
<emphasis id="B907771AFF9DFFA3FE1B3C59E09B3F22" box="[393,424,1536,1561]" italics="true" pageId="7" pageNumber="8">est</emphasis>
|
||
situated distal to
|
||
<emphasis id="B907771AFF9DFFA3FDE13FA6E3B23F22" box="[627,641,1535,1561]" italics="true" pageId="7" pageNumber="8">b</emphasis>
|
||
(
|
||
<figureCitation id="1348B78DFF9DFFA3FD003FA6E3DA3F21" box="[658,745,1535,1562]" captionStart="FIGURE 5" captionStartId="9.[151,250,1894,1918]" captionTargetBox="[197,1391,181,1870]" captionTargetId="figure-18@9.[197,1391,181,1870]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius biyunensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg Ⅰ (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.8090062" httpUri="https://zenodo.org/record/8090062/files/figure.png" pageId="7" pageNumber="8">Fig. 5F</figureCitation>
|
||
). A tiny retrolateral lyrifissure present at base of fixed chelal finger (situated distal to
|
||
<emphasis id="B907771AFF9DFFA3FE2A3C7DE0E13F06" box="[440,466,1572,1597]" italics="true" pageId="7" pageNumber="8">ist</emphasis>
|
||
). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 29–30 macrodenticles, slightly retrorse and pointed; movable chelal finger with 27–29 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed (
|
||
<figureCitation id="1348B78DFF9DFFA3FAAA3CD6E4A33F91" box="[1336,1424,1679,1706]" captionStart="FIGURE 5" captionStartId="9.[151,250,1894,1918]" captionTargetBox="[197,1391,181,1870]" captionTargetId="figure-18@9.[197,1391,181,1870]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius biyunensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg Ⅰ (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.8090062" httpUri="https://zenodo.org/record/8090062/files/figure.png" pageId="7" pageNumber="8">Fig. 5F</figureCitation>
|
||
). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (
|
||
<figureCitation id="1348B78DFF9DFFA3FF0D3C8EE1C93FCA" box="[159,250,1751,1777]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[179,1408,196,1919]" captionTargetId="figure-20@8.[179,1408,181,1919]" captionTargetPageId="8" captionText="FIGURE 4. Lagynochthonius biyunensis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.8090058" httpUri="https://zenodo.org/record/8090058/files/figure.png" pageId="7" pageNumber="8">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="1348B78DFF9DFFA3FE943C8EE01E3FC9" box="[262,301,1751,1778]" captionStart="FIGURE 5" captionStartId="9.[151,250,1894,1918]" captionTargetBox="[197,1391,181,1870]" captionTargetId="figure-18@9.[197,1391,181,1870]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius biyunensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg Ⅰ (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.8090062" httpUri="https://zenodo.org/record/8090062/files/figure.png" pageId="7" pageNumber="8">5G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCCAB08FF9DFFA3FF553CA2E44C3EBA" blockId="7.[151,1437,923,1922]" pageId="7" pageNumber="8">
|
||
Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy Ⅰ–XII: 4: 4: 4: 4: 4: 4–5: 4: 4–5: 4–5: 4: T2T: 0, tergites VIII–IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 13–14: 12–14: 9: 9: 8–9: 9: 9: 9: 0: 2. Anterior genital operculum with ten setae, genital opening slit-like, with 11–13 marginal setae on each side,
|
||
<quantity id="4C8B06EDFF9DFFA3FB0E3D3EE5E63EBA" box="[1180,1237,1895,1921]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" pageId="7" pageNumber="8" unit="in" value="34.0">34 in</quantity>
|
||
total (
|
||
<figureCitation id="1348B78DFF9DFFA3FA8F3D3EE4403EBA" box="[1309,1395,1895,1921]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[179,1408,196,1919]" captionTargetId="figure-20@8.[179,1408,181,1919]" captionTargetPageId="8" captionText="FIGURE 4. Lagynochthonius biyunensis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.8090058" httpUri="https://zenodo.org/record/8090058/files/figure.png" pageId="7" pageNumber="8">Fig. 4F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF0CFB80FF92FFACFF053DCEE2B73ECC" ID-DOI="http://doi.org/10.5281/zenodo.8090058" ID-Zenodo-Dep="8090058" httpUri="https://zenodo.org/record/8090058/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1943,1967]" targetBox="[179,1408,196,1919]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8BCCAB08FF92FFACFF053DCEE2B73ECC" blockId="8.[151,1436,1943,2040]" pageId="8" pageNumber="9">
|
||
<emphasis id="B907771AFF92FFACFF053DCEE0273E94" bold="true" box="[151,276,1943,1967]" pageId="8" pageNumber="9">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C73D08BFF92FFACFE893DCEE30F3E94" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[283,572,1943,1967]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="biyunensis" status="sp. nov.">
|
||
