treatments-xml/data/03/BA/1C/03BA1C13FFA4FA5876E7A6C0FC13F81D.xml

<document id="6EF8A4BB74349822AC0847D476FC2C29" ID-DOI="10.5281/zenodo.213789" ID-GBIF-Dataset="dc33ba22-7a28-4dbc-a022-210421785006" ID-ISSN="1175-5326" ID-Zenodo-Dep="213789" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460552124884" checkinUser="plazi" docAuthor="Wang, Ying-Yong, Zhang, Tian-Du, Zhao, Jian, Sung, Yik-Hei &amp; Yang, Jian-Huan" docDate="2012" docId="03BA1C13FFA4FA5876E7A6C0FC13F81D" docLanguage="en" docName="zt03546p067.pdf" docOrigin="Zootaxa 3546" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Xenophrys jinggangensis Wang, sp. nov." docType="treatment" docVersion="8" lastPageNumber="62" masterDocId="FF83646BFFA1FA517670A225FFAEFFF5" masterDocTitle="Description of a new species of the genus Xenophrys Günther, 1864 (Amphibia: Anura: Megophryidae) from Mount Jinggang, China, based on molecular and morphological data" masterLastPageNumber="67" masterPageNumber="53" pageNumber="58" updateTime="1698318489320" updateUser="plazi">
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<mods:title id="1AB81DFD20658787401BC329DB9729CD">Description of a new species of the genus Xenophrys Günther, 1864 (Amphibia: Anura: Megophryidae) from Mount Jinggang, China, based on molecular and morphological data</mods:title>
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<mods:namePart id="AF2DA68B15D4B57BBAE83B5E88BA1B24">Wang, Ying-Yong</mods:namePart>
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<mods:namePart id="A46ECE5446BD6973DBA4F3393D66B031">Zhang, Tian-Du</mods:namePart>
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<mods:namePart id="45DA0CDE84AC5864AB13A811CED52A72">Zhao, Jian</mods:namePart>
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<mods:namePart id="C870219A2F6C5A6A5D7B65489C078A9C">Sung, Yik-Hei</mods:namePart>
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<mods:namePart id="96260535A0EB9963B9A3F8B58E44C760">Yang, Jian-Huan</mods:namePart>
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<mods:date id="AA1A2B023037047EBDB750E88FAB1B77">2012</mods:date>
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<treatment id="03BA1C13FFA4FA5876E7A6C0FC13F81D" ID-DOI="http://doi.org/10.5281/zenodo.5678529" ID-GBIF-Taxon="119533456" ID-Zenodo-Dep="5678529" LSID="urn:lsid:plazi:treatment:03BA1C13FFA4FA5876E7A6C0FC13F81D" httpUri="http://treatment.plazi.org/id/03BA1C13FFA4FA5876E7A6C0FC13F81D" lastPageId="9" lastPageNumber="62" pageId="5" pageNumber="58">
<subSubSection id="C309FE8EFFA4FA5476E7A6C0FDD0FB0A" box="[151,638,1253,1279]" pageId="5" pageNumber="58" type="nomenclature">
<paragraph id="8BACAD05FFA4FA5476E7A6C0FDD0FB0A" blockId="5.[151,638,1253,1279]" box="[151,638,1253,1279]" pageId="5" pageNumber="58">
<heading id="D0E41A69FFA4FA5476E7A6C0FDD0FB0A" bold="true" box="[151,638,1253,1279]" fontSize="11" level="1" pageId="5" pageNumber="58" reason="1">
<emphasis id="B9677117FFA4FA5476E7A6C0FDD0FB0A" bold="true" box="[151,638,1253,1279]" pageId="5" pageNumber="58">
<taxonomicName id="4C13D686FFA4FA5476E7A6C0FDB6FB0A" ID-CoL="5CB26" authority="Wang" authorityName="Wang" box="[151,536,1253,1279]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="jinggangensis" status="sp. nov.">
<emphasis id="B9677117FFA4FA5476E7A6C0FE66FB0A" bold="true" box="[151,456,1253,1279]" italics="true" pageId="5" pageNumber="58">Xenophrys jinggangensis</emphasis>
Wang
</taxonomicName>
<taxonomicNameLabel id="A254CC6CFFA4FA547450A6C0FDD0FB0A" box="[544,638,1253,1279]" pageId="5" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C309FE8EFFA4FA5476E7A709FC7AF9E9" pageId="5" pageNumber="58" type="materials_examined">
<paragraph id="8BACAD05FFA4FA5476E7A709FEFAFA21" blockId="5.[151,1436,1324,2032]" pageId="5" pageNumber="58">
<emphasis id="B9677117FFA4FA5476E7A709FEA5FAB1" bold="true" box="[151,267,1324,1348]" pageId="5" pageNumber="58">
<typeStatus id="54A813A7FFA4FA5476E7A709FEA8FAB1" box="[151,262,1324,1348]" pageId="5" pageNumber="58" type="holotype">Holotype</typeStatus>
.
</emphasis>
Adult female, SYS a001430, collected by Jian Zhao and Run-Lin Li on 
<date id="FFAD8BC5FFA4FA54725FA709FAA4FAB1" box="[1071,1290,1324,1348]" pageId="5" pageNumber="58" value="2011-09-13">13 September 2011</date>
, from Mount Jinggang (
<geoCoordinate id="EE27CBC2FFA4FA54777BA76AFE13FA92" box="[267,445,1359,1383]" direction="north" orientation="latitude" pageId="5" pageNumber="58" precision="1" value="26.55175">26°33'06.30&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE27CBC2FFA4FA5477B8A76AFD2DFA92" box="[456,643,1359,1383]" direction="east" orientation="longitude" pageId="5" pageNumber="58" precision="1" value="114.15489">114°09'17.60&quot; E</geoCoordinate>
; 
<quantity id="4CEB00E0FFA4FA5474FEA76AFD7BFA92" box="[654,725,1359,1383]" metricMagnitude="2" metricUnit="m" metricValue="8.45" pageId="5" pageNumber="58" unit="m" value="845.0">845 m</quantity>
a.s.l.), Jinggangshan City, Jiangxi Province, 
<collectingCountry id="F304ED95FFA4FA5472BDA76AFABEFA92" box="[1229,1296,1359,1383]" name="China" pageId="5" pageNumber="58">China</collectingCountry>
(
<figureCitation id="1328B180FFA4FA54736EA76AFA2EFA92" box="[1310,1408,1359,1383]" captionStart="FIGURE 3. 3 A" captionStartId="6.[151,250,1885,1908]" captionTargetBox="[151,1435,193,1863]" captionTargetId="figure@6.[151,1436,193,1864]" captionTargetPageId="6" captionText="FIGURE 3. 3 A: Dorsal view of the adult female holotype SYS a 001430 of Xenophrys jinggangensis sp. nov. in life. 3 B: Lateral view of the holotype in life. 3 C and 3 D: Hand and foot of the holotype in life. 3 E and 3 F: Dorsal and ventral views of the holotype in preservative. Photo Ying-Yong Wang and Jiang-Mo Zhang." httpUri="https://zenodo.org/record/213792/files/figure.png" pageId="5" pageNumber="58">Figure 3</figureCitation>
). 
<emphasis id="B9677117FFA4FA5476E7A751FEB8FA79" bold="true" box="[151,278,1396,1420]" pageId="5" pageNumber="58">
<typeStatus id="54A813A7FFA4FA5476E7A751FEBFFA79" box="[151,273,1396,1420]" pageId="5" pageNumber="58" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Two adult females: SYS a001413 (
<figureCitation id="1328B180FFA4FA5474D4A751FCAAFA79" box="[676,772,1396,1420]" captionStart="FIGURE 4. 4 A" captionStartId="7.[151,250,1502,1525]" captionTargetBox="[151,1435,193,1480]" captionTargetId="figure@7.[151,1436,193,1481]" captionTargetPageId="7" captionText="FIGURE 4. 4 A: General aspect of the adult female paratype SYS a 001413 in life. 4 B: X. jinggangensis active at night on a rock in a mountain stream on 10 September 2011. 4 C: Dorsal view of X. jinggangensis tadpole at stage 38 of in life on 5 December 2011. 4 D: Ventrolateral view of X. jinggangensis tadpole at stage 37 of in preservative. Photo Ying-Yong Wang and Jian Zhao." httpUri="https://zenodo.org/record/213793/files/figure.png" pageId="5" pageNumber="58">Figure 4</figureCitation>
A) and a001416; two adult males: SYS a001414–1415. All collected from the same locality as 
<typeStatus id="54A813A7FFA4FA547454A7B2FD28FA5A" box="[548,646,1431,1455]" pageId="5" pageNumber="58" type="holotype">holotype</typeStatus>
at 
<quantity id="4CEB00E0FFA4FA5474D8A7B2FC89FA5A" box="[680,807,1431,1455]" metricMagnitude="2" metricUnit="m" metricValue="7.75" metricValueMax="8.5" metricValueMin="7.0" pageId="5" pageNumber="58" unit="m" value="775.0" valueMax="850.0" valueMin="700.0">700–850 m</quantity>
a.s.l. by Jian Zhao, Zhong Zhang and Run-Lin Li on 
<date id="FFAD8BC5FFA4FA5473F0A7B2FEFEFA21" pageId="5" pageNumber="58" value="2011-09-13">13 September 2011</date>
.
