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<document id="49D244572019199D92F5D83F0158B8C9" ID-CLB-Dataset="55245" ID-DOI="10.11646/zootaxa.5309.1.1" ID-GBIF-Dataset="048abd7b-8c78-494e-b466-b614aeef7f6b" ID-ISSN="1175-5326" ID-Zenodo-Dep="8090046" ID-ZooBank="C930242A-041C-4CEC-8B2F-2DB671A2395D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1687949280324" checkinUser="plazi" docAuthor="Hou, Yanmeng, Feng, Zegang &amp; Zhang, Feng" docDate="2023" docId="03DA1A1EFF90FFABFF053E97E3C23BF6" docLanguage="en" docName="zootaxa.5309.1.1.pdf" docOrigin="Zootaxa 5309 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5309.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lagynochthonius guanniuensis Hou &amp; Feng &amp; Zhang 2023, sp. nov." docType="treatment" docVersion="7" lastPageNumber="16" masterDocId="FFE36266FF9AFFA4FF923A59E133393B" masterDocTitle="New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China" masterLastPageNumber="64" masterPageNumber="1" pageNumber="11" updateTime="1699537060991" updateUser="ExternalLinkService">
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<mods:title id="3C4D8F65404F5E586843B3DCFE0E2BB0">New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China</mods:title>
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<mods:namePart id="8D3C6A7D790D9AE12091E646374B2D6B">Hou, Yanmeng</mods:namePart>
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<mods:namePart id="4383C9F4CAC7E1A2F621C570FFA1839C">Feng, Zegang</mods:namePart>
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<mods:namePart id="E24CAC2C05926094294B8D57F4F16A87">Zhang, Feng</mods:namePart>
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<treatment id="03DA1A1EFF90FFABFF053E97E3C23BF6" ID-DOI="http://doi.org/10.5281/zenodo.8090036" ID-GBIF-Taxon="210479511" ID-Zenodo-Dep="8090036" LSID="urn:lsid:plazi:treatment:03DA1A1EFF90FFABFF053E97E3C23BF6" httpUri="http://treatment.plazi.org/id/03DA1A1EFF90FFABFF053E97E3C23BF6" lastPageId="15" lastPageNumber="16" pageId="10" pageNumber="11">
<subSubSection id="C369F883FF90FFAEFF053E97E2013DDC" box="[151,818,1230,1257]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8BCCAB08FF90FFAEFF053E97E2013DDC" blockId="10.[151,818,1230,1293]" box="[151,818,1230,1257]" pageId="10" pageNumber="11">
<heading id="D0841C64FF90FFAEFF053E97E2013DDC" bold="true" box="[151,818,1230,1257]" fontSize="11" level="2" pageId="10" pageNumber="11" reason="4">
<emphasis id="B907771AFF90FFAEFF053E97E2013DDC" bold="true" box="[151,818,1230,1257]" pageId="10" pageNumber="11">
<taxonomicName id="4C73D08BFF90FFAEFF053E97E3263DD2" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[151,533,1230,1257]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="guanniuensis" status="sp. nov.">
<emphasis id="B907771AFF90FFAEFF053E97E3263DD2" bold="true" box="[151,533,1230,1257]" italics="true" pageId="10" pageNumber="11">Lagynochthonius guanniuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234CA61FF90FFAEFD8F3E96E3493DD2" box="[541,634,1231,1257]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
(ΪϮṄṁae)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C369F883FF90FFAEFF053EAAE1CB3C36" box="[151,248,1267,1293]" pageId="10" pageNumber="11" type="description">
<paragraph id="8BCCAB08FF90FFAEFF053EAAE1CB3C36" blockId="10.[151,818,1230,1293]" box="[151,248,1267,1293]" pageId="10" pageNumber="11">
<figureCitation id="1348B78DFF90FFAEFF053EAAE1CB3C36" box="[151,248,1267,1293]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStartId-0="11.[151,250,1224,1248]" captionStartId-1="12.[151,250,1398,1422]" captionStartId-2="13.[151,250,1843,1867]" captionStartId-3="14.[151,250,1889,1913]" captionTargetBox-0="[151,1436,237,1201]" captionTargetBox-1="[290,1297,181,1374]" captionTargetBox-2="[186,1436,185,1819]" captionTargetBox-3="[198,1388,181,1865]" captionTargetId-0="figure-47@11.[151,1436,237,1201]" captionTargetId-1="figure-20@12.[290,1297,181,1374]" captionTargetId-2="figure-18@13.[151,1436,181,1819]" captionTargetId-3="figure-20@14.