<emphasis id="B907771AFF92FFACFE893DCEE30F3E94" box="[283,572,1943,1967]" italics="true" pageId="8" pageNumber="9">Lagynochthonius biyunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B907771AFF92FFACFDD13DC1E3A73E8B" bold="true" box="[579,660,1944,1968]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="A234CA61FF92FFACFDD13DC1E3A73E8B" box="[579,660,1944,1968]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0CFB80FF93FFADFF053D3FE27A3ED1" ID-DOI="http://doi.org/10.5281/zenodo.8090062" ID-Zenodo-Dep="8090062" httpUri="https://zenodo.org/record/8090062/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,250,1894,1918]" targetBox="[197,1391,181,1870]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8BCCAB08FF93FFADFF053D3FE27A3ED1" blockId="9.[151,1436,1894,2026]" pageId="9" pageNumber="10">
|
||
<emphasis id="B907771AFF93FFADFF053D3FE0273E45" bold="true" box="[151,276,1894,1918]" pageId="9" pageNumber="10">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C73D08BFF93FFADFE893D3FE30F3E45" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[283,572,1894,1918]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="biyunensis" status="sp. nov.">
|
||
<emphasis id="B907771AFF93FFADFE893D3FE30F3E45" box="[283,572,1894,1918]" italics="true" pageId="9" pageNumber="10">Lagynochthonius biyunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B907771AFF93FFADFDD13D3FE3A63E45" bold="true" box="[579,661,1894,1918]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="A234CA61FF93FFADFDD13D3FE3A63E45" box="[579,661,1894,1918]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg Ⅰ (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (A–B, E–I); 0.10 mm (C–D).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCCAB08FF90FFAEFF553ACEE5FD385E" blockId="10.[151,1437,150,1149]" pageId="10" pageNumber="11">
|
||
Legs (
|
||
<figureCitation id="1348B78DFF90FFAEFE993ACEE048398A" box="[267,379,150,177]" captionStart="FIGURE 5" captionStartId="9.[151,250,1894,1918]" captionTargetBox="[197,1391,181,1870]" captionTargetId="figure-18@9.[197,1391,181,1870]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius biyunensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg Ⅰ (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.8090062" httpUri="https://zenodo.org/record/8090062/files/figure.png" pageId="10" pageNumber="11">Fig. 5H–I</figureCitation>
|
||
): generally typical, long and slender. Fine granulation present on anterodorsal faces of trochanter IV, femur Ⅰ, IV and patella IV. Femur of leg Ⅰ 1.79–1.86 times longer than patella and with one lyrifissure at the base of femur; tarsus 2.38–2.47 times longer than tibia. Femoropatella of leg IV 4.33–4.38 times longer than deep; tibia 6.55–6.64 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.78–4.00 times longer than deep (TS = 0.32–0.34), tarsus 14.50–14.83 times longer than deep and 2.62–2.72 times longer than metatarsus (TS = 0.27–0.33). Arolium slightly shorter than the claws, not divided; claws simple.
|
||
</paragraph>
|
||
<paragraph id="8BCCAB08FF90FFAEFF553B36E44B3B22" blockId="10.[151,1437,150,1149]" pageId="10" pageNumber="11">
|
||
Adult females (
|
||
<typeStatus id="54C815AAFF90FFAEFEE03B36E0D238B2" box="[370,481,367,393]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
|
||
) (
|
||
<figureCitation id="1348B78DFF90FFAEFE643B36E36238B2" box="[502,593,367,393]" captionStart="FIGURE 3" captionStartId="7.[151,250,781,805]" captionTargetBox="[159,1430,188,757]" captionTargetId="figure-18@7.[151,1436,181,757]" captionTargetPageId="7" captionText="FIGURE 3. Lagynochthonius biyunensis sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8090056" httpUri="https://zenodo.org/record/8090056/files/figure.png" pageId="10" pageNumber="11">Figs 3B</figureCitation>
|
||
,
|
||
<figureCitation id="1348B78DFF90FFAEFDCE3B36E3B038B2" box="[604,643,367,393]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[179,1408,196,1919]" captionTargetId="figure-20@8.[179,1408,181,1919]" captionTargetPageId="8" captionText="FIGURE 4. Lagynochthonius biyunensis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.8090058" httpUri="https://zenodo.org/record/8090058/files/figure.png" pageId="10" pageNumber="11">4G</figureCitation>
|
||
). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, Ⅰ 3, II 3–4, III 5, IV 5; tergal chaetotaxy Ⅰ–XII: 4: 4: 4: 4: 4: 4: 4: 4: 4: 4: T2T: 0; sternal chaetotaxy IV–XII: 12–13: 10: 9: 9: 9: 9: 9: 0: 2; anterior genital operculum with ten setae, posterior margin with 12–14 marginal setae,
|
||
<quantity id="4C8B06EDFF90FFAEFB693BEEE46E38EA" box="[1275,1373,439,465]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" metricValueMax="6.096" metricValueMin="5.588" pageId="10" pageNumber="11" unit="in" value="23.0" valueMax="24.0" valueMin="22.0">22–24 in</quantity>
|
||
total; leg IV with a long tactile seta on both tarsal segments: metatarsus 4.00–4.63 times longer than deep (TS = 0.32– 0.34), tarsus 14.17–16.17 times longer than deep and 2.62–2.66 times longer than metatarsus (TS = 0.22–0.28).