</paragraph>
<paragraph id="8BACAD05FFA4FA5476B7A7FAFC7AF9E9" blockId="5.[151,1436,1324,2032]" pageId="5" pageNumber="58">
<emphasis id="B9677117FFA4FA5476B7A7FAFDA2FA0D" bold="true" box="[199,524,1503,1528]" pageId="5" pageNumber="58">Other examined materials.</emphasis>
Six tadpoles: SYS a001484, from the same locality as 
<typeStatus id="54A813A7FFA4FA5472E1A7FAFB5DFA02" box="[1169,1267,1503,1527]" pageId="5" pageNumber="58" type="holotype">holotype</typeStatus>
at 
<quantity id="4CEB00E0FFA4FA54736AA7FAFA32FA02" box="[1306,1436,1503,1527]" metricMagnitude="2" metricUnit="m" metricValue="7.75" metricValueMax="8.5" metricValueMin="7.0" pageId="5" pageNumber="58" unit="m" value="775.0" valueMax="850.0" valueMin="700.0">700–850 m</quantity>
a.s.l. by Jian Zhao and Run-Lin Li on 
<date id="FFAD8BC5FFA4FA547432A421FCA6F9E9" box="[578,776,1540,1564]" pageId="5" pageNumber="58" value="2011-12-05">5 December 2011</date>
(
<figureCitation id="1328B180FFA4FA547566A421FC2CF9E9" box="[790,898,1540,1564]" captionStart="FIGURE 4. 4 A" captionStartId="7.[151,250,1502,1525]" captionTargetBox="[151,1435,193,1480]" captionTargetId="figure@7.[151,1436,193,1481]" captionTargetPageId="7" captionText="FIGURE 4. 4 A: General aspect of the adult female paratype SYS a 001413 in life. 4 B: X. jinggangensis active at night on a rock in a mountain stream on 10 September 2011. 4 C: Dorsal view of X. jinggangensis tadpole at stage 38 of in life on 5 December 2011. 4 D: Ventrolateral view of X. jinggangensis tadpole at stage 37 of in preservative. Photo Ying-Yong Wang and Jian Zhao." httpUri="https://zenodo.org/record/213793/files/figure.png" pageId="5" pageNumber="58">Figures 4</figureCitation>
C, 4D).
</paragraph>
</subSubSection>
<subSubSection id="C309FE8EFFA4FA5976B7A402FED7FC80" lastPageId="8" lastPageNumber="61" pageId="5" pageNumber="58" type="diagnosis">
<paragraph id="8BACAD05FFA4FA5476B7A402FC0FF805" blockId="5.[151,1436,1324,2032]" pageId="5" pageNumber="58">
<emphasis id="B9677117FFA4FA5476B7A402FEEFF9B5" bold="true" box="[199,321,1575,1600]" pageId="5" pageNumber="58">Diagnosis.</emphasis>
<taxonomicName id="4C13D686FFA4FA547737A40DFDCEF9B5" box="[327,608,1576,1600]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="jinggangensis" status="sp. nov.">
<emphasis id="B9677117FFA4FA547737A40DFDCEF9B5" box="[327,608,1576,1600]" italics="true" pageId="5" pageNumber="58">Xenophrys jinggangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254CC6CFFA4FA547417A40DFD6EF9B5" box="[615,704,1576,1600]" pageId="5" pageNumber="58" rank="species">
<emphasis id="B9677117FFA4FA547417A40DFD6EF9B5" bold="true" box="[615,704,1576,1600]" pageId="5" pageNumber="58">sp. nov.</emphasis>
</taxonomicNameLabel>
is characterized by the combination of following characters: 
<emphasis id="B9677117FFA4FA547318A402FA26F9B5" bold="true" box="[1384,1416,1575,1600]" pageId="5" pageNumber="58">(1)</emphasis>
a small-sized species with 
<quantity id="4CEB00E0FFA4FA5477C0A469FDFAF991" box="[432,596,1612,1636]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.16" metricValueMin="3.84" pageId="5" pageNumber="58" unit="mm" value="40.0" valueMax="41.6" valueMin="38.4">38.4–41.6 mm</quantity>
SVL in adult females, 
<quantity id="4CEB00E0FFA4FA54752BA469FC51F991" box="[859,1023,1612,1636]" metricMagnitude="-2" metricUnit="m" metricValue="3.5900000000000007" metricValueMax="3.6700000000000004" metricValueMin="3.5100000000000002" pageId="5" pageNumber="58" unit="mm" value="35.9" valueMax="36.7" valueMin="35.1">35.1–36.7 mm</quantity>
SVL in adult males; 
<emphasis id="B9677117FFA4FA547281A46EFABCF991" bold="true" box="[1265,1298,1611,1636]" pageId="5" pageNumber="58">(2)</emphasis>
head length approximately equal to head width (HDL/HDW ratio 1); 
<emphasis id="B9677117FFA4FA547525A44AFCD8F97D" bold="true" box="[853,886,1647,1672]" pageId="5" pageNumber="58">(3)</emphasis>
snout almost truncate in plan view, strongly projecting, sloping backward to mouth in profile, protruding well beyond margin of lower jaw; 
<emphasis id="B9677117FFA4FA5472CEA4B6FB71F959" bold="true" box="[1214,1247,1683,1708]" pageId="5" pageNumber="58">(4)</emphasis>
tympanum great and distinct, TMP/EYE ratio 
<quantity id="4CEB00E0FFA4FA54779EA492FD9CF93A" box="[494,562,1719,1743]" metricMagnitude="-2" metricUnit="m" metricValue="2.032" pageId="5" pageNumber="58" unit="in" value="0.8">0.8 in</quantity>
females and males; 
<emphasis id="B9677117FFA4FA547556A492FCE9F925" bold="true" box="[806,839,1719,1744]" pageId="5" pageNumber="58">(5)</emphasis>
vomerine teeth on two weak ridges; 
<emphasis id="B9677117FFA4FA54728FA492FA8EF925" bold="true" box="[1279,1312,1719,1744]" pageId="5" pageNumber="58">(6)</emphasis>
margin of tongue smooth, not notched behind; 
<emphasis id="B9677117FFA4FA547443A4FEFDFAF901" bold="true" box="[563,596,1755,1780]" pageId="5" pageNumber="58">(7)</emphasis>
relative finger length II &lt;I &lt;IV &lt;III; 
<emphasis id="B9677117FFA4FA547262A4FEFB9DF901" bold="true" box="[1042,1075,1755,1780]" pageId="5" pageNumber="58">(8)</emphasis>
lateral fringes on digits minute, toes with a thick, fleshy web at their bases; 
<emphasis id="B9677117FFA4FA5474E3A4DAFD1AF8ED" bold="true" box="[659,692,1791,1816]" pageId="5" pageNumber="58">(9)</emphasis>
large subarticular tubercle at base of each digit; 
<emphasis id="B9677117FFA4FA547298A4DAFAB9F8ED" bold="true" box="[1256,1303,1791,1816]" pageId="5" pageNumber="58">(10)</emphasis>
dorsal skin granulated with tubercles, forming single swollen dorsolateral folds parallel to contra-lateral fold and discontinuous V-shaped fold from above the tympanum to the shoulder, flanks with larger pustules scattered, ventral surface smooth with granules; 
<emphasis id="B9677117FFA4FA547428A54EFD2BF871" bold="true" box="[600,645,1899,1924]" pageId="5" pageNumber="58">(11)</emphasis>
several large tubercles on the upper eyelid, one of them horn-like, remarkably prominent at the edge of the eyelid; 
<emphasis id="B9677117FFA4FA5474DDA5AAFD72F85D" bold="true" box="[685,732,1935,1960]" pageId="5" pageNumber="58">(12)</emphasis>
dorsum light brown, four wide longitudinal dark-brown stripes parallel each other, a dark brown triangle between eyes; dorsal limbs and digits light brown with dark brown transverse bands; ventral surface grayish with black and brown spots.
</paragraph>
<caption id="DF6CFD8DFFA7FA5776E7A578FC21F845" httpUri="https://zenodo.org/record/213792/files/figure.png" pageId="6" pageNumber="59" targetBox="[151,1435,193,1863]" targetPageId="6">
<paragraph id="8BACAD05FFA7FA5776E7A578FC21F845" blockId="6.[151,1436,1885,1969]" pageId="6" pageNumber="59">
<emphasis id="B9677117FFA7FA5776E7A578FEB8F886" bold="true" box="[151,278,1885,1908]" pageId="6" pageNumber="59">FIGURE 3.</emphasis>
3A: Dorsal view of the adult female holotype SYS a001430 of 
<taxonomicName id="4C13D686FFA7FA5775B0A57BFB6CF881" box="[960,1218,1886,1908]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="6" pageNumber="59" phylum="Chordata" rank="species" species="jinggangensis" status="sp. nov.">
<emphasis id="B9677117FFA7FA5775B0A57BFB6CF881" box="[960,1218,1886,1908]" italics="true" pageId="6" pageNumber="59">Xenophrys jinggangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254CC6CFFA7FA5772BBA57BFAB0F886" box="[1227,1310,1886,1907]" pageId="6" pageNumber="59" rank="species">
<emphasis id="B9677117FFA7FA5772BBA57BFAB0F886" bold="true" box="[1227,1310,1886,1907]" pageId="6" pageNumber="59">sp. nov.</emphasis>
</taxonomicNameLabel>
in life. 3B: Lateral view of the holotype in life. 3C and 3D: Hand and foot of the holotype in life. 3E and 3F: Dorsal and ventral views of the holotype in preservative. Photo Ying-Yong Wang and Jiang-Mo Zhang.
</paragraph>
</caption>
<caption id="DF6CFD8DFFA6FA5676E7A7FBFEAFF9A5" httpUri="https://zenodo.org/record/213793/files/figure.png" pageId="7" pageNumber="60" targetBox="[151,1435,193,1480]" targetPageId="7">
<paragraph id="8BACAD05FFA6FA5676E7A7FBFEAFF9A5" blockId="7.[151,1436,1502,1616]" pageId="7" pageNumber="60">
<emphasis id="B9677117FFA6FA5676E7A7FBFEBAFA01" bold="true" box="[151,276,1502,1525]" pageId="7" pageNumber="60">FIGURE 4.</emphasis>
4A: General aspect of the adult female paratype SYS a001413 in life. 4B: 
<taxonomicName id="4C13D686FFA6FA567252A7C5FB63FA00" box="[1058,1229,1504,1525]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="jinggangensis">
<emphasis id="B9677117FFA6FA567252A7C5FB63FA00" box="[1058,1229,1504,1525]" italics="true" pageId="7" pageNumber="60">X. jinggangensis</emphasis>
</taxonomicName>
active at night on a rock in a mountain stream on 10 September 2011. 4C: Dorsal view of 
<taxonomicName id="4C13D686FFA6FA5675F3A7DAFB81F9E1" box="[899,1071,1535,1556]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="jinggangensis">
<emphasis id="B9677117FFA6FA5675F3A7DAFB81F9E1" box="[899,1071,1535,1556]" italics="true" pageId="7" pageNumber="60">X. jinggangensis</emphasis>
</taxonomicName>
tadpole at stage 38 of in life on 5 December 2011. 4D: Ventrolateral view of 
<taxonomicName id="4C13D686FFA6FA56743DA438FD58F9C7" box="[589,758,1565,1586]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="jinggangensis">
<emphasis id="B9677117FFA6FA56743DA438FD58F9C7" box="[589,758,1565,1586]" italics="true" pageId="7" pageNumber="60">X. jinggangensis</emphasis>
</taxonomicName>
tadpole at stage 37 of in preservative. Photo Ying-Yong Wang and Jian Zhao.
</paragraph>
</caption>
<paragraph id="8BACAD05FFA6FA5976B7A458FED7FC80" blockId="7.[151,1436,1661,2006]" lastBlockId="8.[151,1437,151,2020]" lastPageId="8" lastPageNumber="61" pageId="7" pageNumber="60">
<emphasis id="B9677117FFA6FA5676B7A458FEC3F963" bold="true" box="[199,365,1661,1686]" pageId="7" pageNumber="60">Comparisons.</emphasis>
Comparative data of 
<taxonomicName id="4C13D686FFA6FA567418A458FC2AF963" box="[616,900,1661,1686]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="jinggangensis" status="sp. nov.">
<emphasis id="B9677117FFA6FA567418A458FC2AF963" box="[616,900,1661,1686]" italics="true" pageId="7" pageNumber="60">Xenophrys jinggangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254CC6CFFA6FA5675FEA45BFC44F963" box="[910,1002,1662,1686]" pageId="7" pageNumber="60" rank="species">
<emphasis id="B9677117FFA6FA5675FEA45BFC44F963" bold="true" box="[910,1002,1662,1686]" pageId="7" pageNumber="60">sp. nov.</emphasis>
</taxonomicNameLabel>
with 42 known species of 
<taxonomicName id="4C13D686FFA6FA567354A458FA32F960" box="[1316,1436,1661,1685]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="7" pageNumber="60" phylum="Chordata" rank="genus">
<emphasis id="B9677117FFA6FA567354A458FA32F960" box="[1316,1436,1661,1685]" italics="true" pageId="7" pageNumber="60">Xenophrys</emphasis>
</taxonomicName>
were obtained from the literatures (
<bibRefCitation id="EF82D0F4FFA6FA567452A484FD75F94C" author="Boulenger" box="[546,731,1697,1721]" pageId="7" pageNumber="60" refString="Boulenger, G. A. (1908) A revision of the Oriental pelobatid batrachians (genus Megalophrys). Proceedings of the Zoological Society of London, 1908, 407 - 430." type="proceedings paper" year="1908">Boulenger 1908</bibRefCitation>
; 
<bibRefCitation id="EF82D0F4FFA6FA567497A487FC7BF94C" author="Inger" box="[743,981,1697,1722]" pageId="7" pageNumber="60" refString="Inger, R. F. &amp; Romer, J. D. (1961) A new Pelobatid frog of the genus Megophrys from Hong Kong. Fieldiana: Zoology, 39, 533 - 538." type="journal article" year="1961">Inger &amp; Romer 1961</bibRefCitation>
; 
<bibRefCitation id="EF82D0F4FFA6FA567591A484FBC2F94C" author="Taylor" box="[993,1132,1697,1721]" pageId="7" pageNumber="60" refString="Taylor, E. H. (1962) The amphibian Fauna of Thailand. The University of Kansas Science Bulletin, 43, 284 - 302." type="journal article" year="1962">Taylor 1962</bibRefCitation>
; 
<bibRefCitation id="EF82D0F4FFA6FA567209A484FAB1F94C" author="Fei" box="[1145,1311,1697,1721]" pageId="7" pageNumber="60" refString="Fei, L. &amp; Ye, C. - Y. (1992) Two new species of the Megophrys, Pelobatidae (Amphibia: Anura) from China. Zoological Research, 13, 5 - 12." type="journal article" year="1992">
Fei 
<emphasis id="B9677117FFA6FA5672D4A487FB7BF94C" box="[1188,1237,1697,1721]" italics="true" pageId="7" pageNumber="60">et al</emphasis>
. 1992
</bibRefCitation>
, 
<bibRefCitation id="EF82D0F4FFA6FA56735AA484FF7DF929" author="Tian" pageId="7" pageNumber="60" refString="Tian, Y. - Z., Gu, X. - M. &amp; Sun, A. - Q. (2000) A new species of Megophrys in China (Amphibia: Pelobatidae). Acta Zootaxonomica Sinica, 25, 462 - 466." type="journal article" year="2000">
Tian 
<emphasis id="B9677117FFA6FA567314A487FA3BF94C" box="[1380,1429,1697,1721]" italics="true" pageId="7" pageNumber="60">et al</emphasis>
. 2000
</bibRefCitation>
; 
<bibRefCitation id="EF82D0F4FFA6FA5676ADA4E1FE32F929" author="Ohler" box="[221,412,1732,1757]" pageId="7" pageNumber="60" refString="Ohler, A., Swan, S. R. &amp; Daltry, J. C. (2002) A recent survey of the amphibian fauna of the Cardamom Mountains, Southwest Cambodia with descriptions of three new species. Raffles Bulletin of Zoology, 50 (2), 465 - 481." type="journal article" year="2002">
Ohler 
<emphasis id="B9677117FFA6FA567753A4E3FEFAF928" box="[291,340,1733,1757]" italics="true" pageId="7" pageNumber="60">et al</emphasis>
. 2002
</bibRefCitation>
; 
<bibRefCitation id="EF82D0F4FFA6FA5677D7A4E0FD06F929" author="Inger" box="[423,680,1732,1757]" pageId="7" pageNumber="60" refString="Inger, R. F. &amp; Iskandar, D. T. (2005) A collection of amphibians from West Sumatra, with description of a new species of Megophrys (Amphibia: Anura). Raffles Bulletin of Zoology, 53, 133 - 142." type="journal article" year="2005">Inger &amp; Iskandar 2005</bibRefCitation>
; 
<bibRefCitation id="EF82D0F4FFA6FA5674C2A4E1FC32F929" author="Mathew" box="[690,924,1732,1756]" pageId="7" pageNumber="60" refString="Mathew, R. &amp; Sen, N. (2007) Description of two new species of Xenophrys (Amphibia: Anura: Megophryidae) from north-east India. Cobra, 1 (2): 18 - 28." type="journal article" year="2007">Mathew &amp; Sen 2007</bibRefCitation>
; 
<bibRefCitation id="EF82D0F4FFA6FA5675D7A4E1FBE4F929" author="Fei" box="[935,1098,1732,1757]" pageId="7" pageNumber="60" refString="Fei, L., Hu, S. - Q., Ye, C. - Y. &amp; Huang, Y. - Z. (2009) Fauna Sinica, Amphibia Vol. 2, Anura Ranidae. Science Press, Beijing, 485 pp." type="book" year="2009">
Fei 
<emphasis id="B9677117FFA6FA5675A1A4E3FBACF928" box="[977,1026,1733,1757]" italics="true" pageId="7" pageNumber="60">et al</emphasis>
. 2009
</bibRefCitation>
; 
<bibRefCitation id="EF82D0F4FFA6FA567225A4E0FB54F929" author="Mo" box="[1109,1274,1732,1757]" pageId="7" pageNumber="60" refString="Mo, X. - Y., Shen Y. - H., Li, H. - H. &amp; Wu, X. - S. (2010) A new species of Megophrys (Amphibia: Anura: Megophryidae) from the northwestern Hunan Province, China. Current Zoology, 56, 432 - 436." type="journal article" year="2010">
Mo 
<emphasis id="B9677117FFA6FA5672F2A4E3FB1DF928" box="[1154,1203,1733,1757]" italics="true" pageId="7" pageNumber="60">et al</emphasis>
. 2010
</bibRefCitation>
) and shown in Table 3. The new species can be steadily distinguished from all known 42 congeners by the combination of following characters: tympanum great and distinct, dorsal skin granulated with tubercles, swollen dorsolateral folds, large pustules scattered on flanks and the unique color pattern. Further, the new species is significantly smaller in body size from the following 20 congeners: 
<taxonomicName id="4C13D686FFA6FA567571A576FBECF89F" authority="Boulenger, 1903" authorityName="Boulenger" authorityYear="1903" box="[769,1090,1874,1898]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="aceras">
<emphasis id="B9677117FFA6FA567571A576FCC2F89F" box="[769,876,1875,1898]" italics="true" pageId="7" pageNumber="60">X. aceras</emphasis>
(Boulenger, 1903)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA6FA56723EA576FE9BF87B" authority="Ohler, Swan &amp; Daltry, 2002" authorityName="Ohler, Swan &amp; Daltry" authorityYear="2002" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="auralensis">
<emphasis id="B9677117FFA6FA56723EA576FB4DF89F" box="[1102,1251,1874,1898]" italics="true" pageId="7" pageNumber="60">X. auralensis</emphasis>
(
<bibRefCitation id="EF82D0F4FFA6FA567282A577FE83F87B" author="Ohler" pageId="7" pageNumber="60" refString="Ohler, A., Swan, S. R. &amp; Daltry, J. C. (2002) A recent survey of the amphibian fauna of the Cardamom Mountains, Southwest Cambodia with descriptions of three new species. Raffles Bulletin of Zoology, 50 (2), 465 - 481." type="journal article" year="2002">Ohler, Swan &amp; Daltry, 2002</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA6FA567737A552FD16F87B" authority="Boulenger, 1899" authorityName="Boulenger" authorityYear="1899" box="[327,696,1910,1934]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="baluensis">
<emphasis id="B9677117FFA6FA567737A552FE78F87B" box="[327,470,1910,1934]" italics="true" pageId="7" pageNumber="60">X. baluensis</emphasis>
(Boulenger, 1899)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA6FA5674BAA552FB8DF87B" authority="Shen, 1994" authorityName="Shen" authorityYear="1994" box="[714,1059,1910,1934]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="caudoprocta">
<emphasis id="B9677117FFA6FA5674BAA552FCD3F87B" box="[714,893,1910,1934]" italics="true" pageId="7" pageNumber="60">X. caudoprocta</emphasis>
(Shen, 1994)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA6FA567245A552FADCF87B" authority="Mahony, 2011" authorityName="Mahony" authorityYear="2011" box="[1077,1394,1910,1934]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="damrei">
<emphasis id="B9677117FFA6FA567245A552FB06F87B" box="[1077,1192,1910,1934]" italics="true" pageId="7" pageNumber="60">X. damrei</emphasis>
(
<bibRefCitation id="EF82D0F4FFA6FA5672CEA553FAC4F87B" author="Mahony" box="[1214,1386,1910,1934]" pageId="7" pageNumber="60" refString="Mahony, S. (2011) Two new species of Megophrys Kuhl &amp; van Hasselt (Amphibia: Megophryidae), from western Thailand and southern Cambodia. Zootaxa, 2734, 23 - 39." type="journal article" year="2011">Mahony, 2011</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA6FA5673F4A552FDB1F844" authority="Liu, Hu &amp; Yang, 1960" authorityName="Liu, Hu &amp; Yang" authorityYear="1960" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="gigantica">
<emphasis id="B9677117FFA6FA5673F4A552FEAFF844" italics="true" pageId="7" pageNumber="60">X. gigantica</emphasis>
(Liu, Hu &amp; Yang, 1960)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA6FA56745CA5BFFC5CF844" authority="Fei, Ye &amp; Huang, 1990" authorityName="Fei, Ye &amp; Huang" authorityYear="1990" box="[556,1010,1945,1970]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="glandulosa">
<emphasis id="B9677117FFA6FA56745CA5BFFD64F844" box="[556,714,1945,1969]" italics="true" pageId="7" pageNumber="60">X. glandulosa</emphasis>
(Fei, Ye &amp; Huang, 1990)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA6FA56758FA5BFFAD8F844" authority="Fei &amp; Ye, 1983" authorityName="Fei &amp; Ye" authorityYear="1983" box="[1023,1398,1945,1969]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9677117FFA6FA56758FA5BFFB12F844" box="[1023,1212,1945,1969]" italics="true" pageId="7" pageNumber="60">X. jingdongensis</emphasis>
Fei &amp; Ye, 1983
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA6FA5673F4A5BFFF5CF820" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="lekaguli">
<emphasis id="B9677117FFA6FA5673F4A5BFFF5CF820" italics="true" pageId="7" pageNumber="60">X. lekaguli</emphasis>
</taxonomicName>
(Stuart, Chuaynkern, Chan-ard &amp; Inger, 2006), 
<taxonomicName id="4C13D686FFA6FA567540A59BFB3DF820" authority="Boulenger, 1886" authorityName="Boulenger" authorityYear="1886" box="[816,1171,1981,2005]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="longipes">
<emphasis id="B9677117FFA6FA567540A59BFC1AF820" box="[816,948,1981,2005]" italics="true" pageId="7" pageNumber="60">X. longipes</emphasis>
(Boulenger, 1886)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA6FA5972D4A59BFF74FF5A" authority="Boulenger, 1908" authorityName="Boulenger" authorityYear="1908" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" lastPageId="8" lastPageNumber="61" order="Anura" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="major">
<emphasis id="B9677117FFA6FA5672D4A59BFAA2F820" box="[1188,1292,1982,2005]" italics="true" pageId="7" pageNumber="60">X. major</emphasis>
(
<bibRefCitation id="EF82D0F4FFA6FA597350A598FF7CFF5A" author="Boulenger" lastPageId="8" lastPageNumber="61" pageId="7" pageNumber="60" refString="Boulenger, G. A. (1908) A revision of the Oriental pelobatid batrachians (genus Megalophrys). Proceedings of the Zoological Society of London, 1908, 407 - 430." type="proceedings paper" year="1908">Boulenger, 1908</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA59769AA2BCFE19FF45" box="[234,439,152,176]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="mangshanensis">
<emphasis id="B9677117FFA9FA59769AA2BCFE19FF45" box="[234,439,152,176]" italics="true" pageId="8" pageNumber="61">X. mangshanensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA5977B9A2BCFCEAFF5A" authority="Fei &amp; Ye, 1983" authorityName="Fei &amp; Ye" authorityYear="1983" box="[457,836,151,176]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="medogensis">
<emphasis id="B9677117FFA9FA5977B9A2BCFDC1FF45" box="[457,623,152,176]" italics="true" pageId="8" pageNumber="61">X. medogensis</emphasis>
(Fei &amp; Ye, 1983)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA597523A2BCFB29FF5A" authority="Liu, 1950" authorityName="Liu" authorityYear="1950" box="[851,1159,151,176]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="omeimontis">
<emphasis id="B9677117FFA9FA597523A2BCFC59FF45" box="[851,1015,153,176]" italics="true" pageId="8" pageNumber="61">X. omeimontis</emphasis>
(Liu, 1950)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA5972E6A2BCFF74FF26" authority="Boulenger, 1908" authorityName="Boulenger" authorityYear="1908" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="robusta">
<emphasis id="B9677117FFA9FA5972E6A2BCFAA3FF45" box="[1174,1293,152,176]" italics="true" pageId="8" pageNumber="61">X. robusta</emphasis>
(
<bibRefCitation id="EF82D0F4FFA9FA597350A2B2FF7DFF26" author="Boulenger" pageId="8" pageNumber="61" refString="Boulenger, G. A. (1908) A revision of the Oriental pelobatid batrachians (genus Megalophrys). Proceedings of the Zoological Society of London, 1908, 407 - 430." type="proceedings paper" year="1908">Boulenger, 1908</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA597697A299FD03FF26" authority="Jiang, Ye &amp; Fei, 2008" authorityName="Jiang, Ye &amp; Fei" authorityYear="2008" box="[231,685,187,211]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="sangzhiensis">
<emphasis id="B9677117FFA9FA597697A299FE38FF26" box="[231,406,187,211]" italics="true" pageId="8" pageNumber="61">X. sangzhiensis</emphasis>
(Jiang, Ye &amp; Fei, 2008)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA5974CAA299FC58FF26" authority="Liu, 1950" authorityName="Liu" authorityYear="1950" box="[698,1014,187,211]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="shapingensis">
<emphasis id="B9677117FFA9FA5974CAA299FCC5FF26" box="[698,875,187,211]" italics="true" pageId="8" pageNumber="61">X. shapingensis</emphasis>
(Liu, 1950)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA597273A299FF75FF02" authority="Tian, Gu &amp; Sun, 2000" authorityName="Tian, Gu &amp; Sun" authorityYear="2000" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="shuichengensis">
<emphasis id="B9677117FFA9FA597273A299FB63FF26" box="[1027,1229,187,211]" italics="true" pageId="8" pageNumber="61">X. shuichengensis</emphasis>
(
<bibRefCitation id="EF82D0F4FFA9FA5972ADA29EFF7DFF02" author="Tian" pageId="8" pageNumber="61" refString="Tian, Y. - Z., Gu, X. - M. &amp; Sun, A. - Q. (2000) A new species of Megophrys in China (Amphibia: Pelobatidae). Acta Zootaxonomica Sinica, 25, 462 - 466." type="journal article" year="2000">Tian, Gu &amp; Sun, 2000</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA597696A2C5FD8DFF02" authority="Liu &amp; Hu, 1973" authorityName="Liu &amp; Hu" authorityYear="1973" box="[230,547,223,247]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="spinata">
<emphasis id="B9677117FFA9FA597696A2C5FEF9FF02" box="[230,343,224,247]" italics="true" pageId="8" pageNumber="61">X. spinata</emphasis>
(Liu &amp; Hu, 1973)
</taxonomicName>
and 
<taxonomicName id="4C13D686FFA9FA59742AA2C5FC3EFF02" authority="Mahony, 2011" authorityName="Mahony" authorityYear="2011" box="[602,912,223,247]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="takensis">
<emphasis id="B9677117FFA9FA59742AA2C5FD7DFF02" box="[602,723,223,247]" italics="true" pageId="8" pageNumber="61">X. takensis</emphasis>
(
<bibRefCitation id="EF82D0F4FFA9FA597492A2FAFC29FF02" author="Mahony" box="[738,903,223,247]" pageId="8" pageNumber="61" refString="Mahony, S. (2011) Two new species of Megophrys Kuhl &amp; van Hasselt (Amphibia: Megophryidae), from western Thailand and southern Cambodia. Zootaxa, 2734, 23 - 39." type="journal article" year="2011">Mahony, 2011</bibRefCitation>
)
</taxonomicName>
; it has a small body sized (
<quantity id="4CEB00E0FFA9FA5972CCA2FAFACEFF02" box="[1212,1376,223,247]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.16" metricValueMin="3.84" pageId="8" pageNumber="61" unit="mm" value="40.0" valueMax="41.6" valueMin="38.4">38.4–41.6 mm</quantity>
SVL in adult females, 
<quantity id="4CEB00E0FFA9FA597718A327FDBFFEEF" box="[360,529,258,282]" metricMagnitude="-2" metricUnit="m" metricValue="3.5900000000000007" metricValueMax="3.6700000000000004" metricValueMin="3.5100000000000002" pageId="8" pageNumber="61" unit="mm" value="35.9" valueMax="36.7" valueMin="35.1">35.1–36.7 mm</quantity>
SVL in adult males), as opposed to 
<quantity id="4CEB00E0FFA9FA5975A4A327FB27FEEF" box="[980,1161,258,282]" metricMagnitude="-2" metricUnit="m" metricValue="9.09" metricValueMax="11.83" metricValueMin="6.35" pageId="8" pageNumber="61" unit="mm" value="90.9" valueMax="118.3" valueMin="63.5">63.5–118.3 mm</quantity>
SVL in adult females, 
<quantity id="4CEB00E0FFA9FA5976E7A303FE9CFECB" box="[151,306,294,318]" metricMagnitude="-2" metricUnit="m" metricValue="8.115" metricValueMax="11.5" metricValueMin="4.7299999999999995" pageId="8" pageNumber="61" unit="mm" value="81.15" valueMax="115.0" valueMin="47.3">47.3–115 mm</quantity>
SVL in adult males in those 20 large body-sized species. The new species differs from the following thirteen small body-sized congeners (characters in parentheses) by the presence of vomerine teeth (vs. absent): 
<taxonomicName id="4C13D686FFA9FA5973F4A36FFD89FE71" authority="Ye, Fei &amp; Xie, 2007" authorityName="Ye, Fei &amp; Xie" authorityYear="2007" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="baolongensis">
<emphasis id="B9677117FFA9FA5973F4A36FFE85FE70" italics="true" pageId="8" pageNumber="61">X. baolongensis</emphasis>
(Ye, Fei &amp; Xie, 2007)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA597442A34BFC16FE71" authority="Ye &amp; Fei, 1995" authorityName="Ye &amp; Fei" authorityYear="1995" box="[562,952,364,389]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="binchuanensis">
<emphasis id="B9677117FFA9FA597442A34BFD5FFE70" box="[562,753,365,389]" italics="true" pageId="8" pageNumber="61">X. binchuanensis</emphasis>
(Ye &amp; Fei, 1995)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA5975B3A34BFAD7FE71" authority="Jiang, Fei &amp; Ye, 2009" authorityName="Jiang, Fei &amp; Ye" authorityYear="2009" box="[963,1401,364,389]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="binlingensis">
<emphasis id="B9677117FFA9FA5975B3A34BFBC7FE70" box="[963,1129,365,389]" italics="true" pageId="8" pageNumber="61">X. binlingensis</emphasis>
(Jiang, 
<bibRefCitation id="EF82D0F4FFA9FA5972B2A349FADFFE71" author="Fei" box="[1218,1393,364,388]" pageId="8" pageNumber="61" refString="Fei, L., Hu, S. - Q., Ye, C. - Y. &amp; Huang, Y. - Z. (2009) Fauna Sinica, Amphibia Vol. 2, Anura Ranidae. Science Press, Beijing, 485 pp." type="book" year="2009">Fei &amp; Ye, 2009</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA5973F4A34BFF52FE5D" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="B9677117FFA9FA5973F4A34BFF52FE5D" italics="true" pageId="8" pageNumber="61">X. boettgeri</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA59777CA3B4FE19FE5D" box="[268,439,400,424]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="brachykolos">
<emphasis id="B9677117FFA9FA59777CA3B4FE19FE5D" box="[268,439,400,424]" italics="true" pageId="8" pageNumber="61">X. brachykolos</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA5977B8A3B4FD33FE5D" box="[456,669,400,424]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="huangshanensis">
<emphasis id="B9677117FFA9FA5977B8A3B4FD33FE5D" box="[456,669,400,424]" italics="true" pageId="8" pageNumber="61">X. huangshanensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA5974DEA3B4FCFAFE5D" box="[686,852,400,424]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="kuatunensis">
<emphasis id="B9677117FFA9FA5974DEA3B4FCFAFE5D" box="[686,852,400,424]" italics="true" pageId="8" pageNumber="61">X. kuatunensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA597515A3B4FC67FE5D" box="[869,969,401,424]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="minor">
<emphasis id="B9677117FFA9FA597515A3B4FC67FE5D" box="[869,969,401,424]" italics="true" pageId="8" pageNumber="61">X. minor</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA5975A9A3B4FADBFE5D" authority="Liu &amp; Hu, 1966" authorityName="Liu &amp; Hu" authorityYear="1966" box="[985,1397,400,424]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="nankiangensis">
<emphasis id="B9677117FFA9FA5975A9A3B4FB35FE5D" box="[985,1179,400,424]" italics="true" pageId="8" pageNumber="61">X. nankiangensis</emphasis>
(Liu &amp; Hu, 1966)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA5973F4A3B4FDC3FE3E" authority="Shen, Mo &amp; Li, 2010" authorityName="Shen, Mo &amp; Li" authorityYear="2010" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="tuberogranulatus">
<emphasis id="B9677117FFA9FA5973F4A3B4FEF7FE39" italics="true" pageId="8" pageNumber="61">X. tuberogranulatus</emphasis>
(
<bibRefCitation id="EF82D0F4FFA9FA597719A396FDCBFE3E" author="Shen" box="[361,613,435,459]" pageId="8" pageNumber="61" refString="Mo, X. - Y., Shen Y. - H., Li, H. - H. &amp; Wu, X. - S. (2010) A new species of Megophrys (Amphibia: Anura: Megophryidae) from the northwestern Hunan Province, China. Current Zoology, 56, 432 - 436." type="journal article" year="2010">Shen, Mo &amp; Li, 2010</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA59740AA390FBF6FE3E" authority="Fei, Jiang &amp; Zheng, 2001" authorityName="Fei, Jiang &amp; Zheng" authorityYear="2001" box="[634,1112,435,460]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="wawuensis">
<emphasis id="B9677117FFA9FA59740AA390FCBAFE39" box="[634,788,437,460]" italics="true" pageId="8" pageNumber="61">X. wawuensis</emphasis>
(Fei, Jiang &amp; Zheng, 2001)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA597215A390FEA5FE1A" authority="Ye &amp; Fei, 1995" authorityName="Ye &amp; Fei" authorityYear="1995" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="wuliangshanensis">
<emphasis id="B9677117FFA9FA597215A390FAE0FE39" box="[1125,1358,436,460]" italics="true" pageId="8" pageNumber="61">X. wuliangshanensis</emphasis>
(Ye &amp; Fei, 1995)
</taxonomicName>
and 
<taxonomicName id="4C13D686FFA9FA597734A3FDFD6EFE1A" authority="Ye &amp; Fei, 1995" authorityName="Ye &amp; Fei" authorityYear="1995" box="[324,704,471,495]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="wushanensis">
<emphasis id="B9677117FFA9FA597734A3FDFE5DFE1A" box="[324,499,471,495]" italics="true" pageId="8" pageNumber="61">X. wushanensis</emphasis>
(Ye &amp; Fei, 1995)
</taxonomicName>
; The new species differs from the following six small body-size congeners (characters in parentheses) by having distinct subarticular tubercles on toes (vs. lacking): 
<taxonomicName id="4C13D686FFA9FA5973F4A3D9FD9AFDC3" authority="Rao and Yang, 1997" authorityName="Rao and Yang" authorityYear="1997" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="daweimontis">
<emphasis id="B9677117FFA9FA5973F4A3D9FE8BFDC3" italics="true" pageId="8" pageNumber="61">X. daweimontis</emphasis>
(
<bibRefCitation id="EF82D0F4FFA9FA597747A03BFD85FDC3" author="Rao" box="[311,555,542,567]" pageId="8" pageNumber="61" refString="Rao, D. - Q. &amp; Yang, D. - T. (1997) The karyotypes of Megophryinae (Pelobatidae) with a discussion on their classification and phylogenetic relationships. Asiatic Herpetological Research, 7, 93 - 102." type="journal article" year="1997">Rao and Yang, 1997</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA597434A03AFC1DFDC3" authority="Huang, 1981" authorityName="Huang" authorityYear="1981" box="[580,947,542,566]" class="Amphibia" family="Megophryidae" genus="Ophryophryne" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="pachyproctus">
<emphasis id="B9677117FFA9FA597434A03AFD53FDC3" box="[580,765,542,566]" italics="true" pageId="8" pageNumber="61">X. pachyproctus</emphasis>
(Huang, 1981)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA5975B2A03AFADAFDC3" authority="Bourret, 1937" authorityName="Bourret" authorityYear="1937" box="[962,1396,542,566]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="palpebralespinosa">
<emphasis id="B9677117FFA9FA5975B2A03AFB1AFDC3" box="[962,1204,542,566]" italics="true" pageId="8" pageNumber="61">X. palpebralespinosa</emphasis>
(Bourret, 1937)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA5973F4A03AFDEFFDAC" authority="Inger and Iskandar, 2005" authorityName="Inger and Iskandar" authorityYear="2005" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="parallela">
<emphasis id="B9677117FFA9FA5973F4A03AFF53FDAC" italics="true" pageId="8" pageNumber="61">X. parallela</emphasis>
(
<bibRefCitation id="EF82D0F4FFA9FA597760A064FD97FDAC" author="Inger" box="[272,569,577,601]" pageId="8" pageNumber="61" refString="Inger, R. F. &amp; Iskandar, D. T. (2005) A collection of amphibians from West Sumatra, with description of a new species of Megophrys (Amphibia: Anura). Raffles Bulletin of Zoology, 53, 133 - 142." type="journal article" year="2005">Inger and Iskandar, 2005</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA9FA597420A067FC3EFDAC" authority="Boulenger, 1893" authorityName="Boulenger" authorityYear="1893" box="[592,912,577,601]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="parva">
<emphasis id="B9677117FFA9FA597420A067FD1AFDAC" box="[592,692,578,601]" italics="true" pageId="8" pageNumber="61">X. parva</emphasis>
(Boulenger, 1893)
</taxonomicName>
and 
<taxonomicName id="4C13D686FFA9FA5975BEA067FA88FDAC" authority="Ye and Fei, 1992" authorityName="Ye and Fei" authorityYear="1992" box="[974,1318,577,601]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="zhangi">
<emphasis id="B9677117FFA9FA5975BEA067FB92FDAC" box="[974,1084,577,601]" italics="true" pageId="8" pageNumber="61">X. zhangi</emphasis>
(
<bibRefCitation id="EF82D0F4FFA9FA59723EA064FAB0FDAC" author="Ye" box="[1102,1310,577,601]" pageId="8" pageNumber="61" refString="Fei, L. &amp; Ye, C. - Y. (1992) Two new species of the Megophrys, Pelobatidae (Amphibia: Anura) from China. Zoological Research, 13, 5 - 12." type="journal article" year="1992">Ye and Fei, 1992</bibRefCitation>
)
</taxonomicName>
; The new species differs from 
<taxonomicName id="4C13D686FFA9FA5977F3A043FD09FD88" authority="Mahony, 2011" authorityName="Mahony" authorityYear="2011" box="[387,679,613,637]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="dringi">
<emphasis id="B9677117FFA9FA5977F3A043FE46FD88" box="[387,488,613,637]" italics="true" pageId="8" pageNumber="61">X. dringi</emphasis>
(
<bibRefCitation id="EF82D0F4FFA9FA597789A040FD31FD88" author="Mahony" box="[505,671,613,637]" pageId="8" pageNumber="61" refString="Mahony, S. (2011) Two new species of Megophrys Kuhl &amp; van Hasselt (Amphibia: Megophryidae), from western Thailand and southern Cambodia. Zootaxa, 2734, 23 - 39." type="journal article" year="2011">Mahony, 2011</bibRefCitation>
)
</taxonomicName>
(characters in parentheses) in that in that margin of tongue is not notched (vs. notched); from 
<taxonomicName id="4C13D686FFA9FA5977A2A0ACFCFAFD55" authority="Mathew &amp; Sen, 2007" authorityName="Mathew &amp; Sen" authorityYear="2007" box="[466,852,648,672]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="serchhipii">
<emphasis id="B9677117FFA9FA5977A2A0ACFDCEFD55" box="[466,608,648,672]" italics="true" pageId="8" pageNumber="61">X. serchhipii</emphasis>
<bibRefCitation id="EF82D0F4FFA9FA597416A0ADFCFAFD55" author="Mathew" box="[614,852,648,672]" pageId="8" pageNumber="61" refString="Mathew, R. &amp; Sen, N. (2007) Description of two new species of Xenophrys (Amphibia: Anura: Megophryidae) from north-east India. Cobra, 1 (2): 18 - 28." type="journal article" year="2007">Mathew &amp; Sen, 2007</bibRefCitation>
</taxonomicName>
(characters in parentheses) by having relative finger length II &lt;I &lt;IV &lt;III (vs. 2nd and 4th fingers subequal in length in 
<taxonomicName id="4C13D686FFA9FA5975AEA088FBDEFD31" box="[990,1136,684,708]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="serchhipii">
<emphasis id="B9677117FFA9FA5975AEA088FBDEFD31" box="[990,1136,684,708]" italics="true" pageId="8" pageNumber="61">X. serchhipii</emphasis>
</taxonomicName>
); from 
<taxonomicName id="4C13D686FFA9FA5972BEA088FE28FD12" authority="Mathew &amp; Sen, 2007" authorityName="Mathew &amp; Sen" authorityYear="2007" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="zunhebotoensis">
<emphasis id="B9677117FFA9FA5972BEA088FA32FD31" box="[1230,1436,684,708]" italics="true" pageId="8" pageNumber="61">X. zunhebotoensis</emphasis>
<bibRefCitation id="EF82D0F4FFA9FA5976E7A0EAFE28FD12" author="Mathew" box="[151,390,719,743]" pageId="8" pageNumber="61" refString="Mathew, R. &amp; Sen, N. (2007) Description of two new species of Xenophrys (Amphibia: Anura: Megophryidae) from north-east India. Cobra, 1 (2): 18 - 28." type="journal article" year="2007">Mathew &amp; Sen, 2007</bibRefCitation>
</taxonomicName>
(characters in parentheses) in that head length is approximately equal to head width (vs. head length smaller than head width), flank scattered with large tubercles (vs. flank scattered with small tubercles), supratympanic fold not curving on to the anterior region of the tympanum (supratympanic fold curving on to the anterior region of the tympanum to give it a round shape), vomerine ridges relatively separated (vomerine ridges close to each other).
</paragraph>
</subSubSection>
<subSubSection id="C309FE8EFFA9FA5876B7A1A5FB2AFA71" lastPageId="9" lastPageNumber="62" pageId="8" pageNumber="61" type="description">
<paragraph id="8BACAD05FFA9FA5976B7A1A5FB7FFB41" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="61">
<emphasis id="B9677117FFA9FA5976B7A1A5FE47FC6C" bold="true" box="[199,489,896,921]" pageId="8" pageNumber="61">
Description of 
<typeStatus id="54A813A7FFA9FA597705A1A4FE4AFC6C" box="[373,484,897,921]" pageId="8" pageNumber="61" type="holotype">Holotype</typeStatus>
.
</emphasis>
Adult female, SVL 
<quantity id="4CEB00E0FFA9FA5974BAA1A5FC9FFC6D" box="[714,817,896,920]" metricMagnitude="-2" metricUnit="m" metricValue="3.84" pageId="8" pageNumber="61" unit="mm" value="38.4">38.4 mm</quantity>
. Head length approximately equal to head width (HDL/ HDW ratio 1.0); snout short (SNT/HDL ratio 0.3, SNT/SVL ratio 0.1), almost truncate in plan view, sloping backward to mouth in profile, strongly protruding well beyond margin of lower jaw; top of head flat; eye large and convex, eye diameter 32% of head length (EYE/HDL ratio 0.3), upper eyelid slightly wider than interorbital distance; pupil vertical; nostril oblique ovoid with low flap of skin laterally; canthus rostralis sharp; loreal region vertical, not concave; internasal distance larger than interorbital distance; tympanum elevated relative to skin of temporal region, distinctly visible, TMP/EYE ratio 0.8; tympanum-eye distance great, TEY 
<quantity id="4CEB00E0FFA9FA5972CBA670FAB9FB98" box="[1211,1303,1109,1133]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="61" unit="mm" value="2.5">2.5 mm</quantity>
, TEY/TMP ratio 0.9; choanae large, ovoid at the bases of the maxilla; two vomerine ridges weakly, oblique, posteromedial to choanae, each bearing numerous prominent teeth; margin of tongue smooth, not notched behind.
</paragraph>
<paragraph id="8BACAD05FFA9FA5976B7A6E5FB17FA25" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="61">Forelimbs moderately slender; radioulna length 26% SVL, hands without web, moderately longer, 27% of SVL; fingers slender, relative finger length II &lt;I &lt;IV &lt;III; tips of digits round, slightly dilated; a large subarticular tubercle at base of each finger, slight lateral fringes from subarticular tubercles to terminal phalanges; metacarpal tubercle two, inner one significantly enlarged. Hindlimbs relatively long and moderately robust; tibia length 48% of SVL; foot length 71% of SVL; relative toe lengths I &lt;II &lt;V &lt;III &lt;IV; tips of toes round, slightly dilated; toes with a fleshy web at their bases; a large subarticular tubercle at base of each toe, slight lateral fringes from subarticular tubercles to terminal phalanges; tarsal fold absent; but as outer lateral fringes on toe V from hough to terminal phalanges; inner metatarsal tubercle ovoid; outer metatarsal tubercle absent.</paragraph>
<paragraph id="8BACAD05FFA9FA5976B7A7FEFC60F8FA" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="61">Skin on all upper surfaces granulated with tubercles, forming single swollen dorsolateral folds parallel to contralateral fold and discontinuous V-shaped fold from above the tympanum to the shoulder; several large tubercles at the upper eyelid prominent, one of them horn-like, remarkably prominent at the edge of the eyelid; supratympanic fold distinct, curving posteroventrally from posterior corner of eye to a level above insertion of arm where forming swollen glandular pustule; larger pustules scattered on flanks, ventral surface smooth with granules; pectoral gland large, round, prominently elevated relative to ventral surface, closer to axilla than to min-ventral line; single larger femoral gland on rear of thigh, two larger lip glands on anterior part of lower jaw, also round, prominently elevated relative to periphery skin surface; distinct granules on posterior thighs and around cloaca; cloacal opening unmodified, directed posteriorly, at upper level of thighs.</paragraph>
<paragraph id="8BACAD05FFA9FA5976B7A53EFCF1F8A3" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="61">
<emphasis id="B9677117FFA9FA5976B7A53EFDC0F8C6" bold="true" box="[199,622,1818,1843]" pageId="8" pageNumber="61">
Measurements of 
<typeStatus id="54A813A7FFA9FA5977EAA53EFDAFF8C6" box="[410,513,1819,1843]" pageId="8" pageNumber="61" type="holotype">holotype</typeStatus>
(in mm).
</emphasis>
SVL 38.4, HDL 12.2, HDW 12.2, SNT 4.1, 
<collectingCountry id="F304ED95FFA9FA597212A539FB3AF8C6" box="[1122,1172,1820,1843]" name="India" pageId="8" pageNumber="61">IND</collectingCountry>
4.0, IOD 3.7, EYE 3.8, TMP 2.9, TEY 2.5, HND 10.4, RAD 10.0, FTL 27.2, TIB 18.6.
</paragraph>
<paragraph id="8BACAD05FFA9FA5876B7A544FD86FE5D" blockId="8.[151,1437,151,2020]" lastBlockId="9.[151,1437,151,2024]" lastPageId="9" lastPageNumber="62" pageId="8" pageNumber="61">
<emphasis id="B9677117FFA9FA5976B7A544FD9AF88F" bold="true" box="[199,564,1889,1914]" pageId="8" pageNumber="61">
Coloration of 
<typeStatus id="54A813A7FFA9FA597704A547FE75F88F" box="[372,475,1890,1914]" pageId="8" pageNumber="61" type="holotype">holotype</typeStatus>
in life.
</emphasis>
Dorsum light brown, four wide longitudinal dark-brown stripes, outer two along the dorsolateral folds parallel each other, inside two intersect in middle; a dark brown triangle with a light center between the two eyes, apex of triangle over occiput; a dark brown stripe along the body axes on the dorsum of snout, the both sides large dark brown spots; tip of snout dark brown with black spots; upper lip with vertical dark brown bars, the one under the eye largest and distinct; excepting arm, dorsal limbs and digits light brown with dark brown transverse bands; all dark brown regions bordered by black edge lines; a gray-brown stripe on both flanks from the posterior margin of the forelimb insertion to the anterior margin of the hind limb insertion; ventral surface pale grayish with small black spots and dark blotches; lower lip black with vertical white bars, including two lip glands; three longitudinal black stripes on the throat, large lateral black spots on belly, all bordered by white edges; anterior surface of thighs, knee and part of shank red-orange; ventral surface of limbs with large black blotches; ventral surface of digits black, tip of digits orange; pectoral gland, femoral gland white; pupils black; iris whitish with radiating black stripes.
</paragraph>
<paragraph id="8BACAD05FFA8FA5876B7A396FB56FDCD" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="62">
<emphasis id="B9677117FFA8FA5876B7A396FD35FE39" bold="true" box="[199,667,435,460]" pageId="9" pageNumber="62">
Coloration of 
<typeStatus id="54A813A7FFA8FA587701A391FE76FE39" box="[369,472,436,460]" pageId="9" pageNumber="62" type="holotype">holotype</typeStatus>
in preservative.
</emphasis>
Dorsum creamy white, with black triangle and four wide stripes; a black stripe respectively on both flanks parallel to the dorsolateral stripes; dorsal limbs and digits with black transverse bands; ventral surface pale grayish with small black spots and blotches; creamy white substituted the red-orange in anterior surface of thighs, knee and part of shank; tip of digits whitish without orange.
</paragraph>
<paragraph id="8BACAD05FFA8FA5876B7A066FC0EFC7D" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="62">
<emphasis id="B9677117FFA8FA5876B7A066FE7FFDA9" bold="true" box="[199,465,579,604]" pageId="9" pageNumber="62">Description of tadpole</emphasis>
. Body slender, oval, flattened above; tail depth slightly larger than body depth, dorsal fin arising behind origin of tail, maximum depth near mid-length, tapering gradually to narrow, pointed tip; tail 2.3–2.8 times body length in 37th –39th stages, tail depth 19% of tail length in 37th stage, 22% in 38th stage, 24%in 39th stage; maximum body width 33% of body length in 37th stage, 43% in 38th stage, 48% in 39th stage; body depth 33% of body length in 37th stage, 43% in 38th stage, 44% in 39th stage; eyes large, lateral; nostril dorsolateral, slightly closer to umbelliform oral disk than to eyes, rim raised; internasal wider than interorbital; spiracle on left side of the body, closer to eye than to end of body; anal tube extends backward above ventral fin, opening medial; oral disk terminal, lips expanded and directed upwardly into typical 
<taxonomicName id="4C13D686FFA8FA5875D3A16DFBB5FC95" box="[931,1051,840,864]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="62" phylum="Chordata" rank="genus">
<emphasis id="B9677117FFA8FA5875D3A16DFBB5FC95" box="[931,1051,840,864]" italics="true" pageId="9" pageNumber="62">Xenophrys</emphasis>
</taxonomicName>
umbelliform oral disk; transverse width of expanded funnel 41–42% of body length in 37th –39th stages.
</paragraph>
<paragraph id="8BACAD05FFA8FA5876B7A1B7FC2CFBE2" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="62">
<emphasis id="B9677117FFA8FA5876B7A1B7FEF3FC5E" bold="true" box="[199,349,914,939]" pageId="9" pageNumber="62">Color in life.</emphasis>
All upper surfaces brown red with black spots, sides of body pale gray with black marks; ventral surface brown with small black and white marks; sides of tail brown-gray with black and white marks; distal portions of fins orange-red with large black marks; hindlimbs orange-red with gray spots; in preservative, color of body gray-black, tail and hanglimbs creamy white with dark spots.
</paragraph>
<paragraph id="8BACAD05FFA8FA5876B7A603FDF5FB91" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="62">
Measurements of tadpole: 37th stage: 
<quantity id="4CEB00E0FFA8FA587400A600FD7BFBC8" box="[624,725,1061,1085]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="9" pageNumber="62" unit="mm" value="11.4">11.4 mm</quantity>
SVL, 
<quantity id="4CEB00E0FFA8FA587552A600FC29FBC8" box="[802,903,1061,1085]" metricMagnitude="-2" metricUnit="m" metricValue="3.18" pageId="9" pageNumber="62" unit="mm" value="31.8">31.8 mm</quantity>
TaL; 38th stage: 
<quantity id="4CEB00E0FFA8FA587237A600FB03FBC8" box="[1095,1197,1061,1085]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="9" pageNumber="62" unit="mm" value="13.3">13.3 mm</quantity>
SVL, 
<quantity id="4CEB00E0FFA8FA58728AA600FAF1FBC8" box="[1274,1375,1061,1085]" metricMagnitude="-2" metricUnit="m" metricValue="3.01" pageId="9" pageNumber="62" unit="mm" value="30.1">30.1 mm</quantity>
TaL; 39th stage: 
<quantity id="4CEB00E0FFA8FA58777EA669FEDCFB91" box="[270,370,1100,1124]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="9" pageNumber="62" unit="mm" value="13.8">13.8 mm</quantity>
SVL, 
<quantity id="4CEB00E0FFA8FA5877CCA669FD8EFB91" box="[444,544,1100,1124]" metricMagnitude="-2" metricUnit="m" metricValue="3.26" pageId="9" pageNumber="62" unit="mm" value="32.6">32.6 mm</quantity>
TaL.
</paragraph>
<paragraph id="8BACAD05FFA8FA5876B7A655FB90FB7D" blockId="9.[151,1437,151,2024]" box="[199,1086,1136,1161]" pageId="9" pageNumber="62">
<emphasis id="B9677117FFA8FA5876B7A655FE95FB7C" bold="true" box="[199,315,1136,1161]" pageId="9" pageNumber="62">Variation.</emphasis>
Measurements and body proportions of 
<typeStatus id="54A813A7FFA8FA587574A654FC9AFB7C" box="[772,820,1137,1161]" pageId="9" pageNumber="62">type</typeStatus>
series given in 
<tableCitation id="C69198BEFFA8FA587594A655FB94FB7D" box="[996,1082,1136,1160]" captionStart="TABLE 4" captionStartId="12.[151,235,152,173]" captionTargetBox="[151,1436,232,1441]" captionTargetPageId="12" captionText="TABLE 4. Measurements (in mm) of the type series of Xenophrys jinggangensis sp. nov. See Materials and Methods for abbreviations." httpUri="http://table.plazi.org/id/DF6CFD8DFFADFA5D76E7A2BDFE88FF39" pageId="9" pageNumber="62" tableUuid="DF6CFD8DFFADFA5D76E7A2BDFE88FF39">Table 4</tableCitation>
.
</paragraph>
<paragraph id="8BACAD05FFA8FA5876B7A6B3FCBAFAED" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="62">
<taxonomicName id="4C13D686FFA8FA5876B7A6B3FE2AFB58" box="[199,388,1174,1197]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="62" phylum="Chordata" rank="species" species="jinggangensis" status="sp. nov.">
<emphasis id="B9677117FFA8FA5876B7A6B3FE2AFB58" box="[199,388,1174,1197]" italics="true" pageId="9" pageNumber="62">X. jinggangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254CC6CFFA8FA5877FDA6B0FE47FB58" box="[397,489,1173,1197]" pageId="9" pageNumber="62" rank="species">
<emphasis id="B9677117FFA8FA5877FDA6B0FE47FB58" bold="true" box="[397,489,1173,1197]" pageId="9" pageNumber="62">sp. nov.</emphasis>
</taxonomicNameLabel>
presents obvious sexual dimorphism in secondary sexual characters: SVL in adult females slightly greater than in males, SVL of females 
<quantity id="4CEB00E0FFA8FA587578A69DFC03FB25" box="[776,941,1208,1232]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.16" metricValueMin="3.84" pageId="9" pageNumber="62" unit="mm" value="40.0" valueMax="41.6" valueMin="38.4">38.4–41.6 mm</quantity>
, SVL of males 
<quantity id="4CEB00E0FFA8FA58722FA69DFAABFB25" box="[1119,1285,1208,1232]" metricMagnitude="-2" metricUnit="m" metricValue="3.5900000000000007" metricValueMax="3.6700000000000004" metricValueMin="3.5100000000000002" pageId="9" pageNumber="62" unit="mm" value="35.9" valueMax="36.7" valueMin="35.1">35.1–36.7 mm</quantity>
; snout length larger than eye diameter in females, but opposite in males; single subgular vocal sac in adult males; no nuptial pad and nuptial spines were observed in our male specimens.
</paragraph>
<paragraph id="8BACAD05FFA8FA5876B7A701FB2AFA71" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="62">
All five specimens (two males, three females) were very similar in morphology and color pattern. One exception was an individual, found on 
<date id="FFAD8BC5FFA8FA587425A76DFD5AFA95" box="[597,756,1352,1376]" pageId="9" pageNumber="62" value="2010-09">10 September</date>
, 2011, which featured horn-like conical tubercles that were remarkably prominent at the edges of the eyelids, and had coal black dorsum. (
<figureCitation id="1328B180FFA8FA587279A749FBC4FA71" box="[1033,1130,1388,1412]" captionStart="FIGURE 4. 4 A" captionStartId="7.[151,250,1502,1525]" captionTargetBox="[151,1435,193,1480]" captionTargetId="figure@7.[151,1436,193,1481]" captionTargetPageId="7" captionText="FIGURE 4. 4 A: General aspect of the adult female paratype SYS a 001413 in life. 4 B: X. jinggangensis active at night on a rock in a mountain stream on 10 September 2011. 4 C: Dorsal view of X. jinggangensis tadpole at stage 38 of in life on 5 December 2011. 4 D: Ventrolateral view of X. jinggangensis tadpole at stage 37 of in preservative. Photo Ying-Yong Wang and Jian Zhao." httpUri="https://zenodo.org/record/213793/files/figure.png" pageId="9" pageNumber="62">Figure 4</figureCitation>
B)
</paragraph>
</subSubSection>
<subSubSection id="C309FE8EFFA8FA5876B7A7B5FBA9F93D" pageId="9" pageNumber="62" type="distribution">
<paragraph id="8BACAD05FFA8FA5876B7A7B5FB02F9E1" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="62">
<emphasis id="B9677117FFA8FA5876B7A7B5FDD0FA5C" bold="true" box="[199,638,1424,1449]" pageId="9" pageNumber="62">Distribution and biological ecology.</emphasis>
Currently, 
<taxonomicName id="4C13D686FFA8FA587578A7B4FC67FA5D" box="[776,969,1425,1448]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="62" phylum="Chordata" rank="species" species="jinggangensis" status="sp. nov.">
<emphasis id="B9677117FFA8FA587578A7B4FC67FA5D" box="[776,969,1425,1448]" italics="true" pageId="9" pageNumber="62">X. jinggangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254CC6CFFA8FA5875A6A7B4FB9BFA5C" box="[982,1077,1425,1449]" pageId="9" pageNumber="62" rank="species">
<emphasis id="B9677117FFA8FA5875A6A7B4FB9BFA5C" bold="true" box="[982,1077,1425,1449]" pageId="9" pageNumber="62">sp. nov.</emphasis>
</taxonomicNameLabel>
is known only from the 
<typeStatus id="54A813A7FFA8FA58731CA7B4FA32FA5C" box="[1388,1436,1425,1449]" pageId="9" pageNumber="62">type</typeStatus>
locality, Mount Jinggang, located in the middle of the Luoxiao Range, running along the border between Jiangxi and Hunan Provinces, 
<collectingCountry id="F304ED95FFA8FA5877EAA7FDFE4FFA05" box="[410,481,1496,1520]" name="China" pageId="9" pageNumber="62">China</collectingCountry>
. All individuals were found in small, slow-moving montane streams surrounded by moist subtropical evergreen broadleaved forests between 
<quantity id="4CEB00E0FFA8FA58756AA7D9FC37F9E1" box="[794,921,1532,1556]" metricMagnitude="2" metricUnit="m" metricValue="7.75" metricValueMax="8.5" metricValueMin="7.0" pageId="9" pageNumber="62" unit="m" value="775.0" valueMax="850.0" valueMin="700.0">700–850 m</quantity>
elevations (
<figureCitation id="1328B180FFA8FA587250A7D9FB2EF9E1" box="[1056,1152,1532,1556]" captionStart="FIGURE 1" captionStartId="1.[151,250,1802,1825]" captionTargetBox="[151,1435,806,1777]" captionTargetId="figure@1.[151,1436,806,1778]" captionTargetPageId="1" captionText="FIGURE 1. Collecting localities in Southern and Eastern China:? Mt. Jinggang, Jinggangshan City, Jiangxi Province, specimens collected here refer to Xenophrys jinggangensis sp. nov..? and? Mt. Wuyi, Guixi City, Jiangxi Province, specimens collected here refer to X. boettgeri and X. kuatunensis.? Hong Kong, specimens collected here refer to X. brachykolos.? Mt. Nanling, Ruyuan County, Guangdong Province, specimens collected here refer to X. mangshanensis.? Wuhuangling Forest Park, Qinzhou City, Guangxi, specimens collected here refer to Ophryophryne pachyproctus." httpUri="https://zenodo.org/record/213790/files/figure.png" pageId="9" pageNumber="62">Figure 1</figureCitation>
, 
<figureCitation id="1328B180FFA8FA5872FEA7D9FB30F9E1" box="[1166,1182,1532,1556]" captionStart="FIGURE 5" captionStartId="13.[151,250,1071,1094]" captionTargetBox="[151,1435,193,1049]" captionTargetId="figure@13.[151,1436,193,1050]" captionTargetPageId="13" captionText="FIGURE 5. Habitat of X. jinggangensis at the type locality. Photo Jian Zhao on 5 December 2011." httpUri="https://zenodo.org/record/213794/files/figure.png" pageId="9" pageNumber="62">5</figureCitation>
).
</paragraph>
<paragraph id="8BACAD05FFA8FA5876B7A405FBA9F93D" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="62">
All specimens were collected between 
<date id="FFAD8BC5FFA8FA58740BA405FCC7F9CD" box="[635,873,1568,1592]" pageId="9" pageNumber="62">10 and 13 September</date>
, 2011, and no males were heard calling during the survey. The male 
<typeStatus id="54A813A7FFA8FA58772FA460FE6EF9A8" box="[351,448,1605,1629]" pageId="9" pageNumber="62" type="paratype">paratype</typeStatus>
SYS a001415 had mature spermaries in the abdominal cavity, measuring 4.9 × 
<quantity id="4CEB00E0FFA8FA587336A461FA32F9A9" box="[1350,1436,1604,1628]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="9" pageNumber="62" unit="mm" value="2.4">2.4 mm</quantity>
in 
<taxonomicName id="4C13D686FFA8FA5876C5A44DFF59F975" box="[181,247,1640,1664]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="62" phylum="Chordata" rank="species" species="major">major</taxonomicName>
axis and 
<taxonomicName id="4C13D686FFA8FA587715A44DFE06F975" box="[357,424,1640,1664]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="62" phylum="Chordata" rank="species" species="minor">minor</taxonomicName>
axis. Female 
<typeStatus id="54A813A7FFA8FA587437A44CFD09F974" box="[583,679,1641,1665]" pageId="9" pageNumber="62" type="paratype">paratype</typeStatus>
SYS a001416 had a fully developed oviduct without eggs and the ovary fat was fully grown. Tadpoles at Gosner stage 37–44 were found under rocks in the stream on 
<date id="FFAD8BC5FFA8FA587362A4A9FF7CF93D" pageId="9" pageNumber="62" value="2011-12-05">5 December 2011</date>
. Thus, the breeding season of the new species is likely before September.
</paragraph>
</subSubSection>
<subSubSection id="C309FE8EFFA8FA5876B7A4F0FEFDF8E5" pageId="9" pageNumber="62" type="etymology">
<paragraph id="8BACAD05FFA8FA5876B7A4F0FEFDF8E5" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="62">
<emphasis id="B9677117FFA8FA5876B7A4F0FEE1F918" bold="true" box="[199,335,1749,1773]" pageId="9" pageNumber="62">Etymology.</emphasis>
The specific epithet “ 
<taxonomicName id="4C13D686FFA8FA58743AA4F3FD48F918" box="[586,742,1750,1773]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="62" phylum="Chordata" rank="species" species="jinggangensis">
<emphasis id="B9677117FFA8FA58743AA4F3FD48F918" box="[586,742,1750,1773]" italics="true" pageId="9" pageNumber="62">jinggangensis</emphasis>
</taxonomicName>
” is in reference to the 
<typeStatus id="54A813A7FFA8FA58759BA4F0FBB5F918" box="[1003,1051,1749,1773]" pageId="9" pageNumber="62">type</typeStatus>
locality, Mount Jinggang, Jiangxi Province, 
<collectingCountry id="F304ED95FFA8FA587778A4DDFEE1F8E5" box="[264,335,1784,1808]" name="China" pageId="9" pageNumber="62">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C309FE8EFFA8FA5876B7A538FC13F81D" pageId="9" pageNumber="62" type="discussion">
<paragraph id="8BACAD05FFA8FA5876B7A538FC13F81D" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="62">
<emphasis id="B9677117FFA8FA5876B7A538FE95F8C0" bold="true" box="[199,315,1821,1845]" pageId="9" pageNumber="62">Remarks.</emphasis>
Only two species of 
<taxonomicName id="4C13D686FFA8FA58744AA538FD1CF8C0" box="[570,690,1821,1845]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="62" phylum="Chordata" rank="genus">
<emphasis id="B9677117FFA8FA58744AA538FD1CF8C0" box="[570,690,1821,1845]" italics="true" pageId="9" pageNumber="62">Xenophrys</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C13D686FFA8FA5874B3A53BFCE5F8C0" box="[707,843,1821,1845]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="62" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="B9677117FFA8FA5874B3A53BFD7AF8C0" box="[707,724,1822,1845]" italics="true" pageId="9" pageNumber="62">X</emphasis>
. 
<emphasis id="B9677117FFA8FA587496A538FCE5F8C0" box="[742,843,1821,1845]" italics="true" pageId="9" pageNumber="62">boettgeri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C13D686FFA8FA5875F9A53BFC41F8C0" box="[905,1007,1821,1845]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="62" phylum="Chordata" rank="species" species="minor">
<emphasis id="B9677117FFA8FA5875F9A53BFC34F8C0" box="[905,922,1822,1845]" italics="true" pageId="9" pageNumber="62">X</emphasis>
. 
<emphasis id="B9677117FFA8FA5875DCA53BFC41F8C0" box="[940,1007,1822,1845]" italics="true" pageId="9" pageNumber="62">minor</emphasis>
</taxonomicName>
were previously known occurring at Mount Jinggang (
<bibRefCitation id="EF82D0F4FFA8FA587714A565FE4FF8AD" author="Zou" box="[356,481,1856,1880]" pageId="9" pageNumber="62" refString="Zou, D. - L. (1985) Amphibians and their faunistic distribution of Jinggangshan. Journal of Nanchang University (Natural Science), 1, 51 - 55." type="journal article" year="1985">Zou, 1985</bibRefCitation>
). During our herpetological surveys in Mount Jinggang, three apparently different morphotypes of 
<taxonomicName id="4C13D686FFA8FA58773EA540FE68F888" box="[334,454,1893,1917]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="62" phylum="Chordata" rank="genus">
<emphasis id="B9677117FFA8FA58773EA540FE68F888" box="[334,454,1893,1917]" italics="true" pageId="9" pageNumber="62">Xenophrys</emphasis>
</taxonomicName>
were found, including the new species described herein. Morphologically the other two morphotypes could not be identified as any other 
<taxonomicName id="4C13D686FFA8FA5874B7A5ADFC91F855" box="[711,831,1928,1952]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="62" phylum="Chordata" rank="genus">
<emphasis id="B9677117FFA8FA5874B7A5ADFC91F855" box="[711,831,1928,1952]" italics="true" pageId="9" pageNumber="62">Xenophrys</emphasis>
</taxonomicName>
species that were recorded from Eastern 
<collectingCountry id="F304ED95FFA8FA587361A5ADFAFAF855" box="[1297,1364,1928,1952]" name="China" pageId="9" pageNumber="62">China</collectingCountry>
(
<collectingCountry id="F304ED95FFA8FA587314A5ADFAD7F855" box="[1380,1401,1928,1952]" name="Italy" pageId="9" pageNumber="62">Y</collectingCountry>
. 
<collectingCountry id="F304ED95FFA8FA5873F4A5ACFA36F855" box="[1412,1432,1929,1952]" name="Italy" pageId="9" pageNumber="62">Y</collectingCountry>
. Wong 
<emphasis id="B9677117FFA8FA587691A58BFEB5F830" box="[225,283,1965,1989]" italics="true" pageId="9" pageNumber="62">et al.</emphasis>
, unpublished data). This indicates the cryptic diversity of the 
<taxonomicName id="4C13D686FFA8FA5875AEA588FBF8F830" box="[990,1110,1965,1989]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="62" phylum="Chordata" rank="genus">
<emphasis id="B9677117FFA8FA5875AEA588FBF8F830" box="[990,1110,1965,1989]" italics="true" pageId="9" pageNumber="62">Xenophrys</emphasis>
</taxonomicName>
in Mount Jinggang and thus the systematics of the 
<emphasis id="B9677117FFA8FA5877E0A5F5FD8AF81D" box="[400,548,2000,2024]" italics="true" pageId="9" pageNumber="62">
<taxonomicName id="4C13D686FFA8FA5877E0A5F5FDA6F81D" box="[400,520,2000,2024]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="62" phylum="Chordata" rank="genus">Xenophrys</taxonomicName>
in
</emphasis>
Eastern 
<collectingCountry id="F304ED95FFA8FA5874F6A5F5FD67F81D" box="[646,713,2000,2024]" name="China" pageId="9" pageNumber="62">China</collectingCountry>
needs to be revisited.
</paragraph>
</subSubSection>
</treatment>
</document>