[198,1388,181,1865]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="FIGURE 6. Guanniu Cave, type locality of Lagynochthonius guanniuensis sp. nov., A. Entrance; B. A narrow tunnel to the interior of the cave; C. Area where L. guanniuensis sp. nov. specimens were collected; D. Live female of L. guanniuensis sp. nov. in its natural environment." captionText-1="FIGURE 7. Lagynochthonius guanniuensis sp. nov., holotype female, habitus (dorsal view)." captionText-2="FIGURE 8. Lagynochthonius guanniuensis sp. nov., holotype female:A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Right chelicera (dorsal view); F. Female genital area (ventral view); G. Left pedipalp (minus chela, dorsal view)." captionText-3="FIGURE 9. Lagynochthonius guanniuensis sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi-0="http://doi.org/10.5281/zenodo.8090064" figureDoi-1="http://doi.org/10.5281/zenodo.8090068" figureDoi-2="http://doi.org/10.5281/zenodo.8090070" figureDoi-3="http://doi.org/10.5281/zenodo.8090074" httpUri-0="https://zenodo.org/record/8090064/files/figure.png" httpUri-1="https://zenodo.org/record/8090068/files/figure.png" httpUri-2="https://zenodo.org/record/8090070/files/figure.png" httpUri-3="https://zenodo.org/record/8090074/files/figure.png" pageId="10" pageNumber="11">Figs 69</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C369F883FF90FFAEFF053F62E3673CFA" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BCCAB08FF90FFAEFF053F62E3673CFA" blockId="10.[151,1437,1338,2013]" pageId="10" pageNumber="11">
<emphasis id="B907771AFF90FFAEFF053F62E07B3C6F" bold="true" box="[151,328,1338,1365]" pageId="10" pageNumber="11">Type material.</emphasis>
<materialsCitation id="3B1BA155FF90FFAEFEC03F62E2F73CA6" ID-GBIF-Occurrence="4138993310" collectingDate="2020-08-15" collectorName="Zegang Feng &amp; Hongru Xu &amp; Yanmeng Hou" country="China" county="Xifeng County" elevation="853" latitude="27.223333" location="Wenquan Village" longLatPrecision="1" longitude="106.871666" municipality="Wenquan Town" pageId="10" pageNumber="11" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou" typeStatus="holotype">
<typeStatus id="54C815AAFF90FFAEFEC03F62E0893C6E" box="[338,442,1339,1365]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
<specimenCount id="9D756081FF90FFAEFE563F62E0EA3C6E" box="[452,473,1339,1365]" pageId="10" pageNumber="11" type="female"></specimenCount>
(Ps.-MHBU-GZXF-20-02-01):
<collectingCountry id="F364EB98FF90FFAEFCDA3F62E2BD3C6E" box="[840,910,1339,1365]" name="China" pageId="10" pageNumber="11">China</collectingCountry>
,
<collectingRegion id="49B765EAFF90FFAEFC0E3F62E55F3C6E" box="[924,1132,1339,1365]" country="China" name="Guizhou" pageId="10" pageNumber="11">Guizhou Province</collectingRegion>
,
<collectingCounty id="62ADD384FF90FFAEFBE93F62E4153C6E" box="[1147,1318,1339,1365]" pageId="10" pageNumber="11">Xifeng County</collectingCounty>
,
<collectingMunicipality id="6BA83172FF90FFAEFAA13F62E1EA3C42" pageId="10" pageNumber="11">Wenquan Town</collectingMunicipality>
,
<location id="8EACFDD3FF90FFAEFF763F06E09B3C42" LSID="urn:lsid:plazi:treatment:03DA1A1EFF90FFABFF053E97E3C23BF6:8EACFDD3FF90FFAEFF763F06E09B3C42" box="[228,424,1375,1401]" country="China" county="Xifeng County" latitude="27.223333" longLatPrecision="1" longitude="106.871666" municipality="Wenquan Town" name="Wenquan Village" pageId="10" pageNumber="11" stateProvince="Guizhou">Wenquan Village</location>
,
<location id="8EACFDD3FF90FFAEFE203F06E3653C42" LSID="urn:lsid:plazi:treatment:03DA1A1EFF90FFABFF053E97E3C23BF6:8EACFDD3FF90FFAEFE203F06E3653C42" box="[434,598,1375,1401]" country="China" county="Xifeng County" latitude="27.223333" longLatPrecision="1" longitude="106.871666" municipality="Wenquan Town" name="Guanniu Cave" pageId="10" pageNumber="11" stateProvince="Guizhou">Guanniu Cave</location>
, under the detritus in the deep zone [
<geoCoordinate id="EE47CDCFFF90FFAEFC633F39E5AE3C40" box="[1009,1181,1376,1403]" degrees="27" direction="north" minutes="13" orientation="latitude" pageId="10" pageNumber="11" precision="1" seconds="24.00" value="27.223333">27°13'24.00″N</geoCoordinate>
,
<geoCoordinate id="EE47CDCFFF90FFAEFB3B3F39E4533C40" box="[1193,1376,1376,1403]" degrees="106" direction="east" minutes="52" orientation="longitude" pageId="10" pageNumber="11" precision="1" seconds="18.00" value="106.871666">106°52'18.00″E</geoCoordinate>
],
<elevation id="005E4C3BFF90FFAEFAE03F07E1DA3CA6" metricMagnitude="2" metricUnit="m" metricValue="8.53" pageId="10" pageNumber="11" unit="m" value="853.0">
<quantity id="4C8B06EDFF90FFAEFAE03F07E19E3CA6" metricMagnitude="2" metricUnit="m" metricValue="8.53" pageId="10" pageNumber="11" unit="m" value="853.0">853 m</quantity>
a.s.l.
</elevation>
,
<date id="FFCD8DC8FF90FFAEFF673FDBE09A3CA6" box="[245,425,1410,1437]" pageId="10" pageNumber="11" value="2020-08-15">
<collectingDate id="EF897420FF90FFAEFF673FDBE09A3CA6" box="[245,425,1410,1437]" pageId="10" pageNumber="11" value="2020-08-15">15 August 2020</collectingDate>
</date>
,
<collectorName id="2686CEDEFF90FFAEFE213FDAE37A3CA6" box="[435,585,1411,1437]" pageId="10" pageNumber="11">Zegang Feng</collectorName>
,
<collectorName id="2686CEDEFF90FFAEFDC13FDAE3E23CA6" box="[595,721,1411,1437]" pageId="10" pageNumber="11">Hongru Xu</collectorName>
&amp;
<collectorName id="2686CEDEFF90FFAEFD613FDAE2A73CA6" box="[755,916,1411,1437]" pageId="10" pageNumber="11">Yanmeng Hou</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B1BA155FF90FFAEFC583FDAE3633CFA" ID-GBIF-Occurrence="4138993329" collectingDate="2020-08-15" collectorName="Zegang Feng &amp; Hongru Xu &amp; Yanmeng Hou" country="China" county="Xifeng County" elevation="853" latitude="27.223333" location="Wenquan Village" longLatPrecision="1" longitude="106.871666" municipality="Wenquan Town" pageId="10" pageNumber="11" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou" typeStatus="paratype">
<typeStatus id="54C815AAFF90FFAEFC583FDAE5033CA6" box="[970,1072,1411,1437]" pageId="10" pageNumber="11" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D756081FF90FFAEFBA83FDAE5573CA6" box="[1082,1124,1411,1437]" pageId="10" pageNumber="11" type="female">1 ♀</specimenCount>
(Ps.-MSWU-GZXF-20-02- 02), with the same data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C369F883FF90FFAEFF553F92E2633CDE" box="[199,848,1483,1509]" pageId="10" pageNumber="11" type="etymology">
<paragraph id="8BCCAB08FF90FFAEFF553F92E2633CDE" blockId="10.[151,1437,1338,2013]" box="[199,848,1483,1509]" pageId="10" pageNumber="11">
<emphasis id="B907771AFF90FFAEFF553F92E07D3CDE" bold="true" box="[199,334,1483,1509]" pageId="10" pageNumber="11">Etymology.</emphasis>
Named after the
<typeStatus id="54C815AAFF90FFAEFD823F92E3733CDE" box="[528,576,1483,1509]" pageId="10" pageNumber="11">type</typeStatus>
locality, Guanniu Cave.
</paragraph>
</subSubSection>
<subSubSection id="C369F883FF90FFAEFF553FB7E5AF3FA2" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="8BCCAB08FF90FFAEFF553FB7E5AF3FA2" blockId="10.[151,1437,1338,2013]" pageId="10" pageNumber="11">
<emphasis id="B907771AFF90FFAEFF553FB7E0433F37" bold="true" box="[199,368,1518,1548]" pageId="10" pageNumber="11">
Diagnosis (
<specimenCount id="9D756081FF90FFAEFEDE3FA9E0523F37" box="[332,353,1520,1548]" pageId="10" pageNumber="11" type="female"></specimenCount>
).
</emphasis>
Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome pointed and small, triangular; posterior margin of carapace with two setae; tergites IV each with two setae. Pedipalps slender, femur 8.198.47 times longer than broad; chela 6.867.03 times longer than broad; chelal fingers with intercalary teeth; chelal fingers markedly curved in dorsal view; chemosensory setae (
<emphasis id="B907771AFF90FFAEFD6A3C26E23C3FA3" box="[760,783,1663,1688]" italics="true" pageId="10" pageNumber="11">sc</emphasis>
) present on dorsum of chelal hand.
</paragraph>
</subSubSection>
<subSubSection id="C369F883FF90FFABFF553CFBE0463B92" lastPageId="15" lastPageNumber="16" pageId="10" pageNumber="11" type="description">
<paragraph id="8BCCAB08FF90FFAEFF553CFBE5063F86" blockId="10.[151,1437,1338,2013]" box="[199,1077,1698,1725]" pageId="10" pageNumber="11">
<emphasis id="B907771AFF90FFAEFF553CFBE06A3F87" bold="true" box="[199,345,1698,1724]" pageId="10" pageNumber="11">Description.</emphasis>
Females (
<typeStatus id="54C815AAFF90FFAEFE593CFAE31C3F86" box="[459,559,1699,1725]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
and
<typeStatus id="54C815AAFF90FFAEFDF43CFAE3F83F86" box="[614,715,1699,1725]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
), male unknown (
<figureCitation id="1348B78DFF90FFAEFC003CFAE2DC3F86" box="[914,1007,1699,1725]" captionStart="FIGURE 6" captionStartId="11.[151,250,1224,1248]" captionTargetBox="[151,1436,237,1201]" captionTargetId="figure-47@11.[151,1436,237,1201]" captionTargetPageId="11" captionText="FIGURE 6. Guanniu Cave, type locality of Lagynochthonius guanniuensis sp. nov., A. Entrance; B. A narrow tunnel to the interior of the cave; C. Area where L. guanniuensis sp. nov. specimens were collected; D. Live female of L. guanniuensis sp. nov. in its natural environment." figureDoi="http://doi.org/10.5281/zenodo.8090064" httpUri="https://zenodo.org/record/8090064/files/figure.png" pageId="10" pageNumber="11">Figs 6D</figureCitation>
,
<figureCitation id="1348B78DFF90FFAEFC693CFAE51A3F86" box="[1019,1065,1699,1725]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStartId-0="12.[151,250,1398,1422]" captionStartId-1="13.[151,250,1843,1867]" captionStartId-2="14.[151,250,1889,1913]" captionTargetBox-0="[290,1297,181,1374]" captionTargetBox-1="[186,1436,185,1819]" captionTargetBox-2="[198,1388,181,1865]" captionTargetId-0="figure-20@12.[290,1297,181,1374]" captionTargetId-1="figure-18@13.[151,1436,181,1819]" captionTargetId-2="figure-20@14.[198,1388,181,1865]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionText-0="FIGURE 7. Lagynochthonius guanniuensis sp. nov., holotype female, habitus (dorsal view)." captionText-1="FIGURE 8. Lagynochthonius guanniuensis sp. nov., holotype female:A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Right chelicera (dorsal view); F. Female genital area (ventral view); G. Left pedipalp (minus chela, dorsal view)." captionText-2="FIGURE 9. Lagynochthonius guanniuensis sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi-0="http://doi.org/10.5281/zenodo.8090068" figureDoi-1="http://doi.org/10.5281/zenodo.8090070" figureDoi-2="http://doi.org/10.5281/zenodo.8090074" httpUri-0="https://zenodo.org/record/8090068/files/figure.png" httpUri-1="https://zenodo.org/record/8090070/files/figure.png" httpUri-2="https://zenodo.org/record/8090074/files/figure.png" pageId="10" pageNumber="11">79</figureCitation>
).
</paragraph>
<paragraph id="8BCCAB08FF90FFAEFF553C9EE5DC3FDA" blockId="10.[151,1437,1338,2013]" box="[199,1263,1735,1761]" pageId="10" pageNumber="11">Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</paragraph>
<paragraph id="8BCCAB08FF90FFAFFF553CB2E28639EE" blockId="10.[151,1437,1338,2013]" lastBlockId="11.[151,1437,151,213]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Cephalothorax (
<figureCitation id="1348B78DFF90FFAEFEE83CB2E0E73E3E" box="[378,468,1771,1797]" captionStart="FIGURE 8" captionStartId="13.[151,250,1843,1867]" captionTargetBox="[186,1436,185,1819]" captionTargetId="figure-18@13.[151,1436,181,1819]" captionTargetPageId="13" captionText="FIGURE 8. Lagynochthonius guanniuensis sp. nov., holotype female:A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Right chelicera (dorsal view); F. Female genital area (ventral view); G. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8090070" httpUri="https://zenodo.org/record/8090070/files/figure.png" pageId="10" pageNumber="11">Figs 8C</figureCitation>
,
<figureCitation id="1348B78DFF90FFAEFE4D3CB2E3353E3E" box="[479,518,1771,1797]" captionStart="FIGURE 9" captionStartId="14.[151,250,1889,1913]" captionTargetBox="[198,1388,181,1865]" captionTargetId="figure-20@14.[198,1388,181,1865]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius guanniuensis sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090074" httpUri="https://zenodo.org/record/8090074/files/figure.png" pageId="10" pageNumber="11">9A</figureCitation>
): carapace 0.950.97 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome pointed and small, triangular; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa with small, rounded anteromedial process; coxae II with 89 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (
<figureCitation id="1348B78DFF91FFAFFEB53AE2E04C39EE" box="[295,383,187,213]" captionStart="FIGURE 9" captionStartId="14.[151,250,1889,1913]" captionTargetBox="[198,1388,181,1865]" captionTargetId="figure-20@14.[198,1388,181,1865]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius guanniuensis sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090074" httpUri="https://zenodo.org/record/8090074/files/figure.png" pageId="11" pageNumber="12">Fig. 9C</figureCitation>
); intercoxal tubercle absent; without sub-oral seta.
</paragraph>
<caption id="DF0CFB80FF91FFAFFF053E91E0E53C13" ID-DOI="http://doi.org/10.5281/zenodo.8090064" ID-Zenodo-Dep="8090064" httpUri="https://zenodo.org/record/8090064/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,1224,1248]" targetBox="[151,1436,237,1201]" targetPageId="11" targetType="figure">
<paragraph id="8BCCAB08FF91FFAFFF053E91E0E53C13" blockId="11.[151,1436,1224,1320]" pageId="11" pageNumber="12">
<emphasis id="B907771AFF91FFAFFF053E91E0263DDB" bold="true" box="[151,277,1224,1248]" pageId="11" pageNumber="12">FIGURE 6.</emphasis>
Guanniu Cave, type locality of
<taxonomicName id="4C73D08BFF91FFAFFDF63E91E2923DDB" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[612,929,1224,1248]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="guanniuensis" status="sp. nov.">
<emphasis id="B907771AFF91FFAFFDF63E91E2923DDB" box="[612,929,1224,1248]" italics="true" pageId="11" pageNumber="12">Lagynochthonius guanniuensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFF91FFAFFC3B3E91E2C83DDB" bold="true" box="[937,1019,1224,1248]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="A234CA61FF91FFAFFC3B3E91E2C83DDB" box="[937,1019,1224,1248]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Entrance; B. A narrow tunnel to the interior of the cave; C. Area where
<taxonomicName id="4C73D08BFF91FFAFFD913EB4E3973C3F" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[515,676,1261,1284]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="guanniuensis" status="sp. nov.">
<emphasis id="B907771AFF91FFAFFD913EB4E3973C3F" box="[515,676,1261,1284]" italics="true" pageId="11" pageNumber="12">L. guanniuensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFF91FFAFFD393EB5E3CE3C3F" bold="true" box="[683,765,1260,1284]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="A234CA61FF91FFAFFD393EB5E3CE3C3F" box="[683,765,1260,1284]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens were collected; D. Live female of
<taxonomicName id="4C73D08BFF91FFAFFB473EB4E4443C3F" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[1237,1399,1261,1284]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="guanniuensis" status="sp. nov.">
<emphasis id="B907771AFF91FFAFFB473EB4E4443C3F" box="[1237,1399,1261,1284]" italics="true" pageId="11" pageNumber="12">L. guanniuensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFF91FFAFFAEC3EB5E1F03C13" bold="true" pageId="11" pageNumber="12">
<taxonomicNameLabel id="A234CA61FF91FFAFFAEC3EB5E1F03C13" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in its natural environment.
</paragraph>
</caption>
<paragraph id="8BCCAB08FF91FFAFFF553F23E2893FAA" blockId="11.[151,1437,1402,2041]" pageId="11" pageNumber="12">
Chelicera (
<figureCitation id="1348B78DFF91FFAFFED23F23E08F3CAE" box="[320,444,1402,1429]" captionStart="FIGURE 8" captionStartId="13.[151,250,1843,1867]" captionTargetBox="[186,1436,185,1819]" captionTargetId="figure-18@13.[151,1436,181,1819]" captionTargetPageId="13" captionText="FIGURE 8. Lagynochthonius guanniuensis sp. nov., holotype female:A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Right chelicera (dorsal view); F. Female genital area (ventral view); G. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8090070" httpUri="https://zenodo.org/record/8090070/files/figure.png" pageId="11" pageNumber="12">Figs 8DE</figureCitation>
,
<figureCitation id="1348B78DFF91FFAFFE543F22E0D93CAE" box="[454,490,1403,1429]" captionStart="FIGURE 9" captionStartId="14.[151,250,1889,1913]" captionTargetBox="[198,1388,181,1865]" captionTargetId="figure-20@14.[198,1388,181,1865]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius guanniuensis sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090074" httpUri="https://zenodo.org/record/8090074/files/figure.png" pageId="11" pageNumber="12">9B</figureCitation>
): large, about as long as carapace, 2.482.50 times longer than broad; five setae (right chelicera with six setae) and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 1112 teeth, distal one largest; movable finger with 1517 retrorse contiguous small teeth; galea absent (
<figureCitation id="1348B78DFF91FFAFFAFD3C53E18F3F72" captionStart="FIGURE 9" captionStartId="14.[151,250,1889,1913]" captionTargetBox="[198,1388,181,1865]" captionTargetId="figure-20@14.[198,1388,181,1865]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius guanniuensis sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090074" httpUri="https://zenodo.org/record/8090074/files/figure.png" pageId="11" pageNumber="12">Fig. 9B</figureCitation>
). Serrula exterior with 2123 blades and serrula interior with 1314 blades. Rallum with eight blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (
<figureCitation id="1348B78DFF91FFAFFCC13C2FE29E3FAA" box="[851,941,1654,1681]" captionStart="FIGURE 9" captionStartId="14.[151,250,1889,1913]" captionTargetBox="[198,1388,181,1865]" captionTargetId="figure-20@14.[198,1388,181,1865]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius guanniuensis sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090074" httpUri="https://zenodo.org/record/8090074/files/figure.png" pageId="11" pageNumber="12">Fig. 9D</figureCitation>
).
</paragraph>
<paragraph id="8BCCAB08FF91FFA8FF553CC3E0F33FE8" blockId="11.[151,1437,1402,2041]" lastBlockId="12.[151,1437,1505,1891]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Pedipalp (
<figureCitation id="1348B78DFF91FFAFFEA73CC3E0FD3F8E" box="[309,462,1690,1717]" captionStart="FIGURE 8" captionStartId="13.[151,250,1843,1867]" captionTargetBox="[186,1436,185,1819]" captionTargetId="figure-18@13.[151,1436,181,1819]" captionTargetPageId="13" captionText="FIGURE 8. Lagynochthonius guanniuensis sp. nov., holotype female:A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Right chelicera (dorsal view); F. Female genital area (ventral view); G. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8090070" httpUri="https://zenodo.org/record/8090070/files/figure.png" pageId="11" pageNumber="12">Figs8AB, G</figureCitation>
,
<figureCitation id="1348B78DFF91FFAFFE443CC2E32E3F8E" box="[470,541,1691,1717]" captionStart="FIGURE 9" captionStartId="14.[151,250,1889,1913]" captionTargetBox="[198,1388,181,1865]" captionTargetId="figure-20@14.[198,1388,181,1865]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius guanniuensis sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090074" httpUri="https://zenodo.org/record/8090074/files/figure.png" pageId="11" pageNumber="12">9EG</figureCitation>
):long and slender, trochanter1.231.33, femur 8.198.47,patella 2.392.52, chela 6.867.03, hand 2.572.62 times longer than broad; femur 2.472.62 times longer than patella; movable chelal finger 1.651.67 times longer than hand and 0.62 times longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (
<figureCitation id="1348B78DFF91FFAFFE723D5FE30F3E1A" box="[480,572,1798,1825]" captionStart="FIGURE 8" captionStartId="13.[151,250,1843,1867]" captionTargetBox="[186,1436,185,1819]" captionTargetId="figure-18@13.[151,1436,181,1819]" captionTargetPageId="13" captionText="FIGURE 8. Lagynochthonius guanniuensis sp. nov., holotype female:A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Right chelicera (dorsal view); F. Female genital area (ventral view); G. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8090070" httpUri="https://zenodo.org/record/8090070/files/figure.png" pageId="11" pageNumber="12">Figs 8G</figureCitation>
,
<figureCitation id="1348B78DFF91FFAFFDD53D5EE3593E1A" box="[583,618,1799,1825]" captionStart="FIGURE 9" captionStartId="14.[151,250,1889,1913]" captionTargetBox="[198,1388,181,1865]" captionTargetId="figure-20@14.[198,1388,181,1865]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius guanniuensis sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090074" httpUri="https://zenodo.org/record/8090074/files/figure.png" pageId="11" pageNumber="12">9E</figureCitation>
). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria,
<emphasis id="B907771AFF91FFAFFB1A3D17E5AD3E53" box="[1160,1182,1870,1896]" italics="true" pageId="11" pageNumber="12">ib</emphasis>
and
<emphasis id="B907771AFF91FFAFFB4A3D17E5CA3E53" box="[1240,1273,1870,1896]" italics="true" pageId="11" pageNumber="12">isb</emphasis>
situated close together, submedially on dorsum of chelal hand;
<emphasis id="B907771AFF91FFAFFD233D2BE3F83EB7" box="[689,715,1906,1932]" italics="true" pageId="11" pageNumber="12">eb</emphasis>
,
<emphasis id="B907771AFF91FFAFFD4A3D2BE3CE3EB7" box="[728,765,1906,1932]" italics="true" pageId="11" pageNumber="12">esb</emphasis>
and
<emphasis id="B907771AFF91FFAFFCA33D2AE2783EB7" box="[817,843,1907,1932]" italics="true" pageId="11" pageNumber="12">ist</emphasis>
forming an oblique row at base of fixed chelal finger;
<emphasis id="B907771AFF91FFAFFF053DCEE1943E8B" box="[151,167,1943,1968]" italics="true" pageId="11" pageNumber="12">it</emphasis>
slightly distal to
<emphasis id="B907771AFF91FFAFFEFA3DCEE0B43E8B" box="[360,391,1943,1968]" italics="true" pageId="11" pageNumber="12">est</emphasis>
, situated subdistally;
<emphasis id="B907771AFF91FFAFFDE83DCEE3BD3E8B" box="[634,654,1943,1968]" italics="true" pageId="11" pageNumber="12">et</emphasis>
slightly near to tip of fixed chelal finger, very close to chelal teeth;
<emphasis id="B907771AFF91FFAFFA103DCFE4AF3E8B" box="[1410,1436,1942,1968]" italics="true" pageId="11" pageNumber="12">dx</emphasis>
situated distal to
<emphasis id="B907771AFF91FFAFFEC73DE2E05A3EEF" box="[341,361,1979,2004]" italics="true" pageId="11" pageNumber="12">et</emphasis>
;
<emphasis id="B907771AFF91FFAFFEEA3DE3E0A23EEF" box="[376,401,1978,2004]" italics="true" pageId="11" pageNumber="12">sb</emphasis>
situated midway between
<emphasis id="B907771AFF91FFAFFD293DE3E3FA3EEF" box="[699,713,1978,2004]" italics="true" pageId="11" pageNumber="12">b</emphasis>
and
<emphasis id="B907771AFF91FFAFFD6D3DE2E2213EEF" box="[767,786,1979,2004]" italics="true" pageId="11" pageNumber="12">st</emphasis>
;
<emphasis id="B907771AFF91FFAFFCB23DE3E21D3EEF" box="[800,814,1978,2004]" italics="true" pageId="11" pageNumber="12">b</emphasis>
and
<emphasis id="B907771AFF91FFAFFCF73DE2E25E3EEF" box="[869,877,1979,2004]" italics="true" pageId="11" pageNumber="12">t</emphasis>
situated subdistally,
<emphasis id="B907771AFF91FFAFFBC53DE2E56C3EEF" box="[1111,1119,1979,2004]" italics="true" pageId="11" pageNumber="12">t</emphasis>
situated at the same level as
<emphasis id="B907771AFF91FFAFFF053D86E1943EC3" box="[151,167,2015,2040]" italics="true" pageId="11" pageNumber="12">it</emphasis>
and distal to
<emphasis id="B907771AFF91FFAFFEAF3D87E0783EC3" box="[317,331,2014,2040]" italics="true" pageId="11" pageNumber="12">b</emphasis>
;
<emphasis id="B907771AFF91FFAFFECB3D86E04B3EC3" box="[345,376,2015,2040]" italics="true" pageId="11" pageNumber="12">est</emphasis>
situated distal to
<emphasis id="B907771AFF91FFAFFDAF3D87E3783EC3" box="[573,587,2014,2040]" italics="true" pageId="11" pageNumber="12">b</emphasis>
(
<figureCitation id="1348B78DFF91FFAFFDC83D87E39C3EC2" box="[602,687,2014,2041]" captionStart="FIGURE 9" captionStartId="14.[151,250,1889,1913]" captionTargetBox="[198,1388,181,1865]" captionTargetId="figure-20@14.[198,1388,181,1865]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius guanniuensis sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090074" httpUri="https://zenodo.org/record/8090074/files/figure.png" pageId="11" pageNumber="12">Fig. 9F</figureCitation>
). A tiny retrolateral lyrifissure present at base of fixed chelal finger (situated distal to
<emphasis id="B907771AFF96FFA8FEF43FBBE0B33CC0" box="[358,384,1506,1531]" italics="true" pageId="12" pageNumber="13">ist</emphasis>
). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 33 macrodenticles, slightly retrorse and pointed, plus 25
<specimenCount id="9D756081FF96FFA8FC303C70E50E3F78" box="[930,1085,1577,1604]" pageId="12" pageNumber="13" type="intercalary">27 intercalary</specimenCount>
microdenticles,
<quantity id="4C8B06EDFF96FFA8FB6A3C70E4683F78" box="[1272,1371,1577,1604]" metricMagnitude="0" metricUnit="m" metricValue="1.4986" metricValueMax="1.524" metricValueMin="1.4731999999999998" pageId="12" pageNumber="13" unit="in" value="59.0" valueMax="60.0" valueMin="58.0">5860 in</quantity>
total; movable chelal finger with 29 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus
<specimenCount id="9D756081FF96FFA8FEC93C28E0C63FB0" box="[347,501,1649,1676]" pageId="12" pageNumber="13" type="intercalary">25 intercalary</specimenCount>
microdenticles and 67 vestigial, rounded and contiguous basal teeth,
<quantity id="4C8B06EDFF96FFA8FA913C28E4563FB0" box="[1283,1381,1649,1675]" metricMagnitude="0" metricUnit="m" metricValue="1.5367" metricValueMax="1.5493999999999999" metricValueMin="1.524" pageId="12" pageNumber="13" unit="in" value="60.5" valueMax="61.0" valueMin="60.0">6061 in</quantity>
total (
<figureCitation id="1348B78DFF96FFA8FF0D3CCCE1C53F94" box="[159,246,1685,1711]" captionStart="FIGURE 9" captionStartId="14.[151,250,1889,1913]" captionTargetBox="[198,1388,181,1865]" captionTargetId="figure-20@14.[198,1388,181,1865]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius guanniuensis sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090074" httpUri="https://zenodo.org/record/8090074/files/figure.png" pageId="12" pageNumber="13">Fig. 9F</figureCitation>
). Chelal fingers markedly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (
<figureCitation id="1348B78DFF96FFA8FEB43CE0E0B13FE8" box="[294,386,1721,1747]" captionStart="FIGURE 8" captionStartId="13.[151,250,1843,1867]" captionTargetBox="[186,1436,185,1819]" captionTargetId="figure-18@13.[151,1436,181,1819]" captionTargetPageId="13" captionText="FIGURE 8. Lagynochthonius guanniuensis sp. nov., holotype female:A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Right chelicera (dorsal view); F. Female genital area (ventral view); G. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8090070" httpUri="https://zenodo.org/record/8090070/files/figure.png" pageId="12" pageNumber="13">Figs 8B</figureCitation>
,
<figureCitation id="1348B78DFF96FFA8FE1C3CE0E0803FE8" box="[398,435,1721,1747]" captionStart="FIGURE 9" captionStartId="14.[151,250,1889,1913]" captionTargetBox="[198,1388,181,1865]" captionTargetId="figure-20@14.[198,1388,181,1865]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius guanniuensis sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090074" httpUri="https://zenodo.org/record/8090074/files/figure.png" pageId="12" pageNumber="13">9G</figureCitation>
).
</paragraph>
<caption id="DF0CFB80FF96FFA8FF053F2FE5753CB4" ID-DOI="http://doi.org/10.5281/zenodo.8090068" ID-Zenodo-Dep="8090068" box="[151,1094,1398,1423]" httpUri="https://zenodo.org/record/8090068/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,250,1398,1422]" targetBox="[290,1297,181,1374]" targetPageId="12" targetType="figure">
<paragraph id="8BCCAB08FF96FFA8FF053F2FE5753CB4" blockId="12.[151,1094,1398,1423]" box="[151,1094,1398,1423]" pageId="12" pageNumber="13">
<emphasis id="B907771AFF96FFA8FF053F2FE0203CB4" bold="true" box="[151,275,1398,1423]" pageId="12" pageNumber="13">FIGURE 7.</emphasis>
<taxonomicName id="4C73D08BFF96FFA8FE8B3F2EE3673CB5" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[281,596,1399,1423]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="guanniuensis" status="sp. nov.">
<emphasis id="B907771AFF96FFA8FE8B3F2EE3673CB5" box="[281,596,1399,1423]" italics="true" pageId="12" pageNumber="13">Lagynochthonius guanniuensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFF96FFA8FDC83F2EE3983CB4" bold="true" box="[602,683,1399,1423]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="A234CA61FF96FFA8FDC83F2EE3983CB4" box="[602,683,1399,1423]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female, habitus (dorsal view).
</paragraph>
</caption>
<paragraph id="8BCCAB08FF96FFA8FF553C84E2853E58" blockId="12.[151,1437,1505,1891]" pageId="12" pageNumber="13">
Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy XII: 2: 2: 2: 2: 4: 4: 4: 4: 5: 2: T2T: 0, tergite IX with an unpaired median seta. Sternal chaetotaxy IVXII: 1114: 89: 7: 7: 78: 7: 7: 0: 2. Anterior genital operculum with 1011 setae, posterior margin with 1213 marginal setae,
<quantity id="4C8B06EDFF96FFA8FD413D10E23F3E58" box="[723,780,1865,1891]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" pageId="12" pageNumber="13" unit="in" value="23.0">23 in</quantity>
total (
<figureCitation id="1348B78DFF96FFA8FCC63D10E29A3E58" box="[852,937,1865,1891]" captionStart="FIGURE 8" captionStartId="13.[151,250,1843,1867]" captionTargetBox="[186,1436,185,1819]" captionTargetId="figure-18@13.[151,1436,181,1819]" captionTargetPageId="13" captionText="FIGURE 8. Lagynochthonius guanniuensis sp. nov., holotype female:A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Right chelicera (dorsal view); F. Female genital area (ventral view); G. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8090070" httpUri="https://zenodo.org/record/8090070/files/figure.png" pageId="12" pageNumber="13">Fig. 8F</figureCitation>
).
</paragraph>
<caption id="DF0CFB80FF97FFA9FF053D6AE3AC3EA8" ID-DOI="http://doi.org/10.5281/zenodo.8090070" ID-Zenodo-Dep="8090070" httpUri="https://zenodo.org/record/8090070/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1843,1867]" targetBox="[186,1436,185,1819]" targetPageId="13" targetType="figure">
<paragraph id="8BCCAB08FF97FFA9FF053D6AE3AC3EA8" blockId="13.[151,1436,1843,1939]" pageId="13" pageNumber="14">
<emphasis id="B907771AFF97FFA9FF053D6AE0203E70" bold="true" box="[151,275,1843,1867]" pageId="13" pageNumber="14">FIGURE 8.</emphasis>
<taxonomicName id="4C73D08BFF97FFA9FE8B3D6AE3673E70" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[281,596,1843,1867]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="guanniuensis" status="sp. nov.">
<emphasis id="B907771AFF97FFA9FE8B3D6AE3673E70" box="[281,596,1843,1867]" italics="true" pageId="13" pageNumber="14">Lagynochthonius guanniuensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFF97FFA9FDC83D6AE3993E70" bold="true" box="[602,682,1843,1867]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A234CA61FF97FFA9FDC83D6AE3993E70" box="[602,682,1843,1867]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Right chelicera (dorsal view); F. Female genital area (ventral view); G. Left pedipalp (minus chela, dorsal view).
</paragraph>
</caption>
<caption id="DF0CFB80FF94FFAAFF053D38E2BF3EDE" ID-DOI="http://doi.org/10.5281/zenodo.8090074" ID-Zenodo-Dep="8090074" httpUri="https://zenodo.org/record/8090074/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1889,1913]" targetBox="[198,1388,181,1865]" targetPageId="14" targetType="figure">
<paragraph id="8BCCAB08FF94FFAAFF053D38E2BF3EDE" blockId="14.[151,1436,1889,2021]" pageId="14" pageNumber="15">
<emphasis id="B907771AFF94FFAAFF053D38E0263E42" bold="true" box="[151,277,1889,1913]" pageId="14" pageNumber="15">FIGURE 9.</emphasis>
<taxonomicName id="4C73D08BFF94FFAAFE8D3D38E36F3E42" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[287,604,1889,1913]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="guanniuensis" status="sp. nov.">
<emphasis id="B907771AFF94FFAAFE8D3D38E36F3E42" box="[287,604,1889,1913]" italics="true" pageId="14" pageNumber="15">Lagynochthonius guanniuensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFF94FFAAFDF73D38E38B3E42" bold="true" box="[613,696,1889,1913]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A234CA61FF94FFAAFDF73D38E38B3E42" box="[613,696,1889,1913]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD).
</paragraph>
</caption>
<paragraph id="8BCCAB08FF95FFABFF553ACEE52D385E" blockId="15.[151,1437,151,717]" pageId="15" pageNumber="16">
Legs (
<figureCitation id="1348B78DFF95FFABFE9E3ACEE04C398A" box="[268,383,151,177]" captionStart="FIGURE 9" captionStartId="14.[151,250,1889,1913]" captionTargetBox="[198,1388,181,1865]" captionTargetId="figure-20@14.[198,1388,181,1865]" captionTargetPageId="14" captionText="FIGURE 9. Lagynochthonius guanniuensis sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090074" httpUri="https://zenodo.org/record/8090074/files/figure.png" pageId="15" pageNumber="16">Fig. 9HI</figureCitation>
): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur , IV and patella IV. Femur of leg 1.911.95 times longer than patella and with one lyrifissure at the base of femur; tarsus 2.502.54 times longer than tibia. Femoropatella of leg IV 4.324.76 times longer than deep; tibia 6.187.00 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 4.224.63 times longer than deep (TS = 0.260.27), tarsus 14.5715.67 times longer than deep and 2.542.68 times longer than metatarsus (TS = 0.300.31). Arolium slightly shorter than the claws, not divided; claws simple.
</paragraph>
<paragraph id="8BCCAB08FF95FFABFF553B36E0463B92" blockId="15.[151,1437,151,717]" pageId="15" pageNumber="16">Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Females: body length 2.202.55. Pedipalps: trochanter 0.270.28/0.210.22 (1.231.33), femur 1.311.44/0.160.17 (8.19 8.47), patella 0.530.55/0.210.23 (2.392.52), chela 1.922.04/0.280.29 (6.867.03), hand 0.720.76/0.280.29 (2.572.62), movable chelal finger length 1.191.27. Chelicera 0.750.77/0.300.31 (2.482.50), movable finger length 0.40. Carapace 0.630.67/0.660.69 (0.950.97). Leg : trochanter 0.190.22/0.17 (1.121.29), femur 0.780.86/0.090.10 (8.608.67), patella 0.400.45/0.080.09 (5.00), tibia 0.320.35/0.060.07 (5.005.33), tarsus 0.800.89/0.060.07 (12.1713.33). Leg IV: trochanter 0.300.31/0.17 (1.721.82), femoropatella 1.081.19/0.25 (4.324.76), tibia 0.680.77/0.11 (6.187.00), metatarsus 0.370.38/0.080.09 (4.224.63), tarsus 0.941.02/0.06 0.07 (14.5715.67).</paragraph>
</subSubSection>
<subSubSection id="C369F883FF95FFABFF5538EBE3C23BF6" box="[199,753,690,717]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8BCCAB08FF95FFABFF5538EBE3C23BF6" blockId="15.[151,1437,151,717]" box="[199,753,690,717]" pageId="15" pageNumber="16">
<emphasis id="B907771AFF95FFABFF5538EBE0523BF7" bold="true" box="[199,353,690,716]" pageId="15" pageNumber="16">Distribution.</emphasis>
Known only from the
<typeStatus id="54C815AAFF95FFABFDF238EAE3A33BF6" box="[608,656,691,717]" pageId="15" pageNumber="16">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>