|
||
</paragraph>
|
||
<paragraph id="8BCCAB08FF90FFAEFF55387AE0DE3A02" blockId="10.[151,1437,150,1149]" pageId="10" pageNumber="11">Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 2.02–2.10. Pedipalps: trochanter 0.24/0.19–0.20 (1.20–1.26), femur 1.29/0.16 (8.06), patella 0.47/0.20 (2.35), chela 1.84–1.85/0.26–0.27 (6.85–7.08), hand 0.73/0.26–0.27 (2.70–2.81), movable chelal finger length 1.09–1.11. Chelicera 0.72–0.74/0.30 (2.40–2.47), movable finger length 0.39–0.40. Carapace 0.60/0.60 (1.00). Leg Ⅰ: trochanter 0.19/0.16 (1.19), femur 0.69–0.70/0.09 (7.67–7.78), patella 0.37–0.39/0.08 (4.63–4.88), tibia 0.32/0.06 (5.33), tarsus 0.76–0.79/0.05–0.06 (13.17–15.20). Leg IV: trochanter 0.27–0.29/0.16 (1.69–1.81), femoropatella 1.04–1.05/0.24 (4.33–4.38), tibia 0.72–0.73/0.11 (6.55–6.64), metatarsus 0.32–0.34/0.08–0.09 (3.78–4.00), tarsus 0.87–0.89/0.06 (14.50–14.83).</paragraph>
|
||
<paragraph id="8BCCAB08FF90FFAEFF55391AE3063D62" blockId="10.[151,1437,150,1149]" pageId="10" pageNumber="11">Females: body length 2.19–2.27. Pedipalps: trochanter 0.24–0.27/0.20–0.21 (1.20–1.29), femur 1.33– 1.42/0.17–0.18 (7.82–7.89), patella 0.46–0.51/0.20–0.22 (2.30–2.32), chela 1.89–2.08/0.26–0.28 (7.27–7.43), hand 0.75–0.82/0.26–0.28 (2.88–2.93), movable chelal finger length 1.13–1.24. Chelicera 0.76–0.83/0.33–0.34 (2.30– 2.44), movable finger length 0.41–0.45. Carapace 0.64/0.64–0.72 (0.89–1.00). Leg Ⅰ: trochanter 0.18–0.19/0.16 (1.13–1.19), femur 0.69–0.80/0.09 (7.67–8.89), patella 0.39–0.43/0.08 (4.88–5.38), tibia 0.32–0.36/0.06–0.07 (5.14–5.33), tarsus 0.79–0.86/0.06 (13.17–14.33). Leg IV: trochanter 0.28–0.30/0.16 (1.75–1.88), femoropatella 1.03–1.11/0.23–0.24 (4.48–4.63), tibia 0.69–0.75/0.11–0.12 (6.25–6.27), metatarsus 0.32–0.37/0.08 (4.00–4.63), tarsus 0.85–0.97/0.06 (14.17–16.17).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C369F883FF90FFAEFF553E3BE3C23D46" box="[199,753,1122,1149]" pageId="10" pageNumber="11" type="distribution">
|
||
<paragraph id="8BCCAB08FF90FFAEFF553E3BE3C23D46" blockId="10.[151,1437,150,1149]" box="[199,753,1122,1149]" pageId="10" pageNumber="11">
|
||
<emphasis id="B907771AFF90FFAEFF553E3BE0523D47" bold="true" box="[199,353,1122,1148]" pageId="10" pageNumber="11">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54C815AAFF90FFAEFDF23E3AE3A33D46" box="[608,656,1123,1149]" pageId="10" pageNumber="11">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |